taxonID	type	description	language	source
03F187EDFFF3FFF8E1D2F975E7E4FCE2.taxon	description	(Figs 1 – 2, 35 A)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF3FFF8E1D2F975E7E4FCE2.taxon	materials_examined	Material examined, 121 specimens. USA: District of Columbia: D. C. (ANSP, 1; CMNH, 13); D. C., Brend., Horn Coll. H 1927 / 8303.1 PARATYPE (MCZC, 1). Piney Bridge, V- 12 - 1905, D. H. Clemons (FMNH, 1). Wash. (ington), VI- 18, Hubbard & Schwarz (USNM, 1). Maryland: Baltimore Co.: Baltimore, VI- 10 - 1909, F. E. Blaisdell (MCZC, H. C. Fall Collection, 1). Hawkins Point, VII- 3 - 1966, E. J. Ford, Jr. (light trap DSCC, 2). Cecil Co.: Cecilton, VI- 2 - 1955, R. S. Howard (FMNH, 1). Charles Co.: Indian Head, V- 12 - 1944, G. B. Vogt, at light (USNM, 1). Kent Co.: Eastern Neck Island, III- 27 - 1956, R. S. Howard (FMNH, 4). Montgomery Co.: near Plummers Island, V- 22 - 1915, R. C. Shannon (LSAM, 1). Only state on label: “ Md. ” (CMNH, 1). Massachusetts: Plymouth Co.: Humarock, V- 8 - 1933, C. E. White (MCZC, 1). New Hampshire: Rockingham Co.: Seabrook backdunes, 1 m, VIII- 3 - 1989, D. S. Chandler, UV light (DSCC, 1). New York: Queens Co.: Rock (away) Beach, L. (ong) I. (sland), VI- 11 - 1922 (USNM, 1). Richmond Co.: Staten Island, III- 17 - 1921, A. Nicolay (MCZC, H. C. Fall Collection, 3; USNM, 13; CNCI, 1; AMNH, 2; ANSP, 8); V- 8 - 1921 (USNM, 21); V- 1 - 1920 (ANSP, 1); III- 17 - 1921 (MCZC, 1); IV- 1906, salt meadow (CMNH, 1); V- 12, salt meadow (MCZC, 1; DSCC, 1); VII (MCZC, 6); V- 18 (MCZC, 2); IV- 26, salt marsh (MCZC, 1); III- 17 (FMNH, 1), IV- 26 (MCZC, 1), XI- 10 (FMNH, 1), host Myrmica sabuleti MT; IV- 1906, salt meadow (MCZC, 6); no date (CUIC, 1; DSCC, 1; FMNH, 14). North Carolina: Mecklenburg Co.: Davidson, X- 22 - 1965, T. Daggy (NCSU, 1). Virginia: Spotsylvania Co.: Fredricksburg, V- 9 - 1891, W. D. Richardson (USNM, 1). Only state on label: “ Va. ” (MCZC, Lectotype; CMNH, Paralectotype).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF3FFF8E1D2F975E7E4FCE2.taxon	description	Description. BODY: Length 2.48 – 2.64 mm; orange-brown to elytra orange-brown and rest of body brown; setae on pronotum short, curved and suberect, decumbent over rest of body. Head: surface lightly reticulate to smooth, shiny, punctures minute. Median vertexal fovea setose usually punctiform but sometimes shallow, one-half width to nearly as large as lateral vertexal foveae. Antennae strongly modified in male. Pronotum: with surface smooth, shiny to lightly microreticulate, with distinct, small punctures separated by about twice their width. Median antebasal fovea one-half to two-thirds width of lateral antebasal foveae. Elytra: surface microreticulate, punctures indistinct; discal stria extending to about four-fifths of elytral length. Abdomen: surface smooth, punctures small but distinct; basal striae of tergite 1 extending nearly to one-third paratergite length, at base striae separated by about two-fifths tergite width, with short and sparse setal brush between striae. MALE: Antennomere III elongate, IV – V transversely rectangular, VI – VIII transversely triangular in lateral view, IX transverse, bluntly pointed on lateral margin, articulation of IX and X offset medially on IX, X elongate, asymmetrically flattened on mesal margin, gently convex on lateral margin, with apex of elongate, thin hyaline tubercle arising near base and with apex abruptly (Figs 2 A – B); XI with asymmetric ventrolateral lobe in basal half, apical half broad and twisted (Fig. 2 C). Protrochanter with small acute ventral spine at base (Fig. 1 C); mesotrochanter with prominent narrow spine with blunt apex (Fig. 1 D); metatrochanter with acute ventral spine at apex (Fig. 1 E). Metaventrite with broad impression at middle, setae in impression denser than laterally, elongate and appressed. Metatibiae narrow basally, slightly widening through length to apex, with dense comb of setae on mesal margin in apical fifth where slightly curved medially (Fig. 1 B). Abdominal tergites unmodified (Fig. 1 A); ventrites 2 – 4 flattened at middle, 5 shallowly impressed at middle. Aedeagus 0.54 mm long; with dorsal plate transversely pentagonal, apex bluntly pointed; parameres short and broad, with three thick setae on outer margin in area of preapical constriction, flat preapical hyaline setae very large, nearly as wide as long; internal sac with three large curved spines (Fig. 2 D). FEMALE: Similar to male except: antennomeres IV – V slightly longer than wide, VI – VII about as long as wide, VIII slightly transverse, IX slightly longer than wide, X longer than wide, swollen ventrally. Metatibiae similar to those of male. Metasternum with disc convex, lacking denser setae posteriorly. Abdominal ventrites convex. Collecting data. With ant, Myrmica sabuleti, in salt marsh; at UV light (Chandler, 1997). Adults have been taken from March to July, with a single record from October in North Carolina. This species has been collected in salt marshes and is attracted to ultraviolet lights.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF3FFF8E1D2F975E7E4FCE2.taxon	distribution	Distribution (Fig. 35 A). Blatchley (1910) recorded this species from Indiana based on two specimens from Putnam and Posey Counties. Examination of these two specimens revealed a female of Batrisodes lineaticollis (Aubé) (Posey Co., IX- 29 - 1909), and an unplaced female of Rybaxis (Putnam Co., X- 10 - 1909). Brachygluta cavicornis is a species of the eastern seaboard salt marshes, and is known from New Hampshire to North Carolina.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF3FFF8E1D2F975E7E4FCE2.taxon	discussion	Comments. Readily separated from the other members of the cavicornis species-group by the elongate discal striae of tergite 1, which are indistinct in the other species, and the large setose median vertexal fovea of the head, which is smaller to lacking in the other species. The range and habitat of B. cavicornis overlaps to a large extent with that of B. luniger, but does not extend to the northern and southern extremes of that species.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF6FFF8E1D2FC3CE034FA34.taxon	distribution	Distribution: New York.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF6FFF8E1D2FC3CE034FA34.taxon	discussion	Comments. The type of this species could not be found in the Moscow State University Collection, the Finnish Museum of Natural History in Helsinki, or the Zoological Institute in St. Petersburg (Sergei Kurbatov pers. comm.). In his description Motschulsky mentions that the antennal club is modified, which places this species in the subgenus Nisa, with both northern species of this subgenus commonly found in coastal New York. Motschulsky’s brief description of the antennal form is suggestive of the more common B. luniger (LeConte), but until the type is found this synonymy is not formalized.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF6FFFAE1D2FA52E206F8F2.taxon	description	(Figs 3, 35 B)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF6FFFAE1D2FA52E206F8F2.taxon	materials_examined	Material examined, 304 specimens. CANADA: New Brunswick: Gloucester Co.: Caraquet, nr. Acadian Historical Village, 47.7873 N 66.0797 W, VIII- 14 - 2005, R. Webster & G. Pohl, salt marsh, intertidal zone, on patches of bare clay at bases of Spartina patens (RPWC, 2). Only country on label: “ Canada ”, Kraatz (MNHN, 4 M). USA: Connecticut: Fairfield Co.: Stamford, III, Angell (MCZC, 3). Delaware: Sussex Co.: Lewes, ca. 400 ' into saltwater marsh from Canary Creek, III- 14 - 1959, R. E. Hillman (FMNH, 1). Rehoboth Beach, V- 12 - 1962, W. A. Connell (UDCC, 1). Vines Creek, Dogsboro, II- 1954 (FMNH, 4); III- 11 - 1954 (FMNH, 37; FSCA, 1), II- 3 - 1954 (MHNG, 1), R. S. Howard. District of Columbia: D. C. (ANSP, 4; CMNH, 7; MCZC, 12; MCZC, LeConte Collection, 5; SEMC, 2); Brend. (MCZC, Horn Collection, 4). Washington, VI- 20 (MCZC, LeConte Collection, 1). Wash. (ington), VI- 23 (USNM, 2); VI- 20, Hubbard & Schwarz (USNM, 1). Florida: Duval Co.: Atlantic Beach (AMNH, 1). Indian River Co.: 1 mi S Vero Beach, X- 6 - 1978, O'Brien & Marshall, UV light in salt marsh (DSCC, 1). Miami-Dade Co.: Miami, IV- 1930 (CMNH, 4). Palm Beach Co.: Lake Worth (CMNH, 2). Volusia Co.: Edgewater, II- 28 - 1939, C. A. Frost (MCZC, 1); II- 20 - 1939, C. A. Frost (FMNH, 1). Only state on label: “ Fla. ” (MCZC, 1); Florida, Brend. (MCZC, Horn Collection, 1). Georgia: Glynn Co.: Brunswick, VI- 5 - 1952, D. G. Kissinger, to light (FMNH, 1). St. Simons Island, VII- 18 - 1931, C. A. Frost (DSCC, 1); VII- 21 - 1931, Quirsfeld (CNCI, 1); St. Simon Island, IV- 22 / V- 12 - 1911, J. C. Bradley (CUIC, 1). Laurens Co.: Dublin, VI- 1960 (MHNG, 11). Liberty Co.: Fort Stewart, III- 28 - 1944 (DSCC, 1). McIntosh Co.: Sapelo Island, VI- 1963, H. Kale, ex: Spartina (DSCC, 1). Only state on label: “ Ga. ” (ANSP, Holotype B. perpunctata; CMNH, 2). Maine: York Co.: Old Orchard (Beach), VIII- 15 - 1912 (MCZC, H. C. Fall Collection, 1). Maryland: Anne Arundel Co.: Skidmore, V- 26 - 1956, R. S. Howard (FMNH, 2). Baltimore Co.: Baltimore, Ordnance Depot, VI- 3 - 1973, E. J. Ford, Jr., light trap (DSCC, 1); Baltimore, VI- 17 - 1975, E. J. Ford, Jr., light trap (DSCC, 1). Butler, V- 23 - 1982, E. J. Ford, light trap (EJFC, 1). Hebbville, V- 21 - 1962, E. J. Ford, Jr., light trap (DSCC, 1). Calvert Co.: Flag Pond, 3 mi S Kenwood Beach, VI- 24 - 1959, P. & P. Spangler, on beach under debris (LSAM, 2). Kent Co.: Eastern Neck Island, III- 27 - 1956, R. S. Howard (FMNH, 7). Montgomery Co.: Minnie Island, nr. Cabin John, VI- 5 - 1931, H. S. Barber, at light (USNM, 1). Plummers Island, VI- 23 - 1906, E. A. Schwarz (LSAM, 1). St. Mary's Co.: Point Look Out, IV- 1986, R. Gordon (USNM, 2). Somerset Co.: Marion, VII- 29 - 1978 (DSCC, 1), VI- 5 - 1971 (DSCC, 1), E. J. Ford, Jr., light trap. Only state on label: “ Md. ” (CMNH, 4; MCZC, 3); Brend. (MCZC, Horn Collection, 1). Massachusetts: Barnstable Co.: West Falmouth (FMNH, 6). Woods Hole (FMNH, 1). Bristol Co.: Dighton, X- 8 - 1910, N. S. Easton (MCZC, 1). Somerset, IV- 29 - 1911, N. S. Easton (MCZC, 1). Essex Co.: Ipswich, V- 22 - 1926, P. J. Darlington (MCZC, 1); VII- 27 - 1926 (CUIC, 2). Middlesex Co.: Cambridge (MCZC, Holotype B. luniger). Malden, V- 17 - 1868, C. P. Whitney (MCZC, 3); V- 17 - 1868, E. P. Colby, (MCZC, LeConte Collection, 1). Stoneham, VI- 1907, F. A. Sheriff (MCZC, 2). New Jersey: Atlantic Co.: Atlantic City (MCZC, 2). Cape May Co.: Anglesea, IV- 7, H. W. Wenzel (MCZC, 1); IV- 27 (USNM, 1); IV- 7, meadows (MCZC, H. C. Fall Collection, 2); IV- 15, meadows (FMNH, 1); IV- 17, meadow (AMNH, 1); IV- 27, Wenzel, meadows (CASC, 2); IV- 27, meadows (FMNH, 2); Anglesea (FMNH, 7). Five Mile Beach, V- 30, (CMNH, 2). Warren Co.: Delaware Water Gap (MCZC, 1). New York: Queens Co.: Jamaica, L. (ong) I. (sland), V- 30 - 1905 (USNM, 1). Rock (away) Beach (FMNH, 1). Suffolk Co.: Cold Springs Harbor, L. (ong) I. (sland), VII- 11 - 1921, H. M. Parshley (CASC, 1); VII- 15 - 1919, H. M. Parshley (CASC, 1). Suffolk Co., VI- 17 - 1960, M. I. Blenderman (AMNH, 25 F, 6 M); VI- 24 - 1960 (AMNH, 4); kept alive, died IX- 22 - 1962 (AMNH, 2 F); kept alive, died X- 21 - 1960 (AMNH, 1 F); kept alive, died X- 10 - 1962 (AMNH, 1 F); kept alive, died XI- 24 - 1960 (AMNH, 1 M); kept alive, died XI- 24 - 1960 (AMNH, 1 F); kept alive, died VII- 1 - 1961 (AMNH, 1 F); kept alive, died VIII- 16 - 1960 (AMNH, 1 F); kept alive, died XII- 10 - 1961 (AMNH, 1 F); kept alive, died VIII- 6 - 1960 (AMNH, 1 F); kept alive, died VIII- 8 - 1960 (AMNH, 1 F); kept alive, died VIII- 23 - 1962 (AMNH, 1 F); kept alive, died II- 1 - 1963 (AMNH, 1 F); kept alive, died V- 19 - 1962 (AMNH, 1 F); kept alive, died VI- 23 - 1961 (AMNH, 1 F); kept alive, died VII- 26 - 1960 (AMNH, 1 F); kept alive, died I- 15 - 1961 (AMNH, 1 M); kept alive, died XI- 14 - 1962 (AMNH, 1 F); kept alive, died XII- 1 - 1960 (AMNH, 1 M); kept alive, died V- 3 - 1961 (AMNH, 1 F); kept alive, died XI- 17 - 1960 (AMNH, 1 F); kept alive, died VIII- 26 - 1960 (AMNH, 1 F); kept alive, died IV- 2 - 1961 (AMNH, 1 M); kept alive, died IV- 24 - 1962 (AMNH, 1 F). Westchester Co.: Katonah (MCZC, 1). Peekskill, Hubbard & Schwarz (USNM, 3); V- 12 (FMNH, 1); V- 16 - 1890 (MCZC, 2); V- 12 - 1899 (MCZC, 2); V- 9 - 1890 (MCZC, 2); VI- 1 - 1891 (CASC, 1); IV- 12 - 1899 (DSCC, 1); VI- 12 - 1898 (CMNH, 1). County unknown: Tecky Kill (spell ??), VI, Schumann (AMNH, 1). Only state on label: “ N. Y. ” (FMNH, 3); Brend. (MCZC, Horn Collection, 1). North Carolina: Mecklenburg Co.: Charlotte, V- 17 / 24 - 2005, J. F. Cornell, UV light trap (LSAM, 1). New Hanover Co.: Marine Research Center, VIII- 15 - 1982, K. R. Ahlstrom, at light trap (NCSU, 1). Pennsylvania: Only state on label: “ Pen. ” (CMNH, 2). South Carolina: Charleston Co.: Charleston, IV- 25 - 1945, R. L. Wenzel, light trap (FMNH, 1). Nr. McClellanville, Wedge Plantation, IV- 16 - 1977, E. G. Munroe, at light (CNCI, 1). Pickens Co.: Calhoun (= Clemson), VIII- 1949, W. F. Chamberlain (TAMU, 1). Only state on label: “ S. C. ” (CMNH, 1). Virginia: Alexandria Co.: St. Elmo, I- 6 (USNM, 1). Fairfax Co.: Burke Center, VII- 13 / 18 - 1987, R. D. Ward (CMNH, 1). Falls Church City: Falls Church, V- 30 (MCZC, 1). Hampton City: Fort Monroe, IV- 19, Hubbard & Schwarz (USNM, 10). Norfolk City: Norfolk, IV- 28 / V- 1 - 2003, UV light (EIUC, 1); V- 20 / 26 - 2003, C. A. Springer, UV light (EIUC, 1); Norfolk (MCZC, H. C. Fall Collection, 2). Spotsylvania Co.: Fredricksburg, V- 19 - 1891 (USNM, 1); IV- 28 - 1890 (USNM, 1); VI- 3 - 1891, W. D. Richardson (USNM, 1).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF6FFFAE1D2FA52E206F8F2.taxon	description	Description. BODY: Length 2.40 – 2.80 mm; orange-brown to brown with orange-brown elytra; setae short, curved and suberect on pronotum, decumbent over rest of body. Head: surface smooth, shining, punctures minute. Median vertexal fovea usually lacking, if present usually represented by small patch of white setae in shallow impression, rarely punctiform. Antennae strongly modified in male, form for sexes discussed below. Pronotum: surface smooth, punctures distinct, separated by about one puncture diameter. Median antebasal fovea about twothirds width of lateral antebasal foveae. Elytra: surface microreticulate, granulate, punctures indistinct; discal striae extending to about four-fifths of elytral length. Abdomen: basal striae of tergite 1 lacking or very short, when visible separated by about one-third tergite width, width of setose brush between strial bases about one-fourth tergite width. Surface smooth, punctures distinct. MALE: Antennomeres III – V slightly longer than wide, VI as wide as long, VII – VIII transverse, VIII with length about one-third width, ventral portion slightly more acutely pointed than dorsal, IX enlarged and broadly cup-shaped, dorsal margin slightly emarginate, X elongate, origin from IX offset ventrally, X widening toward apex and curving dorsally, with thin elongate tubercle arising from basal half of incurved side, XI symmetrical, obconical (Figs 3 E – G). Protrochanter with small acute spine on ventral margin (Fig. 3 B); mesotrochanter with blunt ventral spine at base (Fig. 3 C). Metaventrite with broad impression covering middle, with dense fringe of setae on posterior margin between metacoxae. Metatibiae narrow basally, slightly widening through length to apex, with dense comb of setae on mesal margin in apical fifth where slightly curved medially (Fig. 3 D). Abdomen with tergites gently convex, unmodified (Fig. 3 A); ventrites 2 – 4 flattened at middle, 5 shallowly impressed at middle. Aedeagus 0.44 mm long; with dorsal plate roughly triangular, apex bluntly pointed; parameres short and thick, with three thick setae on outer margin of preapical constriction, preapical hyaline setae transversely oval; internal sac with three large curved spines (Fig. 3 H). FEMALE: Antennomeres lacking modifications, III and V clearly longer than wide, IV and VI – VII slightly longer than wide, VIII as long as wide, IX about as long as wide, half again as wide as VIII, X elongate, rectangular. Legs with simple trochanters, ventral margins smoothly curved. Metatibiae similar to those of male. Female metaventrite broadly convex, setae slightly longer between metacoxae than those on lateral portions of the disc. Abdominal ventrites convex. Collecting data. In salt marshes, in low swampy ground (Brendel & Wickham 1890); at UV light (Chandler 1997); on bare clay patches at base of Spartina patens at upper margins of tidal stream (Webster et al. 2012). LeConte (1849) recorded the type from “ pratis salinis ” (salt meadows). Few specimens have habitat or collecting data indicated, with about half taken at ultraviolet light and the rest collected in salt meadows / marshes or under debris on beaches. Most specimens were taken from February to early July, with only a few records scattered through the rest of summer and in autumn.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF6FFFAE1D2FA52E206F8F2.taxon	distribution	Distribution (Fig. 35 B). A species of the salt marshes of the Atlantic Coast states and provinces, with records ranging from New Brunswick to Florida.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF6FFFAE1D2FA52E206F8F2.taxon	discussion	Comments. LeConte (1849) placed the holotype as a possible female, but his description of the antennae clearly indicates that he had a male before him. Blatchley (1910) recorded this species (as B. perpunctata) from Illinois, but this was either an error or was based on a mislabeled specimen of B. luniger (also, see the Distribution section for B. cavicornis for an apparently similar situation). The holotype female of B. perpunctata is a member of the cavicornis species-group, and lacks a setose median vertexal fovea and lacks discal carinae on tergite 1. These features place the type as a member of B. luniger, which is the only species of the cavicornis species-group known from Georgia.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF5FFE6E1D2F8C9E65AFF47.taxon	description	(Figs 4 – 5, 35 C)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF5FFE6E1D2F8C9E65AFF47.taxon	materials_examined	Material examined, 6 specimens. HOLOTYPE, male: USA: Florida: Miami-Dade Co.: // Cutler, FLORIDA, Matheson Hammock, 28. IV. 62, R. S. Howard leg. / Field Mus. Nat. Hist., Orlando Park Pselaphidae Colln. / [red label] HOLOTYPE Brachygluta miccosukee Bückle // (FMNH). PARATYPES, 5: eutopotypical (FMNH, 1 M, 2 F). Pinellas Co.: St. Petersburg, III- 29 - 1922 (MCZC, H. C. Fall Collection, 1 M). Only state on label: “ Fla. ” (DSCC, 1 M).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF5FFE6E1D2F8C9E65AFF47.taxon	description	Description. BODY: Length 2.60 – 2.68 mm; yellow-brown to orange-brown; setae short, curved and suberect on pronotum, decumbent over rest of body. Head: surface smooth, shining, punctures minute. Median vertexal fovea usually not apparent, to represented by small setose patch or (1 specimen) punctiform and setose, clearly smaller than lateral foveae when present. Antennae strongly modified in male, form for sexes discussed below. Pronotum: surface appearing lightly granular in some due to distinct close punctation, varying to surface smooth with punctures separated by about one puncture diameter. Median antebasal fovea one-half to two-thirds width of lateral antebasal foveae. Elytra: surface microreticulate, opaque, punctures faint; discal striae extending to about four-fifths of elytral length. Abdomen: surface smooth, punctures distinct and dense. Basal striae of tergite 1 lacking, base with small setose brush at middle about one-fifth tergite width, indistinct in female. MALE: Antennomeres III and V slightly longer than wide, IV and VI as wide as long, VII slightly transverse, VIII transverse with length about one-third width, ventral portion slightly more acutely pointed than dorsal, IX enlarged and broadly cup-shaped, apical margin broadly emarginate dorsally, X transverse with stem continuing from ventral side and inserting offset into IX, with conical tubercle arising from basal portion of incurved side, with setal tuft at apex (Figs 5 A – B), XI symmetrical, obconical. Protrochanter with short basal blunt angulation, mesotrochanter at base with elongate ventral spine, spine blunt at apex (Figs 4 C – D). Metaventrite broadly and circularly impressed at middle, with dense elongate setae in impression, particularly near margins. Metatibiae narrow basally, slightly widening through length to apex, with dense comb of setae on mesal margin in apical fifth where slightly curved medially (Fig. 4 B). Abdomen with tergites gently convex, unmodified (Fig. 4 A); ventrites 2 – 4 flattened in middle half, 5 shallowly impressed at middle. Aedeagus 0.48 mm long; parameres with abrupt preapical constriction on lateral margins, with three thick setae on outer margin of preapical constriction that reach paramere apex, preapical hyaline setae inserted near mesal margin, broadly and evenly wide through length; internal sac with three large curved spines, one spine straight, two curved (Fig. 5 C). FEMALE: Antennae lacking modifications, antennomeres III and V longer than wide, IV and VI – VII as long as wide, VIII transverse, IX about as long as wide. Legs with simple trochanters, ventral margins smoothly curved. Metaventrite with disc lightly longitudinally impressed, apex between metacoxae with short fringe interrupted at middle. Metatibiae similar to those of male. Abdominal ventrites convex. Collecting data. Both sites are near or at the Atlantic coast of Florida, with the specimens taken in March and April.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF5FFE6E1D2F8C9E65AFF47.taxon	distribution	Distribution (Fig. 35 C). This species has been taken from two widely separated sites on the Atlantic coast of Florida, indicating a wide distribution in the state.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF5FFE6E1D2F8C9E65AFF47.taxon	discussion	Comments. This is the sister-species of B. luniger based on antennal form in the males, the short or absent discal carinae of tergite 1, and the lack of a spine on the male metatrochanters. It is easily separated from B. luniger by the more quadrate form of antennomere X in the male of B. miccosukee, which is strongly transverse in B. luniger.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFF5FFE6E1D2F8C9E65AFF47.taxon	etymology	Etymology. The name is taken from one of the groups of American Indians that lived in southern Florida.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE8FFE0E1D2FC5DE121F806.taxon	description	(Figs 6, 36 A, 44, 45)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE8FFE0E1D2FC5DE121F806.taxon	materials_examined	Material examined, 501 specimens. CANADA: New Brunswick: Albert Co.: Mary's Pit, IX- 8 - 2002, C. Majka, salt marsh, vegetation, (NSMNH, 1). Shapoody NWA, Mary's Point Section, 4.45.7321 N, 66.6765 W, V- 17 - 2004, R. P. Webster, grass litter resting on salt marsh vegetation (RPWC, 4). USA: Connecticut: Fairfield Co.: Westport, IV- 23 - 1932, IX- 25 - 1931, VII- 16 - 1931, VIII- 5 - 1931, X- 1 - 1932, X- 2 - 1932, L. Lacey (AMNH, 6, 1 each site). New London Co.: Groton, K. P. Jansson (MZLU, 4). Delaware: Sussex Co.: Rehoboth, VI- 14 - 1945, Dieke, beach washup (USNM, 1). County uncertain: Ship John Shoal Light House, Delaware Bay, VIII- 23 - 1936, D. MacCreary (UDCC, 1). District of Columbia: D. C., 1932 (AMNH, 1); D. C. (ANSP, 3; CMNH, 11; FMNH, 6; MCZC, 17; MCZC, LeConte Collection, 6; INHS, 3); Brend. (MCZC, Horn Collection, 2). Eastern Branch, nr. Bennings, II- 23 - 1913, W. L. McAtee (USNM, 1). Wash. (ington), C. V. Riley (USNM, 1); VI- 20, Hubbard & Schwarz (USNM, 1; LSAM, 1). Only state on label: “ Columbia ” (MHNG, 1). Georgia: Glynn Co.: St. Simon Island, IV- 22 / V- 12 - 1911, J. C. Bradley (MCZC, 2). Maine: Hancock Co.: Site 16, Big Moose Island lagoon, SE end, VII- 17 - 2005, D. S. Chandler, sifting-berlesing, salt marsh litter (ANPC, 1). Tremont, X- 5 - 1937 (USNM, 7). Knox Co.: Spruce Head, VI- 16 - 1955, R. S. Howard (FMNH, 26). Sagadahoc Co.: Popham Beach, III- 23 - 1983, R. E. Nelson, edge of salt marsh (RENC, 1). Maryland: Baltimore Co.: Baltimore, Lanton, VI- 6 - 1973, E. J., Ford, Jr., light trap (DSCC, 1). Butler, VII- 1 - 1979, E. J. Ford, light-trap (EJFC, 1). Hawkins, Point, VII- 3 - 1966, E. J. Ford, Jr., light trap (DSCC, 1). Hebbville, VII- 17 - 1999, E. J. Ford, Jr., light trap (DSCC, 1). Calvert Co.: Chesapeake Beach, VII- 20 - 1932, F. C. Bishop (USNM, 1). Scientists Cliff, III- 23 - 1980, J. M. Campbell (CNCI, 5). Harford Co.: Edgewood, IX- 12 - 1918, H. Dietrich (CUIC, 1). Kent Co.: Eastern Neck Island, III- 27 - 1956 (879 - A), R. S. Howard (FMNH, 1). Montgomery Co.: Minnie Island, nr Cabin John, VI- 5 - 1931, H. S. Barber (USNM, 1). Plummers Island, V- 24 - 1908, H. S. Barber (LSAM, 1); IX- 28 - 1913, W. L. McAtee (USNM, 1). Takoma Park, VI- 24 - 1951, G. H. Nelson, to light (DSCC, 1). Prince Georges Co.: Bladensburg, IX- 10 - 1961, E. Van Tassel (FSCA, 1). Only state on label: “ Md. ” (MCZC, H. C. Fall Collection, 3; MCZC, 2). Massachusetts: Bristol Co.: Berkeley, VIII- 15 - 1932, N. S. Eaton, beach litter (MCZC, 1). Fall River, VII- 2 - 1919, N. S. Eaton (MCZC, 4). Somerset, VI- 26 - 1910 (SEMC, 1). Essex Co.: Ipswich, V- 22 - 1926, P. J. Darlington (MCZC, 1). Nahant, VI- 3 - 1935, P. J. Darlington (MCZC, 2). Salisbury Beach, III- 23 - 2009, T. Murray, photo (DSCC, 1); IV- 19 - 2010, T. Murray, photo (DSCC, 2); IV- 20 - 2009, T. Murray, photo (DSCC, 2). Middlesex Co.: Cambridge, IV- 14, Hubbard & Schwarz (USNM, 3). Stoneham, VI- 7, F. A. Sheriff (MCZC, 1). Tyngsboro, IV- 13 - 1901, freshet (MCZC, 1). Nantucket Co.: Nantucket, V- 26 - 1935, C. R. Crosby (CUIC, 1). Plymouth Co.: Marion, VIII- 20 - 1904 (MCZC, 6); VIII- 15 - 1904 (MCZC, 1). Suffolk Co.: Chelsea, salt marsh (MCZC, 1). Revere, V- 16 - 1903 (MCZC, 1). Only state on label: “ Mas. ” (CMNH, 1). New Hampshire: Rockingham Co.: Hampton, IV- 5 - 1923, S. A. Shaw (LSAM, 1); III- 11, S. A. Shaw (MCZC, 1); III- 11 - 1923 (USNM, 15). Hampton, III- 11 (ANSP, 1). Hampton, III- 25 - 1902 (2), IV- 15 - 1927, IV- 5 - 1902, V- 25 - 1935, IV- 28 - 1929, IV- 27 - 1930, V- 3 - 1932, III- 23 - 1902, III- 22 - 1903, IV- 15 - 1917, S. A. Shaw (all UNHC). Hampton (FMNH, 3). Odiorne Point State Park, V- 12 - 1982, sift grass litter in marsh (DSCC, 2; MHNG, 1; UNHC, 1); V- 6 - 1982, sift leaf litter along marsh edge (DSCC, 1; UNHC 1); IV- 30 - 1986, sift leaf litter on marsh edge (DSCC, 4; UNHC, 1); IX- 16 - 2006, sweeping (DSCC, 2); IX- 24 - 1986, sift leaf litter on pond margin (DSCC, 1; MHNG, 1); V- 14 - 1982, under leaf litter around pond (DSCC, 5; UNHC, 11); VI- 15 - 1982, D. S. Chandler, sift leaf litter around pool edge (UNHC, 2); VI- 3 - 1986, D. S. Chandler, sift leaf litter on marsh edge (UNHC, 2). Seabrook, back-dunes, 1 m, VI- 1 / 14 - 1989, D. S. Chandler, FITrap (UNHC, 1); VIII- 10 - 1989, D. S. Chandler, under grass debris (UNHC, 1). Seabrook foredunes, VI- 27 - 1989, D. S. Chandler (DSCC, 1); Strafford Co.: Durham, V- 13 - 1900, Weed & Fiske (INHS, 1). New Jersey: Atlantic Co.: Chelsea, VII- 29 (CMNH, 1). Bergen Co.: Fort Lee District (MCZC, 5); VI- 11 - 1922 (USNM, 1). Cape May Co.: Anglesea, IV- 15, W. D. Richardson, meadows (LSAM, 1); IV- 15, meadows (CMNH, 1; FMNH, 1); IV- 7, Wenzel (ANSP, 11); IV- 7 (CMNH, 11); IV- 15, (MCZC, 5); IV- 1915 (AMNH, 2); III- 11 (USNM, 1); IV- 15, meadows (MNHN, 4 M and 1 F); no date (MCZC, 5; FMNH, 5; MNHN, 5). S. M. (Strathmere?) Beach, IV- 7, H. W. Wenzel (FMNH, 8). Cape May (FMNH, 1). Hudson Co.: Arlington (MCZC, H. C. Fall Collection, 1). Snake Hill (= Laurel Hill) (AMNH, 1); II- 1891 (CASC, 3). Middlesex Co.: S. (outh) Amboy, IV- 16 - 1916, F. M. Schott (FMNH, 1). Monmouth Co.: Spring Lake, VIII- 14 - 1941, A. Nicolay (USNM, 2). Ocean Co.: 3 mi S Tuckerton, VI- 15 - 1983, D. S. Chandler, sift beach drift (DSCC, 5). Barnegat Bay, V- 25 - 1924, J. W. Green (CASC, 1). Manchester, V- 1 (MCZC, 1). Passaic Co.: Greenwood Lake, VI- 7 - 1942, A. Nicolay, sweeping at dusk (USNM, 1). Only state on label: “ N. J. ” (FMNH, 1). New York: Bronx Co.: Kingsbridge, 1923 (LSAM, 1). Kingsbridge, 1928 (USNM, 10). Dutchess Co.: Arlington, V- 22 - 1920, under board, salt marsh (AMNH, 4). Kings Co.: Brooklyn, IV- 16 - 1904 (USNM, 3); no date (FMNH, 1); Coney Island (AMNH, 1). Flatbush, L. (ong) I. (sland), IV- 10 - 19 --- (USNM, 1). Flatlands, L. (ong) I. (sland), XII- 25 - 1915, F. M. Schott (UNHC, 3). New Lots, Brooklyn, L. (ong) I. (sland), IV- 18 - 1914 (USNM, 1). Nassau Co.: Lynbrook, IV- 1 - 1918, C. R. Crosby (CUIC, 3). Wantagh, L. (ong) I. (sland), V- 1 - 1943, A. Nicolay (USNM, 5). Queens Co.: [no other data] (MCZC, 2; AMNH, 5). Aqueduct, L. (ong) I. (sland), V- 16 - 1925, F. R. Mason (ANSP, 5); V- 16 - 1925, Quirsfeld (CNCI, 4). Flushing, L. (ong) I. (sland), K. W. Cooper (USNM, 5). North Beach, L. (ong) I. (sland), II- 16 - 1913 (AMNH, 2). Rock (away) Beach, L. (ong) I. (sland), VI- 11 - 1922 (USNM, 4). Rosedale, L. (ong) I. (sland), III- 17 - 1926 (FMNH, 4). Jamaica, L. (ong) I. (sland) (AMNH, 5; MCZC, 2), Frost (INHS, 1). Richmond Co.: S. I. (Staten Island), salt meadow (MCZC, 10; AMNH, 2); III- 17 - 1921, A. Nicolay (ANSP, 1); V- 24 - 1939, A. Nicolay (USNM, 4); C. W. Leng (FMNH, 3); V- 6, salt meadow (MCZC, 1); V- 12, salt meadow (MCZC, 23); IX- 17 (MCZC, 1); salt meadow (MCZC, H. C. Fall Collection, 2). Staten Island, C. W. I. (CUIC, 1). Rockland Co.: Piermont, V- 27 - 1934, Quirsfeld (CNCI, 1); IV- 13 - 1930, Quirsfeld (CNCI, 1). Suffolk Co.: VI- 17 - 1960, M. I. Blenderman (AMNH, 2); VI- 24 - 1960, M. I. Blenderman (AMNH, 3). Babylon, VI- 7 - 1892 (AMNH, 1). Calverton, L. (ong) I. (sland), V- 14 - 1933, R. Latham (CUIC, 1). Orient, IV- 4 - 1931, C. R. Crosby (CUIC, 1). Southold, IV- 4 - 1931, C. R. Crosby (CUIC, 2). Westchester Co.: Peekskill, V- 30 - 1891 (AMNH, 2; MCZC, 1; CUIC, 1). Peekskill (MCZC, 1). L. (ong) I. (sland), XII- 13 - 1925, F. M. Schott (UAIC, 1); IV- 1909 (LSAM, 1); IV- 19 - 1909 (USNM, 2; FMNH, 1); VII- 3 - 1910 (FMNH, 1); IV- 2 - 1910 (FMNH, 1). No county listed: L (ong) Island (CMNH, 1). Only state on label: “ N. Y. ”, 1932 (AMNH, 2); “ N. Y. ” (AMNH, 4; CMNH; 6; FMNH, 11; MCZC, 4); Brend. (MCZC, Horn Collection, 7); salt meadow (SEMC, 2). North Carolina: Mecklenburg Co.: Charlotte, III- 18 - 1969, J. F. Cornell, leaves in field near house (NCSU, 1). Virginia: Fairfax Co.: Falls Church, XII- 27 (MCZC, 1); IX- 29 - 1929, A. Nicolay (CNCI, 1). Nr. Plummers Island Maryland, IX- 10 (USNM, 1). Only state on label: “ Va. ”, T. Pergande (USNM, 1). LITERATURE RECORDS: Nova Scotia: Cumberland Co.: Amherst, VI- 24 - 1994, J. Ogden, at edge of salt marsh. Maine: Penobscot Co.: Howland, V- 20 - 1978, R. Tracy (both records from Majka & Ogden, 2006).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE8FFE0E1D2FC5DE121F806.taxon	description	Description. BODY: Length 2.28 – 2.72 mm; orange-brown to brown, often with elytra lighter than rest of body; setae on head and pronotum short, curved and suberect, decumbent to nearly appressed over rest of body. Head: surface smooth, shining, punctures minute; setose area of median vertexal fovea about two-thirds width of setose area of lateral foveae. Antennomeres III-VII longer than wide, VII only barely so, VIII – IX transverse, wider than long, VIII about three-fourths width of IX, X as long as wide (Fig. 6 C). Pronotum: surface smooth, usually shining, faintly microreticulate for some, with distinct, small punctures. Setose area of median antebasal fovea slightly smaller than those of lateral antebasal foveae. Elytra: surface lightly microreticulate, punctures faint to indistinct; discal stria extending to about four-fifths of elytral length. Abdomen: tergite 1 with surface lightly punctate; basal striae very short, indistinct, one-fifth of paratergite length at most, sparsely setose brush between striae in both sexes, striae separated by about half tergite width. MALE: Antennae and trochanters lacking modifications, simple. Metaventrite with semicircular patch of dense setae extending from bases of metacoxae to middle of metaventrite. Abdomen with tergites 1 – 3 visible in dorsal view, all strongly modified and clearly visible (Fig. 6 A), 4 modified but not visible; tergite 1 longest, 2 – 3 subequal in length, tergites 1 – 3 medially emarginate, with lateral portions elevated and strongly protruding; tergite 1 deeply emarginate at middle for about one-fourth tergite width, elongate dark median tubercle originating beneath raised margin slightly emarginate at apex, lateral portions of tergite 1 each with rounded tubercle at center; tergite 2 with median emargination more shallowly and broadly emarginate, protruding lateral portions with margins slightly concave, ventral surface concave and densely setose, base of tergite 2 deeply depressed, more deeply depressed ventral to tubercles of tergite 1, apices of protruding portions of tergite 2 densely setose; tergite 3 with lateral thirds strongly protruding, ventral surface concave and densely setose, apex with median emarginate area slightly shallower and broader than for tergite 2, lateral margins of protruding angles shallowly concave but more deeply concave than for tergite 2; tergite 4 with lateral margins protruding, middle almost flat, with indistinct median longitudinal through length; tergite 5 nearly flat, not modified; in lateral view tergites 1 – 2 with outline roughly continuous, 2 strongly projecting out over base of 3 (Fig. 6 B). Abdominal ventrites 2 – 5 broadly convex, 6 with transverse impression at base. Metatibiae narrow basally, slightly widening through length to apex with dense comb of setae on mesal margin in apical sixth where slightly curved medially (Fig. 6 D). Aedeagus 0.39 mm long, with dorsal plate asymmetrically triangular; parameres with preapical constriction, with three thick setae at angle of constriction at lateral margin, with preapical hyaline setae near mesal margin thickened to middle, aciculate to apex; internal sac with three large subequal spines, one spine strongly curved (Fig. 6 E). FEMALE: Metaventrite in apical half somewhat flattened, setae evenly dense over metasternum, not as dense as in male. All tergites evenly convex; ventrites evenly convex; metatibia similar in form to that of male. Collecting data. A coastal marsh species taken from grass and deciduous leaf litters at margins of salt marshes, and from under beach drift and at UV light in these areas (Chandler 1997; Majka & Ogden 2006). LeConte (1849) reports this species as being common in " pratis salinis " (salt meadows) in May. Adults have been taken in all months except January, though there are only a few records from autumn and winter in New England.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE8FFE0E1D2FC5DE121F806.taxon	distribution	Distribution (Fig. 36 A). This is a species characteristic of the Atlantic coastal salt marshes, and is found from Nova Scotia to Georgia.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE8FFE0E1D2FC5DE121F806.taxon	discussion	Comments. Aubé (1833: 27) initially compared his new species with B. haematica Reichenbach of Europe and provided a habitus figure that resembles B. floridana (Brendel), which is quite different from the interpretation of this species that was established by LeConte (1849). To confuse matters, Aubé (1844) had also placed B. abdominalis as a junior synonym of B. dentata (Say) — a third species with a very different male tergal form. A study of the types is necessary to resolve this problem, especially since the number of specimens was never stated and the type series could perhaps hold two species or more. The collection at Paris has been searched by Giorgio Sabella and Mrs. Azadeh Taghavian for the types of this species, which should be in the Dejean Collection (MNHN) according to Aubé (1833). Dejean (1837) indeed placed this name on the list of species in his collection, but the types could not be found after a very thorough search in Paris, nor are any present in the LeConte Collection (MCZC). Current usage is maintained for this species until the type (s) may be found. This is the only species with the first three tergites strongly modified and equally visible; these three tergites all strongly laterally lobed (Fig. 6 A).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFECFFECE1D2FF14E796FF6A.taxon	description	(Figs 7, 37 A)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFECFFECE1D2FF14E796FF6A.taxon	materials_examined	Material examined, 121 specimens. USA: Arkansas: Benton Co.: VII- 18 - 1942, M. W. Sanderson (FMNH, 1). Rogers, VII- 3 - 1942, M. W. Sanderson, at light (FMNH, 4). Clark Co.: 4 mi E Arkadelphia on Hwy. 7, VII- 24 - 1985, D. Bowles, UV light (LSAM, 1). Crawford Co.: Cove Creek, V- 24 - 1986, D. E. Bowles, UV light (LSAM, 1). Mountainburg, VIII- 20 - 1948, F. Werner & W. Nutting, rich creek bottom (UAIC, 1). Garland Co.: FS Camp Clearfork, VI- 22 - 2006, B. Baldwin, UV light trap (BBC, 1); VI- 12 - 2005, B. Baldwin, lights (DSCC, 1). Johnson Co.: Wold Pen on Mulberry River, VII- 9 - 1986, D. Bowles, UV light (LSAM, 1). Madison Co.: 5 mi S Kings River Bridge, Hwy. 74, VI- 2 - 1985, D. Bowles, UV light (LSAM, 2). Newton Co.: Ozark NF, Buck Branch, FSR 1209, 4 mi N Hwy. 123, V- 14 - 1986, J. M. Campbell (CNCI, 1). Polk Co.: S of Board Camp, R 29 WT 35 SE sec. 22, VIII- 17 / 20 - 1985, C. B. & J. E. Barr, MV & BL (LSAM, 4; UCBC, 1). Pulaski Co.: Pinnacle Mountain State Park, Base Trail, V- 11 - 1986, J. M. Campbell, ex flood debris on bank (CNCI, 1). Searcy Co.: 6 mi NW Marshall, VIII- 10 - 1987, E. A. Lisowski, (UV light, LSAM, 2; blacklight, INHS, 4). 1 mi S Leslie, VIII- 10 - 1987, E. Lisowski, blacklight (INHS, 1). Ozark NF FSR 1205, Richland Creek, V- 15 - 1986, J. M. Campbell, splashing stones at edge of creek (CNCI, 2). Stone Co.: 3.3 mi N Allison at bridge, VIII- 9 - 1987, E. A. Lisowski, UV light (LSAM, 1). Washington Co.: IV- 10 - 1940, M. W .. Sanderson (FMNH, 1). VIII- 7 - 1940, along river (INHS, 1). 15 mi S Prairie Grove at Cove Creek, VIII- 6 - 1986, D. Bowles, UV light (LSAM, 1). Fayetteville, VI- 30 - 1942 (FMNH, 1), VIII- 8 - 1942 (FMNH, 1), VII- 24 - 1942 (INHS, 1), VI- 20 - 1942 (INHS, 2), M. W. Sanderson, at light. Yell Co.: Dardanelle, V- 27 - 1977, W. H. Cross, UV trap, at edge of wheat field, (MUIC, 1). District of Columbia: D. C. (CMNH, 1; MCZC, 3). Illinois: Coles Co.: Lake Charleston Dam, VII- 8 - 1988, M. A. Goodrich, blacklight trap (FSCA, 1). Grundy Co.: 4 mi W Coal City at 113 bridge, E. Lisowski, blacklight (INHS, 1). LaSalle Co.: Biley Creek, 2 mi S Oglesby, V- 24 - 1986, UV light (LSAM, 1). Pike Co.: Beebe Creek, 3.4 mi WNW Barry, VII- 17 - 1985, J. Gardner, UV light (LSAM, 1). Union Co.: Hutchens Creek, 3 mi E Wolf Lake, VIII- 24 - 1987, E. A. Lisowski, (UV light, LSAM, 2; blacklight, INHS, 1). 3.3 mi E Wolf Lake, VIII- 24 - 1987, E. Lisowski, blacklight (INHS, 1). Vermilion Co.: Middle Fork of Vermilion River @ unnamed ford, 1.9 mi NNE Collison, VII- 8 - 1987, E. A. Lisowski, UV light trap (LSAM, 6). Wabash Co.: Basil Woods N. Park, VI- 22 / 23 - 1995, M. A. Goodrich, UV light trap (EIUC, 2). Indiana: Warren Co.: VI- 24 - 1902, W. S. Blatchley (PERC, 2). Kansas: Douglas Co.: Breidenthal Res., 2 mi N Baldwin, IX- 2 - 1983, J. Pakaluk, UV light (SEMC, 1). Kentucky: Cumberland Co.: Crocus Creek, VIII- 7 - 1966, T. E. Brooks, on gravel bar (SEMC, 7). Crocus Creek, V- 7 - 1966, T. E. Brooks, on gravel bar (SEMC, 3). Missouri: Barry Co.: Roaring River State Park, VI- 10 - 1977, Riley & Thewke, black light (TAMU, 1). Boone Co.: Columbia, VIII- 18 - 1977, E. G. Riley (DSCC, 1; TAMU, 4). McDonald Co.: Big Sugar Creek, 10 mi E Anderson, VII- 12 / 13 - 1960, E. L. Todd (USNM, 1). Oregon Co.: 2.1 mi N Thayer, VIII. 8.1987, E. Lisowski, blacklight (INHS, 1). Randolph Co.: 1 mi E Moberly, VII- 22 - 1972, E. G. Riley (TAMU, 1). Ohio: Hamilton Co.: Cincinnati, V- 26 (MCZC, 1). Pickaway Co.: Circleville, Scioto River at Bridge Crossing Rt. 22, 39 o 36.30, VIII- 24 - 2008, K. D. Karns, flooding gravel along river bank (KDKC, 1). Oklahoma: Cleveland Co.: Norman, IX- 17 - 1976, W. D. Shepard, light traps (UCBC, 1). Delaware Co.: 5 mi E Kansas, Flint Creek, VI- 2 - 1984, W. D. Shepard (DSCC, 2; UCBC, 1). Latimer Co.: IV- 1989, VIII- 1986, VIII- 1984, K. Stephan (DSCC, 3); VII- 1996, K. Stephan, in blacklight (DSCC, 1); IX- 1990, K. Stephan (LSAM, 1); I- 1988, K. Stephan, in blacklight trap (TAMU, 1); V- 1993, IV- 1994, IX- 1987, III- 1989, V- 1987, V- 1989, VIII- 1985, K. Stephan (TAMU, 7). Le Flore Co.: Muse, VI- 24 - 1939, M. W. Sanderson, at light (FMNH, 1). Mayes Co.: Spavinaw, VI- 18 - 1937, Standish & Kaiser (FMNH, 2). McCurtin Co.: Glover River downstream from St. Rtes. 3 & 7, bridge 1.9 mi NNW Glover, VIII- 13 - 1987, E. A. Lisowski, UV light trap (LSAM, 3). Texas: Gillespie Co.: Lange's Mill, VI- 5 - 1969, Board & Hafernik (TAMU, 1). Real Co.: 6 mi SE Leakey, VII- 1 - 1971, G. H. Nelson, ultraviolet light (DSCC, 1). Uvalde Co.: Nueces River, 18 mi NW Uvalde, 1,100 ', A. Brigham, blacklight trap (FMNH, 1). Victoria Co.: Victoria, IX (USNM, 4). Williamson Co.: Taylor, I- 30 - 1968, J. E. Hafernik (TAMU, 1). Only state on label: “ Tx. ” (MCZC, 3, Lectotype and two Paralectotypes); Collection Belfrage (USNM, 2). West Virginia: Greenbrier Co.: White Sulphur, VIII (CASC, 2).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFECFFECE1D2FF14E796FF6A.taxon	description	Description. BODY: Length 1.84 – 2.04 mm; brown, elytra orange-brown; in comparison to other species the body is slightly more flattened; setae on head and pronotum short, curved and suberect, decumbent over rest of body. Head: surface smooth, shiny, punctures minute. Antennae differing slightly between sexes (see below). Setose area of median vertexal fovea subequal in size to those of lateral foveae. Pronotum: surface smooth, shiny, punctures indistinct. Setose area of median antebasal fovea about two-thirds width of those in lateral antebasal foveae. Elytra: surface lightly microreticulate, punctures faint; discal stria extending to about four-fifths of elytral length. Abdomen: surface smooth, shining, punctures indistinct; basal striae of tergite 1 extending to about half paratergite length, bases separated by about half width of tergite, tergite base with distinct setose brush between striae. MALE: Antennomeres III – VII longer than wide, VIII – X as long as wide, VIII smallest, IX – X progressively larger, VIII about three-fourths width of IX, IX slightly longer than wide, X about as long as wide, twice as wide as IX (Fig. 7 D). Trochanters unmodified, smooth. Metaventrite with scattered setae between bases of metacoxae, lacking setose patch. Tergite 1 modified only at apex, apex with deep U-shaped notch at middle, with a narrow hyaline lamina at middle of notch, lateral angles of notch protruding, apices rounded (Figs 7 A – B); tergite 2 short, about one-third length of tergite 1, with deep, broadly bilobed impression at base, impression with elongate narrow tubercle at middle ventral to tergite 1, fitting into base of tergite 1 notch, apex of tergite 2 angularly projecting into impression at middle, projection with anterior margin smoothly swollen, with V-shaped sulcus immediately posterior to swollen rim, with small granulate mound posterior to this sulcus at tergite apex (Figs 7 A – C); in lateral view tergites 1 – 2 curving confluently. Abdominal ventrites 2 – 6 broadly convex. Metatibiae narrow basally, slightly widening through length to near apex, with dense comb of setae on mesal margin in apical fourth where slightly curved medially (Fig. 7 E). Aedeagus 0.30 mm long, dorsal plate triangular, rounded at apex; parameres with broad preapical constriction, with three thick setae at point of constriction on outer margin, flattened preapical hyaline seta elongate and pointed at apex, inserted at inner margin of constricted area; internal sac with one large evenly curved spine (Fig. 7 F). FEMALE: Antennomeres III – VI longer than wide, VII-VIII as long as wide, VIII smaller, IX – X transverse, VIII two-thirds width of IX. Metaventrite with setae appearing longer and denser toward middle than in male. Tergite 2 with faint semicircular impression arising from base with vague longitudinal median area dividing impression, impression extending to slightly more than half tergite width, all other tergites evenly convex; ventrites broadly convex; metatibiae similar in form to those of male. Collecting data. On gravel bars in streams and in flood debris; at UV light (Chandler 1997). Most specimens with collecting data were taken at ultraviolet light, but a few specimens were taken beneath stones on gravel bars in streams, or from under flood debris. Collected from March to September, with one record from January in Texas.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFECFFECE1D2FF14E796FF6A.taxon	distribution	Distribution (Fig. 37 A). This species is found in the lower Ohio River drainage area starting in West Virginia, and is most common in the lower Mississippi River drainage area starting in Missouri and Illinois. A few specimens were taken from the District of Columbia, which is the only locality from outside the Ohio / Mississippi drainage area.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFECFFECE1D2FF14E796FF6A.taxon	discussion	Comments. Brendel & Wickham (1890: 268) listed B. infinita Casey as a synonym of B. belfragei, with B. infinita being treated as the female of B. belfragei. This is incorrect, and the “ female ” specimen labeled as B. belfragei in the Brendel material at the MCZC is a male of B. infinita, which lacks modifications of the male tergites. Males of B. belfragei are readily recognized by having the disc of tergite 1 simple with a distinct median emargination at the apex, and tergite 2 has a deep transverse basal sulcus. Females have tergite 2 slightly transversely impressed in the middle portion, and the body is more flattened than for the other species of the Midwestern states. Females of all other species have tergite 2 broadly convex, except that females of B. corniventris often have a middle of tergite 2 slightly flattened but is more densely punctate than is found for B. belfragei. LeConte (1880) described B. belfragei from Texas via a brief description in a key, and did not indicate the number of specimens before him. Specimens in the LeConte Collection have the simple locality label “ Tex., ” and those specimens in the LeConte and Horn Collections (MCZC) with this label are treated as syntypes. The male Lectotype is here chosen from the series in the LeConte Collection and bears the labels: // Tex. / ♂ / [red label] Type 6135 / B. Belfragei Lec. / Aug. – Dec. 2004 MCZ Image Database / LECTOTYPE Bryaxis belfragei LeConte 2013 des. DSChandler //. The male and female syntypes in the LeConte Collection and a male from the Horn Collection (all in MCZC) are labeled as Paralectotypes. Brendel & Wickham (1890: 285) listed this species from Louisiana, but the only specimen that we have seen from Louisiana is a single female of B. corniventris (MCZC), and the Brendel & Wickham record may well be based on the misidentification of this old specimen.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE2FFEFE1D2F94CE1B3FCE3.taxon	description	(Figs 8, 36 B)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE2FFEFE1D2F94CE1B3FCE3.taxon	materials_examined	Material examined, 128 specimens. USA: Alabama: Baldwin Co.: Weeks Bay NER Res., 30 25 ' 03 " N, 87 49 ' 50 " W, VI- 20 - 2001, T. L. Schiefer, m. v. lamp in mixed forest near estuary, W. H. Cross Expedition (MEMC, 1). Delaware: New Castle Co.: Liftwood, Wil. (mington), VI- 1 - 1965, Famcd (sic), at light (MHNG, 1). Wilmington, VIII- 26 - 1963, black light (USNM, 1). Sussex Co.: Dewey Beach, VIII- 25 - 1972, L. Knutson (USNM, 1). District of Columbia: “ D. C. ”, X- 20 - 1878, c noll T. Pergande (USNM, 1); “ D. C. ” (CMNH, 11). Wash. (ington), III, Hubbard & Schwarz (USNM, 1); X- 15, Hubbard & Schwarz (USNM, 1). Florida: Alachua Co.: Gainesville, III- 4 - 1967, D. L. Mays, UV light (FMNH, 1). Gulf Co.: Port St. Joe, VII- 23 - 1961, R. C. Graves, under board on beach (FMNH, 1). Highlands Co.: Archbold Biol. Sta., Lake Placid, II- 6 - 1995, M Deyrup, pitfall trap 554 (ABSC, 1); I- 11 - 1975, J. L. Bengtson, at light (FMNH, 1). Sebring, I- 20, C. T. Parsons (MCZC, 1). Indian River Co.: Indian River, Hubbard & Schwarz (USNM, 2). Levy Co.: Seahorse Key, nr. Cedar Key, IV- 1 / 2 - 1988, D. L. Matthews, at light (FSCA, 1). Miami-Dade Co.: Haulover, II- 26, III- 4, III- 1 (MCZC, LeConte Collection, 3); VIII- 20, Hubbard & Schwarz (USNM, 1). Okaloosa Co.: Fort Walton Beach, III- 2 - 1972, E. J. Kiteley (FMNH, 1). Okeechobee Co.: Okeechobee, IV- 7 - 1921, W. S. Blatchley (PERC, 1). Pinellas Co.: Dunedin, IV- 6 - 1922 (MCZC, 2); III- 22 - 1925 (MCZC, 1); II- 15 - 1917, W. S. Blatchley (PERC, 1); I- 4 - 1917, W. S. Blatchley (PERC, 1); XII- 20 - 1916, W. S. Blatchley (PERC, 1); II- 17 - 1926, W. S. Blatchley (PERC, 1). St. Petersburg, III- 27 - 1925 (MCZC, H. C. Fall Collection, 7); IV- 6 - 1923 (MCZC, H. C. Fall Collection, 1). St. Johns Co.: St. Augustine, III- 2 / 7 - 1940 (CASC, 1). Volusia Co.: Edgewater, II- 24 - 1939, C. A. Frost (MCZC, 1). Only state on label: “ Fla. ” (FMNH, 1); “ Fla. ” (MCZC, 3 Paralectotypes; ANSP, 1; CMNH, 5); “ Fla. ” (= St. Johns River settlements; ANSP, Lectotype). “ Florida ”, Dr. Will (MHNG, 1). Georgia: Laurens Co.: Dublin, VI- 1960 (MHNG, 1). Maryland: Anne Arundel Co.: Deale (FMNH, 1). Calvert Co.: Kenwood Beach, 5 mi S Prince Frederick, IX- 17 - 1950, G. H. Nelson, under wash-up log (DSCC, 1). Scientists Cliff, III- 23 - 1980, J. M. Campbell (CNCI, 3). Dorchester Co.: Cambridge, IX- 21 - 1975, E. J. Ford, Jr., light trap (DSCC, 1). Fishing Creek, VIII- 7 - 1956 (987 - B), R. S. Howard (FMNH, 1). Kent Co.: Eastern Neck Island, III- 27 - 1956 (879 - A), R. S. Howard (FMNH, 1). Somerset Co.: Crisfield, VII- 16 - 1932, F. C. Bishop, w mosquito trap (USNM, 1). Only state on label: “ Md. ” (CMNH, 1). New York: Only state on label: “ N. Y. ” (FMNH, 1). North Carolina: Mecklenburg Co.: Charlotte, V- 17 / 24 - 2005, J. F. Cornell, UV trap (LSAM, 2). New Hanover Co.: Marine Research Center, VIII- 15 - 1982, K. R. Ahlstrom, at light trap (NCSU, 1). Pender Co.: New Topsail Beach, VIII- 26 - 1963, P. D. Ashlock (DSCC, 2). South Carolina: Charleston Co.: Charleston, IV- 25 - 1945, R. L. Wenzel, light trap (FMNH, 3). McClellansville, VI- 1 - 1973, T. Daggy (NCSU, 3). Virginia: Hampton City: Fort Monroe, V- 30, Hubbard & Schwarz (USNM, 2). Hampton (FMNH, 1). Hampton, V- 17 - 1944, N. M. Downie (PERC, 1). Lancaster Co.: Weems, IX- 3 / IX- 7 - 2002, C. A. Springer, UV trap (EIUC, 18). Weems, IX- 3 - 2002, C. A. Springer, UV light (DSCC, 1). Norfolk City: Norfolk, VIII- 16 / 18 - 2008, C. A. Springer, UV light (EIUC, 1). Virginia Beach: Sagamore State Park, White Hill Lake Marsh, IV- 25 - 1990, K. A. Buhlmann, pitfall (DSCC, 2). Seashore State Park, XI- 16 - 1989, Va. Nat. Heritage Program, pitfall drift fence site (DSCC, 1); XII- 22 - 1989 (DSCC, 1). Seashore State Park, White Hill Lake marsh, IV- 26 - 1990, K. A. Buhlmann, pitfall (DSCC, 4). York Co.: Cheatham Annex, Naval Supply Center, V- 30 - 1990, R. L. Hoffman (DSCC, 1). Neck of Land, Colonial Nat. Historic Parkway, VIII- 30 - 2000, A. C. Chazal, ultraviolet light trap (DSCC, 1; VMNH, 2).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE2FFEFE1D2F94CE1B3FCE3.taxon	description	Description. BODY: Length 2.12 – 2.24 mm; orange-brown to brown; setae short, curved and suberect on head and pronotum, depressed on rest of body. Head: surface smooth, shining, punctures minute. Setose area of median vertexal fovea as large as those of lateral foveae. Antennomeres VII-X slightly different between sexes. Pronotum: surface smooth, shining, punctures indistinct. Setose area of median antebasal fovea as wide as those of lateral antebasal foveae. Elytra: surface faintly punctate, microreticulation faint to distinct, discal stria extending to about four-fifths of elytral length. Abdomen: tergite 1 with surface lightly punctate; basal striae very short, indistinct in male, barely visible in female, striae separated by about two-fifths tergite width, setal brush between striae longer and distinct in male, shorter and indistinct in female. MALE: Antennae and trochanters unmodified, simple. Antennomeres III – VII longer than wide, VIII barely transverse, IX – X as long as wide, VIII two-thirds as wide as IX, X as long as wide (Fig. 8 D). Metaventrite with triangular patch of dense long setae extending from between metacoxae to mesocoxae. Only tergites 1 – 2 visible in dorsal view (Fig. 8 A), 1 nearly twice as long as 2; 1 with pair of glabrous or slightly setose rounded tubercles near base, tergite apex with two large, widely separated projections, projections bluntly rounded at apex, semicircular emargination between projections, slightly convex flat shelf at middle, shelf obscured by dense elongate yellow setae, area lateral to projections concave and with dense appressed setae; tergite 2 with prominent rounded humps posterior to projections of tergite 1, with broad deep excavation between them, humps obliquely impressed on lateral margins, base of tergite 2 with short broadly bifurcate tubercle ventral to apical shelf of tergite 1 (Figs 8 A, 8 C); tergite 3 with indistinct oblique impressions through length near middle; tergites 4 – 5 convexly rounded; in lateral view outline of tergites 1 – 2 confluent (Fig. 8 B). Abdominal ventrites broadly convex. Metatibiae broad near base, evenly wide to near apex, setal brush present in on mesal margin at apical fifth where tibiae slightly curved (Fig. 8 E). Aedeagus 0.34 mm long; with dorsal plate triangular, apex acutely pointed, symmetrical; parameres broad, abruptly constricted near apex, with three long setae on lateral margin at point of constriction, with preapical hyaline seta near mesal margin, seta flattened and relatively narrow; internal sac with three curved spines, two large and one small (Fig. 8 F). FEMALE: Antennomeres III – VI longer than wide, VII as long as wide, VIII – X transverse. Lacking metasternal patch of setae; all abdominal tergites evenly curved; ventrites broadly convex; metatibiae narrow near base, gradually broadening to apex. Collecting data. Few specimens have habitat or collection data. There are two records of individuals being found beneath washed-up logs and boards on beaches (Chandler 1997). The others with collection data were taken through their attraction to ultraviolet light. Specimens have been taken throughout the year in the southern states.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE2FFEFE1D2F94CE1B3FCE3.taxon	distribution	Distribution (Fig. 36 B). Brendel & Wickham (1890: 279) recorded this species “ from the Gulf [of Mexico] to the Ohio river, [sic] ” but nearly all specimens examined have been from marsh areas along the Atlantic coast, though a few specimens have been taken inland from the coast in North Carolina and Florida. There is a single record from New York (“ N. Y. ”), with the range otherwise extending from Delaware to Florida.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE2FFEFE1D2F94CE1B3FCE3.taxon	discussion	Comments. Similar to B. intermedia in having only tergites 1 – 2 visible in dorsal view and those tergites strongly modified with dorsally projecting lobes. The two are easily separated by B. intermedia having tergite 2 as long as 1 and not prominently bilobed at the apex, while B. floridana has tergite 2 slightly more than half the length of 1, and the apex has two prominent lateral lobes. Brendel mentioned both sexes in his description, but only the female lectotype is present at the ANSP, with the male specimen (s) apparently present in the Horn Collection at the MCZC of Harvard University, where Brendel’s non-primary type specimens may be found. Three specimens of B. floridana, two males and one female, are present in the collection of the MCZC. Two have the labels “ Fla. ” like the Lectotype, and all have “ Brend ” indicating their origin from the Brendel collection. These are here labeled as paralectotypes and bear the label “ PARALECTOTYPE Bryaxis floridana Brendel des. DSChandler 2013. ” Based on the figure of B. abdominalis given by Aubé (1833: pl. 82, fig. 2), B. floridana may be the junior synonym of this name. However, until a type of B. abdominalis can be located, this synonymy cannot be confirmed (see Comments section for B. abdominalis).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE1FFEAE1D2FC3CE778FC1A.taxon	description	(Figs 9 – 10, 36 C)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE1FFEAE1D2FC3CE778FC1A.taxon	materials_examined	Material examined, 23 specimens. USA: Delaware: Only state on label: “ Del. ”, IX- 1938, C. Cottam (USNM, 1). District of Columbia: D. C. (ANSP, 1, Lectotype; CMNH, 3; MCZC, 1). Maryland: Montgomery Co.: Plummers Island, V- 11 - 1905, Barber & Schwarz (USNM, 1). Somerset Co.: Marion, VII- 29 - 1978, E. J. Ford, Jr., UV light (DSCC, 1). New York: Westchester Co.: Peekskill, V- 17 - 1890 (USNM, Casey Collection, 1); Peekskill (FMNH, 3). Only state on label: “ N. Y. ” (USNM, holotype B. labyrinthea; SEMC, 3); “ N. Y. ”, V (FMNH, 2). North Carolina: Mecklenburg Co.: Mint Hill, IV- 1 / 7 - 1994, J. Bryan & J. Cornell, UV light (DSCC, 1). Wake Co.: Raleigh, IV- 12 - 1953, D. M. Weisman (NCSU, 1 M). Tennessee: Only state on label: “ Ten. ” (CMNH, 1, Paralectotype). Virginia: Hampton City: Fort Monroe, V- 30, Hubbard & Schwarz (USNM, 1). Spotsylvania Co.: Fredricksburg, VII- 6 - 1891, W. D. Richardson (USNM, 1).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE1FFEAE1D2FC3CE778FC1A.taxon	description	Description. BODY: Length 2.28 – 2.40 mm; orange to orange-brown; setae appressed or nearly so. Head: surface smooth, shining, punctures minute. Setose area of median vertexal fovea one-third larger than lateral foveae. Antennomeres III – VII longer than wide, VIII – IX slightly transverse, X slightly longer than wide, VIIItwo-thirds width of IX (Fig. 10 B). Pronotum: surface smooth, shining, punctures indistinct. Median antebasal fovea about two-thirds width lateral antebasal foveae. Elytra: surface varying from indistinctly punctate and faintly microreticulate to punctures faint on roughened and microreticulate surface, discal stria extending to about fourfifths of elytral length. Abdomen: tergite 1 with surface lightly punctate; males with basal striae of tergite 1 extending about four-fifths length of paratergite (females unknown), basal striae separated by about two-fifths basal width of tergite 1, with long sparse setal brush between striae. MALE: Antennae and trochanters lacking modifications, simple. Metaventrite transversely impressed as broad oval from between metacoxae to near mesocoxae, impression covered with dense short setae. In dorsal view only tergites 1 – 2 visible (Figs 9 A, 10 A), 2 slightly longer; 1 with basal abdominal carinae distinct, extending posteriorly to prominent rounded preapical mounds whose posterior margins protrude over tergite 2, apical margin of 1 slightly concave between these projections, this margin darkly carinate and briefly toothed ventral to apices of projections, carina with apical row of short setae, lateral margin with oblique dense row of setae extending medially to tergite 2, posterolateral portion of projections convexly impressed and densely setose; tergite 2 with paratergites visible, with broadly rounded median ridge originating between projections of tergite 1, tergite broadening apically to broadly rounded lateral humps at apical margin, apical margin between humps gently concave, lateral margins of median ridge with dense long setae directly laterally over deep impressions lateral to ridge, some setae reach lateral margins of tergite, two divergent black tubercles originate at base of tergite 2 ventral to median carinate apex of 1, typically not visible except in posterior view (Fig. 9 C); tergite 3 broadly truncate at apex, lateral portions dorsally impressed and densely setose; tergites 4 – 5 simple, convex; in lateral view tergite 1 strongly projecting dorsally at apex, posterior margin vertical and dropping to relatively distant flat disc of 2 (Fig. 9 B). Abdominal ventrites all broadly rounded. Metatibiae broad near base and evenly wide through length, with thick setal brush on inner margin where metatibiae curves slightly medially at apical sixth (Figs 10 C – D). Aedeagus 0.40 mm long; with dorsal plate triangular, acutely pointed at apex; parameres broad to abrupt preapical constriction, with three thick setae at point of constriction on lateral margin, mesal margin at constriction with broad, flattened, hyaline seta pointed at apex; internal sac with three large slightly curved spines (Figs 9 D, 10 E). FEMALE: Female not examined. Brendel (1866 b: 194) discusses “ the supposed form ” of the female, and states that it is closer to B. dentata, is more convex than B. floridana, and that the frontal groove (fovea?) is smaller and distinct. Females of B. dentata have very short basal discal carinae on tergite 1, while the males of B. intermedia have long and prominent basal discal carinae. This may be a male sexual feature, but until the sexes are associated, speculation is fruitless. Collecting data. The only collecting data indicate that they will fly to ultraviolet light, and that adults are active from April through September.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE1FFEAE1D2FC3CE778FC1A.taxon	distribution	Distribution (Fig. 36 C). This species ranges from New York to North Carolina. The majority of records are from Atlantic coastal marsh areas, but those from central North Carolina and Tennessee indicate that this species may occur (rarely) well inland from the coast.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE1FFEAE1D2FC3CE778FC1A.taxon	discussion	Comments. Brendel (1895: 184) believed that Bryaxis labyrinthea was at most barely a variety of B. intermedia, and in our opinion the male characters are identical. Bryaxis labyrinthea Casey, 1894 is here placed as a junior synonym of Bryaxis intermedia Brendel, 1866. This species is closest to B. floridana in having only tergites 1 – 2 visible in dorsal view and tergite 1 with dorsally projecting lobes. The two are easily separated by B. intermedia having tergite 2 as long as 1 and not prominently bilobed at the apex, while B. floridana has tergite 2 slightly more than half the length of 1, and the apex has two prominent lateral lobes.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE4FFD4E1D2FC74E6ADFEFF.taxon	description	(Figs 11, 37 B)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE4FFD4E1D2FC74E6ADFEFF.taxon	materials_examined	Material examined, 9 specimens. HOLOTYPE male: USA: Maryland: Montgomery Co.: // at light / Plummers Id., 28. V. 14 Md. / R. C. Shannon / [red label] HOLOTYPE Brachygluta shawnee Chandler // (USNM). PARATYPES, 8, all males: USA: Alabama: Jefferson Co.: Vestavia, VII- 18 - 1981, T. King, at light (FMNH, 1). Walker Co.: Jasper, Devils Ladder, V- 27 - 1979, T. King, at light (FMNH, 1). District of Columbia: D. C., ♂, Brend, Horn Coll H 1919 (MCZC, 1); D. C. (CMNH, 1). Georgia: Clarke Co.: Whitehall Forest, 7 - 14 Aug. 1978, R. Turnbow, bl trap (DSCC, 1). Indiana: Knox Co.: 6 - 28 - 1946, Plot 10, S. A. Summerland Coll. / LSAM 0 267298 (LSAM, 1). Ohio: Vinton Co.: 72660 Gambill Hollow Rd., Albany, 2.27 km West of Rt. 356, 6.5 km SE Zaleski, 39 15.218 N, 82 19.641 W, 30 - July to 06 - August- 2011, K. D. Karns / flight intercept trap, north facing slope, upland hardwood beech, oak, buckeye (DSCC, 1). Virginia: Henrico Co.: 4 km upstream from Bottoms Bridge, Chickahominy River, VII- 4 - 1999, I. T. Wilson, UV trap (DSCC, 1).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE4FFD4E1D2FC74E6ADFEFF.taxon	description	Description. BODY: Length 1.88 – 2.10 mm; brown to dark brown, elytra orange to red-brown; setae short, decumbent. Head: surface smooth, shining, punctures indistinct; setose are of median vertexal fovea slightly smaller than those of lateral foveae. Antennomeres III – VI longer than wide, VII about as long as wide, VIII-X transverse; VIII two-thirds width of IX (Fig. 11 C). Pronotum: surface smooth, shining, punctures indistinct. Setose area of median antebasal fovea slightly smaller than those of lateral antebasal foveae. Elytra: surface smooth, shining to opaque where faintly microreticulate on much of elytral disc, discal stria extending to about four-fifths of elytral length. Abdomen: surface with small distinct punctures, punctures denser in apical half; tergite 1 with divergent basal striae extending about half length of paratergite, basal striae separated by about two-fifths basal width of tergite 1, setose brush between bases of striae short. MALE: Antennae and trochanters lacking modifications. Metaventrite with semicircular patch of denser setae between bases of metatrochanters present but indistinct. Usually only tergites 1 – 2 visible in dorsal view (Fig. 11 A), tergite 1 twice length of 2; tergite 1 broadly and shallowly emarginate at apex, with broad hyaline lamina at middle, lamina apex emarginate, setal fringe along tergite apex densest by lamina; tergite 2 with deep narrow impression at base, impression extending to tergite apex in middle third, flanked by prominent rounded ridges, with band of golden setae between bases of ridges and ventral to lamina (Fig. 11 B); in lateral view outline of tergites 1 – 2 confluent. Abdominal ventrites broadly convex. Metatibiae narrow basally, slightly widening through length to apex, with dense comb of setae on mesal margin in apical fifth where slightly curved medially (Fig. 11 D). Aedeagus 0.30 mm long; with dorsal plate elongate, slightly asymmetrical, with short hook at apex; parameres with three thin setae at posterior margin of preapical constriction, hyaline setae small, flattened, widest at middle and gradually tapering to acute point; internal sac with one large curved spine (Fig. 11 E). FEMALE: unknown. The female is expected to have long basal discal striae on tergite 1. Collecting data. The specimens were either taken by light or ultraviolet light (4), or with a flight intercept trap in an upland deciduous forest (1). Collected from May to August.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE4FFD4E1D2FC74E6ADFEFF.taxon	distribution	Distribution (Fig. 37 B). This uncommon species is widely distributed through the eastern United States from Ohio to Maryland and Georgia; it does not occur in the Atlantic coastal marshes.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE4FFD4E1D2FC74E6ADFEFF.taxon	discussion	Comments. The relatively simple structure of the male abdominal tergites is shared with B. belfragei, with the disc of tergite 1 lacking dorsally projecting lobes, with a median emargination at the apex, and tergite 2 having a deep basal transverse sulcus. These two may be separated by B. shawnee being more densely punctate and having a more opaque body surface; the presence of a broad hyaline lamina at the posterior margin of the emargination of tergite 1; and tergite two having lateral setose lobes. Brachygluta belfragei has sparse punctures on the abdominal tergites with a more glabrous body surface, the hyaline lamina in the emargination of tergite 1 is narrower, and tergite 2 is smoothly convex and glabrous laterally.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFE4FFD4E1D2FC74E6ADFEFF.taxon	etymology	Etymology. The name is that given to an American Indian tribe that was active within the range of this species.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFDAFFD6E1D2FE05E060FDA7.taxon	description	(Figs 12, 36 D)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFDAFFD6E1D2FE05E060FDA7.taxon	materials_examined	Material examined, 28 specimens. USA: District of Columbia: “ D. C. ” (CMNH, 4; AMNH, holotype; MCZC, Horn Collection, 1; MCZC, LeConte Collection, 1); IX- 23 - 1927, Quirsfeld (CNCI, 1); X- 20 - 1878 (USNM, 1). Eastern Branch, I- 5 - 1921, H. S. Barber (USNM, 5); no date (CMNH, 5). Wash. (ington), Hubbard & Schwarz (USNM, 1). Florida: Leon Co.: 1.1 mi S Woodville, I- 22 - 2003, C. W. O'Brien & M. Haseeb, berlese sifted longleaf pine-wiregrass (DSCC, 1). Maryland: Worcester Co.: Snow Hill, IX- 14 - 1976, E. J. Ford, Jr. (DSCC, 1). South Carolina: Charleston Co.: Santee Coastal National Wildlife Refuge, X- 8 - 2006, Paiero & Marshall (DEBU, 2). Virginia: Henrico Co.: Wilson Farm, 3.5 up from Bottoms Bridge, Chickahominy River, VII- 17 - 1999, I. T. Wilson, UV trap (DSCC, 1). Norfolk City: Norfolk, IX- 7 / 14 - 2008, C. A. Springer, UV light (EIUC, 1). Spotsylvania Co.: Fredricksburg, V- 2 - 1900, W. D. Richardson (LSAM, 1). York Co.: Neck of Land, Colonial Nat. Historic Parkway, VIII- 30 - 2000, A. C. Chazal, ultraviolet light trap (VMNH, 1).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFDAFFD6E1D2FE05E060FDA7.taxon	description	Description. BODY: Length 2.32 – 2.44 mm; orange-brown to brown with elytra orange-brown; setae short, decumbent. Head: surface smooth, shining, punctures indistinct. Setose area of middle vertexal fovea smaller than those of lateral vertexal foveae. Antennae elongate when compared with other species, antennomeres III – VII clearly longer than wide, VIII – IX about as wide as long, X longer than wide, VIII about two-thirds width of IX (Fig. 12 D). Pronotum: surface smooth, shining, scattered punctures minute but visible. Setose are of median antebasal fovea about two-thirds width of those of lateral antebasal foveae. Elytra: Surface lightly punctate, microreticulation faint, discal stria extending to about four-fifths of elytral length. Abdomen: tergite 1 with surface lightly microreticulate, punctures distinct, densest in apical half; basal striae of tergite 1 distinct, two-thirds to nearly as long as paratergites, striae separated by about two-fifths tergite width, with short sparse setal brush between bases of striae. MALE: Antennae and trochanters lacking modifications. Metaventrite with setae denser in middle portion from between metacoxae to near mesocoxae. Only tergites 1 – 2 clearly seen in dorsal view (Fig. 12 B), portion of 2 extending beyond apex of 1 less than half length of 1; tergite 1 with disc broadly convex, apex narrowly and deeply emarginate between two acutely pointed spines, emargination about as deep as distance between apices of spines, margins lateral to spines curving ventrally, apical margin with blunt tooth lateral to each apical spine; tergite 2 visible dorsally as truncate extension from beneath spines, with two close, acute, erect spines projecting dorsally from area slightly medial to spines of tergite 1, apex of tergite 2 at middle with portions ventral to 1 impressed to form large space, lateral margins curving ventrally and shallowly concave, apical portion with broad sulcus longitudinally divided by raised lamina, lamina convexly rounded in lateral view; tergite 3 with apex broadly rounded, protruding but covered in dorsal view by tergite 2; tergites 4 – 5 shallowly convex (Fig. 12 C); in lateral view tergite 1 protruding over II, distance between them close (Fig. 12 A). Abdominal ventrites broadly convex. Metatibiae narrow at base, gradually widening near base and then width even to apex, with inner brush in apical fourth, metatibiae only slightly curved near apex (Fig. 12 E). Aedeagus 0.38 mm long, dorsal plate triangular, with apex acute; parameres with preapical constriction present, three thickened setae on lateral margin at point of constriction, broader preapical hyaline seta on mesal margin just distal to constriction, seta pointed at apex; internal sac with three thick spines, one spine nearly straight (Fig. 12 F). FEMALE: Not examined. Brendel (1866 b: 194) states that “ the supposed form of the ♀ favors most B. dentata. ” The basal striae of abdominal segment I should be fairly long, as they are in the male. Collecting data. This species has been taken sifting longleaf pine and wiregrass litter in Florida, and several specimens were collected at ultraviolet lights. It has been sparingly collected throughout the seasons, with the majority of records being from late summer into autumn.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFDAFFD6E1D2FE05E060FDA7.taxon	distribution	Distribution (Fig. 36 D). An Atlantic Coast species that occurs from Maryland to Florida, and is found in estuarine areas.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFDAFFD6E1D2FE05E060FDA7.taxon	discussion	Comments. The general pattern of male modifications is shared with B. belfragei and B. shawnee: tergites 1 - 2 are visible in dorsal view, tergite 1 is convex and has an apical emargination, tergite 2 has a deep transverse emargination, and the basal discal carinae of tergite 1 are half the paratergite length or longer. This species is readily separated by the apical emargination of tergite 1 being deep and bordered by acute prominent spines. Chandler (1994) noted that this species was based on a single specimen, the holotype, and that the lectotype designation for this specimen by Moxey (1962) was incorrect.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD8FFD3E1D2FB97E136FEA2.taxon	description	(Figs 13 – 14, 38 A)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD8FFD3E1D2FB97E136FEA2.taxon	materials_examined	Material examined, 697 specimens. CANADA: Only country on label: “ Canada ” (FMNH, 2; most likely from Ontario). USA: Only continent on label: “ North America ” (MNHN, 9 F as B. illinoiensis); “ America bor. ” (MNHN, 3 F as B. illinoiensis). Alabama: Blount Co.: Blount Springs, VI- 30 - 1984, T. King, at light (FMNH, 1). Jefferson Co.: Vestavia, Hoover, V- 23 - 1983, T. King, UV trap (DSCC, 1). Lauderdale Co.: Florence, IV- 12 - 1955, W. Snow, light (FMNH, 1); II- 28 - 1955, W. Snow, light (FMNH, 1). Mobile Co.: Mobile (ZMUM, Lectotype and two Paralectotypes). Shelby Co.: Vandiver, V- 28 - 1974, T. King, at light (FMNH, 1). Walker Co.: Jasper, Devils Ladder, IV- 30 - 1978, T. King, at light (FMNH, 1). Arkansas: Marion Co.: 14 mi SE Yellville, Buffalo R. N. Park, VI- 1 - 1984, W. D. Shepard, blacklight trap (UCBC, 1). Pulaski Co.: Little Rock, VI- 2 - 2001, B. Baldwin, UV light trap (EIUC, 1). Washington Co.: Fayetteville, X- 22 - 1941, M. W. Sanderson, light trap (LSAM, 1); VII- 15 - 1940, M. W. Sanderson (INHS, 1); VIII- 1 - 1940, M. W. Sanderson (INHS, 1). District of Columbia: “ D. C. ” (CMNH, 14; FMNH, 1; MHNG, 2); “ D. C. ”, Bailey (AMNH, 1); “ D. C. ”, Ulke (MCZC, 4; USNM, 2; MCZC, H. C. Fall Collection, 3; MCZC, 9); “ D. C. ”, Brend. (MCZC, 1, Horn Collection). Wash. (ington), IV- 30 - 1891 (LSAM, 1). Illinois: Champaign Co.: 2.7 mi N Penfield, Middle Fork Vermilion River, VII- 29 - 1987, E. A. Lisowski, UV light (UV light, LSAM, 2; blacklight, INHS, 4). Urbana, VIII- 1946, Hasbrouck & Rapp (INHS, 1). Coles Co.: VII- 27 - 1970, A. Decker, UV trap, flood plain (DSCC, 2; EIUC, 1). Charleston, VI- 13 / 14 - 2013, M. A. Goodrich, UV light (EIUC, 1); VI- 14 - 1964, G. T. Riegel, ex. light trap (EIUC, 1); VII- 20 - 1964, G. T. Riegel, ex. light trap (EIUC, 1). Lakeview Park, M. A. Goodrich, UV light trap (EIUC, 2). Cook Co.: Evanston (FMNH, 1). Evanston Beach, V- 18 - 1956, R. C. Graves, under board (FMNH, 3). Hardin Co.: Rosiclaire, VIII- 22 - 1944, M. W. S. & H. T. L. (INHS, 2). Iroquois Co.: 4 mi N Watseka, pond edge, V- 31 - 1986, E. Lisowski, blacklight (( INHS, 1). Jackson Co.: Fountain Bluff, V- 15 - 1932, Ross & Mohr (FMNH, 1); III- 18 - 1943, H. H. Ross & M. Sanderson, debris on hillside (FMNH, 1). Giant City State Parker, III- 6 - 1945, Ross & Sanderson (INHS, 1). Grand Tower, sand bank Mississippi River, IV- 28 - 1985, D. Hildebrandt (MEMC, 2). Grand Tower, III- 18 - 1943, H. W. Ross & M. Sanderson, stump, mouse nest (FMNH, 1). Jersey Co.: Grafton, along Illinois River shore, X- 1 - 1943 Ross & Sanderson (INHS, 4). Jo Davies Co.: 6 mi S Stockton, VII- 29 / VII- 3 - 1972, Reaves & Hollander, at light (FMNH, 1). Kankahee Co.: E of Bourbonnais, VII- 2 - 1955, H. S. Dybas (FMNH, 1). Lake Co.: Fort Sheridan, VI- 28 - 1911, Liljeblad (ANSP, 1). Livingston Co.: Cornell, 2 mi W, VIII- 12 - 1970 (FMNH, 2), VIII- 13 - 1970 (FMNH, 3), J. Wagner, at light. 2 mi W Cornell, " Serendipity Farm ", VII- 2 / 3 - 1976 (DSCC, 1); VI- 12 - 1981 (FMNH, 45), VII- 23 - 1980 (FMNH, 7), VII- 9 - 1983 (FMNH, 15), VII- 10 - 1967 (FMNH, 7), VI- 27 - 1971 (FMNH, 8), J. Wagner, at light. 2 mi W Cornell, nr. Pond at Vermilion River, VI- 12 - 1981, J. Wagner, blacklight trap (FMNH, 46). Cornell, 2 mi W at Wagner Farm, VII- 27 - 1987, J. Wagner, black light trap (FMNH, 18). Cornell, 2.5 mi W, VII- 4 - 1982 (FMNH, 7), VIII- 5 - 1984 (FMNH, 6), J. A. Wagner, black light trap. 5 mi W Cornell, Wagner Farm, VII- 26 - 1964, J. Wagner, light trap (FMNH, 5). Macoupin Co.: 2 mi NW Hetick, BSA Camp Bunn, VII- 8 / 13 - 2001, J. Wagner, blacklight trap (FMNH, 17). Mason Co.: Havana, VIII- 18 - 1907 (USNM, 3). Havana, III- 16 - 1987, Mills & Ross (INHS, 1); X- 1 - 1943, Ross & Sanderson, on soil at base of sedges etc., rivers edge (INHS, 1); XI- 9 - 1943, Ross & Sanderson, round cover in forest (INHS, 3). Havana, Illinois River, X- 1 - 1943, (INHS, 1); X- 4 - 1943 (INHS, 1), Ross & Sanderson, on soil at base of sedges etc., rivers edge. Havana, Spoon River, V- 2 - 1912, drift (INHS, 2). Sand Ridge St. (ate) For. (est), VIII- 6 - 1988, P. Skelley, blacklight trap (FSCA, 1). W side of Decatur, VIII- 10 - 1987, P. Skelley, at light (FSCA, 1). McLean Co.: Morraine View State Park, V- 27 - 1977, R. W. Lundgren, light (RWLC, 1). 3.8 mi E Hudson, on river, VI- 10 - 1985, Gardne & Ulasah, blacklight (( INHS, 1). Pike Co.: Beebe Creek, 3.4 mi WNW Barry, VII- 17 - 1985, J. Gardner & J. Hofmann, UV light (LSAM, 8). Florence, Illinois River pond edge, VIII- 26 - 1943, Ross & Sanderson (INHS, 5). Pope Co.: Dixon Springs, III- 17 - 1947, Ross & Sanderson (INHS, 1). Dixon Springs, VIII- 22 - 1944, M. W. S. & H. T. L., edge of stream (INHS, 1). Golconda, VIII- 22 - 1944, M. W. S. & H. T. L. (INHS, 17). Putnam Co.: Putnam, IX- 26 - 1933, Frison & Mohr (FMNH, 1). Randolph Co.: Ruma, I- 24 - 1947, Burks et al., debris in woods (INHS, 1). St. Clair Co.: Kahokia, V- 24 - 1922 (MCZC, 12); IV- 12 - 1902, G. W. Bock (DSCC, 1); IV- 15 - 1914, G. W. Bock (DSCC, 1); V- 20 - 1896 (FMNH, 1). Cahokia, X- 31 - 1899, G. W. Bock (CASC, 1); V- 24 - 1894, G. W. Bock (CASC, 1). Union Co.: IV- 4 - 1938, W. F. Turner (USNM, 1). Hutchens Creek, 3 mi E Wolf Lake, VIII- 24 - 1977, E. Lisowski, blacklight (INHS, 1). Vermilion Co.: Middle Fork of Vermilion River @ unnamed ford, 1.9 mi NNE Collison, VII- 8 - 1987, E. A. Lisowski, UV light trap (LSAM, 28); Middle Fork of Vermilion River, 2.6 mi E Oakwood on Rt. 150, VI- 11 - 1973, J. D. Unzicker, UV light (LSAM, 9). 3.8 mi N Catlin at 150 bridge, VI- 28 - 1987, E. Lisowski, blacklight (INHS, 11). Wabash Co.: Basil Woods N. Park, VI- 13 / 14 - 1996, M. A. Goodrich, UV light trap (EIUC, 1); VI- 22 / 23 - 1995, M. A. Goodrich, UV light trap (DSCC, 1). White Co.: Grayville, VI- 13 - 1948, Frison & Sanderson (INHS, 2). Whiteside Co.: Prophetstown, VII- 24 - 1947, Burks & Sanderson (INHS, 2). Only state on label: “ Ill. ” (MCZC, 2; AMNH, 2; FMNH, 1); “ N. Ill. ”, V- 17 - 1889 (MCZC, 1). “ So. Ill. ”, H. Soltau (LSAM, 3). “ Illinois ” (MNHN, 1 M, 1 F “ cotypes ” of B. illinoiensis Brendel). Indiana: Clark Co.: V- 6 - 1909, W. S. Blatchley (PERC, 12). Crawford Co.: Grantsburg, V- 15 - 1965, D. Eckert, blacklight trap (FMNH, 1). Floyd Co.: New Albany, VII- 29 - 1966 (FSCA, 1); VI- 6 - 1966, C. E. White (FSCA, 1). Lake Co.: V- 5 - 1907, W. S. Blatchley (PERC, 1). Marion Co.: Indianapolis, V- 22 - 1965, C. E. White (FSCA, 1); VII- 20 - 1968, C. E. White (FSCA, 1). Camp Belzer, BSA, Indianapolis, VII- 26 - 1966, C. E. White (FMNH, 1). Marshall Co.; Tippecanoe, X- 17 - 1971 (FMNH, 1), VIII- 11 - 1974 (FMNH, 1), V- 20 - 1976 (FMNH, 5), VII- 23 - 1976 (FMNH, 2), N. M. Downie. Monroe Co.: Bloomington, IV- 28 / 30 - 1985, F. N. Young, blacklight trap (FSCA, 1); VIII- 9 / 10 - 1984, F. N. Young, blacklight trap (FSCA, 1); VII- 18 - 1991, F. N. Young, blacklight trap (FSCA, 1); IX- 10 - 1987, F. N. Young, blacklight trap (FSCA, 1). Orange Co.: West Baden, VII- 7 - 1962, T. S. Acker (CASC, 1). Perry Co.: Cannelton, V- 16 - 1908 (FMNH, 1); V- 16 - 1908, decaying wood on sandy soil (PERC, 20). Porter Co.: Dunes State Park, V- 9 - 1939, H. S. Dybas (FMNH, 2). Pine, V- 5 - 1907, A. B. Wolcott (FMNH, 2). Tremont, V- 9 - 1939, H. S. Dybas (FMNH, 1). Posey Co.: Hovey Lake, IV- 23 - 1965 (FMNH, 1), VIII- 14 - 1965, C. E. White, blacklight trap (AMNH, 4); IV- 29 - 1965, C. E. White, blacklight trap (FMNH, 1); V- 5 - 1965, C. E. White, blacklight trap (CNCI, 6); IV- 29 - 1965, C. E. White, blacklight trap (AMNH, 2); VII- 16 - 1965, C. E. White, blacklight trap (FSCA, 1); VII- 21 - 1965 (FMNH, 1). Putnam Co.: III- 20 - 1910, W. S. Blatchley (PERC, 2); X- 10 - 1914, W. S. Blatchley (PERC, 1). Spencer Co.: Pigeon Creek, VI- 13 - 1983, A. V. Provonsha (PERC, 1). Tippecanoe Co.: Lafayette, VII- 9 - 1983 (FMNH, 1), VII- 25 - 1983 (FMNH, 2), VI- 19 - 1984 (PERC, 1), VII- 11 - 1984 (PERC, 1), VII- 15 - 1985 (PERC, 1), VII- 30 - 1985 (PERC, 1), N. M. Downie, UV light. Vanderburg Co.: Evansville, VI- 19 - 1943, H. S. Dybas (FMNH, 1). Iowa: Only state on label: “ Ioa. ” (CMNH, 3; FMNH, 1; MCZC, 1; MCZC, LeConte Collection, 1; USNM, 3). Kansas: Douglas Co.: V- 31 - 1936, L. Lipovsky (FNNH, 5). Lawrence, V- 31 - 1936, M. S. Sanderson, mud flats along river (FMNH, 13). Lawrence, VII- 8 - 1933, M. W. Sanderson (INHS, 1). Lawrence, along Kaw River, VI- 7 - 1936, L. J. Lipovsky (INHS, 4). Riley Co.: VII- 18 - 1968, R. L. Bertwell, white light trap (UAIC, 1). Only state on label: “ Ks. ” (AMNH, 1). Kentucky: Crittenden Co.: 13 mi NW Marion at Tolu, Ohio River, VIII- 3 - 1977, L. & N. Herman, # 1420 (AMNH, 3). Edmonson Co.: Mammoth Cave Hollow, Mammoth Cave National Park, XI- 25 - 1960, D. Reichle, leaf mold (FMNH, 4). Mammoth Cave National Park, Cabin Woods, XI- 22 - 1969, W. Suter, berlese log (FMNH, 5). Hart Co.: 4.8 mi N Horse Cave, Green River, V- 23 - 1987, E. A. Lisowski, UV light (LSAM, 2); V- 23 - 1987, E. Lisowski, blacklight (INHS, 3). Horse Cave, VI- 11 - 1934, H. E. McClure, flying 6: 30 PM (FMNH, 1). Only state on label: “ Ky ” (FMNH, 2). Louisiana: Only state on label: “ La. ” (MCZC, 1 F). Maryland: Garrett Co.: Deer Park, VII- 4, Hubbard & Schwarz (USNM, 1). Montgomery Co.: Plummers Island, VIII- 24 - 1912, H. S. Barber (LSAM, 1); V- 11 - 1905, Hubbard & Schwarz (USNM, 12). Michigan: Berrien Co.: Harbert Dunes, VIII, A. W. Andrews (FMNH, 1). Mississippi: Oktibbeha Co.: Craig Springs, X- 30 - 1981, W. H. Cross, interception trap — plain (MUIC, 1); Starksville, V- 7 - 1958, D. C. Cross (MEMC, 1). Missouri: Marion Co.: West Quincy, VIII- 16 - 1912 (FMNH, 1). St. Louis Co.: Berkeley, VI- 1965 (FMNH, 1), I- 1965 (FMNH, 1). St. Louis, V- 28 - 1892, F. Knab (USNM, 4); IX- 3 -, H. Soltau (USNM, 3); V, Bock (CASC, 2); IX- 3 - 1891 (USNM, 15); no date (FMNH, 2). Taney Co.: Hollister, VI- 14 - 1938, Mrs. Vitae Kite (INHS, 4). Only state on label: “ Mo., ” Brend. (MCZC, Horn Collection, 1). Nebraska: Douglas Co.: Omaha, VII- 24 - 1999, C. A. Springer, UV light (EIUC, 1). New Jersey: Cape May Co.: Anglesea, V- 5, Wenzel (ANSP, 1). Ohio: Hamilton Co.: Cincinnati, V- 26 - 1906 (CNCI, 4; CASC, 4); VI- 1 - 1906 (MCZC, 13); V- 26 - 1922 (MCZC, 4); VIII- 6 - 1906 (CASC, 2); V- 24 - 1916 (PERC, 1). Huron Co.: Ridge Road sand quarry, North Fairfield, VII- 3 - 1974, H. J. Lee & T. C. Sheets (DSCC, 1). Only state on label: “ Ohio ” (MCZC, 1); Brend. (MCZC, Horn Collection, 1). Oklahoma: Latimer Co.: 5 mi W Red Oak, VIII- 1985, K. H. Stephan (LSAM, 1). Latimer Co., VIII- 1988, K. Stephan (DSCC, 1); VII- 1985, K. Stephan, in blacklight trap (DSCC, 1); VIII- 1990, K. Stephan (TAMU, 1); V- 1986, K. Stephan, in blacklight trap (TAMU, 1). South Carolina: Aiken Co.: 1.3 mi N Jct. I- 20 & US- 1, XII- 13 / 31 - 1988, P. Skelley, pig dung and malt pitfall in sandhill scrub (FSCA, 1). Tennessee: Hamilton Co.: Falling Water Cave, IX- 13 - 1949, Jones & Valentine (FMNH, 1). Lake Co.: Reelfoot Lake, VI- 25 - 1966, F. J. Moore, UV light (DSCC, 2). Trousdale Co.: Harksville, II- 3 - 1909, S. E. Crumb, hibernating in leaves (USNM, 1). Only state on label: “ Ten. ” (AMNH, 1; ANSP, 4; CMNH, 5; FMNH, 1; MCZC, H. C. Fall Collection, 2; MCZC, LeConte Coll., 7; USNM, 6); Brend. (MCZC, Horn Collection, 2). Tennessee (MHNG, 1). Virginia: Spotsylvania Co.: Fredricksburg, V- 17 - 1890, W. D. Richardson (USNM, 1). Wisconsin: Grant Co.: Boscobel, VII- 3 - 1953, R. Shenefelt, light (DSCC, 3); VII- 1 - 1952 (FMNH, 4).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD8FFD3E1D2FB97E136FEA2.taxon	description	Description. BODY: Length 1.96 – 2.16 mm. Darkest specimens with elytra orange-brown and rest of body dark brown, head and pronotum variable in color, with lightest specimens having the head and pronotum light brown, head usually darker than pronotum; setae on head and pronotum short, curved and suberect, decumbent over rest of body. Head: surface smooth, shining, median vertexal fovea subequal in size to lateral vertexal foveae. Pronotum: surface smooth, shining, punctures indistinct; setose area of median antebasal foveae about two-thirds width that of lateral antebasal foveae. Elytra: surface lightly microreticulate, punctures indistinct; discal stria extending to about four-fifths of elytral length. Abdomen: surface of tergal disc smooth, shining, punctures small but distinct, separated by about one puncture diameter on tergite 1, denser on following tergites. MALE: Antennae unmodified, antennomeres III – V longer than wide, V slightly swollen, VI – VII as long as wide, VIII smallest, wider than long, IX – X slightly wider than long, X nearly twice as wide as IX. (Fig. 13 D) Tergites 1 – II strongly modified; tergite 1 with apex developed as two widely separated, prominent and acute points, setae and punctures densest on points and along apex of emargination between points (Figs 13 A, 14 A). Tergite 2 with deep, glabrous, rounded-triangular impression in apical half, with two small adjacent circular setose areas at posterior margin of impression, two adjacent elongate oval setose pits on anterior wall of impression, prominent elongate tubercle arises from anterior portion of tergite between protruding subtriangular lateral plates at base of tergite, tubercle extending dorsally to between projections of tergite 1, with circular setose area at apex; swollen oval granular area present forming lateral portions of tergite 2 (Figs 13 C, 14 B); basal striae of tergite 1 distinct, striae extending nearly to half length of paratergites, with dense brush of setae between striae that is half as long as striae. Tergites 3 – 5 convex. In lateral view projections of tergite 1 acute, protruding posteriorly well-above tergite 2 (Figs 13 B, 14 C). Abdominal ventrites slightly flattened to broadly convex at middle. Trochanters lacking ventral modifications, smoothly convex. Metatibiae narrow basally, slightly widening through length to apex, with dense comb of setae on mesal margin in apical fifth where slightly curved medially (Fig. 13 E). Aedeagus 0.35 mm long; with dorsal plate broadly and indistinctly trapezoidal; parameres with three aciculate setae on angulation defining posterior margin of preapical constriction, hyaline setae at center of constricted area, elongate and aciculate; internal sac with large curved spine, with transverse complex bearing multiple short spines (Figs 13 F, 14 D). FEMALE: Antennomeres similar in proportions to those of male but shorter, V slender, IX more distinctly transverse. Tergite 1 convex with basal striae short, at most one-quarter paratergite length, with scattered setae between striae, lacking modifications; tergite 2 often lightly flattened in medial third; abdominal ventrites broadly convex. Metatibia similar to that of male. Collecting data. Under driftwood on mud banks of the Ohio River (Blatchley 1910); under boards; at UV light (Chandler 1997). Many specimens with collecting data were taken at ultraviolet light or by UV light traps. Most specimens with habitat data were taken on mud flats or from around the bases of riparian plants, but a few were found under debris in forests. Adults are active throughout the year in the Southern and Midwestern states.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD8FFD3E1D2FB97E136FEA2.taxon	distribution	Distribution (Fig. 38 A). This species has been taken most commonly in the Midwestern states, but is widespread through the eastern United States. It has been collected only occasionally in Atlantic coastal areas.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD8FFD3E1D2FB97E136FEA2.taxon	discussion	Comments. Males are easily recognized by the widely-separated prominent tubercles at the apex of tergite 1, combined with the stalked tubercle bearing a setose apex that arises from the base of tergite 2. The females often have the median portion of tergite 2 slightly flattened and the disc distinctly punctate, which can be used to separate most female specimens from other species. The only other common species from the Ohio and Mississippi drainages taken at lights is B. belfragei, whose body is more flattened, more sparsely and indistinctly punctate, and the basal half of tergite 2 of the female has a weakly impressed area. The complex multidentate tooth of the internal sac is similar to that of B. infinita, which is otherwise quite different in lacking obvious male characters of the tergites, and has ventrite 3 modified as the only Nearctic species to have modifications in this area. Chandler (1994) noted that the description of B. illinoiensis was based on a single specimen, the holotype, and that the lectotype designation for this specimen by Moxey (1962) was incorrect. Brendel (1866 a: 34) and Brendel & Wickham (1890: 281) erroneously stated that the male mesotrochanters of B. illinoiensis have a spine, but the trochanters are smooth ventrally.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFDDFFDEE1D2FD3EE075FD4F.taxon	description	(Figs 15, 39 A)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFDDFFDEE1D2FD3EE075FD4F.taxon	materials_examined	Material examined, 128 specimens. USA: Only continent on label: “ Amerique ” (MNHN, 1 M). District of Columbia: “ D. C. ” (USNM, 4; MCZC, 6; MCZC, LeConte Collection, 3; SEMC, 2; ANSP, 3; DSCC, 1; CMNH, 11; FMNH, 1); Brend. (MCZC, Horn Collection, 1). Wash. (ington), VI- 20, Hubbard & Schwarz (USNM, 1); M. L. Linell (USNM, 1). Florida: Brevard Co.: Capron, IV- 15 (MCZC, LeConte Collection, Neotype). Titusville, III- 21 / 22 - 1939, F. E. Lutz (AMNH, 6). Duval Co.: Pablo beach, IV- 8, H. W. Wenzel (FMNH, 1). St. Lucie Co.: Capron, IV- 22, Hubbard & Schwarz (USNM, 1). Volusia Co.: Enterprise, V- 19, Hubbard & Schwarz (USNM, 1). Ormond, III- 17 - 1915 (AMNH, 1); III- 19 - 1915 (AMNH, 1); [no date] (AMNH, 3). Only state on label: “ Fla ” (INHS, 4). Georgia: Laurens Co.: Dublin, VI- 1960, light trap (MHNG, 2). Richmond Co.: Brigan’s (MNHN, 1 M, 1 F). Illinois: Only state on label: “ Illinois ”, V- 13 - 1889 (INHS, 1). Maryland: Baltimore Co.: Baltimore, VII- 28, F. E. Blaisdell (CASC, 1); VI- 10, F. E. Blaisdell (CASC, 1); VI- 5 - 1966, E. J. Ford, light trap (EJFC, 1). Calvert Co.: Kenwood Beach, 5 mi S Prince Frederick, IX- 17 - 1950, G. H. Nelson, under wash-up log (DSCC, 1). Plum Point, V- 28 - 1922 (LSAM, 1). Somerset Co.: Somerset, VI- 5 - 1971, E. J. Ford, Jr. (DSCC 1). Marion, VII- 13 - 1978 (DSCC, 1). Crisfield, VIII- 8 - 1956 (989 I), R. D. Howard (FMNH, 1); VIII- 4 - 1932 (LSAM, 1). St. Mary's Co.: Piney Point, Hubbard & Schwarz (USNM, 1); VIII- 26 - 1946, R. H. Beamer (SEMC, 14). Only state on label: “ Md. ” (CMNH, 3); Brend. (MCZC, Horn Collection, 2). New Hampshire: Strafford Co.: Farmington, VIII- 18 - 1896 (MCZC, H. C. Fall Collection, 1). New Jersey: Ocean Co.: Long Beach, V- 14 - 1961, E. J. Ford, Jr. (DSCC 1). New York: Nassau Co.: Long Beach, L. (ong) I. (sland), VI- 7 - 1925, Quirsfeld (CNCI, 1); W. Beutenmuller (MCZC, 1). Queens Co.: Rock (away) Beach, VI- 11 - 1932, L. L. Buchanan (LSAM, 1). Richmond Co.: S. I. (Staten Island) (FMNH, 2). Only state on label: “ N. Y. ” (CMNH, 1); Brend. (MCZC, Horn Collection, 1). North Carolina: Macon Co.: Franklin, V- 8 - 1957, J. R. Vockeroth (CNCI, 1). Mecklenburg Co.: Charlotte, V- 11 / 17 - 2005, J. F. Cornell, UV light trap (LSAM, 1). Wake Co.: Raleigh, IV- 12 - 1953, D. M. Weisman (NCSU, 1); XII- 5 - 1902, F. Sherman, Jr. (NCSU, 1). Ohio: Only state on label: “ Ohio ” (CMNH, 2). Pennsylvania: Philadelphia Co.: Frankford, V- 20, G. M. Greene (LSAM, 1). Only state on label: “ Pen. ” (MCZC, LeConte Collection, 2). South Carolina: Aiken Co.: 1.3 mi N Jct. I- 20 + US- 1, XII- 13 / 31 - 1988, P. Skelly, pig dung and malt pitfall in sandhill scrub (FSCA, 1). Charleston Co.: Charleston, 1945, O. L. Cartwright (USNM, 1); IV- 25 - 1945, R. L. Wenzel, light trap (FMNH, 1). McClellansville, VI- 1 - 1973, T. Daggy (NCSU, 1). Texas: Only state on label: “ Tx. ”, through C. V. Riley (USNM, 1). Virginia: Hampton City: Fort Monroe, IV- 19, Hubbard & Schwarz (USNM, 7); V- 29 (USNM, 4). King George Co.: Mathias Point, V- 22 - 1896 (MCZC, H. C. Fall Collection, 1). Spotsylvania Co.: Fredricksburg, IV- 29 - 1893, W. D. Richardson (LSAM, 1); V- 21 - 1891, W. D. Richardson (USNM, 1); V- 19 - 1991 (MCZC, H. C. Fall Collection, 1). Virginia Beach: 1 st Landing State Park, Chicken Turtle Pond, 36.9154, P. A. Bedel & A. C. Chazal, UV trap (VMNH, 1). Virginia Beach, VII- 17, Hubbard & Schwarz (USNM, 1).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFDDFFDEE1D2FD3EE075FD4F.taxon	description	Description. BODY: Length 1.96 – 2.16 mm; orange-brown to elytra orange-brown and rest of body grading to brown; setae on head and pronotum short, curved and suberect, over rest of body setae appressed. Head: surface smooth, shining, punctures indistinct, setose area of median vertexal fovea as wide as to slightly smaller than those of lateral vertexal foveae. Antennomeres (both sexes) III – VI longer than wide, VII slightly longer than wide in male, as long as wide in female, VIII – X slightly wider than long (Fig. 15 D). Pronotum: surface smooth, shining, punctures small, separated by two puncture diameters, setose area of median antebasal fovea about two-thirds width of setose areas of lateral antebasal foveae. Elytra: surface lightly microreticulate, punctures faint. Discal stria extending to about four-fifths of elytral length. Abdomen: tergite 1 with punctures of basal portion of disc sparse, surface shining, punctures becoming denser in apical half and surface appearing more opaque, particularly in area of male tergal projection. Basal striae of tergite 1 distinct, extending at most one-fourth length of paratergite; striae separated at base by about one-half tergal width, with sparse setal brush between striae, longer and more obvious in male. MALE: Antennae not modified. Metaventrite with denser setae in triangular area from between metacoxae to middle. Tergite 1 protruding posteriorly to form median blunt angulation, in dorsal view tergite 1 covering other tergites (Fig. 15 A); tergite 2 essentially vertical, transversely impressed at middle, setae dense at base in middle half; tergite 3 at middle with low oval tubercle densely setose at apex (Fig. 15 B); in lateral view tergite 1 arcuate and protruding beyond tergite 2 (Fig. 15 C). Trochanters not modified. Metatibiae narrow basally, slightly widening through length to apex, with dense comb of setae on mesal margin in apical fifth where slightly curved medially (Fig. 15 E). Aedeagus 0.39 mm long; with dorsal plate triangular, apex bluntly pointed; parameres with three thick setae at posterior margin of preapical constriction, hyaline setae flattened, wide in apical half and then tapering to elongate acute point; internal sac with three large spines, two strongly curved (Fig. 15 F). FEMALE: Metaventrite with denser setae in triangular area from between metacoxae to near middle. Abdomen with all tergites evenly convex; ventrites broadly convex. Metatibiae similar to those of male.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFDDFFDEE1D2FD3EE075FD4F.taxon	discussion	Comments. The short original description by Say is not adequate for placement, and treats two apparent species: a smaller one whose tergum was " convex, simple, blackish, " which indicates that a female was before him since females lack tergal modifications, while another specimen termed “ var. a ” was briefly described as " reddishbrown. " LeConte’s (1851: 85) initial treatment of this species was vague, with Brendel (1865 b) being the first to briefly but clearly place this species in a key. Brendel worked closely with LeConte and thanked him for all of assistance in his first paper (Brendel 1865 a), with this interpretation of the species continued in all subsequent papers by LeConte and Brendel. Friedrich Ernst Melsheimer acquired and preserved many of the types of Thomas Say's species, which were then made available to LeConte as needed for his studies (Hagen, 1884). LeConte (1875) reported that he had the opportunity to compare his specimens with many of those of Say, with these compared specimens residing in LeConte’s collection which is now held in the Museum of Comparative Zoology at Harvard University (Hagen, 1884). In LeConte's series of this species the first specimen is a small dark female that bears the labels // [white circle] {LeConte color code = northern States, eastern states and Canada} / ♀ / B. dentata (Say) Lec. //. This label is in LeConte's handwriting, and so removes this specimen from consideration as a Say type. The second, third, fourth, and eighth specimens are males of B. dentata in its current sense, which is based on a lighter brown species. It is informative that LeConte would place the dark specimen first in his series of B. dentata, with a series of lighter brown specimens following — paralleling the description of B. dentata and its “ var. a ” by Say. There are no specimens that can be placed as B. dentata in the remnants of the Say Collection at Harvard University, and no specimens of an appropriate age are recorded from the collections at The Natural History Museum, London, or are in the Museum d'Histoire naturelle in Paris. The Say types of this species are thought to be destroyed, and resolution of the placement of this species relies on selection of a neotype (see Mawdsley, 1993 for a discussion on selection of Say neotypes). Say did not indicate a collection locality other than it was taken during the Major Long Expedition of 1819 -- 1820 (James, 1823) that started in Pittsburgh, Pennsylvania and proceeded to the eastern edge of the Rocky Mountains, and offered no explanation for the choice of the specific name “ dentata, ” which is an obvious feature of the male abdomen for this species in the sense of LeConte and Brendel. Careful examination of the dark first specimen in LeConte’s series reveals that it is a member of either B. arguta or B. wickhami, whose females cannot be placed to species. Both B. arguta and B. wickhami are species that typically occur in the Ohio River Valley and upper Mississippi River drainage. Brachygluta dentata in its current sense is more of an eastern coastal plain species, however, it historically was found in the Midwest and Southwest based on two specimens with an Ohio label, one with an Illinois label, and one with a Texas label examined during this study. While LeConte’s small dark specimen would be the best choice to serve as a neotype, it would lead to uncertainty in species identity, while the lighter brown and larger species in his series of B. dentata fits the very brief description of “ var. a, ” and is the species concept used in the papers of Brendel and LeConte starting in 1865. Due to the lack of resolution in species placement based on the dark form, LeConte's interpretation of this species is continued here by selection of a neotype from the major portion of his series for the following reasons: a) LeConte had access to the Say types via the Melsheimer Collection; b) there are two separate species present in LeConte's series of B. dentata that fit the limited descriptions of Say; c) the possibility that Say's reddish-brown specimen could be conspecific with B. dentata as currently defined due to a few specimens being labeled as being taken in Ohio and Illinois; and d) the males have tergite 1 with a conspicuous median “ dentate ” projection. A neotype is here selected from the J. L. LeConte Collection at Harvard University for Pselaphus dentata Say, which is the second in LeConte’s series and the first light brown specimen. It bears the labels: // ♂ / Capron 15.4 Fla / 815 / dentata 2 / NEOTYPE Pselaphus dentatus Say des. DSChandler [yellow label] //. The next three specimens have the labels // D. C. / [male symbol] / dentata 3 [4, and 8] //. Collecting data. Few specimens have habitat or collecting data, but most of these were taken at ultraviolet lights, and one was found beneath a washed-up log on a beach and another was found in a dung pit-trap. Most specimens were collected from March to July, but a few collections were taken during the autumn and fall months, usually in Florida.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFDDFFDEE1D2FD3EE075FD4F.taxon	distribution	Distribution (Fig. 39 A). This species is found in states bordering the Atlantic Coast, from New Hampshire to Florida, and occurs in the broader coastal plain as well as near and in the coastal marshes. However there are a few old specimens with state labels from Ohio, Illinois, and Texas that indicate a broader range than is demonstrated by more recented collected material. Leng (1920) records this species from Massachusetts, but no specimens from this state have been seen, though its occurrence there is expected.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD0FFDFE1D2FCC8E11EFE37.taxon	description	(Figs 16 – 17, 39 B)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD0FFDFE1D2FCC8E11EFE37.taxon	materials_examined	Material examined, Holotype male: USA: South Carolina: Charleston Co.: // USA: SC: Charl. Co., Mt. Pleasant, V- 22 - 2007, R. Lareau, UV lt. / [red label] HOLOTYPE Brachygluta lareaui Chandler (FMNH).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD0FFDFE1D2FCC8E11EFE37.taxon	description	Description. BODY: Length 2.60 mm; orange-brown; setae on head and pronotum short, curved and suberect, depressed to appressed over rest of body. Head: surface smooth, shining, punctures indistinct. Setose area of median vertexal fovea slightly smaller than those of lateral vertexal foveae. Antennomere form treated below under male characters. Pronotum: surface opaque due to light microreticulation, punctures indistinct; setose area of median antebasal fovea about two-thirds width of those of lateral antebasal foveae. Elytra: surface lightly microreticulate, punctures indistinct. Discal stria extending to about four-fifths of elytral length. Abdomen: tergite 1 with surface shining, punctures small but distinct. Basal striae of tergite 1 elongate, extending about one-third paratergite length, striae separated at base by about one-half tergal width, setal brush between bases of striae sparse but distinct. MALE: Antennae relatively elongate, antennomere III longer than wide, IV as long as wide, V – VI enlarged, V dorsoventrally flattened, in dorsal view IV about two-thirds width of V, in lateral view IV about three-fourths width of V, VI as wide as IV, roughly cylindrical and longer than wide, VII clearly longer than wide, VIII slightly longer than wide, IX – X elongate, both about half again as long as wide (antennomeres X – XI of right antenna partially fused) (Fig. 17 A). Metaventrite with apical half at middle slightly impressed, setae slightly denser in impression than laterally. Lacking modifications of trochanters, ventral margins smooth. Tergite 1 with apex protruding blunt medial angulation; margin of angulation extending above tergite 2, with row of setae ventrally longest at middle, area of tergite 2 ventral to angulation with setae short and appressed (Figs 16 A – B); in lateral view tergite 1 horizontal, apex deflexed, tergite 2 flat and nearly level, well-separated from 2 (Fig. 16 C). Abdominal ventrites 2 – 6 flattened medially. Metatibiae narrow at base, evenly thickening to middle and then narrowing slightly to apex, apical third of metatibiae straight, with thick setal comb on mesal margin in apical fifth (Fig. 17 B). Aedeagus 0.46 mm long; with dorsal plate narrowly triangular, apex acutely pointed; parameres with three small thin setae at posterior margin of preapical constriction, hyaline setae large, broadly flattened, widest at middle and gradually tapering to acute point; internal sac with two large and one small spines, one large spine strongly curved (Fig. 17 C). FEMALE: Unknown. Collecting data. The holotype was collected at ultraviolet light.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD0FFDFE1D2FCC8E11EFE37.taxon	distribution	Distribution (Fig. 39 B). Only known from South Carolina near the Atlantic coastal salt marshes.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD0FFDFE1D2FCC8E11EFE37.taxon	discussion	Comments. This is a distinctive species in that the male antennae are modified, while the abdominal tergal modifications are typical of the dentata species-group. The antennomeres are all elongate except for swollen antennomere V, which is a pattern not found in any other New World species, where VIII is always transverse or about as long as wide, and IX – X are never so clearly elongate. The bluntly angular apex of tergite 1 is close in form to that of B. dentata, but the medially swollen metatibiae and modified antennae readily separate B. lareaui from this and all other species.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD0FFDFE1D2FCC8E11EFE37.taxon	etymology	Etymology. Named for Richard Lareau, who brought this specimen to the attention of DSC after he posted a photo of it on the BugGuide website, and then kindly sent it to him when notified that it was the first known specimen of an undescribed species.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD6FFDBE1D2F997E2ABFEA2.taxon	description	(Figs 18 – 19, 39 C)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD6FFDBE1D2F997E2ABFEA2.taxon	materials_examined	Material examined, 3 specimens. HOLOTYPE male: MEXICO: Tamaulipas: // MEX: Tamaulipas: La Pesca, 20 Sept. 1981, W. H. Cross / blacklight trap / [red label] HOLOTYPE Brachygluta tamaulipec Sabella // (FMNH). Non-type specimens, 2 females: Texas: Cameron Co.: 1.5 mi E jct. FM 1419 on Hwy. 4, E of Brownsville, X- 19 - 2002, B. Raber & E. Riley, UV light (DSCC, 1; TAMU, 1).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD6FFDBE1D2F997E2ABFEA2.taxon	description	Description. BODY: Length 2.32 – 2.40 mm; brown, with elytra orange-brown; setae over body short, strongly curved and suberect to decumbent. Head: surface smooth, shining, punctures indistinct. Setose area of median vertexal fovea about smaller than those of lateral foveae; antennomeres (both sexes) III – VII longer than wide, VIII transverse, IX – X as long as wide, VIII two-thirds width of IX (Fig. 19 A). Pronotum: surface smooth, shining, small punctures distinct; setose area of median antebasal fovea smaller, about two-thirds width of those in lateral antebasal foveae. Elytra: surface lightly punctate, opaque due to light microreticulation, discal stria extending to about four-fifths of elytral length. Abdomen: surface smooth, shining, punctures indistinct. Basal striae of tergite 1 short, one-fifth to one-sixth as long as paratergite, striae separated by about one-half tergite width; basal setose brush between striae distinct. MALE: Antennae lacking modifications. Protrochanter with ventral acute spine (Fig. 19 C). Metaventrite with broad impression at middle, setae sparse laterally, denser in impression. Tergite 1 with apex protruding as rounded subtruncate lobe, apex with two close, ventrally directed tufts of setae forming flattened lamina, lateral margins of lobe with dense short setae, appearing densely punctate; tergite 2 base raised at middle as transverse carina, difficult to see, two shallow oval impressions centered on tergite disc; tergite 3 narrowly impressed at base (Figs 18 A-B); in lateral view tergite 1 rising posteriorly to deflexed apex, tergite 2 nearly level, apex of 1 well-elevated above 2 (Fig. 18 C). Abdominal ventrites broadly convex. Metatibiae narrow basally, slightly widening through length to apex, with dense comb of setae on mesal margin in apical fifth where slightly curved medially (Fig. 19 B). Aedeagus 0.44 mm long; dorsal plate indistinct; parameres elongate and sinuous, lacking distinct preapical constriction in dorsal view, with slight mesal angulation in lateral view (Fig. 19 E); with elongate thin mesal hyaline seta, similar in form to three short lateral setae; internal sac with one large straight spine and with thick, strongly curved spine complex (Fig. 19 D). FEMALE: Two probable females have: metaventrite with long setae, setae denser in triangular area from between metacoxae to metasternal base; basal carinae of tergite 1 slightly longer, about one-fifth as long as tergite, basal separation of carinae narrower, about two-fifths width of tergite. Abdominal ventrites broadly rounded. Metatibiae similar to those of male. Collecting data. The holotype was collected in September, with the collecting technique unknown. The two probable females were taken by an ultraviolet light in October.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD6FFDBE1D2F997E2ABFEA2.taxon	distribution	Distribution (Fig. 39 C). The holotype and associated females were taken at sites at or near the Caribbean coast in Texas and Tamaulipas. The two sites are 240 km apart.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD6FFDBE1D2F997E2ABFEA2.taxon	discussion	Comments. The two females match up well with the holotype in terms of coloration, size, antennal form, very short basal carinae of tergite 1, and setation. The male and females differ only in the relative width between the basal carinae, and have slight differences in lengths of the basal carinae. This can vary between sexes in other species, and based on the geographic closeness and coastal habitat, these females are very likely conspecific. The prominent median non-emarginate lobe of tergite 1 of the male, and the relatively simple tergite 2 place this species near B. dentata. The apex of tergite 1 is subtruncate in B. tamaulipec and the metaventrite is broadly impressed, while the lobe apex is bluntly angular and the metaventrite nearly convex (slightly longitudinally impressed at the middle) in B. dentata. The holotype is a specimen from the Mississippi Entomological Museum, Mississippi State University, and is gifted to the FMNH courtesy of Terry Schiefer.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD6FFDBE1D2F997E2ABFEA2.taxon	etymology	Etymology. The name is taken from the group of indigenous people most closely identified with the locality of the holotype specimen.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD5FFC4E1D2FD02E7EEFEFF.taxon	description	(Figs 20, 40 A)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD5FFC4E1D2FD02E7EEFEFF.taxon	materials_examined	Material examined, 11 specimens. HOLOTYPE male: USA: California: Mono Co.: // USA: California: Mono Co., Mono Lake, Navy Beach, 12. VI. 2008, KTE 08 - 156, taken with ants under rock, leg. K. T. Eldredge / [red label] HOLOTYPE Brachygluta eldredgei Chandler // (FMNH). PARATYPES, 10: eutopotypical (DSCC, 2 M 4 F; SEMC, 3 F). Arizona: Cochise Co.: Cochise Stronghold, Dragoon Mts., IX- 3 / 7 - 1969, R. J. Shaw, UV light trap (UAIC, 1 M).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD5FFC4E1D2FD02E7EEFEFF.taxon	description	Description. BODY: Length 1.96 – 2.02 mm; red-brown to brown with red-brown elytra; head and pronotum with setae short, curved and suberect, over rest of body setae decumbent. Head: surface smooth, shining, punctures indistinct. Antennomeres III – VII longer than wide, VIII as long as wide, IX elongate, X transverse, VIII about three-fourths width of IX (Fig. 20 C). Setose area of median vertexal fovea slightly narrower than those of lateral foveae. Pronotum: surface smooth, shining, punctures indistinct. Setose area of median antebasal fovea slightly smaller than those of lateral antebasal foveae. Elytra: with faint microreticulation, punctures indistinct, discal stria extending to about four-fifths of elytral length. Abdomen: tergite 1 with disc smooth, shining, punctures indistinct; basal striae of tergite 1 straight, about one-third paratergite length or less for male, about one-fifth paratergite length for female, separated at base by about two-fifths tergite width, with distinct long but sparse setose brush between bases of striae. MALE: Antennae and trochanters unmodified, simple. Metaventrite impressed from middle to apex, impression covered with setae that are longer and denser. Tergite 1 elongate, length nearly equal to width of disc, broadly emarginate at apex, apex deflexed in narrow zone that has thicker red-brown setae denser than on disc; tergite 2 with middle half lacking setae, with slightly arcuate transverse carina at center of disc, margins of seta-free area slightly converging posteriorly, lateral portions of tergite with short thick golden setae, area opaque due to dense punctation (Figs 20 A – B); in lateral view tergite 1 protruding slightly over base of tergite 2; tergites 3 – 4 with short thick golden setae, setae sparser, not as densely punctate; in lateral view with tergites forming evenly curved outline, but with deep grooves between tergites 1 – 2 and 2 – 3. Ventrites broadly convex. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical fourth with brush present (Fig. 20 D). Aedeagus 0.37 mm long; dorsal plate triangular, with preapical constriction of margins to form acutely pointed apex; parameres with three thick setae on lateral margin at basal angle of preapical constriction, with preapical hyaline seta near mesal margin, thick basally, aciculate apically; internal sac with three thick curved spine (Fig. 20 E). FEMALE: Metaventrite with setae similar over breadth; lacking abdominal modifications, tergites gently convex; ventrites broadly curved. Metatibiae similar to those of male. Collecting data. Found in association with a colony of Pogonomyrmex ants under rocks along the margin of an alkaline lake, and one specimen was collected using an ultraviolet light trap. The specimens were taken in June and September.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD5FFC4E1D2FD02E7EEFEFF.taxon	distribution	Distribution (Fig. 40 A). Only known from California at the western margin of the Great Basin, and also from eastern Arizona.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD5FFC4E1D2FD02E7EEFEFF.taxon	discussion	Comments. The relatively long tergite 1 that is broadly emarginate at the apex, together with the thicker and shorter setae present on the deflexed apex of tergite 1 and also across tergite 2, places this species in the texana species-group. This species is easily separated from the other species by the broad transverse impression of tergite 2 that has an arcuate transverse carina at the middle of the tergite. The other two species in this group lack this feature, but have a small bump at the center of tergite 2 with two oblique sulci to each side.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFD5FFC4E1D2FD02E7EEFEFF.taxon	etymology	Etymology. Named for K. Taro Eldredge, collector of all but one specimen of the type series.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFCBFFC7E1D2FF14E67DFB7A.taxon	description	(Figs 21, 40 B)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFCBFFC7E1D2FF14E67DFB7A.taxon	materials_examined	Material examined, 144 specimens. USA: Arizona: Yuma Co.: Fort Yuma, I- 1 - 1921, Hubbard & Schwarz (USNM, 4). “ Ad Gilam ” [mouth of Gila River] (MCZC, 3, Lectotype + 2 Paralectotypes). California: Fresno Co.: 8 mi SW Coalinga, III- 24 - 1988, F. Andrews & A. R. Hardy, under rocks, grassy hillside (CSCA, 1). 8 mi W Coalinga, Warthan Canyon, IV- 16 - 1980, Andrews & Paddock (DSCC, 1). Coalinga, Los Gatos Canyon, V- 17 - 1957, H. L. Wilson, cottonwood litter (DSCC, 2). Inyo Co.: Deep Springs Valley, Buckhorn Springs, II- 1971, D. Giuliani (CASC, 23); III- 1971 (CASC, 1); IV- 11 - 1975, D. Giuliani (CSCA, 1). Deep Springs, Deep Springs Valley, ca. 19 air mi. E Bishop, II- 1971, D. Giuliani (CASC, 1). Fish Slough, 10 mi N Bishop, I- 8 - 1972, D. Giuliani (CASC, 1). Saline Valley, salt marsh, IX- 1 - 1976, D. Giuliani, UV light (CSCA, 2); VI- 7 - 1976 (DSCC, 1). Tinemaha Reservoir, VII- 12 - 1975, F. Andrews & A. R. Hardy, sand dune E side (CSCA, 4). Westergard Pass cave, Inyo Mountains, 60 ' below surface, V- 27 - 1982, D. Giuliani, berlese from bat guano (CSCA, 1); V- 27 - 1982, D. Giuliani, (CSCA, 1). Kern Co.: Bakersfield, V- 5 - 1931 (DSCC, 1). Los Angeles Co.: Los Angeles, Hubbard & Schwarz (USNM, 4; MCZC, 2). Pasadena, VI, A. Fenyes (CASC, 1). Madera Co.: Wind Gap, VII- 13 - 1946, H. P. Chandler (UCDC 1). Merced Co.: Los Banos Valley, 400 ', V- 17 - 1987, D. Giuliani (CSCA, 1); IV- 12 - 1987, D. Giuliani (CSCA, 1); V- 27 - 1982, D. Giuliani, (CSCA, 1). Mono Co.: Paoha Island, Mono Lake, VII- 1980 / VII- 1981, E. Sugden, traps 1 - 6 (CSCA, 1). Riverside Co.: Morongo Valley, I- 22.1915, G. K. C. (CSCA, 1). Riverside, III- 28 - 1891 (MCZC, 1, H. C. Fall Collection). Temescula, VI- 24 - 1987, Green (CSCA, 1). San Bernardino Co.: Needles, IV- 1 / 6 - 1918, J. C. Bradley (CUIC, 1). Saratoga Springs, Death Valley, IV- 15 - 1966, G. L. Peters, UV light (DSCC 1). Zzyzx Station, poolside, IV- 23 - 1977, R. Aalbu, black light (CSCA, 3; DSCC, 2). San Diego Co.: VI- 14 - 1890, F. E. Blaisdell (CASC, 3; UCDC, 1); X, F. E. Blaisdell (CASC, 1). Borego Palm Canyon, IV- 25 - 1955, R. Schuster (UCDC, 6). Oceanside (CASC, 1). San Diego, 1917, H. C. Fall (MCZC, 1); 1917, (MCZC, 1); VIII- 6 - 1890 (MCZC, H. C. Fall Collection, 1; MCZC, 1; FMNH, 3); II- 26 - 1890 (MCZC, H. C. Fall Collection, 2); II- 26 (FMNH, 1). Only state on label: “ Cal. ” (CASC, 1; CMNH, 12); F. E. Blaisdell (UCDC, 1; DSCC, 1). “ So. Cal. ” (ANSP, 1; MCZC, 4). “ California ” Reichle (MNHN, 1 M). Nevada: Clark Co.: Paradise Valley (L. (as) V. (egas), VII- 25 - 1970 (FMNH, 3), VIII- 4 - 1970, B. W. Miller, UV light (FMNH, 5). Lincoln Co.: Alamo, VII- 10 - 1958, F. D. Parker (DSCC, 1); VII- 11 - 1958, F. D. Parker (DSCC, 1; UCDC, 1); VII- 21 - 1958 (DSCC, 1; UCDC, 1); VII- 12 - 1958 (UCDC, 1). Utah: Tooele Co.: 12 mi S Timpie, IX- 28 - 1953, P. D. Ashlock (BYUC, 2). Utah Co.: Provo, VI (USNM, 2). Salem, VIII- 10 - 1960, G. F. Knowlton, light trap (UCDC, 2). Washington Co.: St. George, Springs Subdivision, VII- 21 - 2003, M. Oldham (BYUC, 1). County unknown: Salt Lake, Soltau (FMNH, 2). Only state on label: “ Ut., ” Brend. (MCZC, Horn Collection, 3). Utah (CMNH, 3 Syntypes of B. mormon).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFCBFFC7E1D2FF14E67DFB7A.taxon	description	Description. BODY: Length 1.80 – 2.04 mm; orange-brown, to brown; setae on head and pronotum short, curved and suberect, on body appressed or nearly so. Head: surface smooth, shining, punctures indistinct. Setose area of median vertexal fovea as large as those of lateral foveae. Antennomeres slightly shorter in female than male, differences indicated below. Pronotum: smooth, shining, punctures indistinct. Setose area of median antebasal fovea about two-thirds width those of lateral antebasal foveae. Elytra: smooth, shining, microreticulation faint; discal stria reaching to four-fifths of elytral length. Abdomen: disc smooth, shining, punctures sparse. Tergite 1 with basal abdominal striae variable, ranging from about one-sixth to nearly one third paratergite length, separated at base by more than half of tergite width, with distinct setose comb between bases of striae. MALE: Antennae lacking modifications, antennomeres III – VI clearly longer than wide, VII slightly longer than wide, VIII – IX as long as wide, X wider than long, VIII about two-thirds width of IX (Fig. 21 C). Trochanters lacking modifications. Metaventrite convex, with dense appressed setae covering most of middle portion of metaventrite, setose zone rounded on anterior margin. Abdomen with tergite 1 disc about as long as wide; posterior margin of tergite 1 shallowly emarginate at middle, short appressed setae and punctures dense in deflexed apical zone of tergite; tergites 2 – 3 with short appressed golden setae on disc, surface microreticulate; tergite 2 with two oblique, dorsally converging sulci, with rounded bump between them at middle (Figs 21 A – B); in lateral view with deflexed apex of tergite 1 curving down just posterior to 2, 2 - 3 short and nearly vertical. Ventrites broadly rounded. Metatibiae narrow, slightly widening through length to near apex, with dense comb of setae on mesal margin in apical fourth where slightly curved medially (Fig. 21 D). Aedeagus 0.34 mm long, dorsal plate triangular, with preapical constriction of margins to form acutely pointed apex; parameres with three thick setae on lateral margin at basal angle of preapical constriction, with preapical hyaline seta near mesal margin, thick basally, aciculate apically; internal sac with two large spines, one straight and one curved (Fig. 21 E). FEMALE: Antennomeres III – V longer than wide, VI – VII slightly longer than wide, VIII – X transverse, VIII about two-thirds width of IX. Metaventrite convex, sparsely setose. Abdominal ventrites broadly rounded. Metatibiae similar to that of male. Collecting data. Under rocks on grassy hills; in cottonwood litter, in bat guano; at UV light (Chandler 1997). Only a few specimens have collection data indicated, but those with such data were taken primarily at ultraviolet light. Most specimens were taken from April to September, but there are records from all months except November and December.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFCBFFC7E1D2FF14E67DFB7A.taxon	distribution	Distribution (Fig. 40 B). Found from the Mojave Desert and grasslands of southern California to the Upper Sonoran desert of Nevada and Utah. Only present in Arizona in the Yuma area.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFCBFFC7E1D2FF14E67DFB7A.taxon	discussion	Comments. This species is very similar to B. texana, and can be reliably separated only by the presence of two thick spines in the internal sac, and by its more western distribution that extends from southern California to Utah. Brachygluta texana has three thick spines in the internal sac, and is found in the Chihuahuan and lower Sonoran Deserts, ranging from Texas through Arizona and into California just across the Colorado River at one point. LeConte (1851) did not indicate the number of specimens before him when he described this species, but for other new species in this paper he would indicate when he had only a single specimen, suggesting that he based B. foveata on more than one specimen. Specimens with labels indicating the mouth of the Gila River in “ California ” (Arizona today), or with LeConte’s gold-colored circle indicating California, were chosen to form the syntype series. The male Lectotype here designated bears the labels: // Cal. [with dot below the “ C ” = mouth of Gila River] / ♂ / [red label] Type 6134 / LECTOTYPE Bryaxis foveata LeConte 2013 Desig. DSChandler //. Two males and a female with the same locality label, as well as a female specimen with a gold disc label are labeled as Paralectotypes. “ California ” at the time of LeConte’s visit included what would later become the state of Arizona, with the mouth of the Gila River into the Colorado River being on the Arizona side of the Colorado River. Brendel (1893: 278) and then Casey (1894: 481) apparently confused B. foveata with B. mormon Bowman, 1934 (not yet described at that time), with Casey describing B. loripes in the place of B. foveata. Casey later added B. jacobina (1908: 259) based on subtle differentiating characters, for a group where examination of the genitalia is necessary to separate the two very similar species. Bryaxis loripes Casey and B. jacobina Casey are here placed as a junior synonyms of B. foveata based on the similar form of the male tergites and the aedeagus. A Lectotype of B. jacobina is here designated, and a Paralectotype recognized and labeled as such. Lectotype male: // San Diego, Cal / Casey bequest 1925 / Brachygluta jacobina Csy / [red label] TYPE USNM 38696 / LECTOTYPE Brachygluta jacobina Casey ♂ ‘ 13 //. The Paralectotype female shares the first three labels and has a Paralectotype label.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC9FFC0E1D2FA94E741F806.taxon	description	(Figs 22, 40 C)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC9FFC0E1D2FA94E741F806.taxon	materials_examined	Material examined, 93 specimens. USA: Arizona: Cochise Co.: 4 km E Willcox, VIII- 20 - 1982, L. D. French, UV light (UCDC, 1); IX- 14 - 1972, D. S. Chandler & K. Stephan (DSCC, 1). Cochise Stronghold, Dragoon Mts., VII- 24 / 27, R. J. Shaw, UV light trap (DSCC, 1), VII- 27 / 30 - 1970, R. J. Shaw, UV light trap (UAIC, 1), VIII- 29 / IX- 3 - 1969, R. J. Shaw, UV light trap (UAIC, 1), IX- 3 / 7 - 1969, R. J. Shaw, UV light trap (UAIC, 1); Graham Co.: 2 mi E Safford, VIII- 19 - 1977, D. S. Chandler, UV light (DSCC, 7; UCDC, 1). Aravaipa Canyon, VIII- 19 - 1977, D. S. Chandler, UV light (UCDC, 1); VIII- 12 - 1975, S. McCleve, at light (UAIC, 1). Aravaipa Canyon, 3050 ', VIII- 24 - 1976, D. S. Chandler, at light (DSCC, 6; UAIC, 2); east end Aravaipa Canyon, VII- 24 / 25 - 1974, S. McCleve, at light (UAIC, 3). Geronimo, VIII- 6 - 1948, Werner & Nutting (DSCC, 1; UAIC, 1). Thatcher, VIII- 5 - 1940, E. J. Taylor (DSCC, 1). La Paz Co.: Ehrenberg, VII- 1938, F. H. Parker (UAIC, 2); VI- 24 - 1938, F. H. Parker (UAIC, 1); III- 20 - 1939, F. H. Parker (UAIC, 2); III- 22 - 1940, F. H. Parker (UAIC, 2); IX- 9 - 1939, F. H. Parker (UAIC, 3). Parker, IX- 10 - 1979, T. King (FMNH, 2). Maricopa Co.: Gillespie Dam, Gila Bend, 1000 ', VIII- 9 - 1948, F. Werner & W. Nutting (UAIC, 1). Wickenburg, VIII- 12 - 1950 (FMNH, 1), VIII- 31 - 1950 (FMNH, 1), H. K. Gloyd. Mohave Co.: Burro Creek Camp, V- 11 - 1988, T. Dworakowska (DSCC, 5). Pima Co.: Tucson (USNM, holotype of B. arizonae). Pinal Co.: Superior, VII- 31 - 1984 (FMNH, 5), VIII- 5 - 1948 (FMNH, 1), H. K. Gloyd. Superior, B (oyce) Thomps (on) Arb (oretum), 2500 ', VII- 30 - 1948, H. K. Gloyd, white light trap with 1 / 4 " screen (DSCC, 1). No County: Grand Can (yon), VII- 4 - 1904 (MCZC, H. C. Fall Collection, 1). California: Riverside Co.: Blythe, VIII- 8 - 1947, J. W. MacSwain (UCBC, 1); VIII- 24 - 1947, J. W. MacSwain (UCBC, 1); VII- 8 - 1947 Barr & MacSwain (UCBC, 1); VI- 8 - 1941 (UCBC, 1); VIII- 6 - 1947, J. W. MacSwain (MCZC, 1; DSCC, 1); VII- 10 - 1947, J. W. MacSwain (UCDC, 1; CASC, 6; DSCC, 1). New Mexico: Bernalillo Co.: Albuquerque, II- 15, H. Soltau (USNM, 4); III- 1, H. Soltau (USNM, 1); XII- 3 - 1901 (MCZC, H. C. Fall Collection, 2). Chaves Co.: Bottomless Lakes State Park, VIII- 7 - 1989, Y. Bousquet, gypsum flats (CNCI, 1). Sandoval Co.: 6 mi S Jemez Springs, VIII- 22 - 1968, F. G. Andrews & G. R. Noonan, blacklight (CSCA, 4; DSCC, 2). Only state on label: “ N. M. ” (MCZC, H. C. Fall Collection, 2). Texas: Brazos Co.: College Station, V- 29 - 1933, H. J. Reinhard (TAMU, 1). El Paso Co.: El Paso (USNM, holotype B. texana). Montgomery Co.: 5 mi N Montgomery, III- 7 - 1987, R. S. Anderson, black walnut litter (DSCC, 1). Presidio Co.: 3 mi NE of Porvenir, IX- 26 - 1946, CNMH Palaeont. Exped. Texas (1946) B. Patterson leg. (FMNH, 1).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC9FFC0E1D2FA94E741F806.taxon	description	Description. BODY: Length 1.80 – 1.88 mm; orange-brown to brown, setae over body short, on head decumbent, over pronotum and body appressed. Head: surface smooth, shining, punctures indistinct. Setose area of median vertexal fovea as wide as those of lateral foveae. Antennomeres slightly shorter in female than male, differences indicated below. Pronotum: smooth, shining, punctures indistinct. Setose area of median antebasal fovea slightly smaller than those of lateral antebasal foveae. Elytra: smooth, shining with microreticulation faint and punctures visible, to microreticulation stronger and obscuring punctures; discal stria reaching to four-fifths of elytral length. Abdomen: tergite 1 disc smooth, shining, punctures sparse; with basal striae variable in length, ranging from about one-sixth to nearly one-third paratergite length, at base distant from each other more than half of tergite width, with distinct but sparse setose brush between bases of striae. MALE: Antennae lacking modifications, antennomeres III – VI clearly longer than wide, VII slightly longer than wide to quadrate, VIII slightly transverse, IX about as wide as long, X transverse, VIII close to one-half width of IX (Fig. 22 C). Trochanters lacking modifications. Metaventrite convex, with dense appressed setae covering middle third of metaventrite. Abdomen with tergite 1 disc nearly as long as wide; posterior margin of tergite 1 shallowly emarginate at middle, short appressed setae and punctures dense on deflexed apical zone of tergite; tergites 2 - 3 with short appressed golden setae on disc, surface microreticulate; tergite 2 with two oblique, dorsally converging sulci, with rounded bump between them at middle (Figs 22 A – B); in lateral view tergite 1 with deflexed apex curving down just posterior to 2, 2 - 3 nearly vertical. Ventrites broadly convex. Metatibiae narrow basally, slightly widening through length to near apex, with dense comb of setae on mesal margin in apical fourth where slightly curved medially (Fig. 22 D). Aedeagus 0.38 mm long; dorsal plate triangular, with acutely pointed apex; parameres with three thick setae on lateral margin at basal angle of preapical constriction, with preapical hyaline seta near mesal margin, thick basally, aciculate apically; internal sac with three spines, one straight and two curved (Fig. 22 E). FEMALE: Antennomeres III – V longer than wide, VI slightly longer than wide, VII – X transverse, VIII about two-thirds width of IX. Metaventrite with setae at middle more abundant and longer than laterally, but not noticeably dense. Abdominal ventrites broadly rounded. Collecting data. One collection was taken from black walnut litter, but nearly all other specimens with collecting data were taken at ultraviolet light. One specimen was attracted to white light. Nearly all individuals were taken from July to September, but a small number were taken during the winter months and May.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC9FFC0E1D2FA94E741F806.taxon	distribution	Distribution (Fig. 40 C). This species is found in the Chihuahuan and lower Sonoran Deserts from central Texas through Arizona.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC9FFC0E1D2FA94E741F806.taxon	discussion	Comments. This species is very close in form to B. foveata, and can be reliably separated only by the presence of three thick spines in the internal sac, and by its more easterly distribution. Brachygluta foveata has only two large spines in the internal sac, and is found basically west of the Colorado River (except the type series), while B. texana is found east of the Colorado River (except for collections at Blythe, California).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFCCFFC3E1D2FF14E666F897.taxon	description	(Figs 23, 41 A)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFCCFFC3E1D2FF14E666F897.taxon	materials_examined	Material examined, 247 specimens. CANADA: Ontario: Essex Co.: Tilbury, III- 24 - 1967 (FMNH, 40), IV- 2 - 1967 (FMNH, 7), IV- 15 - 1967 (FMNH, 12), IV- 21 - 1965 (FMNH, 1), IV- 28 - 1967 (FMNH, 1), V- 2 - 1961 (FMNH, 1), VI- 29 - 1967 (FMNH, 6), VII- 1962 (MHNG, 5), IX- 5 - 1967 (FMNH, 12), K. Stephan, sifting. Wheatley, IV- 8 - 1967 (FMNH, 3), IV- 17 - 1967 (FMNH, 2), IV- 29 - 1967 (FMNH, 1), V- 3 - 1967 (FMNH, 2), VI- 10 - 1967 (FMNH, 4), K. Stephan, sifting. Kent Co.: Merlin, IV- 30 - 1967, K. Stephan, sifting (FMNH, 4). Rondeau Prov. Pk., Beach near entrance, VI- 3 - 1985, A. Davies & J. M. Campbell, in debris on beach at high water line (CNCI, 22). Rondeau Provincial Park, South Beach, VI- 5 - 1985, A. Davies & J. M. Campbell, in debris on beach at high water line (CNCI, 2). Only province on label: “ Ontario ” (MHNG, 1 M). Only country on label: “ Can. ” (CMNH, 1). USA: District of Columbia: “ D. C. ” (CMNH, 2). Illinois: Champaign Co.: University Woods, Urbana, IV- 10 - 1937, M. Farrar (FMNH, 2); X- 16 - 1933, M. D. Farrar, from soil cover in woods (INHS, 2); X- 3 - 1933, M. D. Farrar, ground cover (INHS, 2). Cook Co.: Bowmanville, X- 14 - 1910, Liljeblad (ANSP, 1); X- 16 - 1904 (FMNH, 19). Lions Woods, V- 5 - 1984, L. E. Watrous, oak litter (FMNH, 8). McLean Co.: Funks Grove, IV- 27 - 1981, R. W. Lundgren, at carcass in woods (RWLC, 1; DSCC, 1); IV- 4 - 1981, R. W. Lundgren, under bark of log (RWLC, 1). Only state on label: “ Ill. ” (CMNH, 1). Indiana: Floyd Co.: New Albany, VII- 31 - 1966, C. E. White, blacklight trap (FSCA, 1). Monroe Co.: Bloomington, VIII- 20 - 1985 (FSCA, 1), IX- 10 - 1987 (FSCA, 1), V- 31 / VI- 1 - 1986 (FSCA, 1), F. N. Young, blacklight trap. Parke Co.: 3 mi SE Montezuma, VIII- 3 / 4 - 1992, S. Turner, UV light trap (EIUC, 1). Wells Co.: I- 9 - 1973, N. M. Downie, tree duff (FMNH, 1). Maryland: Montgomery Co.: Plummers Island, VI- 8 / 13, W. L. McAtee (USNM, 1). Michigan: Wayne Co.: Detroit, Hubbard & Schwarz (USNM, 4; ANSP, 1; MCZC, 1); X, Hubbard & Schwarz (USNM, 2). New York: Queens Co.: Rock (away) Beach, L. (ong) I. (sland), VI- 16 - 1922 (USNM, 1). North Carolina: Northampton Co.: 7 km S Jackson, V- 21 / VII- 1 - 1987, H Goulet, FIT Camasia slope (DSCC, 3; CNCI, 1). Ohio: Crawford Co.: Plankton, XI- 12 - 1918, P. S. Bartholomew (CASC, 3). Cuyahoga Co.: Rocky River Reservation, Cleveland, IX- 16 - 2005, H. J. Lee, vernal pool vacuum sedge clumps (DSCC, 6); VIII- 31 - 2005, H. J. Lee, sedge clumps vernal pool area (DSCC, 14). Delaware Co.: 5 mi N Delaware: V- 21 - 1977, L. E. Watrous, sift debris along lake shore (DSCC, 2). Fairfield Co.: 11 mi S Lancaster, V- 8 - 1976, Q. D. Wheeler (DSCC, 1). Franklin Co.: 1 mi N Worthington, III- 25 - 1977, L. E. Watrous, berlese litter near stream (DSCC, 1). Columbus, XI- 20 - 1976, L. E. Watrous, sift leaf litter (DSCC, 1); III- 9 - 1980, L. E. Watrous, damp leaf litter (FMNH, 1). Scioto River, IV- 13 - 1975, L. E. Watrous (DSCC, 1). Sharon Woods Metropolitan Park, VII- 2 / 10 - 1973, A. J. Penniman, pitfall trap (DSCC, 1). West Campus pond, O. S. U., Columbus, IV- 20 - 1976, D. S. Chandler (DSCC, 1). Greene Co.: John Bryan State Park, VI- 1 - 1975, D. S. Chandler, sifting river drift (DSCC, 2). Hocking Co.: 2 mi S Gibisonville, VII- 8 - 1978, L. E. Watrous, sift litter in swamp (DSCC, 1). Rockbridge, IX- 21 - 1975, D. S. Chandler, sifting forest litter (DSCC, 1). Lawrence Co.: Ald. Twp., Wayne National Forest, Sharp's Creek, V- 23 / VI- 7 - 1995, P. Kovarik, F. I. T. (LSAM, 2). Pickaway Co.: Circleville, Scioto River at Bridge Crossing Rt. 22, 39 o 36.30, VIII- 24 - 2008, K. D. Karns, flooding gravel along river bank (KDKC, 1). Portage Co.: Eagle Lake Bog, Nelson Twp., IX- 11 - 2004, H. J. & C. J. Lee, berl. Sphagnum moss (DSCC, 1). Pennsylvania: Jefferson Co.: Cook State Forest, 1.2 mi N Cooksburg, Troutman Run, VI- 6 - 1997, D. S. Chandler, leaf litter along stream (DSCC, 1). Westmoreland Co.: Beatty, IX- 11 - 1896 (FMNH, 2); St. Vincent, IV- 26 (MCZC, 1). Only state on label: “ Pen. ” (CMNH, 3; MNHN, 2). West Virginia: Cabell Co.: Wolfpen Hollow, 3 mi S Culloden, V- 15 - 1998, S. M. & A. T. Clark (WVDA, 1). Grant Co.: Miser, V- 25 - 1994, S. M. Clark (WVDA, 1). Kanawha Co.: Guthrie, VII- 14 - 1986, M. C. Thomas, UV trap (DSCC, 2); V- 29 - 1987, M. C. Thomas, UV trap (DSCC, 1; WVDA, 6). Mason Co.: McClintic Wildlife station, VII- 26 - 1986, M. C. Thomas, UV trap (WVDA, 1; DSCC, 2). Morgan Co.: Widmeyer Wildlife Management Area, VIII- 21 - 1997, S. M. Clark (DSCC, 1). Tucker Co.: Dolly Sods Scenic Area, IX- 16 - 1992, S. M. Clark (DSCC, 1).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFCCFFC3E1D2FF14E666F897.taxon	description	Description. BODY: Length 1.84 – 2.04 mm; usually brown, a few specimens with elytra lighter, or hind body orange-brown; setae on head and pronotum short, curved and suberect, longer and decumbent over rest of body. Head: surface smooth, shining, punctures indistinct. Antennomeres III – VI longer than wide, VII barely longer than wide (as long as wide in female), VIII – X distinctly transverse, VIII about two-thirds width of IX (Fig. 23 C). Setose area of median vertexal fovea slightly smaller than those of lateral foveae. Pronotum: surface smooth, shining, punctures indistinct. Setose area of median antebasal fovea slightly narrower than those of lateral antebasal foveae. Elytra: surface varying from smooth with indistinct punctures to microreticulate and with distinct punctures, discal stria extending to about four-fifths of elytral length. Abdomen: disc of tergite 1 smooth and shining, lightly punctate. Basal striae of tergite 1 long, divergent, one-half length of paratergite or more in male, one-third to one half paratergite length in female, separated at base by about two-fifths tergite width; with sparse but distinct setose comb at base between striae in males, fringe less distinct in females. MALE: Metaventrite with transverse band of long dense setae between metacoxae. Only tergites 1 – 2 visible in dorsal view, tergite 1 twice as long as 2; tergite 1 with disc smoothly and shallowly convex, apex shallowly emarginate at middle, with row of appressed golden setae forming elongate flattened lamina, lamina shallowly emarginate at apex and directly posteroventrally (Fig. 23 A); tergite 2 with semicircular impression at middle extending from base to or near to apex, narrow median strip glabrous, lateral portions of impression with setae suberect, directed obliquely medially, denser and more elongate than on lateral portions of tergite; lateral portions of tergite 2 with flattened or lightly impressed area, setae in this area much shorter and appressed; tergites 3 – 5 evenly convex, not modified (Fig. 23 B); in lateral view with tergite 1 rising toward deflexed apex, 2 clearly lower, sloping ventrally from base of 1. Ventrites broadly rounded. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical fifth with brush present (Fig. 23 D). Aedeagus 0.37 mm long; with parameres curved laterally in apical portion, lacking distinct preapical constriction, with three aciculate preapical setae on lateral margin, with broad, flat, hyaline seta on inner margin near apex; dorsal plate asymmetric, abruptly narrowed near base, thin to apex, apical portion with margins about same width until curving slightly to right at apex or nearly straight (Fig. 23 I); internal sac with two large isolated spines and complex thick tooth with cluster of 3 - 6 small spines on apical portion (Figs 23 E – H). FEMALE: Metaventrite with setae in middle portion denser than laterally; abdominal tergites evenly convex; ventrites broadly convex. Metatibiae similar to those of male. Collecting data. Harry J. Lee has collected many specimens from sedge clumps in and around vernal pools in the Cleveland area of Ohio. Specimens have also been taken from river drift, and from litter on pond and lake shores, as well as being collected by ultraviolet lights (Chandler 1997). This species appears to prefer leaf litter in and around forested freshwater swamps, as well as litter along the margins of streams. Collected from March to October, with one record from January.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFCCFFC3E1D2FF14E666F897.taxon	distribution	Distribution (Fig. 41 A). This is a species of the mid-western and the western portion of the northeastern United States that has also been commonly taken from southern Ontario in Canada. There is only one record from east of this area, with a single specimen being taken from Long Island (New York).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFCCFFC3E1D2FF14E666F897.taxon	discussion	Comments. This species is similar in appearance to B. wickhami, in being small and dark, and the male modifications of tergite 2 are similar in general form. Brachygluta arguta has the slightly asymmetrical median lobe of the aedeagus long and thin, a thick spine in the internal sac with 3 – 6 (usually 4) small spines at the apex, and the impression of tergite 2 has the apex semicircular with the lateral portions of the impression not densely setose. Brachygluta wickhami has the symmetrical median lobe of the aedeagus shorter and broadly triangular, and the more longitudinal impression of tergite 2 extends to or close to the tergite apex, with the lateral portions of the impression being densely setose and a median longitudinal glabrous strip present through most of the length of the impression. Both B. arguta and B. wickhami have the setose portion of the median pronotal fovea narrower than for the lateral foveae, while in B. terebrata the setose portion of all pronotal foveae are similar in size. Lectotype is here designated, and Paralectotypes recognized and labeled as such. Lectotype male: // Mich / Casey bequest 1925 / TYPE USNM 38691 / arguta / LECTOTYPE Bryaxis arguta Casey ♂ ‘ 13, des. DSChandler //. Two Paralectotypes designated: // Mich / Casey bequest 1925 /? arguta - 2 PARATYPE USNM 38691 / PARALECTOTYPE Bryaxis arguta Casey ♀ ‘ 13, des. DSChandler //; // St. Vinc. Penn / 68 / Casey bequest 1925 /? arguta - 3 PARATYPE USNM 38691 / PARALECTOTYPE Bryaxis arguta Casey ♂ ‘ 13, des. DSChandler //.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC2FFCEE1D2F8BBE1E8FBAF.taxon	description	(Figs 24, 41 D)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC2FFCEE1D2F8BBE1E8FBAF.taxon	materials_examined	Material examined, 5 specimens. USA: Arizona: Cochise Co.: Chiricahua Nat. Mon., VIII- 8 - 58, R. O. Schuster, L. M. Smith Colrs. (UCDC, 1). Coronado N (ational) F (orest), Chiricahua Mts., VIII- 5 - 2013, C. Carlton & B. Owens, sifted litter-Berlese (LSAM, 2). Coconino Co.: Williams (holotype, ANSP, 1). Only state on label: “ Ariz. ” (CMNH, 1).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC2FFCEE1D2F8BBE1E8FBAF.taxon	description	Description. BODY: Length 1.76 – 1.84 mm; orange-brown to red-brown; head and pronotum with setae short, curved and suberect, over rest of body decumbent to appressed. Head: surface smooth, shining, punctures indistinct. Antennomeres III – VI longer than wide, VII as long as wide, VIII – X strongly transverse, VIII about twothirds width of IX (Fig. 24 C). Setose area of median vertexal fovea as large as those of lateral foveae, setose areas smaller than in other species. Pronotum: surface smooth, shining, punctures indistinct. Setose area of median antebasal fovea about two-thirds width of lateral antebasal foveae, setose areas relatively small when compared to other species. Elytra: with microreticulation faint to distinct, punctures indistinct, discal stria extending to about four-fifths of elytral length. Abdomen: disc smooth, shining, punctures minute; basal striae of tergite 1 elongate, one-half paratergite length, slightly divergent, separated at base by about one-half tergite width, setose brush distinct between bases of striae. MALE: Antennae and trochanters unmodified, simple. Metaventrite with area of longer setae only between metacoxae. Tergite 1 with shallow small emargination at apex; tergite 2 with short, shallow arcuate impression at base, at base small median triangular acutely pointed tubercle arises from short transverse carina ventral to apex of tergite 1; disc of 2 and following tergites smooth, shining, impunctate (Figs 24 A – B). In lateral view 1 nearly level, projecting posteriorly and above base of 2, 2 – 4 confluently curved. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical third with brush present (Fig. 24 D). Ventrites broadly convex. Aedeagus 0.33 long mm; with dorsal plate broadly triangular, apex bluntly rounded; parameres with three thin setae at posterior margin of preapical constriction, hyaline setae flattened, elongate, tapering from near base to acute point; internal sac with three thick spines, two strongly curved (Fig. 24 E). FEMALE: Microreticulation on elytra distinct. Metaventrite with setae sparse. Abdomen lacking modifications; ventrites broadly convex. Metatibiae similar to that of male. Collecting data. Three specimens were taken in August, with two of these being sifted from litter.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC2FFCEE1D2F8BBE1E8FBAF.taxon	distribution	Distribution (Fig. 41 D). Only known from higher forested elevations in two distant areas of Arizona.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC2FFCEE1D2F8BBE1E8FBAF.taxon	discussion	Comments. Brendel originally placed this species in the cavicornis species-group (Nisa) due to its relatively small median prothoracic fovea. However, this species lacks the punctate dorsum typical of the cavicornis speciesgroup, and resembles more closely B. arguta and others of this species group in the shining cuticle, long basal discal striae of tergite 1, the simple apex of tergite 1, and its small size. It is separated from the other species in this group by the comparatively simple tergite 2, where the median basal impression is faint and the median triangular spine at the base is clearly visible in posterior view.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC0FFC9E1D2FBE8E036FBE7.taxon	description	(Figs 25, 41 C)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC0FFC9E1D2FBE8E036FBE7.taxon	materials_examined	Material examined, 224 specimens. CANADA: Ontario: Carleton Co.: Osgood, X- 28 - 1967, A. Smetana (MHNG, 4); X- 20 - 1967, A. Smetana (MHNG, 2). Hastings Co.: Trenton, VII- 18 - 1908, Evans (CNCI, 1). Kent Co.: Rondeau Provincial Park, beach nr. Entrance, VI- 3 - 1985, A. Davies & J. M. Campbell, in debris on beach at high water line (CNCI, 3). Lanark Co.: Pakenham Mountains, nr. White Lake, V- 13 - 1987, A. Davies & L. Lesage, ex muskrat nest (DSCC, 2); V- 13 - 1987, A. Davies & L. Lesage (CNCI, 4). Leed & Greenville Co.: Chaffeys Locks, X- 16 - 1986, A. Davies, alder & sphagnum litter in marsh (CNCI, 1), alder / birch litter, marsh edge (DSCC, 2). Chaffeys Locks Biol. Station, VI- 18 - 1986, A. Davies, sifted moss & litter (CNCI, 5). Nipissing District: Algonquin Provincial Park nr. Brent, VIII- 20 - 1980, R. Baranowski (DSCC, 1). Norfolk Co.: Longpoint, VI- 18 - 1972, A. & A. Morgan (JPC, 1). Normondale, VI- 14 - 1931, W. J. Brown (CNCI, 1; CUDB, 1). Simcoe Co.: Simcoe, VI- 13 - 1931, W. J. Brown (CNCI, 1). Sudbury District: Sudbury, Laurentian University, VI- 10 / 16 - 1982, R. DeClerck, FITrap (CNCI, 1). Toronto City: Toronto, R. J. Crew (LEMC, 1). Welland Co.: Ridgeway (MCZC, 1; SEMC, 2). Wentworth Co.: Ancaster, IX- 1 - 1957, J. E. H. Martin (CNCI, 1). York Region: Wilcox Lake, III- 28 - 1902 (LEMC, 1). Only province on label: “ Ontario ”, Sherman (CASC, 1). Quebec: Bellechasse MRC (Municipalité Régionale de Comté): St. - Damien, XI- 9 - 1975, F. Liard (LLC, 1); St. - Damien Berth., V- 4 - 1977, C. Chantal, hutte à castor (FMNH, 24). Brome-Mississiquoi MRC: Mississiquoi, St. - Armand O., V- 11 - 2001, C. Chantal (CCC, 1). L’Érable MRC: 4 mi W Marsham, nr. Mud Lake, VIII- 17 - 1967, J. M. Campbell, berlese sample ex. lining old beaver lodge (CNCI, 2); X- 25 - 1967, berlese sample ex. lining old beaver lodge (CNCI, 2); X- 24 - 1967, J. M. Campbell & A. Smetana, berlese sample ex. lining old beaver lodge (CNCI, 3). Le Collines-de-l’Outaouais MRC: Fortune Lake, Gatineau Park, XI- 7 - 1967, J. M. Campbell & A. Smetana, ex: beaver lodge (CNCI, 1). Gatineau Park, Fortune Lake, XI- 11 - 1967, A. Smetana (MHNG, 2). Gatineau Park, near Mud Lake, X- 24 - 1967, A. Smetana (MHNG, 1). Lac Fortune, Parc Gatineau, VIII- 22 - 1976 (DSCC, 1); VIII- 15 - 1976, R. Sexton, hutte à castor (DSCC, 1); VIII- 29 - 1976 (DSCC, 1). Pontiac MRC: Blind Lake, Gatineau Park, XI- 11 - 1970, J. M. Campbell & S. Peck (CNCI, 1). Lac Holly, Parc Gatineau, IX- 11 - 1976, R. Sexton, hutte à castor (DSCC, 1). Ramsay Lake Area, IX- 1 - 1968, J. M. Campbell (CNCI, 1). Témiscamingue MRC: Laniel, VIII- 22 - 1932, W. J. Brown (CNCI, 3). Saskatchewan: Tp. 59 Rge. 22, W. 3 Meridian, VII- 30 - 1984, B. F. & J. L. Carr (CNCI, 1). Only country on label: “ Can. ” (CMNH, 6). USA: District of Columbia: “ D. C. ” (CMNH, 4; MCZC, 1). Illinois: Lake Co.: Sayer Bog, Volo, IV- 7 - 1959, W. Suter (FMNH, 4); IV- 21 - 1962, berlese leaf mold (FMNH, 1). Antioch, X- 27 - 1943, Ross & Sanders, sphagnum moss, tamarck bog (INHS, 15); Antioch bog, VI- 3 - 1960, W. Suter, sphagnum (FMNH, 1). Indiana: Only state on label: “ In. ” (FMNH, 1). Kentucky: Only state on label: “ Ky. ” (MCZC, 1). Maryland: Baltimore Co.: Butler, II- 12 - 1979, E. J. Ford, Jr., light trap (DSCC, 1). Montgomery Co.: Takoma Park, VIII- 29 - 1970, M. Druckenbrod, UV light (USNM, 1). Massachusetts: Middlesex Co.: Drac (ut), VII- 24 - 1888 (MCZC, 1). Lowell (MCZC, 1). Spencer Circle, 1 km E West Groton, VIII- 21 - 2011, T. Murray, light, photo (DSCC, 1). Only state on label: “ Mas. ” (DSCC, 1; MCZC, 1). New Hampshire: Coos Co.: White Mtns., Peabody River, VII- 11 - 1925, Quirsfeld (CNCI, 1). New Jersey: Bergen Co.: Hillsdale, XII- 4 - 1930 (CNCI, 1). Hudson Co.: Snake Hill (= Laurel Hill), VIII- 30 (FMNH, 1). Morris Co.: Towaco, XI- 7 - 1944, A. Nicolay, sifting leaves (USNM, 5). Passaic Co.: Mountainview, IV- 22 - 1939, A. Nicolay (USNM, 1). Union Co.: Elizabethtown (FMNH, 1). Roselle Park, XII- 9 - 1923, A. Nicolay (USNM, 5; ANSP, 1); IV- 27 - 1924 (MCZC, H. C. Fall Collection, 2); XI- 25 - 1923 (USNM, 10); IV- 4 - 1926 (USNM, 8); XI- 1 - 1925 (MCZC, H. C. Fall Collection, 3; UAIC, 1); X- 30 - 1927, Quirsfeld (CASC, 2; MCZC, H. C. Fall Collection, 1); XI- 29 - 1924 (CNCI, 1); XI- 9 - 1924, Quirsfeld (CNCI, 8); IV- 16 - 1911 (AMNH, 1); XII- 9 - 1923, Quirsfeld (CUIC, 1); XI- 1 - 1925, A. Nicolay (CUIC, 3). County unknown: Linton?, XI- 25 - 1923, sifting in swamp (AMNH, 2). New York: Albany Co.: Rensselaerville, SW E. N. Huyck Preserve, VIII- 27 - 1974, W. Suter, lake shore debris 4 - fern (DSCC 1; FMNH, 1). Erie Co.: Buffalo, VI- 23 - 1901 (DSCC, 1); Buffalo (SEMC, 1; INHS, 7). Greene Co.: Catskill, IV, Angell (MCZC, 1). Ontario Co.: 3 mi N Geneva, VII- 11 - 1983, M. T. Quinn, sift beech litter in wet depressions (DSCC, 1). Queens Co.: Flushing, L. (ong) I. (sland), VI- 3 - 1931, K. W. Cooper, sweeping (FMNH, 1). Tompkins Co.: Ithaca (USNM, 3); VI- 11 - 1939; P. P. Bably (MHNG, 1). McLean Bogs Preserve, Mud Pond, V- 8 / 15 - 2000, J. K. Liebherr, pitfall trap (CUIC, 1). Only state on label: “ N. Y. ” (CMNH, 3); Brend. (MCZC, Horn Collection, 1). Ohio: Ashtabula Co.: Trumbull, VI- 5 - 1976, D. S. Chandler, sift leaf litter along stream (DSCC, 1). Cuyahoga Co.: Fairview Park, Macbeth Avenue, VI- 29 - 1974, H. J. Lee, UV light (AMNH, 1). Huron Co.: Ridge Road sand quarry, North Fairfield, VII- 13 - 1974, H. J. Lee & T. C. Sheets, taken at ultraviolet light (DSCC, 1). Only state on label: “ Ohio ” (CMNH, 6; USNM, 2). Pennsylvania: Westmoreland Co.: Jeannette, X, H. G. Klages (CMNH, 1); X- 12 (CMNH, 1); X- 6 (CMNH, 1). West Virginia: Mineral Co.: Piney Swamp, 5 mi SW Keyser, V- 27 - 1993, S. M. Clark (WVDA, 1). Wisconsin: Grant Co.: VII- 1 - 1952 (FMNH, 1). Boscobel, VII- 11 - 1953, R. Shenefelt (DSCC, 2).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC0FFC9E1D2FBE8E036FBE7.taxon	description	Description. BODY: Length 1.68 – 1.80 mm; brown, some with elytra red-brown; setae on head, pronotum, and elytra short, curved and suberect, decumbent over rest of body. Head: surface smooth, shining, punctures indistinct. Antennomere III-VI longer than wide, VII as long as wide, VIII – X transverse, VIII about two-thirds width of IX (Fig. 25 E). Median vertexal fovea as wide as lateral foveae. Pronotum: surface smooth, shining, punctures indistinct. Median antebasal fovea with setose area as wide as those of lateral antebasal foveae. Elytra: with faint microreticulation, punctures indistinct, discal stria extending to about four-fifths of elytral length. Abdomen: basal striae of tergite 1 long, about one-half length of paratergites, separated at base by two-fifths tergite width at base, basal comb of setae between striae slightly denser in males than in females; disc smooth, shining, punctures indistinct. MALE: Antennae and trochanters not modified. Metaventrite with short triangular patch of longer, appressed setae between metacoxae. Tergite 1 with apex angled to become prominent at middle, appearing humped in lateral view, apical margin with shallow emargination at middle, lacking setal lamina; tergite 2 with deep V-shaped impression extending from tergite base to near apex, apical portion of impression with prominent margins, strongly protruding dorsally in lateral view, posterior margin of protuberance vertical; with two appressed setal tufts at apex of protruding margin, complex tubercle extending from base of impression at angle to meet apical margin of tergite 1; lateral portions of tergite 2 slightly flattened and coarsely punctate (Figs 25 A – D); in lateral view tergite 1 rising to apex, 2 gently curving dorsally with posterior face nearly vertical, base of 2 well-separated from 1. Abdominal ventrites broadly curved. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical sixth with brush present (Fig. 25 F). Aedeagus 0.32 mm long; dorsal plate triangular, margins evenly converging to acute point; parameres truncate at apex, three thick setae on lateral margin at angle of preapical constriction; hyaline seta on mesal margin, small, thickened to middle, with acute apex; internal sac with three thick spines, one short, none strongly curved (Fig. 25 G). FEMALE: Metaventrite with denser setae in area between metacoxae, sparse over rest of metaventrite. Abdominal tergites gently convex, lacking modifications; ventrites broadly convex. Metatibiae similar to that of male. Collection data: From alder and sphagnum litter in marshes; in beaver houses and muskrat nests; in beach debris; at UV light (Chandler 1997); also from leaf litter along streams. Habitat data from specimens indicates that this species is found in leaf litter along lake margins and in swamps / marshes. Several series have been taken from debris in beaver huts. This is a species of the Northeast and Midwest regions, with all collections but one being taken from March to November.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC0FFC9E1D2FBE8E036FBE7.taxon	distribution	Distribution (Fig. 41 C). In Canada found from Quebec to Saskatchewan, and in the United States documented from New Hampshire to West Virginia and Wisconsin. This species was reported from Connecticut (Bowman 1934) and Rhode Island (Sikes 2004), but the specimens upon which these records are based have not been examined.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC0FFC9E1D2FBE8E036FBE7.taxon	discussion	Comments. This species is similar to B. arguta and B. wickhami in sharing the small size, dark color, the long basal discal carinae of tergite 1, and have the male characters expressed at the middle of tergite 2. This species may be separated from the other two by the prominent, bluntly rounded projection that extends to the middle of tergite 2 (best seen in lateral view), with the area anterior to the tubercle deeply impressed and bearing a basal median projection, and the posterior margin nearly vertical. The other two species lack the prominently projecting median portion of the disc. Also, the median pronotal fovea is subequal in size to the lateral foveae in B. terebrata, while in the other two eastern species the median fovea is about two-thirds the size of the lateral foveae. Casey (1894: 479) proposed the name B. intricata for the smaller dark variety of B. dentata that Say (1824) mentioned in his description, while proposing B. terebrata as a replacement name in the same paper for the junior homonym B. perforata Brendel. Since B. terebrata was proposed to replace an older name, as first revisers we choose it as the most appropriate valid name to be used for this species.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC7FFCBE1D2FB20E7EAFE4A.taxon	description	(Figs 26, 41 B)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC7FFCBE1D2FB20E7EAFE4A.taxon	materials_examined	Material examined, 70 specimens. HOLOTYPE male: USA: Iowa: Johnson Co.: // Iowa City, Wickham, V- 22 - 98 / [yellow circle] 277. / F. C. Bowditch Coll. / [red-bordered label] Brachygluta arguta Csy. / [red label] HOLOTYPE Brachygluta wickhami Bückle // (MCZC). PARATYPES, 64: USA: District of Columbia: “ D. C. ” (CMNH, 1). Illinois: DuPage Co.: Argonne Natl. Lab., VI- 13 - 1967 (FMNH, 1 M), VIII- 1 - 1967 (FMNH, 1 M), W. Suter, at light. Iowa: Johnson Co.: Iowa City, IV- 3 - 1898 (FMNH, 2 M), IV- 7 - 1900 (DSCC, 2; FMNH, 2 M, 1 F; CUIC, 1), IV- 16 - 1899 (FMNH, 2 M, 4 F), V- 15 - 1898 (DSCC, 1 M; FMNH, 1 M, 1 F; MCZC, 1), V- 22 - 1898 (AMNH, 2; UNHC, 1 F; MCZC, H. C. Fall Collection, 2 M, 1 F; FMNH, 7 M, 4 F; MCZC, 2 M, 6 F; SEMC, 1 F), VI- 11 - 1898 (CUIC, 1; MCZC, H. C. Fall Collection, 1; FMNH, 1 M; MCZC, 1 F), Wickham; VI- 16 - 1898, W. (MCZC, 1 M); specimen without collector (FMNH, 1 M). Pennsylvania: Bedford Co.: Shellburg, VIII- 16 (CMNH, 6 M; DSCC, 1 M). Jefferson Co.: Cook State Forest, 1.2 mi N Cooksburg, Troutman Run, VI- 6 - 1997, D. S. Chandler, leaf litter along stream (DSCC, 1 M). West Virginia: Grant Co.: Hiser, V- 25 - 1994, S. M. Clark (WVDA, 1). Mason Co.: McClintic Wildlife station, VII- 26 - 1986, M. C. Thomas, UV trap (WVDA, 1). Morgan Co.: Widmeyer Wildlife Management Area, VIII- 21 - 1997, S. M. Clark (DSCC, 1). Females that lack associated males collected on the same day are: Iowa: Johnson Co.: Iowa City, IV- 10 - 1898 (FMNH, 1 F; MCZC, 2 F), IV- 24 - 1898 (DSCC, 1 F; FMNH, 1 F; SEMC, 1 F). These are not designated as paratypes, but are almost certainly members of this species.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC7FFCBE1D2FB20E7EAFE4A.taxon	description	Description. BODY: Length 1.72 – 2.00 mm; brown; setae on head and pronotum short, curved and suberect, over rest of body longer and decumbent. Head: surface smooth, shining, punctures indistinct. Antennomere III – VI longer than wide, VII as long as wide, VIII – X transverse, VIII about two-thirds width of IX. Setose area of median vertexal fovea as wide as those of lateral foveae. Pronotum: surface smooth, shining, punctures minute. Setose area of median antebasal fovea slightly smaller than those of lateral antebasal foveae. Elytra: surface smooth to with faint microreticulation, punctures indistinct, discal stria extending to about four-fifths of elytral length. Abdomen: basal striae of tergite 1 long, divergent, close to two-thirds length of paratergite in male, about half length of paratergite in female, separated at base by about two-fifths tergite width, with distinct brush of spaced setae between striae in male, brush indistinct with only sparse setae present in female; disc lightly punctate. MALE: Antennae and trochanters not modified. Metaventrite with transverse band of dense setae between metacoxae. Only tergites 1 – 2 visible in dorsal view, tergite 1 twice as long as 2; tergite 1 with disc smoothy and shallowly convex, apex slightly emarginate at middle, with row of appressed golden setae forming elongate flattened lamina, lamina shallowly emarginate at apex and directly posteroventrally; tergite 2 with semicircular impression at middle extending from base to or near apex, narrow median strip glabrous, lateral portions of impression with setae suberect, directed obliquely medially, denser and more elongate than on lateral portions of tergite; lateral portion of tergite 2 with flattened or lightly impressed coarsely punctate area, setae in this area much shorter and appressed; apex of tergite straight posterior to impression; tergites 3 – 5 evenly convex, not modified; in lateral view tergite 1 rising to deflexed apex, 2 directed posteriorly, 2 – 4 curved confluently, with distinct gap between base of 2 and apex of 1. Abdominal ventrites broadly convex. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical fourth with brush present. Aedeagus 0.36 long mm; with parameres curved laterally in apical portion, with distinct preapical constriction, with three aciculate preapical setae on lateral margin, with narrow, flat, hyaline seta on inner margin near apex; dorsal plate usually slightly asymmetric, roughly triangular to narrowing to acute apex, apex curving slightly to either side; internal sac with two large isolated and slightly curved spines (Figs 26 A – C). FEMALE: Metaventrite convex, setae sparse over disc, but denser at middle than on lateral portions of metaventrite. Abdominal tergites evenly convex; ventrites broadly convex. Metatibiae similar to those of male. Collecting data. There are few habitat / collection records for this species, with one specimen being sifted from leaf litter by a stream, and three specimens were collected at light. Taken from April to August, with most of the specimens collected in April and May.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC7FFCBE1D2FB20E7EAFE4A.taxon	distribution	Distribution (Fig. 41 B). A species of the Midwestern and Northeastern states, and found in two general areas at this time: Iowa and Illinois, and from Pennsylvania to West Virginia plus the District of Columbia. Most of the specimens were taken in the Iowa City area by H. F. Wickham over a three year period, where 57 of the 71 specimens examined were collected.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC7FFCBE1D2FB20E7EAFE4A.taxon	discussion	Comments. This species is similar to B. arguta, and only the males can be separated by using subtle differences in the form of tergite 2, and by examining the male genitalia. The median impression of tergite 2 tends to reach the apex of the tergite, and the setae on the inner margins of the impression are denser and longer. The median lobe of the genitalia is shorter and broader, and the internal sac lacks a multi-dentate tooth.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC7FFCBE1D2FB20E7EAFE4A.taxon	etymology	Etymology. Named for H. F. Wickham, who collected most of the type series, and whose collaboration with Emil Brendel produced one of the most important treatments of the North American Pselaphinae.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC5FFB5E1D2FC06E253FD4F.taxon	description	(Figs 27, 38 B)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC5FFB5E1D2FC06E253FD4F.taxon	materials_examined	Material examined, 22 specimens. USA: Texas: Cameron Co.: Brownsville, V- 10 - 1908 (MCZC, 2); Esperanza Ranch, Brownsville, VIII (USNM, 1). Harlingen, VIII- 30 - 1945, D. E. Hardy & V. L. Wooley, at light (FMNH, 2). Collingsworth Co.: 7 mi N Wellington, VI- 26 - 1995, Godwin, Marqua & Wolfe, UV light (TAMU, 1). San Patricio Co.: Lake Corpus Christi, 3 mi S Swinney Switch, VIII- 22 - 1974, C. W. O'Brien, UV trap (DSCC, 2). Travis Co.: Tex [dot under " e "] (= Austin; Lectotype and 6 Paralectotypes, USNM). Val Verde Co.: Del Rio, 955 ', VII- 23 / 24, Wickham (AMNH, 1). Only state on label: “ Tex. ” (CMNH, 1); Brend, Horn Coll H 9383 (MCZC, 3, possible syntypes); Casey, (MCZC, 1, Paralectotype). MEXICO: Nuevo León: 29 km W Linares, Santa Rosa Can. (yon), VI- 3 - 1983, S. Peck & M. Kaulbars, UV light (DSCC, 1).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC5FFB5E1D2FC06E253FD4F.taxon	description	Description. BODY: Length 1.76 – 2.12 mm; orange-brown; setae over dorsum short, curved and suberect, appressed on ventral surface. Head: surface shining and smooth to opaque, microreticulate and appearing roughened in a few, punctures indistinct. Antennomeres III – VII longer than wide, VIII as long as wide, IX elongate, X slightly longer than wide, VIII about one-half width of IX (Fig. 27 C). Setose area of median vertexal fovea as wide as those of lateral foveae. Pronotum: surface smooth and shining with punctures indistinct, to opaque due to dense punctation. Setose area of median antebasal fovea same size as those of lateral antebasal foveae. Elytra: with faint microreticulation, punctures indistinct, discal stria extending to about four-fifths of elytral length. Abdomen: tergite 1 disc smooth, shining, punctures indistinct; basal striae about one-fourth paratergite length, apices difficult to distinguish, striae separated at base by one-half tergite width, with distinct setose brush between bases of striae. MALE: Antennae and trochanters lacking modifications. Metaventrite with triangular patch of dense longer setae extending anteriorly from between metacoxae to middle of metaventrite. Tergite 1 with disc convex, apex straight, lacking modifications (Fig. 27 A), in lateral view evenly curving to meet base of tergite 2; tergite 2 lacking obvious modifications, across most of base with low carina that is covered by apex of 1, not visible in dorsal view and challenging to recognize in posterodorsal view. Ventrites broadly convex except for ventrite 3, ventrite 3 convex in middle portion, laterally with posterior half of forming smooth ridge, anterior to ridge with deep transverse sulcus (Fig. 27 B). Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical fifth with brush present (Fig. 27 D). Aedeagus 0.32 mm long; with dorsal plate triangular; parameres curved laterally in apical portion, margins even in width to apex, with three preapical large setae on lateral margin, preapical hyaline seta near mesal margin, thickened to middle and aciculate; internal sac with three short spines, and complex broad multispined structure (Figs 27 E – F). FEMALE: Similar to male; metaventrite with apical triangular setose patch present but less obvious than in male. Metatibiae similar to that of male. Ventrites broadly curved. Collection data: Three of the records indicate collection by use of ultraviolet light. Taken from May to August.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC5FFB5E1D2FC06E253FD4F.taxon	distribution	Distribution (Fig. 38 B). There is one record from the northern portion of Nuevo León, Mexico, with the rest of the records being from central and southern Texas.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFC5FFB5E1D2FC06E253FD4F.taxon	discussion	Comments. This species is unique in lacking both antennal and dorsal abdominal modifications in males, and is the only species where the male characters are ventrally placed. In males the lateral portion of ventrite 3 is smoothly ridged posteriorly and transversely sulcate at the base. Also, the metasternal triangular setose patch is distinct with dense setae; in females the metasternal setose patch is not as obvious. Casey (1894: 478) lamented that the specimens in his type series seemed to be all females, but ironically five of the 7 present at the USNM are males. The odd large multispined structure of the internal sac is shared with B. corniventris, a species that has complex male characters on tergites 1 – 2, and is otherwise quite dissimilar from B. infinita. Casey indicated that he had 14 specimens from Austin, Texas as the basis for his original description, with only seven of these now present at the USNM. From these a male is here designated as the Lectotype: // Tex [dot under " e "] / Casey bequest 1925 / [red label] TYPE USNM 38692 / infinita Casy / [red label] LECTOTYPE Bryaxis infinita Casey ♂ ‘ 13 des. DSChandler //. Five other specimens with similar label data, and one lacking a “ Tex ” and USNM type labels are labeled as Paralectotypes. Seven specimens of Casey’s type series are presumed to have been dispersed to other collections. Brendel & Wickham (1890: 268) commented on B. infinita, indicating that they had seen some of Casey’s type material, and placed his species as a junior synonym of B. belfragei. One male is present in the MCZC collection, where the major portion of Brendel’s collection is held (Brendel’s primary type specimens reside in the ANSP), with a “ Casey ” label and identified as B. belfragei. Three other specimens, two males and a female, are present in the Horn Collection at the MCZC, bearing the labels “ Tex ” and “ Brend ”, and identified as B. infinita. The specimen with the “ Casey ” label is here labeled as a paralectotype, and the other three are labeled as possible syntypes.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBBFFB7E1D2FBB2E16AFEFF.taxon	description	(Figs 28, 42 D)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBBFFB7E1D2FBB2E16AFEFF.taxon	materials_examined	Material examined, HOLOTYPE male: MEXICO: Baja California: // MEXICO: Baja Calif. Norte, Isla San Esteban, main canyon wash, east side, VII- 28 / 29 - 86, R. L. Aalbu., BL / [red label] HOLOTYPE Brachygluta seri Chandler (CSCA).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBBFFB7E1D2FBB2E16AFEFF.taxon	description	Description. BODY: Length 2.20 mm; brown with elytra and appendages orange-brown; setae over body decumbent, longest on abdomen. Head: surface smooth, shining, lightly microreticulate, punctures indistinct. Antennomeres III – VI longer than wide, VII as long as wide, VIII – X transverse, VIII about two-thirds width of IX (Fig. 28 C). Setose area of median vertexal fovea as large as those of lateral foveae. Pronotum: surface shining, lightly microreticulate, punctures indistinct. Setose area of median antebasal fovea about two-thirds width of lateral antebasal foveae. Elytra: microreticulate, surface granulate, punctures indistinct, discal stria faint, extending to about three-fourths of elytral length. Abdomen: disc with small, distinct punctures, basal two tergites opaque due to distinct microreticulation, apical tergites more shining; basal striae of tergite 1 short / indistinct, separated at base by about three-fifths tergite width; setose comb at base between basal striae sparse. MALE: Antennae and trochanters lacking modifications, simple. Metaventrite with rounded-triangular area extending anteriorly from metacoxae to anterior portion of disc, area with dense long setae, metasternal disc laterally with setae distinct but becoming sparser. Tergite 1 distinctly microreticulate across disc, apical margin slightly concave, with broad impression in basal third of disc (Fig. 28 A); in lateral view tergite apex slightly protruding out over tergite 2; tergite 1 only slightly longer than tergite 2. Tergite 2 with semicircular basal carina in median fourth just covered by apex of tergite 1; tergites 2 - 3 slightly and broadly impressed at middle across width (Fig. 28 B). Ventrites broadly rounded. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical fifth with brush present (Fig. 28 D). Aedeagus 0.40 mm long; dorsal plate rounded triangular, margins evenly converging to bluntly rounded point; parameres rounded at apex, three thick setae on lateral margin at angle of distinct preapical constriction; thickened hyaline seta on mesal margin, longer than preapical setae, thickened to middle, with acute apex; internal sac with four curved large spines (Fig. 28 E). FEMALE: Unknown. Collecting data. The holotype was taken at blacklight in July.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBBFFB7E1D2FBB2E16AFEFF.taxon	distribution	Distribution (Fig. 42 D). Found on a desert island near the middle of the Gulf of California.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBBFFB7E1D2FBB2E16AFEFF.taxon	discussion	Comments. This species shares with B. cochimi the relatively long tergite 2, which in that species is clearly two-thirds the length of tergite 1, and their occurrence on islands in the Gulf of California. It may be separated from B. cochimi by the distinct basal depression of tergite 1 and the lack of a discal impression on tergite 2; for B. cochimi the disc of tergite 1 is smoothly convex to the base, and tergite 2 has a visible transverse basal impression.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBBFFB7E1D2FBB2E16AFEFF.taxon	etymology	Etymology. The name originates from the Seri indigenous people, which historically have lived on San Esteban Island in the Gulf of California.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB9FFB7E1D2FD5AE746F8D2.taxon	description	(Figs 29, 42 A)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB9FFB7E1D2FD5AE746F8D2.taxon	materials_examined	Material examined, HOLOTYPE male: USA: Texas: Brewster Co.: // Big Bend N. P., TEX. 1850 ', Boquillas, May 17, 1959. Light, Howden & Becker / [red label] HOLOTYPE Brachygluta chisos Sabella // (CNCI).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB9FFB7E1D2FD5AE746F8D2.taxon	description	Description. BODY: Length 1.84 mm; orange-brown with elytra lighter; head and pronotum with setae short, decumbent, over rest of body longer. Head: surface smooth, shining, punctures indistinct. Antennomeres III – VI longer than wide, VII, VIII slightly transverse, IX as long as wide, X wider than long, VIII about four-fifths width of IX (Fig. 29 C). Setose area of median vertexal fovea as large as those of lateral foveae. Pronotum: surface smooth, shining, punctures indistinct. Setose area of median antebasal fovea about two-thirds width of lateral antebasal foveae. Elytra: with faint microreticulation, punctures indistinct, discal stria extending to about fourfifths of elytral length. Abdomen: disc smooth, shining, punctures indistinct. Basal striae of tergite 1 about onethird paratergite length, slightly divergent, separated at base by one-half tergite width, setose brush between bases of striae distinct, long. MALE: Antennae and trochanters unmodified, simple. Metaventrite with middle of disc flattened through length, with dense long setae in this area; tergite 1 with disc smoothly swollen to middle, apex broadly and smoothly convex (Fig. 29 A), in lateral view apex of tergite curving down to disc of tergite 2; tergite 2 with mediobasal area semicircular, roughened and slightly impressed, about one-third width of disc, with low transverse carina at base just below apex of tergite 1, carina about one-third tergite width (Fig. 29 B), in lateral view with disc of 1 confluently curved with following tergites. Abdominal ventrites broadly rounded. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical fourth with brush present (Fig. 29 D). Aedeagus 0.40 mm long; dorsal plate with broad triangular base, margins in apical half constricted to form sharply pointed curved apex; parameres with distinct preapical constriction, three thick setae on lateral margin at basal angle of constriction, flattened preapical hyaline setae near mesal margin widening to middle, apex aciculate; internal sac with two large spines, both only slightly curved (Fig. 29 E). FEMALE: Unknown. Collecting data. The holotype was collected in May at light.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB9FFB7E1D2FD5AE746F8D2.taxon	distribution	Distribution (Fig. 42 A). This species is known only from western Texas (Big Bend National Park).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB9FFB7E1D2FD5AE746F8D2.taxon	discussion	Comments. This species shares with B. kiowa the indistinct / flattened / impressed roughened area in the basal portion of the disc of tergite 2, but in B. chisos this area is narrower and extends only to the middle of the disc, while in B. kiowa it extends nearly the length of the tergite and two impressions are often distinct at the middle of this area.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB9FFB7E1D2FD5AE746F8D2.taxon	etymology	Etymology. The name is that of one of the indigenous groups known from this area.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBEFFB2E1D2F947E3D0FBAF.taxon	description	(Figs 30, 42 C)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBEFFB2E1D2F947E3D0FBAF.taxon	materials_examined	Material examined, 6 specimens. HOLOTYPE male: MEXICO: Baja California: // MEXICO: Baja Calif. Norte, Isla San Lorenzo, SW cyn. wash, VII- 27 / 28 - 86, Rolf L. Aalbu col., BL / [red label] HOLOTYPE Brachygluta cochimi Bückle // (CSCA). PARATYPES, 5 males: eutopotypical (CSCA, 3; DSCC, 2). A female from Baja California, Bahía de los Ángeles, VII- 19 - 1978, D. W. Anderson (UCBC) compares well with the type series.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBEFFB2E1D2F947E3D0FBAF.taxon	description	Description. BODY: Length 2.00 – 2.16 mm; orange-brown with elytra sometimes lighter; head and pronotum with setae short, curved and suberect, over rest of body setae appearing denser, longer and decumbent. Head: surface smooth, shining, punctures indistinct. Antennomeres III – VI longer than wide, VII as long as wide, VIII transverse, IX as long as wide, X slightly wider than long, VIII about two-thirds width of IX (Fig. 30 C). Setose area of median vertexal fovea as large as those of lateral foveae. Pronotum: surface smooth, shining, punctures indistinct. Setose area of median antebasal fovea about two-thirds width of lateral antebasal foveae. Elytra: with microreticulation, punctures varying from indistinct to small and distinct, discal stria extending to about four-fifths of elytral length. Abdomen: disc with small, distinct punctures, opaque in apical half due to distinct microreticulation, basal half shining; basal striae of tergite 1 short, one-fifth to one-fourth paratergite length, divergent, separated at base by about one-half tergite width; setose comb at base between basal striae distinct. MALE: Antennae and trochanters lacking modifications, simple. Metaventrite with circular patch of dense short silvery setae extending from between metacoxae to middle. Tergite 1 distinctly microreticulate in apical half, apical margin convex; in lateral view slightly protruding out over tergite 2. Tergite 2 with weak slightly arcuate carina in middle third ventral to apex of tergite 1, base impressed anterior to carina, shallow longitudinal impression posterior to carina extending to tergite apex, impression shining but smoothly roughened (Fig. 30 A); tergites 3 – 4 with punctures sparse and distinct; tergite 4 with lateral margins protruding, bluntly tuberculate, with tuft of long setae at apices, tergite apex nearly subtruncate; tergite 5 with long setae directed posteromedially from lateral thirds (Figs 30 A – B); in lateral view tergite 1 rising to apex, projecting out above base of 2, 2 – 4 confluently curved. Ventrites broadly rounded. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical fifth with brush present (Fig. 30 D). Aedeagus 0.34 mm long; dorsal plate triangular, margins evenly converging to bluntly rounded point; parameres rounded at apex, three thick setae on lateral margin at angle of preapical constriction; thickened hyaline seta on mesal margin, longer than preapical setae, thickened to middle, with acute apex; internal sac with two curved large spines (Fig. 30 E). FEMALE: [one female tentatively placed as this species] Lacking modifications of abdomen, otherwise similar to male; metaventrite sparsely and evenly setose, setae similar to others on abdomen. Collecting data. The type series was taken at blacklight in July on a desert island.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBEFFB2E1D2F947E3D0FBAF.taxon	distribution	Distribution (Fig. 42 C). Found on the east coast of Baja California near the middle of the Gulf of California.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBEFFB2E1D2F947E3D0FBAF.taxon	discussion	Comments. This species is unusual in having tergites 4 – 5 modified. Tergite 4 has a bluntly protruding and apically setose tubercle at each lateral margin. Tergite 5 has long setae on the lateral portions of this sclerite directed posteromedially. The gently rounded apical margin of tergite 1, and the impression of tergite 2 defined posteriorly by a slight carina further serve to identify this species. It shares with B. seri a relatively long tergite 2, and its collection location from an island in the middle of the Gulf of California, but B. seri has tergites 1 and 2 subequal in length, and lacks the modifications of tergites 2, 4, and 5.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBEFFB2E1D2F947E3D0FBAF.taxon	etymology	Etymology. Named for an American indigenous group of central Baja California where this species was found.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBCFFBCE1D2FBE8E20FFD3B.taxon	description	(Figs 31, 43 A)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBCFFBCE1D2FBE8E20FFD3B.taxon	materials_examined	Material examined, 18 specimens. HOLOTYPE male: USA: Kansas: Meade Co.: // KANSAS: Meade Co., Jct. Cimarron Riv. & Hwy, 23. VI- 25 - 1992, E. Riley, night sweep / [red label] HOLOTYPE Brachygluta kiowa Sabella / / (TAMU). PARATYPES, 17: eutopotypical (DSCC, 5 M, 1 F; TAMU, 5 M). Colorado: Fremont Co.: Canon City, IV- 2 - 1900 (FMNH, 2 M). New Mexico: Chaves Co.: Bottomless Lakes S. (tate) P. (ark), VIII- 7 - 1989, Y. Bousquet, gypsum flats (CNCI, 1 M). Texas: Comanche Co.: Proctor and nearby farms, VI- 24 - 1976, J. W. Smith & A. R. Hardy (CSCA, 1 M). Hemphill Co.: Gene Howe WMA, V- 20 - 2006, R. Turnbow, mv + bl (DSCC, 1 M). Randall Co.: Palo Duro Cyn. St. Pk., VI- 25 - 1995, Godwin, Marqua, & Wolfe, UV (TAMU, 1 M).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBCFFBCE1D2FBE8E20FFD3B.taxon	description	Description. BODY: Length 1.72 – 2.00 mm; orange-brown to brown; setae on head and pronotum short, curved and suberect, over rest of body longer and decumbent. Head: surface smooth, shining, punctures indistinct. Antennomeres III-VI longer than wide, VII as long as wide, VIII slightly transverse, IX as long as wide, X slightly longer than wide, VIII about two-thirds width of IX (Fig. 31 D). Setose are of median vertexal fovea slightly smaller than those of lateral foveae. Pronotum: surface smooth, shining, punctures indistinct. Setose area of median antebasal fovea varying from about same size to about two-thirds size of lateral antebasal foveae. Elytra: with faint microreticulation, punctures indistinct, discal stria extending to about four-fifths of elytral length. Abdomen: disc smooth, shining, punctures indistinct; basal striae of tergite 1 about one-fourth paratergite length, slightly divergent, separated at base by one-half tergite width; setose brush between bases of striae short. MALE: Antennae and trochanters not modified. Metaventrite with triangular setose patch arising between metacoxae present but not distinct. Tergite 1 with apex deflexed in middle third, appearing slightly emarginate at middle, setae along apex longest in areas of deflexed area and obscuring it (Figs 31 A – B), in lateral view tergite 1 only slightly protruding above base of tergite 2; tergite 2 with slightly arcuate transverse carina at base below apex of tergite 1, area anterior to carina impressed, disc at middle posterior to carina with two shallow longitudinal impressions that diverge slightly toward apex, impressions vary from distinctly bifoveate to obscure (Figs 31 A – C); in lateral view tergite 1 with disc convex, 2 flat, 1 – 4 close to confluently curved. Abdominal ventrites broadly rounded. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical fifth with brush present (Fig. 31 E). Aedeagus 0.42 mm long; with dorsal plate abruptly narrowed in apical half to form elongate thin finger-like lobe; parameres with distinct preapical constriction, with three thickened setae on lateral margin at anterior angle of constriction, flattened hyaline seta near mesal margin, thickened to middle and aciculate to apex; internal sac with two large curved spines (Fig. 31 F). FEMALE: Lacking modifications of the abdominal tergites, otherwise similar to the male, including presence of the metasternal patch of setae. Collecting data. The one series was collected by sweeping at night along a river; individuals were taken due to their attraction to ultraviolet light, and one was found in an area of gypsum flats. Specimens were taken from April to August.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBCFFBCE1D2FBE8E20FFD3B.taxon	distribution	Distribution (Fig. 43 A). Collected in the shortgrass prairie area of the southern portion of the plains states from Kansas to west Texas.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBCFFBCE1D2FBE8E20FFD3B.taxon	discussion	Comments. This species is distinct in the form of male tergites 1 and 2. Tergite 1 has the apical margin slightly deflexed at the middle, and tergite 2 has a quadrate impression of varying distinctness, defined laterally by the two shallow longitudinal impressions. The two lateral impressions in the broader impression of the basal portion of male tergite 2, plus its distribution in the high plains near the Rocky Mountains leads to the ready recognition of this species. Closest is B. chisos, which has a semicircular basal area of tergite 2 roughened and indistinct flattened / modified, but this area reaches only to the midpoint of the disc and the discal impressions of B. kiowa are suggested but are much shorter.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFBCFFBCE1D2FBE8E20FFD3B.taxon	etymology	Etymology. The name is based on the name of one of the native American groups that lived in the area that this species occurs.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB2FFBEE1D2FD41E7CBFEDB.taxon	description	(Figs 32, 42 B)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB2FFBEE1D2FD41E7CBFEDB.taxon	materials_examined	Material examined, 33 specimens. HOLOTYPE male: USA: Oregon: Klamath Co.: // Oregon: Kla. Co., Lower Klamath L., II- 4 / 22 – 1963 / JSchuh, under rock pile w / ants / [red label] HOLOTYPE Brachygluta klamath Bückle // [with ant on point] (AMNH). PARATYPES, 32: USA: Oregon: Harney Co.: Harney Lake Hot Springs, 28. V. 1957, H. S. Dybas leg., along margin (UCDC, 1 M). Klamath Co.: eutopothypical, (DSCC, 6 M, 18 F); Lower Klamath Lk., IV- 16 - 1962, J. Schuh, rock outcrop (DSCC, 1 M, 2 F). Lake Co.: SE shore L. (ake) Abert, fresh pool, 17 - VII- 50, H. B. Leech Collector (UCDC, 1 M). Abert Lake, VI- 7 - 1958, Vertrees & Schuh (UCDC, 2 M). California: Siskiyou Co.: Indian Tom Lake, 17 mi SW of Klamath Falls (Oregon), June 19, 1978, L. & N. Herman # 1483 (AMNH, 1 M).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB2FFBEE1D2FD41E7CBFEDB.taxon	description	Description. BODY: Length 2.16 – 2.28 mm; red-brown with orange-brown elytra; head and pronotum with setae short, curved and suberect, over rest of body longer and decumbent. Head: surface smooth, shining, punctures indistinct. Antennomeres III-VI longer than wide, VIII – IX as long as wide, X longer than wide, VIII about twothirds width of IX (Figs 32 D – E). Setose area of median vertexal fovea as large as those of lateral foveae. Pronotum: surface smooth, shining, punctures indistinct. Setose area of median antebasal fovea about two-thirds width of lateral antebasal foveae. Elytra: with faint microreticulation, punctures indistinct, discal stria extending to about four-fifths of elytral length. Abdomen: disc smooth, shining, punctures indistinct. Basal striae of tergite 1 about one-fourth paratergite length, straight, separated at base by one-half tergite width, setose brush between bases of striae composed of thin sparse setae in both sexes. MALE: Antennae and trochanters unmodified, simple. Metaventrite with elongate-arcuate area of dense setae projecting anteriorly from between metacoxae, setose area lightly depressed; tergite 1 broadly and shallowly emarginate, middle third slightly deflexed, setae along apex relatively sparse, not denser at emargination (Fig. 32 A), in lateral view apex of tergite 1 only slightly protruding over base of tergite 2; tergite 2 with low carina at base, carina appearing slightly sinuate, carina beneath apex of tergite 1 and not visible in dorsal view, middle portion of carina lacking, disc with flattened longitudinal area at middle extending from carina to tergite apex, sometimes more distinct as slight impression (Figs 32 B – C); in lateral view with apex of 1 slightly elevated above base of 2, all confluently curved. Abdominal ventrites broadly rounded. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical fourth with brush present (Fig. 32 F). Aedeagus 0.40 mm long; with dorsal plate triangular, margins in apical half concave, apex bluntly pointed; parameres with distinct preapical constriction, three thick setae on lateral margin at basal angle of constriction, flattened preapical hyaline setae near mesal margin widening to middle, apex aciculate; internal sac with two large spines, one strongly curved, one straight except at base (Fig. 32 G). FEMALE: Metaventrite impressed between metacoxae, but setae not dense, similar to rest of metaventrite; lacking abdominal modifications. Otherwise similar to male. Collecting data. Found around the margins of lakes and hot springs; one large series was taken with Pogonomyrmex ants from beneath rocks. Taken from February through July.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB2FFBEE1D2FD41E7CBFEDB.taxon	distribution	Distribution (Fig. 42 B). South-central Oregon and adjacent California, primarily within the Klamath River drainage.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB2FFBEE1D2FD41E7CBFEDB.taxon	discussion	Comments. The lack of a distinct transverse basal impression places this species near B. chisos. The medianly flattened tergite 2, together with the basal carina that is lacking medially is distinct and slightly lobed laterally, serves to distinguish this species from all others.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB2FFBEE1D2FD41E7CBFEDB.taxon	etymology	Etymology. Named for the group of American Indians that live in this area of the United States.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB0FFBEE1D2FE34E100F82C.taxon	description	(Figs 33, 43 C)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB0FFBEE1D2FE34E100F82C.taxon	materials_examined	Material examined, 20 specimens. HOLOTYPE male: MEXICO: Sinaloa: // MEXICO: Sinaloa, 2 mi. E San Blas, VII- 5 - 82, F. Andrews, Blacklight / [red label] HOLOTYPE Brachygluta mayo Sabella // (CSCA). PARATYPES, 19 males: MEXICO: Baja California Sur: Isla Magdalena, Howlands Lagoon, VII- 6 - 1983, D. Faulkner (CSCA, 1). Nayarit: San Blas, VI- 15 - 55, B. Malkin (UCDC, 1). Sinaloa: eutopotypical (CSCA, 6; DSCC, 2); same data but Fred G. Andrews, coll. blacklight (CSCA, 4; DSCC, 1). Culiacán, 19 mi. S., VI- 22 - 63, J. Doyen Collector (UCBC, 1). Sonora: San Carlos Bay, VIII- 10 - 60, P. H. Arnaud, Jr., E. S. Ross, D. C. Rentz (CASC, 2).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB0FFBEE1D2FE34E100F82C.taxon	description	Description. BODY: Length 1.88 – 2.00 mm. Body orange-brown to brown, when brown often with elytra orange-brown; head and pronotum with setae short, curved and suberect, on rest of body longer and decumbent. Head: surface smooth, shining, punctures small, distinct. Antennomeres III – VII longer than wide, VIII slightly transverse, IX about as wide as long, X about as long as wide, VIII about two-thirds width of IX (Fig. 33 C). Setose area of median vertexal fovea as large as those of lateral foveae. Pronotum: surface smooth, shining, punctures small, distinct. Setose of median antebasal fovea about two-thirds width of lateral antebasal foveae. Elytra: microsculpture mostly lacking, surface smooth, shining, punctures small and distinct, discal stria extending to about four-fifths of elytral length. Abdomen: disc smooth, shining, punctures distinct; basal discal striae of tergite 1 at most one-fourth paratergite length, straight, separated at base by greater than one-half tergite width; setose brush between bases of striae long, setae sparse. MALE: Antennae and trochanters unmodified, simple. Metaventrite covered with long setae that are densest in arcuate area curving anteriorly from between metacoxae. Tergite 1 with small shallow emargination at middle of apex to apex appearing flattened; tergite 2 with arcuate impression extending from base to near apex, impression about three-fifths width of tergite, with line of anteriorly directed thick setae in impression positioned near lateral margins in impression, rounded carinate tubercle projecting postero-dorsally ventral to apex of tergite 1, seldom visible in dorsal view (Figs 33 A – B); in lateral view tergite 1 slightly protruding over base of tergite 2; in lateral view with tergite 1 disc slightly rising to deflexed apex, distinct drop from apex to flat tergite 2. Abdominal ventrites broadly rounded. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical third with brush present (Fig. 33 D). Aedeagus 0.44 mm long; with apex of dorsal plate broadly rounded; parameres with preapical constriction weakly developed, three thick setae on lateral margin at basal angle of constriction, flattened preapical hyaline setae near mesal margin gradually widening to middle, apex aciculate; internal sac with three large spines, one strongly curved (Fig. 33 E). FEMALE: Unknown. Collecting data. Taken at blacklight. Present from June to August.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB0FFBEE1D2FE34E100F82C.taxon	distribution	Distribution (Fig. 43 C). Found in coastal areas of Mexico from northern Sonora to Nayarit, and with one specimen taken in Baja California Sur.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB0FFBEE1D2FE34E100F82C.taxon	discussion	Comments. The male abdominal characters serve to place this species near B. mormon. They share the form of tergites 1 and 2, but the impression of tergite 2 is semicircular and a bit smaller, while the impression for B. mormon is wider and more transverse. The dorsal plate of the aedeagus is also different, with the apex broadly rounded in B. mayo, and acutely triangular in B. mormon.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB0FFBEE1D2FE34E100F82C.taxon	etymology	Etymology. The name is derived from the Mayo indigenous group centered in the range of this coastal species that extends from Sonora to Nayarit.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB1FFB9E1D2F890E2D2FE6F.taxon	description	(Figs 34, 43 B)	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB1FFB9E1D2F890E2D2FE6F.taxon	materials_examined	Material examined, 191 specimens. USA: Arizona: Yavapai Co.: Kirkland, VII- 14 - 1970, J. E. May, UV trap (UAIC, 1). County unknown: Grand Can (yon), VII- 4 - 1904 (MCZC, H. C. Fall Collection, 6). California: Inyo Co.: Panamint Mts., IV- 1891, Koebele (CASC, 36; FMNH, 2). Panamint V (alley) (CASC, 48). Panamint Valley, IV- 1891, Koebele (CASC, 4; USNM, 11). Panamint Mountains, IV- 1982, Koebele (USNM, 26). Surprise Canyon, V- 6 - 1966, Irwin & Cavagnaro (UCDC, 2). Nevada: Lincoln Co.: Alamo, VII- 10 - 1958, F. D. Parker (UCDC, 1); VII- 13 - 1958 (UCDC, 1). New Mexico: Only state on label: “ N. M. ” (CMNH, 1). Texas: El Paso Co.: El Paso, II- 27 - 1889 (MCZC, H. C. Fall Collection, 1); II- 21 - 1889 (MCZC, H. C. Fall Collection, 1); II- 19 - 1889 (MCZC, H. C. Fall Collection, 1). Utah: Cache Co.: Logan, VII- 4 - 1960, G. F. Knowlton (DSCC, 1). Juab Co.: Mills, V- 31 - 1993, B. F. & J. L. Carr, Lot 9 (DSCC, 2). Salt Lake Co.: Salt Lake (CMNH, 3 types); VI- 14, Hubbard & Schwarz (USNM, 4); VI- 25, Hubbard & Schwarz (USNM, 7); (USNM, 1, female, ID not confirmed). Salt Lake City (MNHN, 1). Tooele Co.: Black Pond, Dugway Proving Ground, V- 27 - 2003, R. W. Baumann & R. L. Johnson (BYUC, 1). Utah Co.: American Fork Canyon, VI- 21, Hubbard & Schwarz (USNM, 1). Goshen Pond, III- 23 - 1968, D. L. Hayes (BYUC, 1); IV- 9 - 1968, T. L. Pay (BYUC, 1); IV- 27 - 1968, R. E. Coombs, under rock (DSCC, 1); IV- 29 - 1967, W. Clark, under rock (TAMU, 3). Goshen Springs, II- 20 - 1967, W. Clark (TAMU, 2). Provo, III- 14 - 1965, J. Leetham (BYUC, 1). Utah Lake, VI- 19, Hubbard & Schwarz (USNM, 2); VI- 20, Hubbard & Schwarz (USNM, 2). Utah Lake, west shore, V- 23 - 1979, S. L. Wood & S. M. Clark (BYUC, 2). Utah Lake shore, II- 14 - 1966, D. Nichols (BYUC, 1). West edge Utah Lake, IV- 17 - 1965, W. J. Despain (BYUC, 3); IV- 17 - 1965, D. G. Gray (DSCC, 1; CUIC, 1). West shore Utah Lake, IV- 16 - 1965, J. Leetham (BYUC, 2). West Utah Lake, IV- 17 - 1965, S. L. Wood (BYUC, 3). Only state on label: “ Utah ” (CMNH, 1; FMNH, 1); Brendel (DSCC, 2).	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB1FFB9E1D2F890E2D2FE6F.taxon	description	Description. BODY: Length 1.68 – 1.92 mm; orange-brown, to some brown with orange-brown elytra; head and pronotum with setae short, curved and suberect, on rest of body longer and decumbent to appressed. Head: surface smooth, shining, punctures indistinct. Antennomeres III-VII longer than wide, VIII – IX transverse, VIII barely so (distinctly so in female), X transverse, VIII about two-thirds width of IX (Fig. 34 C). Setose area of median vertexal fovea as wide as those of lateral foveae. Pronotum: surface smooth, shining, punctures indistinct. Setose area of median antebasal fovea about two-thirds size of those of lateral antebasal foveae. Elytra: with faint microreticulation, punctures indistinct, discal stria extending to about four-fifths of elytral length. Abdomen: tergite 1 disc smooth, shining, punctures indistinct. Basal striae of tergite 1 about one-third paratergal length in males (one-half length in one male), about one-fifth in females, separated at base by one-half tergite width; setal brush between bases of striae distinct and thicker in males, indistinct and sparse in females. MALE: Antennae and trochanters not modified. Metaventrite with anteriorly rounded patch of denser setae in depressed area between bases of metacoxae, patch extending to middle of sternum. Tergite 1 slightly emarginate at middle, setae along apical margin longer and denser in area of emargination, in lateral view tergite 1 gently curving ventrally so that apical portion protrudes above base of tergite 2; tergite 2 semicircularly impressed from base, about three-fifths tergite width, impression not quite reaching apex of tergite 2, arcuate tubercle at base of tergite 2 ventral to emargination of tergite 1 and not visible in dorsal view (Figs 34 A – B); in lateral view disc of tergite 1 rising to apex, distinct drop from apex of 1 to flat tergite 2. Abdominal ventrites broadly rounded. Metatibiae thin, narrow at base, gradually widened to near apex, mesal margin of apical fourth with brush present (Fig. 34 D). Aedeagus 0.36 mm long; with dorsal plate irregularly triangular; parameres with distinct preapical constriction, with three thick setae on lateral margin at basal angle of constriction, flattened hyaline seta near mesal margin, thickened to middle and aciculate; internal sac with three thick spines, one curved (Fig. 34 E). FEMALE: Lacking tergal modifications; antennomeres VIII – IX slightly shorter than in male; otherwise similar to male. Collecting data. Taken beneath rocks by a pond. Collected from February through July.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB1FFB9E1D2F890E2D2FE6F.taxon	distribution	Distribution (Fig. 43 B). Present in high deserts from eastern California to northern Utah and southern New Mexico.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
03F187EDFFB1FFB9E1D2F890E2D2FE6F.taxon	discussion	Comments. This species is similar to B. mayo and shares the form of the male tergites. The dorsal plate of the aedeagus is triangular in B. mormon and the impression of tergite 2 is more transverse with the posterior margin nearly straight. In B. mayo the dorsal plate of the aedeagus is broadly rounded, and the posterior margin of the impression on tergite two is clearly semicircular. Antennomere X is transverse for B. mormon, while it is about as long as wide for B. mayo. In his description of this species Bowman (1934) stated there were seven females in the type series, and that males were unknown. However, the type series consists of two species, both with males present. Since one of the species (B. foveata) is already described, a Lectotype male was selected from the other species in the series. Lectotype male, here designated: // Salt Lake, Ut. / [light green label] Henry Ulke Beetle Coll. CMNH Acc. No. 1648 / [red label] LECTOTYPE Brachygluta mormon Bowman ♂ ‘ 13, des. DSChandler //. Three syntypes are members of this species, and two bear the data label “ Salt Lake, Ut., ” while the other is simply labeled “ Utah. ” Three other syntypes have a “ Utah ” label, and are a male and two females of Brachygluta foveata. One of the females of B. foveata has a label with “ Brachygluta mormon Bowman TYPE ” handwritten in pencil. This type designation was never published and is not valid. Since the note is attached to one of the females of B. foveata, this unpublished selection of Bowman is here noted but not followed. All six specimens are now labeled as paralectotypes of B. mormon.	en	Chandler, Donald S., Sabella, Giorgio, Bückle, Christoph (2015): A revision of the Nearctic species of Brachygluta Thomson, 1859 (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 3928 (1): 1-91, DOI: 10.11646/zootaxa.3928.1.1
