identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
296B422E59F70E7C2EA921B93C3B4CD7.text	296B422E59F70E7C2EA921B93C3B4CD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptobrachella wuhuangmontis Wang, Yang & Wang	<div><p>Leptobrachella wuhuangmontis Wang, Yang &amp; Wang sp. n. Figures 6, 7</p><p>Holotype.</p><p>SYS a003486, adult male, collected on 29 March 2015 by JW, ZTL, YYW and ZYL from Mt. Wuhuang (MWH hereafter) (22°08'30.77"N, 109°24'43.90"E; 500 m a.s.l.), Pubei County, Qinzhou City, Guangxi Province, China.</p><p>Paratypes.</p><p>Adult males SYS a000578, 581 and an adult female SYS a000580, collected on 28 April 2009 by Jian-Huan Yang (JHY hereafter) and Run-Lin Li (RLL hereafter), adult males SYS a003487-3489, 3505-3506, SYS a003500 / CIB107274 and adult females SYS a003485, 3499, 3504,, collected from 29-30 March 2015 by JW, ZTL, YYW and ZYL, all from the same locality as the holotype.</p><p>Diagnosis.</p><p>(1) small size (SVL 25.6-30.0 mm in males, 33.0-36.0 mm in females), (2) dorsal surface rough with skin ridges and dense conical tubercles, (3) iris bicolored, coppery yellow on upper half and silver on lower half, (4) tympanum distinctly discernible, slightly concave, dark brown, distinct black supratympanic line present, (5) dorsal surface greyish purple background with dark brown markings and scattered with orange-yellow blotches and white speckling, (6) distinct dark blotches on flanks, (7) ventral surface greyish white mixed by tiny white and black dots, (8) lateral fringes on fingers absent, (9) toes with narrow lateral fringes and rudimentary webbing, (10) longitudinal ridges under toes not interrupted at the articulations, and (11) dense conical spines on lateral and ventral surface of tarsus, dorsal surface of tibia-tarsal and surface of inner-side shank and surface around cloacal region.</p><p>Comparisons.</p><p>Comparative morphological data of Leptobrachella wuhuangmontis sp. n. with the 66 recognized Leptobrachella species were obtained from examination of museum specimens (see Appendix 1) and from the references listed in Table 2. All comparative data were shown in Tables 4, 5, 6.</p><p>Compared with the 24 known congeners of the genus Leptobrachella occurring south of the Isthmus of Kra, by the presence of supra-axillary and ventrolateral glands, L. wuhuangmontis sp. n. can be easily distinguished from L. arayai, L. dringi, L. fritinniens, L. gracilis, L. hamidi, L. heteropus, L. kajangensis, L. kecil, L. marmorata, L. melanoleuca, L. maura, L. picta, L. platycephala, L. sabahmontana, and L. sola, all of which lacking supra-axillary and ventrolateral glands; and by the significantly larger body size, SVL 25.6-30.0 mm in males, 33.0-36.0 mm in females, L. wuhuangmontis sp. n. differs from the smaller L. baluensis (14.9-15.9 mm in males), L. brevicrus (17.1-17.8 mm in males), L. itiokai (15.2-16.7 mm in males), L. juliandringi (17.0-17.2 mm in males and 18.9-19.1 mm in females), L. mjobergi (15.7-19.0 mm in males), L. natunae (17.6 mm in male), L. parva (15.0-16.9 mm in males and 17.8 mm in female), L. palmata (14.4-16.8 mm in males), L. serasanae (16.9 mm in female) and Dring’s (1983) Leptobrachella sp. 3 “baluensis” (15.0-16.0 mm in males).</p><p>Leptobrachella wuhuangmontis sp. n. significantly differs from L. yunkaiensns sp. n. by a large genetic divergence (p=10.2-11.1%), lateral fringes on toes narrow (vs. wide), black supratympanic line distinct (vs. weak), dorsal surface of body rough and scattered with dense conical tubercles (vs. shagreened with short skin ridges and raised warts), belly greyish white mixed by tiny white and black dots (vs. belly pink with distinct or indistinct speckling).</p><p>From the rest 42 known congeners (Table 5), with SVL 25.6-30.0 mm in nine males and 33.0-36.0 mm in four females, Leptobrachella wuhuangmontis sp. n. differs from the larger L. bourreti (females 42.0-45.0 mm), L. eos (males 33.1-34.7 mm, female 40.7 mm), L. lateralis (female 36.6 mm), L. nahangensis (male 40.8 mm), L. nyx (females 37.0-41.0 mm), L. sungi (males 48.3-52.7 mm, females 56.7-58.9 mm), L. tamdil (male 32.3 mm) and L. zhangyapingi (males 45.8-52.5 mm); and from the smaller L. aerea (females 28.8-28.9 mm), L. ardens (female 24.5 mm), L. alpinus (females 32.1-32.5 mm in), L. applebyi (males 19.6-20.8 mm, female 21.7 mm), L. bidoupensis (males 18.5-25.4 mm), L. botsfordi (females 30.0-31.8 mm), L. kalonensis (females 28.9-30.6 mm), L. laui (female 28.1 mm), L. liui (females 23.0-28.0 mm), L. maculosa (female 27.0 mm), L. maoershanensis (female 29.1 mm), L. melica (males 19.5-22.7 mm), L. oshanensis (female 31.6 mm), L. pluvialis (males 21.0-22.0 mm), L. puhoatensis (females 27.3-31.5 mm), L. rowleyae (females 27.0-27.8 mm), L. tadungensis (female 32.1 mm), and L. tengchongensis (females 28.9-28.9 mm). Having head longer than wide in the new species (vs. head wider than long in L. bourreti, L. khasiorum, L. lateralis and L. sungi, and head width equal to or wider than long in L. nokrekensis). By having narrow fringes on toes, the new species differs from L. applebyi, L. ardens, L. crocea, L. kalonensis, L. lateralis, L. maculosa, L. macrops, L. melica, L. minima, L. nahangensis, L. nyx, L. oshanensis, L. pallida, L. pluvialis, L. pyrrhops, L. rowleyae, L. tadungensis, L. tuberosa, and L. ventripunctata, all of which have no lateral fringes on toes; L. bidoupensis, L. bourreti, L. fuliginosa, and L. sungi, all of which have weak lateral fringes on toes; L. alpinus, L. firthi and L. isos, all of which have wide lateral fringes only in males; L. aerea, L. eos, L. khasiorum, L. laui, L. liui, L. purpura, L. tamdil, L. yingjiangensis, and L. zhangyaping, all of which have wide lateral fringes both in males and females. By having rudimentary webbing on toes, the new species differs from L. ardens, L. kalonensis, L. maculosa, L. oshanensis, L. pallida, L. petrops, L. rowleyae, and L. tadungensis, all of which have no webbing on toes; L. pelodytoides, L. sungi, and L. tamdil, all of which have wide webbing on toes. By having black spots on flanks, the new species differs from L. aerea, L. botsfordi, L. eos, L. firthi, L. isos, L. pallida, L. petrops, L. tuberosa, and L. zhangyapingi, all of which have no black spots on flanks. By having rough dorsal skin with skin ridges and dense conical tubercles, the new species differs from L. applebyi, L. bidoupensis, L. kalonensis, L. melica, L. minima, L. nahangensis, and L. tadumgensis, all of which have smooth dorsal skin, and from L. purpura, L. tengchongensis, and L. yingjiangensis, all of which have shagreened dorsal skin with small tubercles.</p><p>Description of holotype.</p><p>Adult male. Body size small, SVL in 30.0 mm. Head slightly longer than wide, HDL/HDW 1.04; snout rounded in dorsal view, nostril rounded, closer to tip of snout than to eye; canthus rostralis distinct; lores slightly concave; eye large, diameter equal to snout length, in 4.3 mm; tympanum distinct, rounded and slightly concave, its diameter significantly shorter than eye, TMP/EYE 0.56; distinct black supratympanic line present; vomerine teeth absent; vocal sac opening slit-like; tongue deeply notched behind; supratympanic ridge distinct, running from eye towards supra-axillary gland with raised tubercles.</p><p>Tips of fingers rounded, slightly swollen; relative finger lengths I = II &lt;IV &lt;III; nuptial pad absent; subarticular tubercles absent; a large, rounded inner palmar tubercle distinctly separated from small, round outer palmar tubercle; finger webbing absent and lateral fringes absent. Tips of toes like fingers; relative toe length I &lt;II &lt;V &lt;III &lt;IV; subarticular tubercles absent; dermal ridges undeveloped but present under the 3rd to 5th toes; large, oval inner metatarsal tubercle present, outer metatarsal tubercle absent; toes webbing rudimentary; narrow lateral fringes present on all toes. Tibia 45% of snout-vent length; tibiotarsal articulation reaches to middle of eye; heels just meeting each other when thighs are appressed at right angles with respect to body.</p><p>Skin on dorsum body and limbs rough with skin ridges and dense conical tubercles, ventral skin smooth; pectoral gland and femoral gland large, oval, slightly elevated; femoral gland situated on posterovertral surface of thigh, closer to knee than to vent; supra-axillary gland raised. Ventrolateral gland distinct, forming an incomplete line. Dense conical spines present on surface of lateral and ventral tarsus, surface of tibia-tarsal, inner-side surface of shank and surface around cloacal region.</p><p>Measurements of holotype</p><p>(in mm).SVL 30.0, HDL 10.9, HDW 10.5, SNT 4.3, EYE 4.3, IOD 2.9, IND 3.0, TMP 2.4, TEY 0.8, TIB 13.5, ML 7.8, PL 13.0, LAHL 15.4, HLL 44.9.</p><p>Coloration of holotype in life.</p><p>Dorsal surface greyish purple with distinct dark brown markings and scattered with yellow blotches; distinct small white speckling present on edges of dark markings. A distinct dark brown triangle pattern between eyes, connected to the incomplete W-shaped dark brown marking between axillae. Tubercles on dorsum of body and limbs brown, those on lower flanks somewhat whitish; anterior upper lip with distinct blackish brown patches; transverse dark brown bars on dorsal surface of limbs; distinct dark brown blotches on flanks from groin to axilla; elbow and upper arms coppery orange and with distinct dark bars; fingers and toes with distinct dark brown blotches.</p><p>Ventral surface greyish-white mixed with tiny white and black dots. Supra-axillary, femoral, and ventrolateral glands white, pectoral gland greyish white as the color of ventral surface. Iris bicolored, coppery yellow on upper half and silver on lower half.</p><p>Coloration of holotype in preservative.</p><p>Dorsal of body dark with greyish white dots on flanks, while dorsal of limbs dark brown, transverse bars on dorsal of forelimbs become more distinct, and indistinct on dorsal of hindlimbs, dark brown patterns, markings and spots on back become indistinct. Ventral surface light yellow with brown speckling. Supra-axillary, femoral, ventrolateral and pectoral glands light yellow (Figure 7).</p><p>Sexual dimorphism.</p><p>Females with a larger body size than males, SVL 33.0-36.0 mm (34.9 ± 1.4) (vs. SVL 25.6-30.0 mm (28.5 ± 1.5) in males); presence of a single vocal sac in males (vs. absent in females); dense conical spines on lateral and ventral surface of tarsus, surface of tibia-tarsal, inner-side surface of shank and surface around cloacal region distinct in males (vs. barely visible in females); pectoral gland and femoral gland large, oval, slightly elevated in males (vs. indistinct in females).</p><p>Variations.</p><p>All paratypes match the overall characters of the holotype except that: tibiotarsal articulation reaches to posterior corner of eye in female paratypes SYS a003499, 3504 and reaches to anterior corner of eye in male paratypes SYS a003487 and SYSa 003500 / CIB 107274; pectoral gland large, oval, slightly elevated in all individuals in life, and become indistinct in preservation. Yellow blotches and white speckling present on dorsum in the holotype (vs. indistinct in the female paratype SYS a003499). Elbow and upper arms coppery orange and with distinct dark bars in the holotype (vs. elbow and upper arms light orange, dark bars indistinct in the male paratypes SYS a003488, 3505 and the female paratype SYS a003499) (Figure 6).</p><p>Etymology.</p><p>The specific epithet, wuhuangmontis, is in reference to the type locality, Mt. Wuhuang of Guangxi Province, China. For the common name, we suggest "Mt. Wuhuang’s Leaf Litter Toad", and for the Chinese name "Wu Huang Shan Zhang Tu Chan ( 五皇山掌突蟾)” .</p><p>Distribution and habits.</p><p>Currently, Leptobrachella wuhuangmontis sp. n. is only known from its type locality MWH from Guangxi Province of China (Figure 1). The new species was found along a clear-water rocky streams and small steep rocky streams in well-preserved montane evergreen broadleaf forest (500 m a.s.l.) (Figure 8). During field surveys in March, males were found calling exposed on the rocks or hiding in the rock seams; gravid female collected on March and April bear pure white oocytes.</p></div>	https://treatment.plazi.org/id/296B422E59F70E7C2EA921B93C3B4CD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Jian;Yang, Jianhuan;Li, Yao;Lyu, Zhitong;Zeng, Zhaochi;Liu, Zuyao;Ye, Youhua;Wang, Yingyong	Wang, Jian, Yang, Jianhuan, Li, Yao, Lyu, Zhitong, Zeng, Zhaochi, Liu, Zuyao, Ye, Youhua, Wang, Yingyong (2018): Morphology and molecular genetics reveal two new Leptobrachella species in southern China (Anura, Megophryidae). ZooKeys 776: 71-103, DOI: http://dx.doi.org/10.3897/zookeys.776.22925, URL: http://dx.doi.org/10.3897/zookeys.776.22925
114FCDB24EA2DA8C00A45DDFAD0CE54F.text	114FCDB24EA2DA8C00A45DDFAD0CE54F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptobrachella yunkaiensis Wang, Li, Lyu & Wang	<div><p>Leptobrachella yunkaiensis Wang, Li, Lyu &amp; Wang sp. n. Figures 3, 4C1-C3</p><p>Holotype.</p><p>SYS a004665, adult male, collected on 15 April 2016 by Jian Wang (JW hereafter), Zhao-Chi Zeng (ZCZ hereafter), Ying-Yong Wang (YYW hereafter), Zu-Yao Liu (ZYL hereafter), Hai-Long He (HLH hereafter) and Zhi-Tong Lyu (ZTL hereafter) from Dawuling Forest Station (DWL hereafter) (22°16'32.9"N, 111°11'42.87"E; 1600 m a.s.l.), Maoming City, Guangdong Province, China.</p><p>Paratypes.</p><p>Collectors and locality data of paratypes were the same as holotype: adult males, SYS a004664 / CIB107272, SYS a004666-4669 and an adult female SYS a004663, collected on 15 April 2016, the other adult female, SYS a004690, collected on 16 April 2017.</p><p>Diagnosis.</p><p>(1) small size (SVL 25.9-29.3 mm in males, 34.0-35.3 mm in females), (2) dorsal skin shagreened with short skin ridges and raised warts, (3) iris bicolored, coppery orange on upper half and silver on lower half, (4) tympanum distinctly discernible, slightly concave, weakly black supratympanic line present, (5) dorsal surface yellowish-brown grounding, with distinct darker brown markings and rounded spots and scattered with irregular orange patches, (6) flanks with several dark blotches, (7) surface of belly pinkish, with distinct or indistinct light dark brown speckling, (8) supra-axillary, femoral, pectoral and ventrolateral glands distinctly visible, (9) absence of webbing and presence of distinct lateral fringes on fingers, toes with rudimentary webbing and wide lateral fringes, (10) longitudinal ridges under toes not interrupted at the articulations, and (11) dense conical spines present on lateral and ventral surface of tarsus, surface of tibia-tarsal, inner-side surface of shank and surface around cloacal region.</p><p>Comparisons.</p><p>Comparative morphological data of Leptobrachella yunkaiensis sp. n. with 66 recognized Leptobrachella species were obtained from examination of museum specimens (see Appendix 1) and from the references listed in Table 2. All comparative data were shown in Tables 4, 5.</p><p>Compared with the 24 known congeners of the genus Leptobrachella occurring south of the Isthmus of Kra, by the presence of supra-axillary and ventrolateral glands, L. yunkaiensis sp. n. can be easily distinguished from L. arayai, L. dringi, L. fritinniens, L. gracilis, L. hamidi, L. heteropus, L. kajangensis, L. kecil, L. marmorata, L. melanoleuca, L. maura, L. picta, L. platycephala, L. sabahmontana and L. sola, all of which lacking supra-axillary and ventrolateral glands; and by the significantly larger body size, SVL 25.9-29.3 mm in males, 34.0-35.3 mm in females, L. yunkaiensis sp. n. differs from the smaller L. baluensis (14.9-15.9 mm in males), L. brevicrus (17.1-17.8 mm in males), L. itiokai (15.2-16.7 mm in males), L. juliandringi (17.0-17.2 mm in males and 18.9-19.1 mm in females), L. mjobergi (15.7-19.0 mm in males), L. natunae (17.6 mm in male), L. parva (15.0-16.9 mm in males and 17.8 mm in female), L. palmata (14.4-16.8 mm in males), L. serasanae (16.9 mm in female) and Dring’s (1983) Leptobrachella sp. 3 “baluensis” (15.0-16.0 mm in males).</p><p>Leptobrachella yunkaiensis sp. n. is most similar to L. laui and L. liui, but it can be distinguished by the larger body sized, SVL 34.0-35.3 mm in females (vs. SVL 28.1 mm in a single female of L. laui; SVL 23.0-28.0 mm in females of L. liui), presence of short skin ridge and raised warts on dorsum (vs. absent in L. laui), black supratympanic line weak (vs. black supratympanic line distinct in L. liui), longitudinal ridges under toes not interrupted at the articulations (vs. interrupted in L. liui) (Figure 4), belly pinkish with distinct or indistinct speckling (vs. belly creamy white with dark brown dusting on margins in L. laui; belly creamy white with dark brown spots on chest and margins in L. liui).</p><p>From the remaining 40 known congeners (Table 5), with SVL 25.9-29.3 mm in six males, SVL 34.0-35.3 mm in two females in Leptobrachella yunkaiensis sp. n., it can be distinguished from the larger L. eos (males 33.1-34.7 mm, female 40.7 mm), L. nahangensis (male 40.8 mm), L. pyrrhops (males 30.8-34.3 mm), L. sungi (males 48.3-52.7 mm, females 56.7-58.9 mm) and L. zhangyapingi (males 45.8-52.5 mm), and the smaller L. applebyi (males 19.6-22.3 mm, females21.7-25.9 mm), L. melica (males 19.5-22.7 mm), and L. pluvialis (males 21.3-22.3 mm). By having wide fringes on toes, the new species differs from L. applebyi, L. ardens, L. crocea, L. kalonensis, L. lateralis, L. maculosa, L. macrops, L. melica, L. minima, L. nahangensis, L. nyx, L. oshanensis, L. pallida, L. pluvialis, L. pyrrhops, L. rowleyae, L. tadungensis, L. tuberosa, and L. ventripunctata, all of which have no lateral fringes on toes; L. bidoupensis, L. bourreti, L. fuliginosa, and L. sungi, all of which have weak lateral fringes on toes; L. botsfordi, L. maoershanensis, L. pelodytoides, L. petrops, L. puhoatensis, and L. tengchongensis, all of which have narrow lateral fringes on toes; L. alpinus, L. firthi, and L. isos, all of which have wide lateral fringes only in males. With rudimentary webbing on toes, the new species differs from L. ardens, L. kalonensis, L. maculosa, L. oshanensis, L. pallida, L. petrops, L. rowleyae, and L. tadungensis, all of which have no webbing on toes; L. pelodytoides, L. sungi, and L. tamdil, all of which have wide webbing on toes. By having black spots on flanks, the new species differs from L. aerea, L. botsfordi, L. eos, L. firthi, L. isos, L. pallida, L. petrops, L. tuberosa, and L. zhangyapingi, all of which have no black spots on flanks. With belly pink with distinct or indistinct speckling, the new species differs from L. bourreti, L. eos, L. firthi, L. khasiorum, L. lateralis, L. minima, L. nahangensis, and L. nokrekensis, all of which have creamy white belly without patterns; from L. macrops, which have greyish-violet with white speckling; from L. purpura, which have dull white belly with indistinct grey dusting; and from L. yingjiangensis, which have creamy white belly with dark brown flecks on chest and margins. By dorsal skin shagreened with short skin ridges and raised warts, the new species differs from L. purpura, L. yingjiangensis and L. tengchongensis, all of which have shagreened dorsal skin with small tubercles, and from L. macrops, which have no skin ridges dorsally.</p><p>Description of holotype.</p><p>Adult male. Body size small, SVL in 28.7 mm. Head length slightly larger than head width, HDL/HDW 1.03; snout slightly protruding, projecting slightly beyond margin of the lower jaw; nostril equidistance between snout and eye; canthus rostralis gently rounded; loreal region slightly concave; interorbital space flat, larger internarial distance; pineal ocellus absent; vertical pupil; snout length slightly larger than eye diameter, SNT/EYE 1.03; tympanum distinct, rounded, and slightly concave, diameter smaller than that of the eye and larger than tympanum-eye distance, TMP/EYE 0.46 and TEY/TMP 0.47; weakly black supratympanic line present; vomerine teeth absent; vocal sac openings slit-like, located posterolaterally on floor of mouth in close proximity to the margins of the mandible; tongue deeply notched behind; supratympanic ridge distinct, extending from posterior corner of eye to supra-axillary gland; tubercles present on supratympanic ridge.</p><p>Tips of fingers rounded, slightly swollen; relative finger lengths I = II = IV &lt;III; nuptial pad absent; subarticular tubercles absent; a large, rounded inner palmar tubercle distinctly separated from small, round outer palmar tubercle; absence of webbing and presence of distinct lateral fringes on fingers. Tips of toes like fingers; relative toe length I &lt;II &lt;V &lt;III &lt;IV; subarticular tubercles absent; distinct dermal ridges present under the 3rd to 5th toes; large, oval inner metatarsal tubercle present, outer metatarsal tubercle absent; toes webbing rudimentary; wide lateral fringes present on all toes. Tibia 43% of snout-vent length; tibiotarsal articulation reaches to middle of eye; heels just meeting each other when thighs are appressed at right angles with respect to body.</p><p>Skin on dorsum shagreened and scattered with fine, round tubercles; short skin ridges and raised warts on dorsum surface present; ventral skin smooth; pectoral gland and femoral gland large, oval; pectoral glands greater than tips of fingers and femoral glands; femoral gland situated on posteroventral surface of thigh, closer to knee than to vent; supra-axillary gland raised. Ventrolateral gland distinctly visible, forming an incomplete line. Dense conical spines on lateral and ventral surface of tarsus, surface of tibia-tarsal, inner-side surface of shank and surface around cloacal region present.</p><p>Measurements of holotype</p><p>(in mm).SVL 28.7, HDL 10.3, HDW 10.0, SNT 3.8, EYE 3.7, IOD 2.9, IND 2.8, TMP 1.7, TEY 0.8, TIB 12.4, ML 7.2, PL 12.1, LAHL 12.3, HLL 38.3.</p><p>Coloration of holotype in life.</p><p>Dorsal surface orange-brown with distinct dark brown blotches edged distinct light orange pigmentation. A dark brown triangular pattern between eyes, connected to the dark brown W-shaped marking between axillae. Tympanum black. Orange-brown tubercles present on dorsum of body and limb, those on flanks much distinct and dense; anterior upper lip with distinct blackish brown patches; transverse dark brown bars on dorsal surface of limbs; indistinct dark brown blotches on flanks from groin to axilla; elbow and upper arms without dark bars but with distinct coppery orange coloration; fingers and toes with indistinct dark brown blotches.</p><p>Surface of throat creamy white and scattered with small whitish dots; belly pinkish and scattered with small brown speckling; ventral surface of thighs pinkish and scattered with small light orange-brown spots. Supra-axillary coppery orange; femoral, pectoral and ventrolateral glands whitish orange. Iris bicolored, coppery orange on upper half and silver on lower half.</p><p>Coloration of holotype in preservative.</p><p>Dorsum of body and hindlimbs dark brown while dorsum of forelimbs yellowish brown; transverse bars on limbs become more distinct, dark brown patterns, markings and spots on back become indistinct. Ventral surface of body yellowish brown, with brown marbling on sides and chest. Orange supra-axillary, femoral, pectoral and ventrolateral glands fade to greyish white (Figure 4C1-C3).</p><p>Sexual dimorphism.</p><p>Females with a larger body size than males, SVL 34.0-35.3 mm (34.7 ± 0.9) (vs. SVL 25.9-29.3 mm (27.6 ± 1.4) in males); presence of a single vocal sac in males (vs. absent in females); dense conical spines on lateral and ventral surface of tarsus, surface of tibia-tarsal, inner-side surface of shank, surface of thighs and surface around cloacal region distinct in males, and barely visible in females.</p><p>Variations.</p><p>All paratypes match the overall characters of the holotype except that: the heels just meeting each other when thighs are appressed at right angles with respect to body, tibiotarsal articulation reaches to middle of eye in holotype SYS a004665 (vs. tibiotarsal articulation reaches to anterior corner of eye in SYS a004666, reaches the posterior corner of eye in SYS a004669). Surface of belly scattered with distinctly dark brown speckling in holotype (vs. such speckling indistinct in female paratypes SYS a004663, 4690. Tympanum black in the holotype (vs. tympanum black grounding with orange speckling in SYS a004667-4668). Distinct black spots present on dorsum in the female paraype SYS a004690 (Figure 3).</p><p>Etymology.</p><p>The specific epithet, yunkaiensis, is in reference to the type locality, DWL of Guangdong, China located in the Yunkai Mountains Range. For the common name, we suggest "Yunkai Mountain’s Leaf Litter Toad", and Chinese name "Yun Kai Zhang Tu Chan ( 云开掌突蟾)” .</p><p>Distribution and habits.</p><p>Currently, Leptobrachella yunkaiensis sp. n. is known only from its type locality DWL of Guangdong Province (Figure 1). The new species was found along a clear-water rocky stream (ca. 2-3 m in width and ca. 20-30 cm in depth) and small nearby seeps in well-preserved montane evergreen broadleaf forest (1600 m a.s.l.) (Figure 5). During April and June, males were found calling mainly hidden under leaf litter, and some were found calling perching on the rocks or under rocks by the side of the stream. Females collected on April bear pure white oocytes.</p></div>	https://treatment.plazi.org/id/114FCDB24EA2DA8C00A45DDFAD0CE54F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Wang, Jian;Yang, Jianhuan;Li, Yao;Lyu, Zhitong;Zeng, Zhaochi;Liu, Zuyao;Ye, Youhua;Wang, Yingyong	Wang, Jian, Yang, Jianhuan, Li, Yao, Lyu, Zhitong, Zeng, Zhaochi, Liu, Zuyao, Ye, Youhua, Wang, Yingyong (2018): Morphology and molecular genetics reveal two new Leptobrachella species in southern China (Anura, Megophryidae). ZooKeys 776: 71-103, DOI: http://dx.doi.org/10.3897/zookeys.776.22925, URL: http://dx.doi.org/10.3897/zookeys.776.22925
