identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
AFFA2F3D7128607210CEFA2DA6835501.text	AFFA2F3D7128607210CEFA2DA6835501.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gnathusa Fenyes	<div><p>Gnathusa Fenyes</p><p>Gnathusa Fenyes 1909: 197. Type species: Gnathusa eva Fenyes.</p><p>Diagnosis.</p><p>Body dark brown to black, compact and robust, subparallel (Figs 1a, 2a, b, 3a, 4a), length 3.5-4.3 mm; integument with distinct meshed microsculpture; head large with mandibles extremely long and sickle-shaped, apices very slender, each crossing the other in resting position, each mandible bearing long spine or tooth (Figs 1h, i, 2i, j, 3h, i, 4h, i); infraorbital carina strong and complete; ligula more or less deeply bilobed (Figs 1l, 2m, 3l, 4l); last article of maxillary palpus needle-shaped (Figs 1k, 2l, 3k, 4k); frontal suture of head absent; anterior margin of mesosternum with short V-shaped basal carina, remaining mesosternum uncarinated; mesosternal process long, triangular basally and then narrowly produced and extending to about ¾ length of mesosternal cavities, metasternal process triangular in shape and short, isthmus short; median lobe of aedeagus with large crista apicalis, internal sac structures variable in shape (Figs 1b, 2c, 3b, 4b), paramere with narrowly elongate apical lobe bearing four macrosetae; spermatheca pipe-shaped with spherical or tubular capsule and elongate narrow stem (Figs 1e, 2d, 3e, 4e).</p><p>Key to Canadian species of Gnathusa</p><p>New provincial and territorial records are indicated in boldface font.</p></div>	https://treatment.plazi.org/id/AFFA2F3D7128607210CEFA2DA6835501	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
9C5BFBA0604103C7880E4B27D3672679.text	9C5BFBA0604103C7880E4B27D3672679.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gnathusa eva Fenyes	<div><p>1. Gnathusa eva Fenyes Figure 1 a–l, Map 1</p><p>Gnathusa eva Fenyes 1909: 198, 1920: 352, Moore and Legner 1975: 458, Majka and Klimaszewski 2008: 88.</p><p>Diagnosis.</p><p>Body length 3.5-4.0 mm, sides subparallel; body colour light brown to dark brown, antennae and tarsi rust-brown, head and abdomen often dark brown; integumental microsculpture dense and surface moderately glossy; head round and almost as wide as pronotum with labrum bearing long spines; pronotum transverse, angular, wider than maximum width of elytra; elytra at suture subequal in length to pronotum; abdomen subparallel; antennal articles 6-10 moderately-to-strongly transverse, last article short and broadly oval (Fig. 1a). MALE: tergite VIII widely truncate apically (Fig. 1c); sternite VIII slightly pointed at apex (Fig. 1d); median lobe of aedeagus with tubus arcuate and apex pointed and produced ventrally in lateral view (Fig. 1b). FEMALE: tergite VIII truncate apically (Fig. 1f); sternite VIII rounded apically (Fig. 1g); spermatheca pipe-shaped, with short sac-shaped capsule and long, thin and sinuate stem, spermathecal neck weakly sclerotized and neck to capsule angle variable (Fig. 1e).</p><p>Distribution.</p><p>The Canadian distribution of this native Nearctic species includes Alberta [new provincial record], British Columbia (Majka and Klimaszewski 2008), and Yukon Territory (Map 1). It is also known from California (Fenyes 1909).</p><p>Bionomics.</p><p>Adults were captured in clear-cut Sitka spruce forest on Vancouver Island and in moss and gravel at the edge of small pools at other localities in the interior of British Columbia (Klimaszewski and Winchester 2002). Additional specimens were found in British Columbia in a 1-year-old harvested Douglas-fir stand. In west-central Alberta, adults were collected in pitfall traps deployed in Upper Cordilleran coniferous forests, including subxeric lodgepole pine forests, mesic white spruce and lodgepole pine stands and spruce-dominated subhygric and hygric forests, but not in deciduous-dominated forest or in grassy or shrubby meadows. In Alberta, adults also emerged from lodgepole pine trees infested by bark beetles. In the Yukon Territory, adults were found in a squirrel midden in spring, probably overwintering, and in a coniferous woodchip pile.</p><p>Locality data.</p><p>CANADA: Alberta: Lusk Creek, Kananaskis F.E.S., 14.VII.1971, J.M. &amp; B.A. Campbell (CNC) 1 male, 4 females; vicinity of Swan Hills, 54°42'N, 115°23'W, Picea / Pinus forest, 15.VI.1990, D.W. Langor (NoFC) 2 males; Grande Prairie, 64 km S, 54.5597°N, 118.6633°W, emergence trap on MPB infested lodgepole pine, 14 July 2011, col. Bleiker (NoFC) 1 female; same data except 15 July 2011, 1 female, 1 sex undetermined; Grande Prairie, 75 km S, 54.4706°N, 118.6560°W, 13 Aug. 2011, emergence trap on MPB infested lodgepole pine, col. Bleiker (NoFC) 1 sex undetermined; Fox Creek, 24 km E, 54.4575°N, 116.4377°N, emergence trap on MPB infested lodgepole pine, 9 Aug. 2011, col. Bleiker (NoFC) 1 female; Fox Creek, 7 km SW, 54.3241°N, 116.8335°W, emergence trap on MPB infested lodgepole pine, 18 July 2011, col. Bleiker (NoFC) 1 sex undetermined; 20 km NW Hinton, 4 km NW of Jarvis Lake, 53.484°N, 117.854°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite I1, Stand I103, pitfall trap # 4, 19. VI– 3.VII.2004, J. Hammond et al. coll. (NoFC) 1 sex undetermined; 20 km S Hinton, 31.V.1990, pitfall trap, D. Langor (NoFC) 1 sex undetermined; 21.3 km NW Hinton, W.A. Switzer Prov. Pk., 53.529°N, 117.824°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite E1, Stand E129, pitfall trap # 5, 3-17.VII.2004, J. Hammond et al. coll. (NoFC) 1 female; 23 km NW Hinton, 1.7 km W of Gregg Lake, 53.545°N, 117.821°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite H1, Stand H101, pitfall trap # 1, 2-16.VII.2004, J. Hammond et al. coll. (NoFC) 1 female; 25 km NW Hinton, west side Hay River Rd., 53.502°N, 117.909°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite C1, Stand C101, pitfall trap # 4, 2-17.VII.2004, J. Hammond et al. coll. (NoFC) 1 male; same data except: pitfall trap # 6, 17-31.VII.2004, J. Hammond et al. coll. (NoFC) 1 sex undetermined; 26 km SE Hinton, 7 km S of Gregg River Rd., 53.220°N, 117.343°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite H1, Stand H104, pitfall trap # 1, 16. V– 1.VI.2004, J. Hammond et al. coll. (NoFC) 2 sex undetermined; 32 km NW Hinton, 1 km W of Rock Lake Rd., 53.561°N, 117.998°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite F1, Stand F104, pitfall trap # 1, 2-16.VI.2004, J. Hammond et al. coll. (NoFC) 1 male, 5 sex undetermined; 33 km NW Hinton, 3.75 km N of Highway 40, 53.594°N, 117.964°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite E1, Stand E108, pitfall trap # 4, 4-18.VI.2004, J. Hammond et al. coll. (NoFC) 1 female, 1 sex undetermined; 34 km NW Hinton, 0.5 km W of Highway 40, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite F1, Stand F102, pitfall trap # 4, 4-18.VI.2004, J. Hammond et al. coll. (NoFC) 3 sex undetermined; 35 km NW Hinton, 3.75 km N of Highway 40, 53.596°N, 118.002°W, Ecoregion: UF, Ecosite D1, Stand D109, pitfall trap # 4, 2-16.VII.2004, J. Hammond et al. coll. (NoFC) 1 female; 36 km NW Hinton, 3.75 W of Rock Lake Rd., 53.564°N, 118.046°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite I1, Stand I107, pitfall trap # 4, 14. V– 4.VI.2004, J. Hammond et al. coll. (NoFC) 1 female; 59 km NW Hinton, 3.5 km N of Polecat Rd., 53.902°N, 117.911°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite H1, Stand H105, pitfall trap # 2, 3-17.VI.2004, J. Hammond et al. coll. (NoFC) 1 sex undetermined; 62 km N Hinton, 5 km W of J. Wright Rd., 53.969°N, 117.668°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite F1, Stand F105, pitfall trap # 2, 12. V– 2.VI.2004, J. Hammond et al. coll. (NoFC) 1 female, 1 sex undetermined; 63 km N Hinton, 3.75 km S of J. Wright Rd., 53.974°N, 117.449°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite D1, Stand D102, pitfall trap # 2, 30. VI– 14.VII.2004, J. Hammond et al. coll. (NoFC) 1 male, 1 female; 65 km N Hinton, 5 km W of J. Wright Rd., 53.995°N, 117.656°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite E1, Stand E118, pitfall trap # 5, 30. VI– 14.VII.2004, J. Hammond et al. coll. (NoFC) 1 sex undetermined; 67 km N Hinton, north side of J. Wright Rd., 53.998°N, 117.435°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite D1, Stand D104, pitfall trap # 2, 2-16.VI.2004, J. Hammond et al. coll. (NoFC) 1 sex undetermined. British Columbia: Fort St. James, PG13B-trap 4, 10.V. 1995 (CNC) 1 female; same data except: GP 11 km-2, 1 year post harvest (CNC) 1 male; GP 115, 30.V.1996, 1 year post harvest, R. Felix (CNC) 1 male; 20.VI.1920, GP 11 km, 2 years post harvest, D. Rodriguez (CNC) 1 female; 4.VIII.1996, Tachie-Pinchi, M. Cloet, trap 5 (CNC) 1 female; GP 11 km-2, 10.V.1995, 1 year post harvest (CNC) 3 females; 21 km SW Campbell River, 49°51'55"N, 125°27'51"W, 22.V-6.VI.1996, Balsam Cr., LT 1-T, 1-E (LFC) 2 males; Monashee Mountain near Cherryville, 12.VIII.1982, R. Baranowski (LFC, MZLU) 1 female, 1 sex undetermined [published record Majka and Klimaszewski 2008]. Yukon Territory: Whitehorse, Paddy’s Pond, 15.V.2010, 60.7067°N, 135.0917°W, 649 m, soil sifting, squirrel midden, B. Godin (ECW) 2 males, 3 females; Whitehorse, Granger subdivision, coniferous woodchip pile, 2.IX.2007, 60.7097°N, 135.0996°W, 661 m, pitfall trap, B. Godin (ECW) 1 male; same data as before except: 3.V.2008 (ECW) 2 males.</p><p>UNITED STATES OF AMERICA: California: Mono Co., 6 mi SW Toms Place, 9000', 8.VIII.1969, A. Smetana (CNC) 3 males, 1 female [not shown in Map 1].</p></div>	https://treatment.plazi.org/id/9C5BFBA0604103C7880E4B27D3672679	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
4AFD9A27A7FE701A1589C9B8CE56DE83.text	4AFD9A27A7FE701A1589C9B8CE56DE83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gnathusa tenuicornis Fenyes	<div><p>2. Gnathusa tenuicornis Fenyes Figure 2 a–m, Map 2</p><p>Gnathusa tenuicornis Fenyes 1921: 26, Moore and Legner 1975: 458, Klimaszewski and Winchester 2002: 58.</p><p>Diagnosis.</p><p>Body length 2.5-3.7 mm, sides subparallel; body colour light brown to almost black, with antennae, tarsi and often elytra and apical part of abdomen rust-brown; integumental microsculpture dense and surface strongly glossy; head round (Fig. 2a) to somewhat quadrate (Fig. 2b) and almost as wide as pronotum, labrum with short fine setae but lacking coarse spines; pronotum transverse, subequal to slightly narrower than maximum width of elytra, corners somewhat angular; elytra at suture subequal in length to pronotum; abdomen subparallel; antennal articles 6-10 subquadrate, last article short and broadly oval (Fig. 2a, b). MALE: tergite VIII widely truncate apically (Fig. 2e); sternite VIII slightly pointed at apex (Fig. 2f); median lobe of aedeagus with tubus almost straight and apex pointed ventrally in lateral view (Fig. 2c). FEMALE: tergite VIII truncate apically (Fig. 2g); sternite VIII rounded apically (Fig. 2h); spermatheca pipe-shaped, with spherical capsule and long, thin, and slightly sinuate stem, neck weakly sclerotized, and neck to capsule angle variable (Fig. 2d).</p><p>Distribution.</p><p>This native Nearctic species was described from specimens captured in Glacier, British Columbia, later recorder from Yokon, and is herein recorded for the first time from Alberta and New Brunswick (four female specimens tentatively identified as this species) (Map 2). In the United States, this species was previously known from California (Fenyes 1921, Moore and Legner 1975, Klimaszewski and Winchester 2002), and is herein recorded for the first time from Oregon.</p><p>Bionomics .</p><p>Adults were captured in a clear-cut Sitka spruce forest on Vancouver Island and in moss and gravel at the edge of small pools in British Columbia (Klimaszewski and Winchester 2002). Other adults were found in a subalpine meadow at 3000 feet, in cold moss and gravel along the edges of streams. The Yukon specimens were taken from mixed aspen and spruce forest by sifting litter. In west-central Alberta, adults were collected in pitfall traps deployed in Upper Cordilleran coniferous forests, including subxeric lodgepole pine forests, mesic white spruce and lodgepole pine stands and and spruce-dominated subhygric and hygric forests, but not in deciduous-dominated forest or in grassy or shrubby meadows. The New Brunswick specimens were taken from moss and leaves under alders near a brook in an eastern white-cedar swamp and from under cobblestones and gravel in sand on a partially shaded cobblestone bar near the outflow of a brook into a river. Adults were captured from May through August.</p><p>Locality data.</p><p>CANADA: Alberta: Waterton Lakes National Park, Cameron Lake, 5450', 4.VIII.1976, J.M. Campbell (CNC) 1 male; 20 km S Hinton, 20.VII.1989, pitfall trap, D. Langor coll., site C, trap 6, conifer study (NoFC) 1 male; 20 km S Hinton, 26.VI.1989, D. Langor coll., site C, trap 6, conifer study (NoFC) 1 male; 26 km SE Hinton, 7 km S of Gregg River Rd., 53.220°N, 117.343°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite H1, Stand H104, pitfall trap # 4, 15-29.VI. J. Hammond et al. (NoFC) 2 males, 1 female; 31 km SE Hinton, 3 km of Highway 40, 53.593°N, 117.925°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite D1, Stand D108, pitfall trap # 6, 11.V-3.VI. J. Hammond et al. (NoFC) 1 male; 32 km NW Hinton, 3 km W of Highway 40, 53.586°N, 117.954°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite E1, Stand E103, pitfall trap # 5, 11.V-4.VI. J. Hammond et al. (NoFC) 1 male; 32 km NW Hinton, 1 km W of Hay River Rd., 53.760°N, 117.652°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite I102, Stand I102, pitfall trap # 2, 13. V– 3.VI. 2004, J. Hammond et al. (NoFC) 1 male, 1 female; 43 km SE Hinton, 1.5 km N Coalspur, 53.194°N, 117.046°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite B1, Stand B103, pitfall trap # 6, 1-15.VI.2004, J. Hammond et al. (NoFC) 1 male; 55 km N Hinton, north side of Polecat Rd., 53.855°N, 117.926°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite H1, Stand H103, pitfall trap # 3, 3-17.VI.2004, J. Hammond et al. (NoFC) 1 male, 1 female; 65 km N Hinton, 5 km W of Wright Rd., 53.995°N, 117.656°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite E1, Stand E118, pitfall trap # 5, 2-16.VI.2004, J. Hammond et al. (NoFC) 2 females; 69.5 km N Hinton, 0.7 km NW of JV Haul Rd., 54.017°N, 117.618°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite D1, Stand D103, pitfall trap # 5, 1-16.VI.2004, J. Hammond et al. (NoFC) 1 female. British Columbia: Glacier, Fenyes collection (CAS) 1 female [holotype]; Nitinat, Heather Mtn., subalpine meadow at 3000', 14.VII.1979, I.M. Smith, moss on seepage slope (CNC) 1 female; Forbidden Plateau, Murray Meadows, 3400', 21.VII.1975, J.M. and B.A. Campbell (CNC) 1 female; Queen Charlotte Islands, 10.5 km NW Rennell, Sound Rd., Ghost Main Rd., 900', J.M. Campbell, cold moss along stream (CNC) 1 male; Queen Charlotte Islands, Moresby Is., Mt. Moresby, 25.VII.1983, 2100', J.M. Campbell, ex gravel at edge of stream (CNC) 1 male; 20 mi E Hope, Manning Pk., 21.VI.1968, Campbell and Smetana (CNC) 1 female; Copper River Valley, A37574/P4-1-1, 6. VI– 5.VII.1996, pitfall trap, J. Lemieux (LFC) 1 female; same data except: 5. VII– 12.VIII.1996, (LFC) 1 male; Upper Carmanah Valley, UTM: 10U CK 803005, 16.VII-30.VII.1991, CC MT3, N. Winchester (LFC) 2 females [additional records from the same locality Klimaszewski and Winchester 2002]. Yukon Territory: Whitehorse, Paddy’s Pond, 6.V.2007, 60.7067°N, 135.0917°W, 649 m, litter sifting, mixed aspen and spruce forest, B. Godin (ECW) 1 male [record from Klimaszewski et al. 2012]. New Brunswick: Restigouche Co., MacFarlane Brook Protected (Natural) Area, 47.6018°N, 67.6263°W, 25.V.2007, R.P. Webster // old growth eastern white cedar swamp, in moss &amp; leaves under alders near stream (RWC) 2 females; Jacquet River Gorge PNA, 47.8257°N, 66.0779°W, 24.V.2010, R.P. Webster // partially shaded cobblestone bar near outflow of brook at Jacquet River, under cobblestones &amp; gravel on sand (RWC) 1 female; Mount Atkinson, 441 m elev., 41.8192°N, 68.2618°W, 7.VII.2011, R.P. Webster // Boreal forest, small shaded spring-fed brook with mossy margin, sifting moss (LFC) 1 female.</p><p>UNITED STATES OF AMERICA: Alaska: Kenai Peninsula, 2 mi NE Soldotna, 10.VI.1978, Smetana and Becker (CNC) 1 female. Oregon: Mt. Hood, Timberline Lodge Road, 4500 –5000’, 28.VI.1974, A. and D. Smetana (CNC) 1 male.</p><p>Comments.</p><p>We have tentatively included the females from New Brunswick as belonging to this species. The difference in body colour, the slightly different shape of pronotum and the temples of the head in the New Brunswick and western specimens we attribute to infraspecific variations because the shape of spermatheca and the tergites and sternite VIII are similar in females of both populations. The study of males from New Brunswick is critical to confirm our identification. The specimens from the north usually are darker than the specimens from more southern localities in many species of aleocharines.</p></div>	https://treatment.plazi.org/id/4AFD9A27A7FE701A1589C9B8CE56DE83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
EA5D8ACEA50F34F91551BF0712C9279C.text	EA5D8ACEA50F34F91551BF0712C9279C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gnathusa caribou Lohse	<div><p>3. Gnathusa caribou Lohse Figure 3 a–l, Map 2</p><p>Gnathusa caribou Lohse, in Lohse et al. 1990: 146; Klimaszewski et al. 2011: 55.</p><p>Diagnosis.</p><p>Body length 2.8-3.6 mm, sides narrowly subparallel; body colour dark brown to almost black, with antennae bright yellow and tarsi rust-brown to yellowish; integumental microsculpture dense and surface strongly glossy; head round, about the same size as the pronotum, labrum lacking stout spines but with fine setae of unequal length; pronotum small, transverse, angular, slightly narrower than elytra; elytra at suture subequal in length to pronotum; abdomen subparallel; antennal articles 6-10 subquadrate to slightly transverse, last article short and broadly oval (Fig. 3a). MALE: tergite VIII widely truncate apically (Fig. 3c); sternite VIII slightly pointed at apex (Fig. 3d); median lobe of aedeagus with tubus almost straight and apex pointed ventrally in lateral view (Fig. 3b). FEMALE: tergite VIII truncate apically (Fig. 1f); sternite VIII rounded apically (Fig. 3g); spermatheca pipe-shaped, with spherical capsule and long and straight stem, neck well sclerotized (Fig. 3e).</p><p>Distribution.</p><p>This native Nearctic species is known in Canada from the Northwest Territories and Yukon Territory (Map 2), and from Alaska (Lohse et al. 1990).</p><p>Bionomics.</p><p>Adults were captured from June to July in tundra by sifting organic litter under Salix, moss, and a pile of leaves stored by a rodent.</p><p>Locality data.</p><p>CANADA: Northwest Territories: Lac Maunoir, North shore, 19-27.VII.1969, G.E. Shewell (CNC) 1 male; Reindeer Sta., Caribou Hills, 2.VII.1972, A. Smetana (CNC) 1 male, 2 females, 1 sex undetermined [paratypes]; same label data except: 30.VI.1972 (CNC) 1 female [paratype]. Yukon Territory: British Mts., Firth River, 250 m, 69°13'N, 140°04'W, 25.VI.1984, 84-31, tundra, sifting litter under Salix (CNC) 1 male [holotype]; same label data (CNC) 1 female, 3 sex undetermined [paratypes]; British Mts., Windy Ridge, 550 m, 69°27'N, 140°25'W, 2.VII.191984, 84-46, sifting moss, J.M. Campbell (CNC) 4 sex undetermined [paratypes]; British Mts., Fish Creek, 200 m, 69°27'N, 140°23'W, 5.VII.1984, 84-58, sifting moss and arctic willow on tundra, J.M. Campbell (CNC) 1 female, 4 sex undetermined [paratypes]; British Mts., Sunday Mts., 680 m, 69°14'N, 140°05'W, 24.VI.1984, sifting pile of leaves stored by rodent, J.M. Campbell (CNC) 1 male, 2 females, 4 sex undetermined [paratypes]; Dawson City, 11.VII.1968, Campbell and Smetana (CNC) 1 male, 1 female; Dempster Hwy., mi 53, North Fork Pass, 24.VII.1978, 4200', A. Smetana and J.M. Campbell (CNC) 1 female, 1 sex undetermined [paratypes].</p><p>UNITED STATES OF AMERICA: Alaska: Prudhoe Bay Rd., 9 mi N Atigun Pass, 68°14'N, 149°25'W, 6.VII.1978, 3100', J.M. Campbell and A. Smetana (CNC) 1 female, 3 sex undetermined [paratypes]; mi 104.5 Denali Hwy., Brushkana Cr., 15.VII.1978, A. Smetana and J.M. Campbell (CNC) 1 sex undetermined [paratype]; mi 110 Denali Hwy., Seattle Cr., 15.VII.1978, J.M. Campbell and A. Smetana (CNC) 1 male [paratype]; mi 24, Wales Hwy., Hess Cr., 600', 65°40'N, 149°10'W, 10.VII.1978, J.M. Campbell and A. Smetana (CNC) 1 male [non-paratype].</p></div>	https://treatment.plazi.org/id/EA5D8ACEA50F34F91551BF0712C9279C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
9787AD1130EC0554932C0B6F143B7496.text	9787AD1130EC0554932C0B6F143B7496.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gnathusa alfacaribou Klimaszewski & Langor	<div><p>4. Gnathusa alfacaribou Klimaszewski &amp; Langor Figure 4 a–l, Map 3</p><p>Gnathusa alfacaribou Klimaszewski et al. 2011: 55-56.</p><p>Diagnosis.</p><p>Body length 3.0-3.4 mm, sides subparallel; body colour dark brown, with tarsi lighter and antennae brown and often with reddish tinge; head round and almost as wide as pronotum or at most as wide as pronotum, equal in size to pronotum; pronotum transverse, angular, about as wide as maximum width of elytra; abdomen subparallel, at base as wide as elytra, widest in apical half; antennal articles 5-10 quadrate to slightly transverse (Fig. 4a). MALE: tergite VIII pointed apically (Fig. 4c); sternite VIII slightly pointed at apex (Fig. 4d); median lobe of aedeagus with tubus strongly produced ventrally in lateral view, apex pointed (Fig. 4b). FEMALE: tergite VIII truncate apically (Fig. 4f); sternite VIII rounded apically (Fig. 4g); spermatheca pipe-shaped, with small spherical capsule and long, thin and almost straight stem (Fig. 4e).</p><p>Distribution.</p><p>This native Nearctic species is known only from Labrador (Map 3).</p><p>Bionomics.</p><p>This epigaeic species was collected from June to October using pitfall and flight intercept traps in black spruce-lichen, spruce-moss and old fir forests.</p><p>Locality data.</p><p>CANADA: Newfoundland: Labrador, Middle Brook, Lake Melville, Plot: MID 4, 17.VI.2005 (LFC) 1 male [holotype]; Labrador, Middle Brook, Lake Mel ville, Plot: MID 4, 17.VI.2005 (MUN) 1 male [paratype]; same data except: Plot: MID 3, 4.VII.2005 (MUN) 1 female [paratype], Plot: MID 4 (MUN) 1 female [paratype], Plot: MID 5, 18.VII.2005 (MUN) 1 female [paratype]; Labrador, Ossak Camp, Station 1, lichen-black spruce forest, 8.X.2004 (MUN) 1 female [paratype]; SW Labrador, 72 km E Labrador City, Rt. 500, km 93, 53° 08.6 N, 66° 05.9 W, 12-27.VIII.2001, S. and J. Peck, FIT, 600 m, spruce-moss forest 2001-34 (LFC) 1 female [paratype].</p></div>	https://treatment.plazi.org/id/9787AD1130EC0554932C0B6F143B7496	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
A7056D0E9B22482908D3E0BF2899A539.text	A7056D0E9B22482908D3E0BF2899A539.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mniusa Mulsant & Rey	<div><p>Mniusa Mulsant &amp; Rey</p><p>Mniusa Mulsant &amp; Rey, 1875. Type species: Homalota incrassata Mulsant &amp; Rey.</p><p>Diagnosis.</p><p>Body dark brown to black, compact, sides subparallel or body narrowly oval in outline (Figs 5a, 6a, 7a), length 2.0-3.2 mm; integument with distinct meshed microsculpture and moderately dense punctation and pubescence; head large with mandibles broad and long, left mandible with a small tooth (Figs 5h, 6h, 7h), and right one with a slightly larger tooth at the base of arcuate cutting edge of mandible, apices strongly narrowly elongate [more than in Ocyusa and less than in Gnathusa] (Figs 5h, i, 6h, i, 7h, i); infraorbital carina strong and complete; ligula shallowly split apically (Figs 5l, 6l, 7l); labial palpus with three articles, second article minute, last one needle-shaped (Figs 5l, 6l, 7l), and lacinia and galea as illustrated (Figs 5k, 6k, 7k); labrum narrow and transverse, apical edge entire (Figs 5j, 6j, 7j); frontal suture of head absent; pronotal pubescence along midline directed anteriad or obliquely anteriad in about apical third and posteriad or obliquely posteriad medio-basally; anterior margin of mesosternum without longitudinal carina; mesosternal process triangular basally and then produced and extending from 1/3 to almost 2/3 length of mesosternal cavities, metasternal process triangular in shape and short; isthmus short to long; median lobe of aedeagus strongly produced ventrally, internal sac structures as illustrated (Figs 5b, 6b, 7b); spermatheca with spherical or tubular capsule narrowed posteriorly into L-shaped neck, invagination narrow; stem narrowly elongate (Figs 5e, 6e, 7e).</p><p>Key to Canadian species of Mniusa</p><p>New provincial and territorial records are indicated in boldface font.</p></div>	https://treatment.plazi.org/id/A7056D0E9B22482908D3E0BF2899A539	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
B5A5DD0F50BC8AB4B4BE3135082072F8.text	B5A5DD0F50BC8AB4B4BE3135082072F8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mniusa minutissima (Klimaszewski & Langor 2011) Klimaszewski & Langor 2011	<div><p>1. Mniusa minutissima (Klimaszewski &amp; Langor, 2011) Figure 5 a–l, Map 3</p><p>Gnathusa minutissima Klimaszewski et al. 2011: 55.</p><p>Diagnosis.</p><p>Body length 2.0-2.3 mm, sides subparallel; body colour dark brown, with tarsi and often tibiae rust-brown, antennae brown; forebody with dense microsculpture, glossy and with moderately dense punctation and pubescence; head round, narrower than pronotum; pronotum transverse, rectangular in shape with sides feebly arcuate, and as wide as elytra; elytra at suture as long as pronotum (Fig. 5a); abdomen subparallel, narrower than elytra with deep basal impressions on first three visible tergites; antennae with articles V-X strongly transverse, with the outer segments at least twice as wide as long (Fig. 5a). MALE: male tergite VIII widely truncate apically (Fig. 5c); sternite VIII slightly produced at apex (Fig. 5d); median lobe of aedeagus with straight venter of tubus slightly arched laterally and internal sac with band-formed, subapical structure (Fig. 5b). FEMALE: tergite VIII truncate apically (Fig. 5f); sternite VIII rounded apically (Fig. 5g); spermatheca with small spherical capsule with small invagination, short L-shaped neck, and long, thin and broadly curved stem (Fig. 5e).</p><p>Distribution.</p><p>This native Nearctic species was described from Newfoundland and is herein recorded for the first time from New Brunswick (Map 3).</p><p>Bionomics.</p><p>Adults were collected from May to July using pitfall traps in an old boreal balsam fir forest in Newfoundland; by sifting moss near a brook, sifting deep conifer litter at base of large red spruce in a mature red spruce forest, and from Lindgren funnel traps in a rich Appalachian hardwood forest in New Brunswick.</p><p>Locality data.</p><p>CANADA: Newfoundland: Little Grand L., 2 km E. Martin Pond, 24. VI– 15.VII.1992, old fir forest, pitfall 13, (LFC) 1 male [holotype]; same data except: pitfall 19, (CFS-CB) 1 male and 1 female [paratypes]; same data except: pitfall 20, 2 males and 1 female [paratypes]; pitfall 16, 1 female [paratype]; pitfall 13, 1 female [paratype]; pitfall 20, (LFC) 1 female paratype; Little Grand L., Bakeapple Brook, 24. VI– 15.VII.1992, old fir forest, pitfall 1, (CFS-CB) 1 male [paratype]; same data except: pitfall 4, 1 male; pitfall 11, 1 female [paratype]; pitfall 3, (LFC) 1 female [paratype]; Manuals R., 8 km W. St. John’s, 10.VI.1984, D. Langor, Lot, (CFS-CB) 1 female [paratype]. New Brunswick: Sunbury Co., Acadia Research Forest, 45.9799°N, 66.3394°W, 14.V.2007, 18.VI.2007 // mature red spruce and red maple forest, sifting moss near brook, R.P. Webster (LFC, RWC) 3 males, 2 females; same locality data and forest type but 14.V.2007 // sifting deep conifer litter at base of large red spruce (RWC) 2 females: Carleton Co., Jackson Falls, "Bell Forest", 46.2200°N, 67.7231°W, 4-12.VI.2008, R.P. Webster // Rich Appalachian Hardwood Forest with some conifers, Lindgren funnel trap (RWC) 1 female; same locality data and forest type but 1-8.VI.2009, 8-16.VI.2009, R. Webster &amp; M.-A. Giguère, Lindgren funnel trap (RWC) 2 males.</p></div>	https://treatment.plazi.org/id/B5A5DD0F50BC8AB4B4BE3135082072F8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
F660C9F7D4144843F15B69DF3C0BADF6.text	F660C9F7D4144843F15B69DF3C0BADF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mniusa yukonensis Klimaszewski & Godin	<div><p>2. Mniusa yukonensis Klimaszewski &amp; Godin Figure 6 a–l, Map 6</p><p>Ocyusa yukonensis Klimaszewski &amp; Godin, in Klimaszewski et al. 2012: 218.</p><p>Diagnosis.</p><p>Body length 2.8-3.0 mm, narrowly elongate and broadest at elytra; body dark brown to almost black, sometimes with reddish tinge on elytra, reddish-brown antennae and legs; forebody with dense microsculpture, dense punctation and pubescence, and strongly glossy; head round and narrower than pronotum; pronotum transverse, with sides strongly arcuate and narrowed anteriad, slightly narrower than elytra; elytra at suture about as long as pronotum or slightly longer (Fig. 6a); abdomen broadly arcuate laterally, slightly narrower than elytra and with basal impressions on first three visible tergites; antennal articles V-X slightly transverse (Fig. 6a). MALE: male tergite VIII truncate apically (Fig. 6c); sternite VIII rounded apically and slightly produced medially (Fig. 6d); median lobe of aedeagus with tubus straight and slightly produced ventrally and with complex structures of internal sac (Fig. 6b). FEMALE: tergite VIII truncate apically with slightly acute lateral angles (Fig. 6f); sternite VIII rounded apically and slightly produced apically (Fig. 6g); spermatheca with narrowly elongate sac-shaped capsule connected with L-shaped neck and narrow and long stem (Fig. 6e).</p><p>Distribution.</p><p>This native Nearctic species was recently described from the Yukon Territory (Klimaszewski et al. 2012), and is here newly reported from Nova Scotia, New Brunswick, Quebec, and British Columbia, which constitute new provincial records (Map 6).</p><p>Bionomics.</p><p>Adults were collected from May to July in pitfall traps, flight intercept traps, and Lindgren funnel traps in various forest types: rich Appalachian hardwood forest with some conifers, old-growth white spruce/balsam fir forest, balsam fir and maple sugar stands, mature white spruce with feather moss, and a red spruce forest.</p><p>Locality data.</p><p>CANADA: Nova Scotia: Colchester Co., Debert, 6.V.1993, J. Ogden (NSPM) 1 male; Black Duck Lake, NS, 22.VI.2003, 4U: Funnel 16: 844 WPi/RSp (40-80 y), P. Dollin (NSPM) 1 male. New Brunswick: Carleton Co., Jackson Falls, "Bell Forest", 46.2200°N, 67.7231°W, 4-12.VI.2008//, Rich Appalachian hardwood forest with some conifers, Lindgren funnel traps, R.P. Webster (RWC) 1 male, 2 females; same data except 12-19.VI.2008 (RWC) 1 female; same data except 19 -27.VI.2008 (RWC) 1 female: same data except 26. V– 1.VI.2009, R. Webster &amp; M.-A. Giguère (RWC) 1 male; Restigouche Co., Dionne Brook P.N.A., 47.9064°N, 68.3441°W, 31. V– 15.VI.2011 // old-growth white spruce and balsam fir forest, flight intercept trap, M. Roy &amp; V. Webster (RWC) 1 male; same data except 31. V– 1.VI.2011, Lindgren funnel trap (RWC) 3 males; same data except 15-27.VI.2011, Lindgren funnel trap (RWC) 1 female; same data except 21. VI– 14.VII,2011, Lindgren funnel trap (RWC) 1 female. Quebec: Dosquet Co., Lotb. Quebec, 27.IV.1984, Claude Chantal (LFC) 1 male; Pelegrin, North of Chandler, 48°32'N, 64°54'W, SAP Lindgren, 21.06.1994 (LFC) 1 male; Tremblant, SAP Lindgren, 28.VI.1994 (LFC) 1 female; La Tuque, SAP Lindgren, 11.VII.1994 (LFC) 1 female; St-Jacques-de-Leeds, Sapinière Quebec, 31.V.1993, 9.VI.1993, 16.VI.1993, 28.VI.1993, 30.VI.1993, Lindgren 1, 2, 4 (LFC) 6 females. British Columbia: 25 km SW Campbell River, 49°50'21"N, 125°28'34"W, 23.V-6.VI.1996, Balsam Cr. LT 5-D 10, J. Lemieux (LFC) 4 males, 1 female, 1 sex undetermined; Copper River Valley, A36435/04-1-1, 07.VI-6.VII.1996, pitfall trap, J. Lemieux (LFC) 1 male. Yukon Territory: EMAN Plot (Ecological Monitoring and Assessment Network), mature white spruce and feather moss forest, 60.5963°N, 134.9522°W, 8.VII.2003, 738 m, yellow pitfall trap (LMKM31Y), (LFC) 1 male [holotype]; EMAN Plot, 60.5963°N, 134.9522°W, 24.VII.2003, 738 m, black pitfall trap (LMKM31B), (ECW) 1 male [paratype].</p></div>	https://treatment.plazi.org/id/F660C9F7D4144843F15B69DF3C0BADF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
0FB603F29313E156E3934E689B816F6F.text	0FB603F29313E156E3934E689B816F6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mniusa odelli Klimaszewski & Webster	<div><p>3. Mniusa odelli Klimaszewski &amp; Webster sp. n. Figure 7 a–l, Map 5</p><p>Holotype</p><p>(female). CANADA: New Brunswick, York Co., Fredericton, Odell Park, 45.9571°N, 66.6650°W, 15.V-1.VI.2012 // Old-growth eastern hemlock forest, Lindgren funnel trap, 1 m high under Betula alleghaniensis, C. Alderson and V. Webster (LFC). PARATYPES: New Brunswick, York Co., Odell Park, 45.9539°N, 66.6666°W, 10-24.VI.2013 // Hardwood stand, Lindgren funnel trap, 1 m high under trees (RWC) 2 females.</p><p>Diagnosis.</p><p>Body length 2.8-3.2 mm, narrowly subparallel with head slightly narrower than pronotum; body colour dark brown, and with tibiae, tarsi and often basal antennal articles reddish-brown; forebody with dense microsculpture, and moderately dense punctation and pubescence, and strongly glossy; head round and slightly narrower than pronotum; pronotum slightly transverse, with sides rounded, and as wide as elytra; elytra at suture about as long as pronotum or slightly longer (Fig. 7a); abdomen subparallel, as wide as elytra and with basal impressions on first three visible tergites; antennal articles V-X slightly transverse (Fig. 7a). MALE [description of male is based on poorly preserved specimen and should be consider as tentative]: male tergite VIII truncate apically (Fig. 7c); sternite VIII rounded apically and slightly produced medially (Fig. 7d); median lobe of aedeagus with tubus straight and with complex structures of internal sac (Fig. 7b). FEMALE: tergite VIII truncate apically with sharp lateral angles (Fig. 7f); sternite VIII rounded apically (Fig. 7g); spermatheca with narrowly elongate sac-shaped capsule connected with L-shaped neck and narrow and long stem slightly sinuate posteriorly at apex (Fig. 7e).</p><p>Distribution .</p><p>This native Nearctic species is here described from Nova Scotia, New Brunswick, and Quebec (Map 5).</p><p>Bionomics.</p><p>Adults were collected from May to July in Lindgren traps in an old-growth eastern hemlock stand, an old hardwood stand, and in a sugar maple forest and a red spruce forest.</p><p>Etymology.</p><p>This species is named after Odell Park in Fredericton, New Brunswick, where the holotype was found. This park was originally the estate of Reverend Jonathan Odell whom the park was named after. This park was established in 1954.</p><p>Other locality data</p><p>[specimens in poor condition and tentatively identified as this species].</p><p>CANADA: Nova Scotia: Pictou Co., Marshy Hope, 17.V.1995, ethanol lure, M. LeBlanc (NSPM) 1 sex undetermined; Antigonish Co., Fairmont Tower Road, 17.V.1995, 3-Component Lure w/+/- mcol, M. LeBlanc (NSPM) 1 female; Fairmont Tower Road, 25.V.1995, 3-Component Lure w/+/- mcol, M. LeBlanc (NSPM) 1 male, 1 female; Antigonish Co., Eigg Mountain, 25.V.1995, ethanol lure, M. LeBlanc (NSPM) 1 male, 1 female; Melopseketch Lake, Guy, 14. V– 2.VI.1997, young red spruce, D.J. Bishop 201 (NSPM) 1 male; Halifax, Lake Little, 14. V– 2.VI.1997, regenerating red spruce, D.J. Bishop 127 (NSPM) 1 male. Quebec: St-Jacques-de-Leeds, Erablière Québec, 28.VI.1993, Lindgren 2 (LFC) 1 female; Mont Orford, 29.VI-6.VII.1999, Lindgren 1, Erablière, 99-3-1061 (LFC) 1 male.</p></div>	https://treatment.plazi.org/id/0FB603F29313E156E3934E689B816F6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
A28EAD4F026BCDE251279B7A5C5AAF35.text	A28EAD4F026BCDE251279B7A5C5AAF35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ocyusa Kraatz	<div><p>Ocyusa Kraatz</p><p>Ocyusa Kraatz, 1856. Type species: Oxypoda maura Erichson.</p><p>Diagnosis.</p><p>Body dark brown to almost black, compact, sides subparallel or body narrowly oval in outline (Figs 8a, 9a), length 2.5-3.5 mm; integument with distinct meshed microsculpture and moderate to dense punctation and pubescence; head large with mandibles broad and long, left mandible with a small tooth (Figs 8h, 9h), and right one with a slightly larger tooth at the base of arcuate cutting edge of mandible, apices moderately narrowly elongate [less than in Mniusa] (Figs 8i, 9i); infraorbital carina strong and complete; ligula shallowly split apically (Figs 8l, 9l); labial palpus with three articles, second article minute, last one needle-shaped and lacinia and galea as illustrated (Figs 8k, 9k); labrum narrow and transverse, apical edge entire (Figs 8j, 9j); frontal suture of head present; pronotal pubescence along midline directed posteriad or obliquely posteriad; anterior margin of mesosternum without longitudinal carina; mesosternal process triangular basally and then produced and extending to about 2/3 length of mesosternal cavities, metasternal process narrowly triangular in shape and short; isthmus short; median lobe of aedeagus strongly produced ventrally, internal sac structures as illustrated (Figs 8b, 9b); spermatheca S-shaped with spherical capsule, short neck and elongate sinuate stem (Figs 8e, 9e).</p><p>Key to Canadian species of Ocyusa</p></div>	https://treatment.plazi.org/id/A28EAD4F026BCDE251279B7A5C5AAF35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
CE09EF65A6E5CBBD9808959E7D2070D7.text	CE09EF65A6E5CBBD9808959E7D2070D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ocyusa canadensis Lohse	<div><p>1. Ocyusa canadensis Lohse Figure 8 a–l, Map 4</p><p>Ocyusa canadensis Lohse, in Lohse et al. 1990: 147; Brunke et al. 2012: 134.</p><p>Diagnosis.</p><p>Body length 2.5-3.0 mm, narrowly elongate and broadest at elytra; body colour dark brown to almost black, with tarsi, two basal antennal articles and tibiae rust-brown, rest of antennal articles brown; forebody with dense microsculpture, moderate punctation and pubescence, and strongly glossy; head round and narrower than pronotum; pronotum transverse, rectangular in shape with sides strongly arcuate, and narrower than elytra; elytra at suture about as long as pronotum (Fig. 8a); abdomen broadly arcuate laterally, slightly narrower than elytra and with basal impressions on first three visible tergites; antennae with articles V-X subquadrate to slightly elongate (Fig. 8a). MALE: male tergite VIII broadly rounded apically (Fig. 8c); sternite VIII rounded apically (Fig. 8d); median lobe of aedeagus with tubus strongly bent ventrally and with pronounced structures of internal sac (Fig. 8b). FEMALE: tergite VIII truncate apically (Fig. 8f); sternite VIII rounded apically (Fig. 8g); spermatheca with semi-spherical capsule with long invagination, and S-shaped broad stem (Fig. 8e).</p><p>Distribution.</p><p>This native Nearctic species is known from Alaska, Yukon Territory and Ontario (Brunke et al. 2012), and is here newly reported from the island of Newfoundland and New Brunswick [new provincial records] (Map 4).</p><p>Bionomics.</p><p>Some adults were collected from June to July at lake margins, on moist soil/gravel among sedges and by treading Carex and grasses.</p><p>Locality data.</p><p>CANADA: Newfoundland: George’s Lake, Corner Brook, 29.VII.1972, J.M. Campbell (CNC) 1 male; New Brunswick: Restigouche Co., Wild Goose Lake, 419 m elevation, 47.8540°N, 68.3200°W, 21.VII.2010 // lake margin, on moist soil/gravel among sedges, R.P. Webster (RWC) 1 female; same data except: 420 m elevation, 47.8543°N, 68.3219°W, 7.VI.2011 // lake margin with emergent Carex and grasses, treading Carex and grasses, R.P. Webster (RWC) 1 male; same data except 20.VI.2011, (RWC) 2 males, 4 females. Ontario: Timiskaming Distr., 52 km South Armstrong, 27.VI.1973, R. Parry and J.M. Campbell (CNC) 4 females, 7 sex undetermined; 52 km S Armstrong, 27.VI.1973, R. Parry and J.M. Campbell (CNC) 1 female. Yukon Territory: Dempster Hwy., mi. 122, 20.VII.1978, 2000 feet, A. Smetana and J.M. Campbell (CNC) 2 males, holotype and paratype; Dempster Hwy., mi. 147, 1900 feet, 22.VII.1978, J.M. Campbell and A. Smetana (CNC) 1 sex undetermined.</p><p>UNITED STATES OF AMERICA: Alaska: Prudhoe Bay Rd., Bonanza Creek, 900 feet, 66°40'N, 150°40'W, 2.VII.1978, A. Smetana and J.M. Campbell (CNC) 1 paratype sex undetermined; Nutirwick Creek, 67°55'N, 149°45'W, 2300 feet, 8.VII.1978, J.M. Campbell and A. Smetana (CNC) 2 sex undetermined.</p></div>	https://treatment.plazi.org/id/CE09EF65A6E5CBBD9808959E7D2070D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
0579CE7AC747EFE6B27729AF21563CA0.text	0579CE7AC747EFE6B27729AF21563CA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ocyusa asperula Casey	<div><p>2. Ocyusa asperula Casey Figure 9 a–l, Map 5</p><p>Ocyusa asperula Casey 1894: 305 [often cited as 1893], Webster et al. 2009: 192.</p><p>Ocyusa brevipennis Bernhauer 1906: 344. Moore and Legner 1975: 458. Synonymy confirmed.</p><p>Diagnosis.</p><p>Body length 2.8-3.0 mm, sides subparallel; body colour dark brown, with tarsi, two basal antennal articles and legs rust-brown, rest of antennal articles dark brown; forebody with moderately dense microsculpture, punctation and pubescence, and strongly glossy; head round and about as wide as pronotum; pronotum transverse, with sides strongly arcuate, widest in apical third, and as wide as elytra; elytra at suture much shorter than pronotum (Fig. 9a); abdomen broadly arcuate laterally, slightly broader than elytra at middle and with basal impressions on first three visible tergites; antennae with articles V-X subquadrate (Fig. 9a). MALE: male tergite VIII with apical margin slightly pointed medially (Fig. 9c); sternite VIII rounded apically (Fig. 9d); median lobe of aedeagus with tubus convex basally and then strongly bent ventrally and with complex structures of the internal sac (Fig. 9b). FEMALE: tergite VIII truncate apically with small projections laterally (Fig. 9f); sternite VIII rounded apically (Fig. 9g); spermatheca with small spherical capsule with long and broad invagination, and S-shaped broad stem slightly swollen posteriorly (Fig. 9e).</p><p>Distribution.</p><p>This native Nearctic species was described from Rhode Island by Casey 1894 [often cited as 1893]. It was recorded also from New Brunswick in Canada (Map 5) and from Iowa and Massachusetts in the United States (Casey 1894, Bernhauer 1906 [as brevipennis], Moore and Legner 1975, Webster et al. 2009).</p><p>Bionomics.</p><p>Some adults were collected from April to July at lake margins, on moist soil/gravel among sedges, and by treading emergent Carex and grasses. Webster et al. (2009) collected adults by sifting grass litter and mosses (usually sphagnum) near small pools in eastern white-cedar swamps, red maple ( Acer rubrum L.) swamps with eastern white-cedar, and in alder swamps. Others were collected by treading green sphagnum, Carex, and grasses in a black spruce bog and by treading cattails and sedges in a boggy marsh.</p><p>Locality data.</p><p>CANADA: New Brunswick: Charlotte Co., 3 km SW of King Brook Lake, 45.3194°N, 67.4414°W, 27.V.2007 (RWC) 1 sex undetermined; 3.0 km NW of Pomeroy Ridge, 45.3059°N, 67.4343°W, 5.VI.2008 (RWC) 1 sex undetermined; Gloucester Co., ca. 1.5 km NE of Six Roads, off Paleot Rd., 47.6292 °N, 64.8565°W, 32.V.2010, R.P. Webster (RWC) 1 sex undetermined; Northumberland Co., Goodfellow Brook PNA, 46.8943°N, 65.3796°W, 23.V.2007 (BM) 1 sex undetermined; York Co., New Maryland, off Hwy 2, E of Baker Brook, 45.8760°N, 66.6252°W, 6.IV.2005 (RWC) 1 male; near Mazerolle Settlement, 45.8987 °N, 66.7903°W, 9.IV.2006, R.P. Webster (LFC, RWC, NBM) 6 males, 3 females; 9.2 km W of Tracy off Rt. 645, 45.6837°N, 66.8809°W, 22.V.2008 (RWC) 1 female; ca. 14 km SW of Tracy, S of Rt. 645, 45.6603°N, 66.8603°W, 2.VII.2010, R.P. Webster (RWC) 1 sex undetermined.</p></div>	https://treatment.plazi.org/id/0579CE7AC747EFE6B27729AF21563CA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Bourdon, Caroline;Hammond, H. E. James;Pohl, Greg R.;Godin, Benoit	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R., Godin, Benoit (2014): Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant & Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 412: 9-40, DOI: http://dx.doi.org/10.3897/zookeys.412.7282, URL: http://dx.doi.org/10.3897/zookeys.412.7282
