taxonID	type	description	language	source
03FC87D3EB00FF8401B2FB4B836540E0.taxon	materials_examined	Type species. Mesoleptobasis incus Sjöstedt 1918 by monotypy. [NOTE: Davies & Tobin (1984) followed by Tsuda (2000), Steinmann (2007), and Heckman (2008) listed this species as Mesoleptobasis inca apparently to conform to Art. 31.2.1 of the Code. The species epithet incus is a noun in apposition (f., Latin: anvil), probably referring to the shape of the pronotum, and therefore should not be changed [Arts. 32. 2.2 and 31.2.3]. Other species included. M. acuminata Santos 1961, M. cantralli Santos 1961, M. cyanolineata (Wasscher 1998), M. elongata sp. nov.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB00FF8401B2FB4B836540E0.taxon	description	Characterization. Medium sized (36 – 45 mm), largely pale coenagrionids with a long abdomen (ratio of 5.1 – 6.36 to length of head plus thorax); head and dorsum of abdomen orange to brown or black with some metallic iridescence, pale areas yellow or light blue. Blue postocular spots present or absent and pale occipital bar absent. Pterothorax lacking dark mid-dorsal and metapleural stripes; in mature specimens brown mesepisternal and metepisternal stripes and pale antehumeral, mesepimeral, and metepisternal stripes present (Fig. 8 a); in immature specimens pterothorax entirely pale brown or orange. Frons rounded; location of most posterior point of head at level of eyes (Fig. 1 a). Posterior lobe of prothorax in male with long projections, which may be represented by lateral processes (Fig. 4 a) or by a medial bifurcate process (Figs. 4 b – e); pronotum in female projected as in male (in M. cyanolineata; Fig. 5 d) or slightly trilobate (in M. acuminata, M. cantralli, M. elongata, and M. incus; Figs. 5 a – c, e – g). Hind femur short, not reaching anterior margin of S 1; metatibial spurs shorter than twice intervening spaces (Fig. 8 a); pretarsus with vestigial supplementary tooth represented by an obtuse low prominence (Fig. 9 a). CuA extending from two to four (Figs. 10 a) or six or seven cells (in M. cyanolineata and females of M. acuminata; Figs. 11 a, 12 a, 13 a) distal to vein descending from subnodus to one or two cells proximal to vein descending from subnodus (Figs. 10 b, 11 b, c); CuP reaching posterior margin of wing; vein descending from quadrangle forming an unbroken line to wing margin (Figs. 10 – 13). Genital ligula distal segment with inner fold; with sclerotized areas present as spine-like projections on lateral margins (Fig. 15); with two pairs of lateral lobes, a smaller latero-basal one and a larger latero-medial one; apex transverse (M. acuminata; Fig. 14 a) or deeply bifid (Figs. 14 b – e). Postero-dorsal margin of male S 10 projected caudally and with a pair of medial lobe-like projections (M. cantralli, M. elongata, and M. incus; Figs. 17 b, d, e, 18 b, d, e, 19 a – c) or emarginate and lacking lobe-like projections (M. acuminata and M. cyanolineata; Figs. 17 a, c, 18 a, c). Male cercus entire, approximately horizontal with tip bent ventrally, shorter than to subequal to S 10 (Figs. 17 – 19), with an elongate membranous depression on dorsal surface only in M. acuminata (Fig. 17 a); male paraproct forcipate, much longer than S 10, with tip recurved medio-ventrally (Figs. 17 – 19). Female lacking vulvar spine on S 8 (Figs. 16 a, b) or with a very small spine in some females of M. cyanolineata (reported by Wasscher 1998 for allotype) and M. incus (Fig. 16 e); postero-dorsal margin of S 9 with denticles; ovipositor extending beyond tips of cerci for a distance shorter (Figs. 16 a, c, e) to subequal (Fig. 16 d) or longer (Fig. 16 b) than cerci length.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB00FF8401B2FB4B836540E0.taxon	diagnosis	Diagnosis. Within New World Coenagrionidae, Mesoleptobasis shares only with Aceratobasis and Metaleptobasis the combination of a pretarsus with vestigial supplementary tooth represented by an obtuse low prominence or else absent (Fig. 9) and a vein descending from quadrangle forming an unbroken line to wing margin (Figs. 10 – 13). However, it differs from both Aceratobasis and Metaleptobasis by its rounded frons (Fig. 1 a), highly modified pronotum with long processes at least in males (Figs. 3, 4, 5 d), pterothoracic color pattern lacking dark mid-dorsal stripe (Fig. 8 a), and genital ligula with a small inner fold and with spinelike projections on lateral margins distal to flexure (Fig. 15), this last character being unique for the genus. In Aceratobasis and Metaleptobasis the frons is angled (Fig. 1 b), the pronotum is devoid of long projections (Fig. 2), and the distal segment of genital ligula lacks an inner fold, lateral lobes, and any sclerotized projections distal to flexure (Cumming 1954; Rácenis 1955; von Ellenrieder & Garrison 2008). CuA in Metaleptobasis extends more than seven cells posterior to the vein descending from the subnodus and males of all species have long mesepisternal horns (Fig. 8 b), which in females are similar in size and length (Figs. 6 b, c) or may be reduced to tubercles in some species (Fig. 6 d). In Mesoleptobasis CuA never extends more than 7 cells distal to the vein descending from the subnodus (Figs. 10 – 13) and the mesepisterna are always smooth (Figs. 6 a, 7). Mesoleptobasis further differs from Aceratobasis by having male cerci entire and shorter than long, forcipate paraprocts (Figs. 17 – 19); cerci in Aceratobasis are branched with a ventro-basal spur or spine and are longer than the paraprocts.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB00FF8401B2FB4B836540E0.taxon	distribution	Distribution. Amazon forest in Venezuela, Guyana, Surinam, Brazil, and Peru (Fig. 21). Specimens are infrequently collected, and it is therefore likely that distribution ranges are larger than those indicated by existing specimens. Habitat. Adults found within forest, in dark or shaded areas flying near the ground; some specimens of M. cyanolineata were captured at night attracted to lights (Wasscher 1998); breeding habitat and larvae unknown.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB07FF8801B2FCF0817040DE.taxon	description	Specimens examined. Total: 3 3, 4 Ƥ. Peru, Loreto Department: 1 3, Explorama Lodge, 80 km NE of Iquitos on Amazon River at junction with Yanamono River (3 ° 21 ' 59 " S, 72 ° 47 ' 56 " W), 14 viii 1989, leg. S. W. Dunkle (RWG); 2 3, same but 13 viii 1989 (RWG); 2 Ƥ, same but 17 viii 1989 (RWG); 2 Ƥ, same but 31 viii 1989 (RWG); Explornapo Camp at junction of Sucusari River and Napo River, ca 160 km NE of Iquitos (3 ° 16 ' 33 " S, 72 ° 56 ' 18 " W), 27 viii 1989, leg. S. W. Dunkle (SWD).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB07FF8801B2FCF0817040DE.taxon	diagnosis	Diagnosis. Male prothorax with a pair of lateral projections separated at base (Fig. 4 a; unique) and with acuminate apices directed antero-laterally; female prothorax lacking processes, with posterior margin slightly trilobate, with smoothly convex medial lobe slightly projected posteriorly beyond level of lateral lobes (Fig. 5 a; unique). Costal side of FW pt shorter than basal side (unique), its posterior margin strongly convex in male (Fig. 10 a), moderately convex in female (Fig. 12 a). CuA relatively long (shared with M. cyanolineata), extending one and a half to four cells distal to vein descending from subnodus in male, three to seven cells in female (Figs. 10 a, 12 a). Genital ligula in ectal view with distal margin transverse and lateral sub-apical deep emarginations (Fig. 14 a; unique); in lateral view with a large triangular lateral lobe bearing a spine at its posterior base (Fig. 15 a; unique), and with an inconspicuous low latero-basal lobe (shared with M. cyanolineata). Posterior margin of male S 10 recessed and with a medio-dorsal emargination, lacking posterolateral processes (Figs. 17 a, 18 a; shared with M. cyanolineata). Male cercus subtriangular and attenuate dorsally with a membranous central area (Fig. 17 a; unique); in lateral view smoothly curved, with tip directed postero-ventrally (Fig. 18 b; shared with M. cantralli and M. incus); male paraproct about as high as half of S 10 height at base in lateral view (Fig. 18 a; shared with M. cyanolineata); base of paraproct lacking a thumblike tubercle (Figs. 17 a, 18 a; shared with M. cyanolineata and M. incus). Ovipositor surpassing tip of cerci for a distance shorter than length of cerci (Fig. 16 a; shared with M. cyanolineata and M. incus). Dimensions. Males (n 3; mean in parenthesis): Hw 18.5 – 19.0 (18.8); abdomen 32.0 – 33.0 (32.7); total length 38.0 – 39.0 (38.3). Females (n 4): Hw 20.0 – 21.0 (20.6); abdomen 32.0 – 33.0 (32.25); total length 37.0 – 39.0 (38.0).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB07FF8801B2FCF0817040DE.taxon	discussion	Remarks. Santos (1961) described this species from one male lacking locality data, which he designated as holotype, and two females from Porto Velho, Rondônia, Brazil. His description and figures of the holotype show no central membranous area on the dorsal surface of male cercus characteristic of the males from Peru we examined (Fig. 17 a). Unfortunately the holotype is incomplete and only its wings and hind legs remain (J. M. Costa pers. comm.). J. M. Costa kindly illustrated a pair of its wings for us, which had not been figured by Santos (1961). Her drawing (J. M. Costa in litt.) shows Fw Pt as less markedly convex along posterior margin than in the male we illustrated (Fig. 10 a), and Hw Pt rectangular, with costal and posterior sides slightly longer than basal and distal sides, rather than rhomboidal, with costal and posterior sides slightly shorter than basal and distal sides, as in our illustration (Fig. 10 a). The remainder of Santos’ (1961) description and illustrations of the holotype fully agree with males we ascribe to this species, and we consider them conspecific. The females described by Santos (1961) as M. acuminata (allotype and paratype) are still in the MNRJ and not in the UMMZ as stated in the description (J. M. Costa pers. comm.). J. M. Costa sent us illustrations of one pair of wings, posterior lobe of pronotum, and S 8 - 10 of female allotype (J. M. Costa in litt.). According to her drawings, pterostigmata are unmodified and rectangular and ovipositor is short, not surpassing tip of cerci. The posterior lobe of the pronotum has lateral processes similar to those of males of M. acuminata, but the posterior margin between the lateral processes is bilobate lacking a medial lobe, rather than trilobate with a medial lobe as in the holotype and in our males (Fig. 4 a). We believe these females are not conspecific with the holotype male, and belong instead to an undescribed species (M. sp. in the key). We have females collected at the same locality as males we ascribe to M. acuminata, and they lack the long lateral prothoracic processes mentioned by Santos (1961). They have instead a smooth pronotum with slightly trilobate posterior margin (Fig. 5 a), similar to the trilobate margin of male (Fig. 4 a). They also differ by their ovipositor, which is longer (surpassing tip of cerci; Fig. 16 a) compared to the illustrations by Santos (1961, fig. 12) and J. M. Costa (in. litt.). We believe we have correctly associated the female sex of M. acuminata since it shares the characteristic modified pterostigma of male (unique for this species within Mesoleptobasis), although the modification is less pronounced than in male. The identity of the two females described by Santos (1961) as M. acuminata will remain uncertain until more of these females are found in association with males.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB07FF8801B2FCF0817040DE.taxon	distribution	Distribution. Amazonian region of Peru (Fig. 21).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0AFF8901B2F8ED8719447E.taxon	description	Specimens examined. Total: 5 3, 8 Ƥ. Brazil, Rondônia State: 1 paratype 3, Porto Velho (8 ° 46 ' S, 63 ° 53 ' W), 22 ii 1922, leg. J. H. Williamson & J. W. Strohm (RWG); 1 paratype Ƥ, same but 24 iv 1922 (RWG); 1 paratype Ƥ, same but (UMMZ); 1 paratype 3, same but 27 ii 1922 (RWG); 1 paratype Ƥ, same but 0 4 v 1922 (RWG); 2 paratypes 3, 1 paratype Ƥ, same but 14 v 1922 (UMMZ); 3 paratypes Ƥ, same but 16 v 1922 (UMMZ); Amazonas State: 1 3, 1 Ƥ, Rio Uaupés, Taraquá (3 ° 27 ' 15 '' S, 62 ° 51 ' 5 '' W), 14 viii 1964, leg. A. B. M. Machado & Pereira (ABMM).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0AFF8901B2F8ED8719447E.taxon	diagnosis	Diagnosis. Male prothorax with a medial bifurcate process with arms forming a transverse line between them (Fig. 4 b; shared with M. elongata and M. incus) and apices directed anteriorly (Fig. 3 b; shared with M. acuminata and M. cyanolineata); female prothorax lacking processes, with posterior margin slightly trilobate and medial lobe not surpassing lateral lobes posteriorly; lateral lobes not bent anteriorly (Figs. 5 b, c; shared with M. elongata). Costal side of FW pt longer than basal side, its posterior margin slightly convex in both sexes (Figs. 10 b, 12 b; shared with M. cyanolineata, M. elongata, and M. incus). WF CuA relatively short (shared with M. elongata and M. incus), ending from one cell proximal to vein descending from subnodus (Fig. 10 b) to level of vein descending from subnodus (Fig. 12 b) in both sexes. Genital ligula in ectal view with distal margin deeply bifid and lacking lateral emarginations (Fig. 14 b; shared with M. cyanolineata, M. elongata, and M. incus); in lateral view with basal lobe sclerotized, long, pointed, and directed posteriorly, and dorsal margin of lateral lobe with a single small sclerotized spine (Fig. 15 b; shared with M. incus). Posterior margin of S 10 projected medio-dorsally, with a pair of postero-lateral small lobe-like processes (Fig. 17 b, 19 a; shared with M. elongata and M. incus). Male cercus lacking a membranous area dorsally (shared with M. cyanolineata, M. elongata, and M. incus), oblong, longer than wide (Figs. 17 b, 19 a; unique), and usually armed at tip with a small externally recurved tooth (Figs. 17 b, 19 a); in lateral view smoothly curved, with tip directed postero-ventrally (Fig. 18 b, shared with M. acuminata, M. elongata, and M. incus); paraproct slender and narrower than half of S 10 height at base in lateral view (Fig. 18 b, shared with M. elongata and M. incus); base of paraproct usually with well developed thumb-like process (Figs. 17 b, 18 b, 19 a; shared with M. elongata), which can be vestigial. Ovipositor surpassing tip of cerci for a distance subequal to length of cerci (Fig. 16 b; shared with M. elongata). Dimensions. Males (n 5; mean in parenthesis): Hw 17.0 – 18.0 (17.6); abdomen 33.0 – 36.0 (34.6); total length 39.0 – 41.0 (39.8). Females (n 8): Hw 18.5 – 20.0 (19.1); abdomen 30.0 – 34.0 (31.9); total length 35.0 – 39.0 (37.5).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0AFF8901B2F8ED8719447E.taxon	discussion	Remarks. The pair of M. cantralli from Taraquá differs from the type series as follows: male lacks the small externally recurved tooth (as in Figs. 17 b, 19 a), and the thumb-like process at the base of paraproct is vestigial; female middle lobe of hind lobe of pronotum is more strongly developed, but still does not surpass lateral lobes. Wing venation, shape of male pronotal process, male cerci, and morphology of hind lobe of female pronotum as well as length of ovipositor (surpassing tip of cerci for a distance subequal to length of cerci as in Fig. 16 b) match those for the type series and indicate that this pair is correctly placed here.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0AFF8901B2F8ED8719447E.taxon	distribution	Distribution. Amazonas and Rondônia State in Brazil; sympatric with M. incus in the latter (Fig. 21).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0BFF8C01B2F99B86CA4511.taxon	description	Metaleptobasis (?), sp. — Williamson 1915: 604 – 607 (partial description and illustrations of Ƥ from Rockstone, Guyana).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0BFF8C01B2F99B86CA4511.taxon	diagnosis	Diagnosis. Male prothorax with medial bifurcate process with arms directed latero-posteriorly forming a v-shaped line between them (Fig. 4 c; unique); female prothorax with similar processes (Fig. 5 d; unique). Costal side of FW pt longer than basal side, its posterior margin slightly convex in both sexes (Fig. 11 a; shared with M. cantralli, M. elongata, and M. incus). CuA relatively long (shared with M. acuminata), extending four to six cells distal to vein descending from subnodus in male (Fig. 11 a), two to six cells in female (Fig. 13). Genital ligula in ectal view with distal margin deeply bifid and lacking lateral emarginations (Fig. 14 c; shared with M. cantralli, M. elongata, and M. incus); in lateral view with a small triangular lateral lobe bearing a series of denticles along its dorsal margin (Fig. 15 c; unique), and with an inconspicuous low latero-basal lobe (shared with M. acuminata). Posterior margin of male S 10 recessed and with medio-dorsal emargination, lacking postero-lateral processes (Fig. 17 c; shared with M. acuminata). Male cercus strongly arched, curled over itself with tip directed antero-ventrally (Figs. 17 c, 18 c; unique); male paraproct about as high as half of S 10 height at base in lateral view (Fig. 18 c; shared with M. acuminata); base of paraproct lacking a thumb-like tubercle (Figs. 17 c, 18 c; shared with M. acuminata and M. incus). Ovipositor only slightly surpassing tip of cerci, for a distance shorter than cerci length (Fig. 16 c; shared with M. acuminata and M. incus). Dimensions. Males (n 1): Hw 19.5; abdomen 35.0; total length 38.0. Females (n 1): Hw 19.5; abdomen 30.5; total length 36.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0BFF8C01B2F99B86CA4511.taxon	discussion	Remarks. Wasscher (1998) did not justify placement of M. cyanolineata in Metaleptobasis. He noted it differed from all Metaleptobasis species by the presence of pale blue pterothoracic stripes (Fig. 8 a), and named the species according to that feature. The presence in M. cyanolineata of well developed male prothoracic processes (Figs. 3 c, 4 c, 8 a) and of pale stripes in pterothorax is shared with all known species of Mesoleptobasis, as is its rounded frons (as in Fig. 1 a) and presence of affixed sclerotized processes in genital ligula (Fig. 15 c), none of which is found within Metaleptobasis. All Metaleptobasis species have a pair of well developed horns on the anterior portion of the male mesepisterna and a dark metallic-green mid-dorsal stripe on the pterothorax (Fig. 9 b), absent in M. cyanolineata and in all known Mesoleptobasis. For these reasons we transfer M. cyanolineata from Metaleptobasis to Mesoleptobasis.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0BFF8C01B2F99B86CA4511.taxon	distribution	Distribution. Guyana and Saramacca and Marowijne Districts in Surinam (Fig. 21).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0DFF9101B2FA19832743BB.taxon	etymology	Etymology. From elongatus (Lat.), referring to its long abdomen which distinguishes it from its congeners (Fig. 20 a). Types. Holotype 3 (from Boven Coesewijne, Para, Surinam, 5 ° 22 ' N, 55 ° 32 ' W) in RMNH (examined).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0DFF9101B2FA19832743BB.taxon	description	Specimens examined. Total: 2 3, 1 Ƥ. Surinam, Para District: 1 3 holotype, 1 Ƥ allotype, Boven Coesewijne (5 ° 22 ' N, 55 ° 32 ' W), 0 6 viii 1960, leg. J. Belle (RMNH). Brazil, Amazonas State: 1 3 paratype, Paraná Costa da Ilha de Curari (Rio Solimões), canopy fogging (with pyrethrum), project TRS # 04, tray # 322, white water inundation forest, varzea (3 ° 25 ' S, 60 ° 15 ' W), 0 3 viii 1914, leg. Adis, T. Erwin, Montgomery, et al. (RWG). Description. Male holotype (Fig. 20 a). — Head: Labium ivory, remainder of head including epicranium dull olive green, paler on rear of head, dark brown along base of postfrons, base of antennae, postfrontal suture, and irregular diffuse spot between ocelli and antenna; postocular spots absent. — Thorax: Prothorax dull olive blue, middle lobe light brown; pterothorax entirely olive blue with diffuse brown thoracic stripes along medial portion of mesepisternum and mesopleural suture, remainder of thorax becoming paler laterally; hind lobe of prothorax with a medial bifurcate process (Fig. 4 d), with apices directed laterally (Fig. 3 e) forming a transverse line between them. Costal side of Fw pt longer than basal side, its posterior margin slightly convex. CuA relatively short, ending one cell proximal to vein descending from subnodus. Wings hyaline, Px Fw 10; Px Hw 8 (left) / 9 (right); RP 2 originating at Px 5 in Fw, at Px 4 in Hw; pterostigma 0.4 mm long. Legs ivory with wash of brown at flexor surfaces of femora, spines black — Abdomen: S 1 - 7 brown dorsally, pale olive blue laterally, S 3 - 7 with pale basal ring, S 8 - 10 dull orange brown. Genital ligula in ectal view (Fig. 14 d) with distal margin deeply bifid and lacking lateral emarginations; in lateral view (Fig. 15 d) with basal lobe sclerotized, long, pointed, and directed posteriorly, and dorsal margin of lateral lobe lacking a single small sclerotized spine. Posterior margin of S 10 (Figs. 17 d, 18 d) projected medio-dorsally, with a pair of postero-lateral small lobe-like processes. Male cercus lacking a membranous area dorsally, approximately semicircular, as long as wide; in lateral view (Fig. 18 d) smoothly curved, with tip directed postero-ventrally; paraproct slender and narrower than half of S 10 height at base in lateral view; base of paraproct with a thumblike tubercle (Fig. 18 d). Dimensions. Hw 18.4; abdomen 40; total length 45. Female allotype. Head: similar to male but head entirely dark olive becoming paler along genae and lacking dark suture and epicranial spots. — Thorax: as in male except hind lobe of pronotum (Fig. 5 e) weakly trilobate, transverse, gently concave medially in dorsal view and with middle lobe erect; mesostigmal plates as in Fig. 7 d. Costal side of FW pt longer than basal side, its posterior margin slightly convex. CuA relatively short, ending one cell proximal to vein descending from subnodus in Fw, at level of subnodus in Hw (Fig. 13 b). Wings hyaline, Px Fw 11 (left) / 10 (right); Px Hw 9 (left) / 10 (right); RP 2 originating at Px 5 in Fw, at Px 4 in Hw; pterostigma 0.5 mm long. — Abdomen: as in male but shorter, more robust, and S 8 - 10 dark brown above and pale green laterally; ovipositor surpassing tip of cerci for a distance subequal to length of cerci (Fig. 16 d). Dimensions. — Hw 19.2; abdomen 32; total length 38. Variation in male paratype. As in holotype but overall body coloration paler with more of an orange tinge on olivaceous areas, CuA relatively short, ending two cells proximal to vein descending from subnodus in Fw, one cell proximal in Hw (Fig. 11 b), Px Fw 9; Px Hw 8 (left) / 7 (right); RP 2 originating at Px 5 in Fw, at Px 4 in Hw; pterostigma 0.4 mm long. Dimensions. Hw 18.0; abdomen 37; total length 42.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0DFF9101B2FA19832743BB.taxon	diagnosis	Diagnosis. Morphology of genital ligula, male cerci and paraprocts, and female ovipositor ally this species to M. cantralli and M. incus, but the longer abdomen of male (Fig. 20 a) relative to the other two species (as in Fig. 20 b) readily separate them. It can be further diagnosed from its congeners by: male prothorax with a medial bifurcate process with arms directed laterally forming a transverse line between them (Fig. 3 e; shared with M. cantralli and M. incus); female prothorax lacking processes, with posterior margin slightly trilobate, with medial lobe not surpassing lateral lobes posteriorly; lateral lobes not bent anteriorly (Fig. 5 e; shared with M. cantralli). Costal side of Fw pt longer than basal side, its posterior margin slightly convex in both sexes (Figs. 11 b, 13 b; shared with M. cantralli, M. cyanolineata, and M. incus). CuA relatively short (Figs. 11 b, 13 b; shared with M. cantralli and M. incus), ending one or two cells proximal to vein descending from subnodus in male and one cell proximal to vein descending from subnodus in female. Genital ligula in ectal view with distal margin deeply bifid and lacking lateral emarginations (Fig. 14 d; shared with M. cantralli, M. cyanolineata, and M. incus); in lateral view with basal lobe sclerotized, long, pointed, and directed posteriorly (Fig. 15 d; shared with M. cantralli and M. incus). Posterior margin of S 10 projected medio-dorsally, with a pair of postero-lateral small lobe-like processes (Figs. 17 d, 19 b; shared with M. cantralli and M. incus). Male cercus lacking a membranous area dorsally (shared with M. cantralli, M. cyanolineata, and M. incus), approximately semicircular, as long as wide (Figs. 17 d, 19 b, d; shared with M. incus); in lateral view smoothly curved, with tip directed postero-ventrally (Fig. 18 d); paraproct slender and narrower than half of S 10 height at base in lateral view (Fig. 18 d; shared with M. cantralli and M. incus); base of paraproct with a thumb-like tubercle (Fig. 18 d; shared with M. cantralli). Ovipositor surpassing tip of cerci for a distance as long as length of cerci (Fig. 16 d; shared with M. cantralli).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0DFF9101B2FA19832743BB.taxon	discussion	Remarks. Despite careful examination we did not observe any spines such as those present in M. cantralli (Fig. 15 b) and M. incus (Fig. 15 e) on the dorsal margin of lateral lobe of genital ligula; this area appears to be slightly darker and thickened in the holotype.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB0DFF9101B2FA19832743BB.taxon	distribution	Distribution. Para State in Surinam and Amazonas State in Brazil (Fig. 21). The paratype male was canopy-fogged.	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB13FF9401B2FDAD804D45C0.taxon	description	Specimens examined. Total: 8 3, 15 Ƥ. Brazil, Amazonas State: 1 3 lectotype, Rio Autaz, W to lower Madeira River, S. Amelia, forest (4 ° 24 ' 51 " S, 59 ° 56 ' 21 " W), viii 1914, leg. A. Roman (NHRS); 2 3, Tefé (3 ° 22 ' S, 64 ° 42 ' W, 43 m), i 1962, leg. A. Carvalho (ABMM); 1 3, same but Amana Lake, i 1991, leg. J. Ribeiro (ABMM); 1 3 paralectotype, 2 Ƥ paralectotypes, same but ix 1914 (NHRS); Rondônia State: 1 Ƥ, Porto Velho (8 ° 46 ' S, 63 ° 53 ' W), 0 3 ii 1922, leg. J. H. Williamson & J. W. Strohm (UMMZ); 2 Ƥ, same but 0 6 ii 1922 (UMMZ); 2 Ƥ, same but (FSCA); 2 Ƥ, same but (IORI); 1 3, 1 Ƥ, same but 13 ii 1922 (UMMZ); 1 3, 1 Ƥ, same but 21 ii 1922 (UMMZ); 1 3, 1 Ƥ, same but 22 ii 1922 (FSCA); 2 Ƥ, same but 27 ii 1922 (UMMZ); 1 Ƥ, same but 14 v 1922 (RWG).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB13FF9401B2FDAD804D45C0.taxon	diagnosis	Diagnosis. Male prothorax with medial bifurcate process with arms directed laterally forming a transverse line between them (Fig. 3 f; shared with M. cantralli and M. elongata); female prothorax lacking processes, with posterior margin slightly trilobate, with medial lobe not surpassing lateral lobes posteriorly; lateral lobes bent anteriorly (Figs. 5 f, g; unique). Costal side of FW pt longer than basal side, its posterior margin slightly convex in both sexes (Figs. 11 c, 13 c; shared with M. cantralli, M. cyanolineata, and M. elongata). FW CuA relatively short (Figs. 11 c, 13 c; shared with M. cantralli and M. elongata), ending at vein descending from subnodus to two cells distal in male and four cells distal in female. Genital ligula in ectal view with distal margin deeply bifid and lacking lateral emarginations (Fig. 14 d; shared with M. cantralli, M. cyanolineata, and M. elongata); in lateral view with basal lobe sclerotized, long, pointed, and directed posteriorly, and dorsal margin of lateral lobe with a single small sclerotized spine (Fig. 15 e; shared with M. cantralli). Posterior margin of S 10 projected medio-dorsally, with a pair of postero-lateral small lobe-like processes (Figs. 17 e, 19 c; shared with M. cantralli and M. elongata). Male cercus lacking a membranous area dorsally (shared with M. cantralli, M. cyanolineata, and M. elongata), approximately semicircular, as long as wide (Figs. 17 e, 19 c, e; shared with M. elongata); in lateral view smoothly curved, with tip directed posteroventrally (Fig. 18 e); paraproct slender and narrower than half of S 10 height at base in lateral view (Fig. 18 e; shared with M. cantralli and M. elongata); base of paraproct lacking a thumb-like tubercle (shared with M. acuminata and M. cyanolineata). Ovipositor surpassing tip of cerci for a distance shorter than length of cerci (Fig. 16 e; shared with M. acuminata and M. cyanolineata). Dimensions. Males (n 6; mean in parenthesis): Hw 16.9 – 19.5 (17.8); abdomen 31.0 – 33.0 (32.2); total length 37.0 – 38.0 (37.5). Females (n 10): Hw 19.0 – 20.0 (19.4); abdomen 30.0 – 32.0 (30.3); total length 35.0 – 37.0 (36.4).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB13FF9401B2FDAD804D45C0.taxon	discussion	Remarks. Sjöstedt (1918) described M. incus based on a syntype series of four males and three females. We designate here a male labeled (all printed unless noted otherwise) ' Amazon / Roman, Rio / Autaz, aug., Mesoleptobasis / 3 incus n. sp. [written] / Yngve Sjöstedt det. ' as lectotype, and provide illustrations of its diagnostic structures (Figs 1 a, 3 f, 4 e, 11 c, 14 e, 15 e, 17 e, 18 e, 19 c). The small series collected by J. H. Williamson and J. S. Strohm in 1922 was identified by E. B. Williamson as M. incus although there is no evidence that he ever compared them directly with Sjöstedt's types. The series of males from Rondônia agree with the lectotype (Fig. 19 c) except that the cercus in dorsal view is more strongly excavated laterally (Fig. 19 e).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
03FC87D3EB13FF9401B2FDAD804D45C0.taxon	distribution	Distribution. Amazonas State in Venezuela and Amazonas and Rondônia States in Brazil, sympatric with M. cantralli in the latter (Fig. 21).	en	Garrison, Rosser W., Ellenrieder, Natalia Von (2009): Redefinition of Mesoleptobasis Sjöstedt 1918 with the inclusion of Metaleptobasis cyanolineata (Wasscher 1998) comb. nov. and description of a new species, Mesoleptobasis elongata (Odonata: Coenagrionidae). Zootaxa 2145: 47-68, DOI: 10.5281/zenodo.188653
