taxonID	type	description	language	source
03FC963FBA1BFFBFFF49FF7E6E2CFC3D.taxon	diagnosis	Diagnosis [adapted from Skevington & Marshall (1998)]. Small to medium-sized (3.2 – 6.0 mm). Eyes holoptic in males, dichoptic in females. Pospedicel obtuse, acute or acuminate. Ocellar setae absent. Wing with vein M 2 present or absent. Pterostigma present. Proepisternum with fan of setae. At least the anterior third of scutum and all abdominal segments covered with distinct setae. Abdominal tergites with at least some brownish or grayish pruinosity. Syntergosternite 8 with a membranous area. Surstyli symmetrical to subsymmetrical (rarely asymmetrical), covered with many conspicuous setae (rarely with inconspicuous setae). Apex of phallic guide with tip thin hook-shaped and lobes apically and ventrally (rarely absent). Ejaculatory apodeme fan-shaped, parasol-shaped wide or narrow (rarely bottle-shaped). Phallus trifid, simple or coiled. Species of Pipunculus from Colombia	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1BFFB9FF49FC4A6E65FAD5.taxon	description	Figs 1 – 13, 40	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1BFFB9FF49FC4A6E65FAD5.taxon	materials_examined	Type material. (3 ♂). HOLOTYPE. Male: COLOMBIA, Antioquia, Belmira, Páramo de Sta. [Santa] Inés, El Morro, 06 º 37 ’ 57.5 ’’ N / 75 º 38 ’ 42.9 ’’ W, 3100 – 3300 m [eters], 25. III – 05. IV. 2017, Proyecto moscas de las flores, A. L. Montoya, C. Rodríguez, JP. Carmona leg., (1 ♂, CEUA- 101617) (photographed specimen). PARATYPES. idem Boyacá, Cab. [Cabaña] Chaina, 05 º 25 ’ N / 73 º 27 ’ W, 2550 m [eters], 01 – 14. feb. [II]. 2001, A. Roberto leg., M 1271 (1 ♂, IAvH); idem 2600 m [eters], M 2662 (1 ♂, LEUA). Holotype with left wing mounted on a microslide with Canada balsam. Left antenna and terminalia placed in a microvial with glycerin, both pinned along the specimen.	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1BFFB9FF49FC4A6E65FAD5.taxon	diagnosis	Diagnosis. Male. Antenna brown; postpedicel with acuminate apex. Wing with anal lobe narrowed. Coxae opaque dark brown. All tibiae without distinct apical spines and outstanding setae anteromedially. Abdomen velvety dark blue, gray-brown pruinose. Syntergosternite 8 brown with a large membranous area. Surstyli subsymmetrical, with inner margins sinuous, apices slightly truncated and downwards directed, never directed outward. Apex of phallic guide stout and shorter without depression apically and without lobe ventrally. Ejaculatory apodeme narrowed, needle-shaped. Phallus with ejaculatory ducts coiled, completely separated.	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1BFFB9FF49FC4A6E65FAD5.taxon	description	Description. MALE (holotype). Body length 4 mm. Head (Figs 1 – 2). Eyes contiguous for 17 facets. F, EM, V (mm) = 0.3, 0.4, 0.2. Frontal triangle gray-brown pruinose, with conspicuous dark brown callus. Occiput brown, brown pruinose dorsally and upper half laterally, gray pruinose in lower half laterally and ventrally. Antenna (Fig. 3) brown; scape with one seta dorsally; pedicel with two setae dorsally and three ventrally; postpedicel with acuminate apex. LPP / WPP = 2.6. Thorax (Figs 1 – 2, 4). Postpronotal lobe brown, gray-brown pruinose. Scutum dark brown, gray-brown pruinose, with dorsocentral setae; long and brown setae covering basal third. Notopleuron brown, gray pruinose. Scutellum concolorous with scutum. Mesopleuron concolorous with notopleuron. Mediotergite concolorous with mesopleuron. Wing (Fig. 5). Length 6.1 mm. LW / MWW = 3.8; LTC / LFC = 1.3. Membrane brown infuscate; anal lobe narrowed. Halter dark brown. Legs (Fig. 1). Coxae opaque dark brown, with apices yellowish brown; fore and hind coxae gray pruinose; mid coxa gray-brown pruinose; trochanters opaque dark brown, gray-brown pruinose; femora opaque dark brown, gray-brown pruinose, with apices yellowish brown, ventral ctenidia and a row of long and fine yellow setae posterolaterally; tibiae opaque dark brown, with bases yellowish brown, without distinct apical spines and outstanding setae anteromedially; fore and hind tibiae with a patch of short and fine setae yellow, giving fluffy appearance in distal two thirds posteriorly; hind tibia without an acute process posteroapically; tarsomeres 1 – 5 dark brown; pulvilli brownish yellow. Abdomen (Figs 1 – 2, 6). Ground color velvety dark blue, gray-brown pruinose, with few inconspicuous setae scattered dorsally and laterally, equal in length; tergite 1 gray-brown pruinose dorsally and laterally; tergites 2 – 5 dark blue, brown pruinose in basal third; tergites and sternites 6 and 7 as in Fig. 7. Syntergosternite 8 brown, gray-brown pruinose, clearly longer than tergite 5, with large membranous area (Fig. 8). Terminalia (Figs 7 – 1). Epandrium and surstyli brown (Fig. 8). Surstyli (Figs 8 – 10) subsymmetrical, slightly shorter than epandrium, with short setae in outer margins. Both surstyli thickened basally, thin apically, with inner margins sinuous, apex slightly truncated and downwards directed (Fig. 8); left surstylus slightly thinner than right, left surstylus with outer margin slightly straight, right surstylus with outer margin slightly sinuous (Fig. 8); both surstyli with upper margin sinuous and acute apex; left surstylus with lower margin slightly straight; right surstylus with lower margin slightly sinuous when seen in lateral view (Figs 8 – 10). Gonopods asymmetrical; left gonopod thinner and shorter than right (Fig. 11). Apex of phallic guide stout and shorter, with tip thin slightly hook-shaped (Fig. 12). Ejaculatory apodeme narrowed, needle-shaped (Fig. 13). Phallus trifid, coiled, with ejaculatory ducts completely separated (Fig. 12). FEMALE. Unknown. Geographical distribution. Colombia (Antioquia, Boyacá) (Fig. 40).	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1BFFB9FF49FC4A6E65FAD5.taxon	etymology	Etymology. From the Latin ‘ caeruleum ’ (= blue) and refers to the predominant color in tergites of the males.	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1BFFB9FF49FC4A6E65FAD5.taxon	biology_ecology	Habitat. The specimens were collected in the Páramo Santa Inés, with areas of very humid premontane forest of the Oriental and Central cordillera of the Northwest region of Colombia and Santuario de Fauna y Flora Iguaque reserve, where the vegetation is composed of Andean cloud forests of the cordillera of the Northeast region of Colombia.	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1BFFB9FF49FC4A6E65FAD5.taxon	description	Taxonomic notes. Pipunculus caeruleus sp. nov. runs to P. abnormis Skevington, 1998 in Skevington & Marshall (1998) in couplet 10 of the key presented by Skevington & Marshall (1998). It differs from P. abnormis by postpedicel with acuminate apex (Fig. 3) (versus postpedicel with obtuse apex [see figure 1, presented by Skevington & Marshall (1998)]; trochanters opaque dark brown, gray-brown pruinose (Fig. 1) (versus trochanters entirely yellow); tibiae opaque dark brown, with bases yellowish brown (Fig. 1) (versus tibiae entirely yellow); abdomen with velvety dark blue reflections, gray-brown pruinose; tergite 1 gray-brown pruinose dorsally and laterally (Figs 1 – 2, 6) (versus tergite 1 entirely gray pruinose to bare anterodorsally; tergites 2 – 3 brown pruinose except anterolaterally; tergites 4 – 5 gray pruinose posterolaterally); both surstyli with apices slightly downwards directed (Fig. 8) (versus both surstyli with apices clearly outwards directed [see figure 18 e, presented by Skevington & Marshall (1998)]; apex of phallic guide without lobe apically (Fig. 12) (versus apex of phallic guide with lobe with tip hook-shaped apically [see figure 18 f, presented by Skevington & Marshall (1998)]; ejaculatory apodeme needle-shaped (Fig. 13) (versus ejaculatory apodeme fan-shaped [see figure 18 d, presented by Skevington & Marshall (1998)]; phallus with ejaculatory ducts coiled (Fig. 12) (versus phallus with ejaculatory ducts not coiled [see figure 18 g, presented by Skevington & Marshall (1998)].	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1DFFBBFF49FA0768F4F9F5.taxon	description	Figs 14 – 26, 40	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1DFFBBFF49FA0768F4F9F5.taxon	materials_examined	Type material. (4 ♂). HOLOTYPE. Male: COLOMBIA, Boyacá, SFF [Santuario de Fauna y Flora] Iguaque, Cab. [Cabaña] Chaina, 05 º 25 ’ N / 73 º 27 ’ W, 2600 m [eters], 14. II – 05. III. 2001, P. Reina leg., Malaise, M 1359 (1 ♂, IAvH) (photographed specimen). PARATYPES. idem (1 ♂, IAvH); idem 01 – 16. XII. 2001, P. Roberto leg., M 2682 (1 ♂, INPA); idem Qda. [Quebrada] Los Mudos, 05 º 44 ’ N / 73 º 26 ’ W, 2840 m [eters], 27. II – 17. III. 2003, P. Reina leg., Malaise, M 3563 (1 ♂, LEUA). Holotype with left wing mounted on a microslide with Canada balsam. Left antenna and terminalia placed in a microvial with glycerin, both pinned along the specimen.	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1DFFBBFF49FA0768F4F9F5.taxon	diagnosis	Diagnosis. Male. Antenna brown; postpedicel with acuminate apex. Wing with anal lobe narrowed basally. Coxae opaque dark brown. Fore and mid tibiae with distinct apical spines; hind tibia without an acute process posteroapically and outstanding setae anteromedially. Abdomen dark brown, gray-brown pruinose, with many conspicuous setae scattered dorsally and laterally equal in length. Syntergosternite 8 with membranous area apically. Surstyli subsymmetrical, with inner margins slightly straight, outer margin sinuous, and apices rounded downwards directed. Apex of phallic guide with rounded and translucent lobe ventrally. Ejaculatory apodeme fan-shaped. Phallus with ejaculatory ducts not coiled, completely separated.	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1DFFBBFF49FA0768F4F9F5.taxon	description	Description. MALE (holotype). Body length 4.2. mm. Head (Figs 14 – 15). Eyes contiguous for 18 facets. F, EM, V (mm) = 0.3, 0.4, 0.1. Frontal triangle gray-brown pruinose, with conspicuous dark brown callus. Occiput brown, brown pruinose dorsally and upper half laterally, gray pruinose in lower half laterally and ventrally. Antenna (Fig. 16) brown; scape with one seta dorsally; pedicel with three setae dorsally and three ventrally; postpedicel with acuminate apex. LPP / WPP = 2.5. Thorax (Figs 14 – 15, 17). Postpronotal lobe brown, gray-brown pruinose. Scutum dark brown, gray-brown pruinose, with dorsocentral setae, long and brown setae covering basal third. Notopleuron brown, gray pruinose. Scutellum concolorous with scutum. Mesopleuron concolorous with notopleuron. Mediotergite concolorous with mesopleuron. Wing (Fig. 18). Length 6 mm. LW / MWW = 3.6; LTC / LFC = 0.3. Membrane brown infuscate; anal lobe narrowed basally. Halter dark brown. Legs (Figs 14 – 15). Coxae opaque dark brown, with apices yellowish brown; fore and hind coxae gray pruinose; mid coxa gray-brown pruinose; trochanters yellowish brown; femora shiny dark brown to black, with basal third and apices yellowish brown, ventral ctenidia and a row of long and fine yellow setae posterolaterally; tibiae yellowish brown; fore and mid tibiae with distinct apical spines; fore and hind tibiae with a patch of short and fine setae yellow, giving fluffy appearance in distal two thirds posteriorly; hind tibia without an acute process posteroapically and outstanding setae anteromedially; tarsomeres 1 – 3 yellowish brown; tarsomeres 4 – 5 dark brown; pulvilli yellowish brown. Abdomen (Figs 14 – 15, 19). Ground color dark brown, gray-brown pruinose, with many conspicuous setae scattered dorsally and laterally equal in length; tergite 1 completely covered by gray-brown pruinosity; tergites 2 – 4 shiny dark brown dorsally, with an I-shaped brown pruinose spot centrally; tergites and sternites 6 and 7 as in Fig. 20. Syntergosternite 8 brown, gray-brown pruinose, clearly shorter than tergite 5, with membranous area apically (Fig. 21). Terminalia (Figs 20 – 26). Epandrium and surstyli dark brown (Fig. 21). Surstyli (Figs 21 – 23) subsymmetrical, slightly equal to epandrium length, with short setae in inner and outer margins. Both surstyli thickened basally and medially, thin apically, with inner margins slightly straight and outer margin sinuous, apices rounded downwards directed (Fig. 21); both surstyli with upper and lower margin sinuous and apices rounded downwards directed when seen in lateral view (Figs 22 – 23). Gonopods subsymmetrical, left gonopod longer than right (Fig. 24). Apex of phallic guide stout, with tip hook-shaped and rounded and translucent lobe ventrally (Fig. 25). Ejaculatory apodeme fan-shaped (Fig. 26). Phallus trifid, not coiled, ejaculatory ducts completely separated (Fig. 25). FEMALE. Unknown. Geographical distribution. Colombia (Boyacá) (Fig. 40).	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1DFFBBFF49FA0768F4F9F5.taxon	etymology	Etymology. The specific epithet ‘ chiminiguagua’ is derived from the indigenous language “ Chibcha ”, of the indigenous people that inhabited the type locality, the meaning of ‘ Chiminiguagua’ is: “ supreme god, creator ”, very significant for their culture. The name is to be treated as a noun apposition. Habitat. The specimens were collected in the Santuario de Fauna y Flora Iguaque reserve, where the vegetation is composed of Andean cloud forests of the cordillera of the Northeast region of Colombia. Taxonomic notes. Based on males, Pipunculus chiminiguagua sp. nov. runs to P. ferepauculus Hardy, 1965 in couplet 24 of the key presented by Skevington & Marshall (1998). It differs from P. ferepauculus by having the postpedicel with acuminate apex (versus postpedicel with obtuse apex); frontal triangle gray-brown pruinose (versus frontal triangle black, not pruinose); occiput brown pruinose dorsally and upper half laterally, gray pruinose in lower half laterally and ventrally (versus occiput silver-grey pruinose ventrally and laterally, not pruinose dorsally); fore and mid tibiae with distinct apical spines (versus all tibiae without distinct apical spines); surstyli subsymmetrical (Fig. 21) (versus surstyli asymmetrical [see figure 4, presented by Rafael (1991)]; apex of phallic guide with rounded and translucent lobe ventrally and without lobe apically (Fig. 25) (versus apex of phallic guide with triangular lobe ventrally and long, stout and rounded lobe apically [see figure 5, presented by Rafael (1991)].	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1FFFBAFF49F9676FA3F8F1.taxon	description	Figs 27 – 39, 40	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1FFFBAFF49F9676FA3F8F1.taxon	materials_examined	Type material. (5 ♂). HOLOTYPE. Male: COLOMBIA, Boyacá, SFF [Santuario de Fauna y Flora] Iguaque, Qda. [Quebrada] Los Mudos, Malaise, 05 º 44 ’ N / 73 º 26 ’ W, 2840 m [eters], 14. II – 14. III. 2003, P. Reina leg., M 3637 (1 ♂, IAvH) (photographed specimen). PARATYPES. idem Cab. [Cabaña] Chaina, 05 º 25 ’ N / 73 º 27 ’ W, 2550 m [eters], 01 – 14. feb. [II]. 2001, A. Roberto leg., M 1271 (2 ♂, LEUA); idem (1 ♂, LEUA); idem (1 ♂, INPA). Holotype with left wing mounted on a microslide with Canada balsam. Left antenna and terminalia placed in a microvial with glycerin, both pinned along the specimen.	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1FFFBAFF49F9676FA3F8F1.taxon	diagnosis	Diagnosis. Male. Antenna dark brown; postpedicel with acute apex. Wing with anal lobe large. Coxae shiny dark brown. Mid tibia with distinct apical spines; hind tibia with an acute process posteroapically and outstanding setae anteromedially. Abdomen velvety dark brown, brown pruinose dorsally, giving the appearance of longitudinal bands dorsally. Syntergosternite 8 with membranous area large. Surstyli subsymmetrical, with inner and outer margins curved, apices rounded and slightly inward directed. Apex of phallic guide with translucent and distinct lobe and one row of inconspicuous setae laterally. Ejaculatory apodeme narrowed, bottle-shaped. Phallus with ejaculatory ducts coiled, distinctly separated in distal half.	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1FFFBAFF49F9676FA3F8F1.taxon	description	Description. MALE (holotype). Body length 4.5 mm. Head (Figs 27 – 28). Eyes contiguous for 18 facets. F, EM, V (mm) = 0.4, 0.4, 0.1. Frontal triangle brown pruinose, with conspicuous dark brown callus. Occiput brown, brown pruinose dorsally and upper half laterally, gray pruinose in lower half laterally and ventrally. Antenna (Fig. 29) dark brown; scape with one seta dorsally; pedicel with three setae dorsally and two ventrally; postpedicel with acute apex. LPP / WPP = 2.2. Thorax (Figs 27 – 28, 30). Postpronotal lobe brown, brown pruinose. Scutum dark brown, brown pruinose, with dorsocentral setae; long and brown setae covering basal third. Notopleuron concolorous with scutum. Scutellum concolorous with scutum. Mesopleuron brown, gray-brown pruinose. Mediotergite brown, brown pruinose. Wing (Fig. 31). Length 5.8 mm. LW / MWW = 3.5; LTC / LFC = 1.3. Membrane brown infuscate; anal lobe large. Halter stem brown; knob yellowish brown. Legs (Fig. 27). Coxae shiny dark brown, gray-brown pruinose, with apices yellowish brown; trochanters shiny brown, brown pruinose; femora shiny dark brown, with apices yellowish brown, ventral ctenidia and a row of long and fine yellow setae posterolaterally; tibiae shiny dark brown, with bases yellowish brown; fore and hind tibiae with a patch of short and fine setae yellow, giving fluffy appearance in distal two thirds posteriorly; fore tibia without distinct apical spines; mid tibia with distinct apical spines; hind tibia with a process acute posteroapically and outstanding setae anteromedially; tarsomeres 1 – 4 brown, 5 dark brown; pulvilli yellowish brown. Abdomen (Figs 27 – 28, 32). Ground color velvety dark brown, brown pruinose dorsally, giving the appearance of longitudinal bands dorsally, shiny posterolaterally, with inconspicuous setae scattered dorsally and long and fine setae brownish yellow laterally, about 3 times longer than dorsal seta; tergites and sternites 6 and 7 as in Fig. 33. Syntergosternite 8 dark brown, brown pruinose, slightly shorter than tergite 5, with membranous area large (Fig. 34). Terminalia (Figs 33 – 39). Epandrium and surstyli dark brown (Fig. 34). Surstyli (Figs 34 – 36) subsymmetrical, shorter than length of epandrium, with long setae in outer margins. Both surstyli thickened basally and medially, thin apically, with inner and outer margins curved, apex rounded, and slightly inward directed (Fig. 34); left surstylus slightly thinner than right (Fig. 34); both surstyli with apices forward directed, left surstylus with apex rounded, right surstylus with an acute apex when seen in lateral view (Figs 35 – 36). Gonopods asymmetrical; left gonopod thicker than right (Fig. 37). Apex of phallic guide stout, with tip thin hook-shaped, translucent distinct lobe and one row of inconspicuous setae laterally (Fig. 38). Ejaculatory apodeme narrowed, bottle-shaped (Fig. 39). Phallus trifid, with ejaculatory ducts coiled, distinctly separated in distal half (Fig. 38). FEMALE. Unknown. Geographical distribution. Colombia (Boyacá) (Fig. 40).	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
03FC963FBA1FFFBAFF49F9676FA3F8F1.taxon	etymology	Etymology. From the Latin ‘ planus ’ (= flat), referring to the dorsoventral flat areas of the ejaculatory ducts of the phallus in the male terminalia. Habitat. The specimens were collected in the Santuario de Fauna y Flora Iguaque reserve, where the vegetation is composed of Andean cloud forests of the cordillera of the Northeast region of Colombia. Taxonomic notes. Based on males, P. planus sp. nov. is similar in appearance to P. kotaneni Skevington, 1998. It differs from P. kotaneni by having postpedicel with acute apex (versus postpedicel with obtuse apex); frontal triangle brown pruinose (versus frontal triangle silver grey pruinose); occiput brown pruinose dorsally and upper half laterally, gray pruinose in lower half laterally and ventrally (versus occiput silver-grey pruinose ventrally and laterally, not pruinose dorsally); tibiae shiny dark brown, with bases yellowish brown (Fig. 27) (versus tibiae brownish black); mid tibia with distinct apical spines (versus mid tibiae without distinct apical spines); hind tibia without distinct apical spines (versus hind tibia with distinct apical spines); abdomen dark brown, shiny posterolaterally (Figs 27 – 28, 32) (versus abdomen black, sparsely gray pruinose laterally); apex of phallic guide with lobe and one row of inconspicuous setae laterally (Fig. 38) (versus apex of phallic guide without lobe laterally and with tuft of setae apically [see figure 33 f, presented by Skevington & Marshall (1998)].	en	Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A., Rafael, José Albertino (2023): Pipunculus Latreille, 1802 (Diptera: Pipunculidae) of Colombia, with description of three new species and an updated key to males of the Neotropical species. Zootaxa 5389 (4): 459-472, DOI: 10.11646/zootaxa.5389.4.4, URL: https://www.mapress.com/zt/article/download/zootaxa.5389.4.4/52530
