identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FD87C7EC107A13FF6CFD1875B5FAEA.text	03FD87C7EC107A13FF6CFD1875B5FAEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mannerheimia cachemirica Coiffait 1982	<div><p>Mannerheimia cachemirica Coiffait, 1982</p><p>(Figs 1–2, 6–8)</p><p>Mannerheimia cachemirica Coiffait, 1982: 146; Shavrin 2021: 307</p><p>Mannerheimia vicina Coiffait, 1982: 146 syn. n.</p><p>Mannerheimia vicina: Shavrin 2021: 307</p><p>Type material examined. Holotype of Mannerheimia cachemirica Coiffait, 1982 ♂ (Fig. 1; dissected): ‘Kashmir, Aru | lg.H.Franz, Okt. 1977 ’ &lt;printed&gt;, ‘TYPE’ &lt;red, printed&gt;, ‘ Mannerheimia | cachemirica | H. COIFFAIT det. 19[printed]76’ &lt;handwritten in black&gt; (NMW). Paratype ♂ (dissected; aedeagus missing): first labels as that in the holotype, with red printed label: ‘PARATYPE’ (NMW). All types with additional printed label: ‘ Mannerheimia | cachemirica Coiffait, 1982 | Shavrin A. V. det. 2024’.</p><p>Holotype of Mannerheimia vicina Coiffait, 1982 ♂ (Fig. 2; dissected): ‘Kashmir, Chandanwari | b.Pahalgam, 2800-3100 m | lg.H.Franz, Okt.1977 ’ &lt;printed&gt;, ‘TYPE’ &lt;red, printed&gt;, ‘ Mannerheimia | vicina | H. COIFFAIT det. 19[printed]76’ &lt;handwritten in black&gt;, ‘ Mannerheimia | cachemirica Coiffait, 1982 | Shavrin A. V. det. 2024’ &lt;printed&gt; (NMW).</p><p>Redescription. Measurements (n=3): HW: 0.47–0.55; HL: 0.32–0.42; OL: 0.12; AL (holotype): 0.92; PL: 0.37; PW: 0.62–0.72; ESL: 0.75–0.90; EW: 0.84–0.96; MTbL (holotype): 0.54; MTrL (holotype): 0.30 (MTrL 1–4: 0.18; MTrL 5: 0.12); AW: 0.85–1.03; AedL: 0.35–0.39; BL: 2.55 (holotype of M. cachemirica)–2.65 (holotype of M. vicina).</p><p>Habitus as in Figs 1–2. Body reddish-brown, with distinctly darker head and abdomen (pronotum and elytra of holotype of M. cachemirica distinctly paler; paratergites of abdomen paler); antennomeres 6–11 brown; mouthparts, antennomeres 1–5, legs and paratergites of abdomen yellow. Body shiny, without microsculpture except for middle portion of head (holotype of M. cachemirica) with very indistinct longitudinal meshes and abdomen with fine and dense transverse microreticulation (other specimens without meshes on head), distinctly coarser in abdominal tergite III–IV. Head with irregular and sparse, moderately large and deep punctation, denser in middle; middle portion of neck with very sparse and fine punctation; punctation of pronotum denser, larger and deeper than that on head, finer and sparser in middle (holotype of M. cachemirica), with impunctate middle and mediobasal portion; scutellum without (holotype of M. cachemirica) or with several very fine punctures in middle; punctation of elytra about as that on pronotum or denser, larger and deeper (holotype of M. cachemirica), finer and sparser in middle along suture, punctures not forming disitnct longitudinal rows except for paratype with fused five rows of serial punctures; abdominal tergites with very fine and sparse punctation.</p><p>Head 1.3–1.4 times as broad as long, indistinctly elevated in middle, with very short and shallow anteocellar foveae about as long as diameter of ocellus; anterior portion between antennal insertion and anterior margins of eyes slightly concave. Eyes very large, convex. Ocelli large, well visible, situated at about level of posterior margins of eyes; distance between ocelli slightly greater than distance between ocellus and posterior margin of eyes. Apical segment of maxillary palpi distinctly more than twice as long as preceding segment, from widest basal third gradually narrowed toward subacute apex. Antennae reaching basal portion of elytra when reclined; basal antennomere ovoid, slightly less than three times as long as broad, antennomere 2 distinctly shorter than basal antennomere, 3 narrower and slightly longer than 2, 4 small, about as long as broad, 5–6 distinctly longer and broader than 4, 7–8 slightly broader than 6, 9–10 indistinctly longer and broader than 8, apical antennomere about as long as two preceding antennomeres, from middle gradually narrowed toward subacute apex.</p><p>Pronotum 1.6–1.9 times as broad as long, 1.3 times as broad as head, widest in about middle, markedly more narrowed posteriad than anteriad; apical margin rounded, about as long as somewhat straight posterior margin; anterior angles widely rounded, indistincly protruded anteriad; posterior angles obtuse, with slightly sinuate laterobasal margins; each laterobasal portions with distinct wide impressions.</p><p>Elytra slightly broader than long, slightly widened apicad, twice longer than pronotum; apical margins straight or rounded. Surface of elytra in middle with distinct, irregular and transverse elevations between punctures (holotype of M. cachemirica). Hind wings fully developed.</p><p>Metatarsus 1.8 times as long as metatibia.</p><p>Abdomen about as broad as elytra or slightly broader, with two small transverse tomentose wing-folding spots in middle of abdominal tergite V; apical margin of abdominal tergite VII with distinct narrow palisade fringe.</p><p>Male. Apical margins of abdominal tergite VIII and sternite VIII widely and rather deeply concave. Aedeagus with wide basal portion, gradually narrowed apically, with narrow median lobe and small rounded apex; parameres moderately wide, distinctly broadened in apical portions, slightly not reaching apex of median lobe; internal sac narrow and moderately long (Fig. 6, 8). Lateral aspect of aedeagus as in Fig. 7.</p><p>Female unstudied.</p><p>Comparative notes. Based on the general shape of the body and the aedeagus, M. cachemirica is similar to Nepalese M. emodensa, from which can be distinguished by the slightly narrower elytra, finer punctation of the pronotum and the elytra, slightly longer preapical antennomeres, and shorter parameres with markedly narrower apical portions.</p><p>Distribution. Mannerheimia cachemirica is known from two localities in Kashmir, India.</p><p>Bionomics. The holotype of M. vicina was collected at altitudes from 2800 to 3100 m a.s.l. Detailed bionomical data are unknown.</p><p>Remarks. Mannerheimia cachemirica was originally described based on three specimens from “ Inde, Cachemire, Aru…”. Mannerheimia vicina was originally described based on a male and a female from “ Inde, Cachemire, Chandanwari, près Pahalgam…”. I studied the holotype and the paratype of M. cachemirica and the holotype of M. vicina, and during the study I have not found sufficient morphological differences between them. The holotype of M. cachemirica has slightly more transverse pronotum and distinctly elevated surface between punctures in the middle of the elytra, but the aedeagus of it is similar to the type of M. vicina . The body of the paratype of M. cachemirica is almost identical to that of the holotype of M. vicina . Thus, I synonymize M. vicina with M. cachemirica, the species which has page priority in the same article (Coiffait 1982).</p></div>	https://treatment.plazi.org/id/03FD87C7EC107A13FF6CFD1875B5FAEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2024): On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5474 (5): 584-592, DOI: 10.11646/zootaxa.5474.5.8, URL: http://dx.doi.org/10.11646/zootaxa.5474.5.8
03FD87C7EC127A15FF6CFAB8734BFB22.text	03FD87C7EC127A15FF6CFAB8734BFB22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mannerheimia emodensa Coiffait 1977	<div><p>Mannerheimia emodensa Coiffait, 1977</p><p>(Figs 3, 9–10)</p><p>Mannerheimia emodensa Coiffait, 1977: 266; Coiffait 1982: 150, Shavrin 2021: 307</p><p>Type material examined. Holotype ♂ (Fig. 3; dissected prior to the present study: the aedeagus and the apical genital segment are glued on the same card with the specimen): ‘Umg.Alm Darghari | b. Maharigaon, 4000m’ &lt;printed&gt;, ‘Gebiet von Jumla | Westnepal,lg.H.Franz’ &lt;printed&gt;, ‘TYPE’ &lt;red, printed&gt;, ‘ Mannerheimia | emodensa | H. COIFFAIT det. 19[printed]76’ &lt;handwritten in black&gt; (NMW). Paratypes: 3 ♂♂ (one specimen dissected), 2 ♀♀: first two labels as that in the holotype, with additional red printed label: ‘PARATYPE’ (NMW). All types with additional printed label: ‘ Mannerheimia | emodensa Coiffait, 1977 | Shavrin A. V. det. 2024’.</p><p>Additional material examined. NEPAL: 9 ♂♂, 3 ♀♀: same data as the holotype (NMW); 1 ♂: ‘ Dzunda Khola-Tal b. Talphi 3000-3500m’, ‘ Gebiet von Jumla Westnepal,lg. H.Franz’ (NMW); 1 ♀: ‘ Dampelek b. Jumla’, ‘ Gebiet des Rarasees Westnepal, lg. H. Franz’ (NMW) .</p><p>Redescription. Measurements (n=20): HW: 0.47–0.53; HL: 0.31–0.33; OL: 0.14–0.15; AL (holotype): 0.85; PL: 0.33–0.37; PW: 0.61–0.68; ESL: 0.74–0.78; EW: 0.90–1.05; MTbL (holotype): 0.52; MTrL (holotype): 0.30 (MTrL 1–4: 0.17; MTrL 5: 0.13); AW: 0.95–1.03; AedL: 0.36–0.40; BL: 1.98–2.65 (holotype: 2.37).</p><p>Habitus as in Fig. 3. Body yellow-brown to reddish-brown, with distinctly darker head (abdomen of some specimens slightly darker than forebody; lateral and basal portions of pronotum and apical portion of elytra slightly paler in holotype and in some paratypes); antennomeres 5–11 or 6–11 brown; mouthparts, antennomeres 1–4 or 1–5 and paratergites yellow. Forebody without microsculpture; abdomen with fine and dense transverse microreticulation. Head with moderately fine and sparse punctation, distinctly denser in middle; punctation of pronotum denser, larger and deeper than that in middle portion of head, finer and sparser in middle, and sometimes with narrow impunctate portion in middle or in mediobasal third; scutellum with one to four fine punctures in middle or without them; punctation of elytra denser, slightly larger and deeper than that on pronotum, punctures in middle of each elytron forming five to six vague and tangled rows of serial punctures, but some specimens with markedly coarse surface without longitudinal rows between punctures.</p><p>Head 1.5–1.6 times as broad as long, slightly elevated in middle, with very short and sometimes indistinct anteocellar foveae as long as diameter of ocellus. Antennomere 3 narrower and slightly shorter than 2, 4 distinctly shorter than 3 and slightly longer than broad, 5–6 indistinctly broader than 4, 7–8 slightly longer and distinctly broader than 6, 9–10 slightly longer and broader than 8, apical antennomere about as long as two preceding antennomeres or slightly shorter.</p><p>Pronotum 1.8 times as broad as long, 1.2 times as broad as head, widest slightly above middle, evenly narrowed toward obtuse posterior angles, sometimes with indistinctly sinuate laterobasal margins; lateral portions slightly impressed, distinctly broader and deeper in laterobasal portions.</p><p>Elytra 1.2–1.3 times broader than long, slightly or strongly widened apicad.</p><p>Metatarsus 1.7 times as long as metatibia.</p><p>Abdomen about as broad as elytra, with two small oval or transverse tomentose wing-folding patches in middle of abdominal tergite V.</p><p>Male. Apical margin of abdominal tergite VIII straight or slightly sinuate. Apical margin of abdominal sternite VIII widely sinuate. Aedeagus with wide basal portion, gradually narrowed apically, with relatively narrow median lobe and small rounded apex; parameres with wide apical portions, slightly exceeding apex of median lobe; internal sac narrow and moderately long (Fig. 9). Lateral aspect of aedeagus as in Fig. 10.</p><p>Female. Apical margin of abdominal tergite VIII straight or somewhat rounded. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Based on the general shape of the body and the aedeagus, M. emodensa is similar to M. cachemirica, from which can be distinguished by the slightly broader elytra, larger and denser punctation on the pronotum and elytra, slightly shorter preapical antennomeres, and longer parameres with significantly broader apical portions.</p><p>Distribution. Mannerheimia emodensa is known from four localities in western Nepal.</p><p>Bionomics. Specimens were collected at altitudes from 3000 to 4000 m a.s.l. Detailed bionomical data are unknown.</p><p>Remarks. Mannerheimia emodensa based on several specimens was originally described from four localities in “..région de Jumla”, western Nepal. I studied the holotype and five paratypes from NMW, and several specimens without paratype label, some of which are apparently belonging to the type serie (see Additional material section above). Coiffait (1982) included it to the key of Mannerheimia of the Himalayan Region. Shavrin (2021) included this species to the key of Mannerheimia .</p></div>	https://treatment.plazi.org/id/03FD87C7EC127A15FF6CFAB8734BFB22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2024): On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5474 (5): 584-592, DOI: 10.11646/zootaxa.5474.5.8, URL: http://dx.doi.org/10.11646/zootaxa.5474.5.8
03FD87C7EC147A17FF6CFAF47481FD2A.text	03FD87C7EC147A17FF6CFAF47481FD2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mannerheimia obscura Coiffait 1982	<div><p>Mannerheimia obscura Coiffait, 1982</p><p>(Figs 4, 12)</p><p>Mannerheimia obscura Coiffait, 1982: 148; Coiffait 1983: 274, Shavrin 2021: 307, 324, 2022: 4</p><p>Type material examined. Holotype (Fig. 4; dissected prior to the present study: the aedeagus and the apical genital segment are glued on the same card with the specimen): ‘Kashmir,Chandanwari | b. Pahalgam, 2800-3100m | lg.H.Franz, Okt.1977 ’ &lt;printed&gt;, ‘TYPE’ &lt;red, printed&gt;, ‘ Mannerheimia | obscura | H. COIFFAIT det. 19[printed]79’ &lt;handwritten in black&gt; (NMW). Paratypes: 2 ♂♂, 1 ♀: the first label as that in the holotype (NMW); 4 ♂♂ (one specimen dissected), 1 ♀: ‘Kashmir,Zagipal b. | Pahalgam, 3500-3600m | lg. H.Franz, Okt.1977 ’ &lt;printed&gt; (NMW); 1 ♂, 1 ♀: ‘Kashmir, Pahalgam | lg.H.Franz, Okt.1977 ’ &lt;printed&gt; (NMW); 1 ♀: ‘Kashmir, Aru | lg.H.Franz, Okt. 1977 ’ &lt;printed&gt; (NMW). All paratypes with additional red printed label: ‘PARATYPE’. All types with additional printed label: ‘ Mannerheimia | obscura Coiffait, 1982 | Shavrin A. V. det. 2024’.</p><p>Additional material examined. INDIA: KASHMIR: 1 ♂, 2 ♀♀: ‘ Kashmir, Pahalgam lg. H. Franz, Okt. 1977 ’ (NMW) ; HIMACHAL PRADESH: 16 ♂♂, 16 ♀♀: ‘ Rohtangpass S-Hang 2500-3500 m Himachal-Pradesh Indien, lg. H.Franz’ (cSh, NMW) ; 1 ♂, 1 ♀: ‘ Umg. Katrain b. Kulu Himachal-Pradesh Indien, lg. H. Franz’ (NMW) ; NEPAL: 1 ♂: ‘ Godawari b. Kathmandu, Nepal, lg. Franz’ (NMW) .</p><p>Redescription. Measurements (n=90): HW: 0.53–0.60; HL: 0.30–0.39; OL: 0.15–0.17; AL (holotype): 1.07; PL: 0.48–0.54; PW: 0.83–0.90; ESL: 0.95–0.97; EW: 0.98–1.17; MTbL (holotype): 0.57; MTrL (holotype): 0.31 (MTrL 1–4: 0.19; MTrL 5: 0.12); AW: 1.05–1.25; AedL: 0.66–0.72; BL: 2.55–3.40 (holotype: 3.37).</p><p>Body convex, elongate (Fig. 4 and Fig. 6 in Shavrin (2021)). Body yellowish-brown to reddish-brown, with distinctly darker head (lateral and basal portions of pronotum, laterobasal and apical portions of elytra, and paratergites usually paler); antennomeres 4–11 or 5–11 brown; mouthparts, antennomeres 1–3 or 1–4 and legs yellow. Body without microsculpture except for abdomen with very fine transverse or isodiametric dense microreticulation. Head with irregular and sparse, moderately fine punctation; punctation of pronotum realtively dense, slightly larger and deeper than that in middle portion of head, sparser and finer in middle, sometimes with impunctate portions in middle or in mediobasal third; scutellum with several very fine punctures in middle or without them; punctation of elytra about as that on pronotum, but slightly larger and deeper, sometimes finer along suture, punctures in middle of each elytron forming five to six vague and tangled rows of serial punctures, indistinct in some specimens.</p><p>Head 1.7 times as broad as long, with very short and indistinct anteocellar foveae about as long as diameter of ocellus. Antennomere 3 distinctly longer and narrower than 2, 4 about twice shorter and slightly broader than 3, 5–6 slightly longer and distinctly broader than 4, 7–9 slightly longer and broader than 6, 10 slightly shorter than 9, apical antennomere slightly shorter than two preceding antennomeres.</p><p>Pronotum 1.6–1.7 times as broad as long, 1.5 times as broad as head, widest in middle, sometimes with slightly sinuate laterobasal margins.</p><p>Elytra slightly or strongly widened apicad, 1.7–1.9 times as long as pronotum. Surface of elytra in middle with irregular longitudinal elevations in some specimens.</p><p>Male. Apical margins of abdominal tergite VIII and sternite VIII widely concave. Aedeagus with wide basal portion, gradually narrowed apically toward widely rounded apex; parameres wide, significantly exceeding apex of median lobe, widened apically, with row of small distinct thorns along inner margins of apical lobes; internal sac moderately wide and long (Fig. 61 in Shavrin (2021)). Lateral aspect of aedeagus as in Fig. 62 in Shavrin (2021).</p><p>Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Mannerheimia obscura can be easily distinguished from the remaining species of the genus by the completely different external morphology of the aedeagus, with widely rounded apex of the median lobe (somewhat resembles some specimens of widespread M. brevipennis (Motschulsky, 1860)), and the shape of the apical portion of the parameres with the presence of small thorns along each latero-apical margin (unique feature).</p><p>Distribution. Mannerheimia obscura is distributed from Pakistan and Kashmir to Himachal Pradesh (India) and Nepal (Fig. 11).</p><p>Bionomics. Specimens were collected at altitudes from 2100 to 3600 m a.s.l. (Shavrin 2021, the present study). One specimen from Kashmir was sifted from moss (Shavrin 2021).</p><p>Remarks. Mannerheimia obscura based on several specimens was originally described from Kashmir, India. Coiffait (1983) recorded it from “Pahalgam, Shashnag, 3000 m ” (Kashmir). Shavrin (2021) recorded it from Pakistan and Kashmir, provided the photograph of the habitus (Shavrin 2021: Fig. 6) and figures of the aedeagus (Shavrin 2021: Figs. 61–62), and included this species to the key of the genus. Later, the same author recorded additional material from Pakistan (Shavrin 2022). Mannerheimia obscura is the most widely distributed species of the genus in the Himalayan Region (Fig. 11). It is here recorded from Himachal Pradesh (India) and Nepal for the first time.</p></div>	https://treatment.plazi.org/id/03FD87C7EC147A17FF6CFAF47481FD2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2024): On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5474 (5): 584-592, DOI: 10.11646/zootaxa.5474.5.8, URL: http://dx.doi.org/10.11646/zootaxa.5474.5.8
03FD87C7EC167A17FF6CFCFC706DFACC.text	03FD87C7EC167A17FF6CFCFC706DFACC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mannerheimia scutellata Coiffait 1982	<div><p>Mannerheimia scutellata Coiffait, 1982</p><p>(Fig. 5)</p><p>Mannerheimia scutellata Coiffait, 1982: 149; Shavrin 2021: 307</p><p>Type material examined. Holotype ♂ (Fig. 5; dissected prior to the present study: the aedeagus glued on the same card with the specimen): ‘ INDIA: Kashmir: Sonamarg | Himalaya-Hauptkette | Nichinai-Tal, 9. VI.1976 | 3200 – 3100 m | MARTENS &amp; SCHAWALLER leg.’ &lt;photo label&gt;, ‘TYPE’ &lt;red, printed&gt;, ‘Holo – [handwritren in black] | Typus’ &lt;red, printed&gt;, ‘ Mannerheimia | scutellata | H. COIFFAIT det. 19 [printed]39’ &lt;handwritten&gt;, ‘SNGMuseum | Frankfurt/Main | Loan-000297’ &lt;printed&gt;, ‘ Mannerheimia | scutellata Coiff. ’ &lt;handwritten in blue&gt;, ‘SNG-Museum | Frankfurt/Main | Loan-000297’ &lt;printed&gt;, ‘ Mannerheimia | scutellata Coiffait, 1982 | Shavrin A. V. det. 2013’ &lt;printed&gt; (SF).</p><p>Remarks. Mannerheimia scutellata was described based on the holotype from “ Inde, Cachemire, Sonamarg…”. I study the holotype (Fig. 5) from SF, but aedeagus is slightly damaged, thus I decide to not redescribe this species in the present study.</p></div>	https://treatment.plazi.org/id/03FD87C7EC167A17FF6CFCFC706DFACC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.	Shavrin, Alexey V. (2024): On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5474 (5): 584-592, DOI: 10.11646/zootaxa.5474.5.8, URL: http://dx.doi.org/10.11646/zootaxa.5474.5.8
