identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FD136EB94BA93FFED5EEA4FCDDFD90.text	03FD136EB94BA93FFED5EEA4FCDDFD90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cercyon Leach 1817	<div><p>Genus Cercyon Leach, 1817</p> <p>Diagnosis. Body variously oval, rather flat to strongly convex; pronotum and elytra forming a continuous curve in lateral view. Dorsal surface glabrous, with punctation and sometimes with microsculpture or indistinct pubescence; pronotum without longitudinal grooves or pit-like impressions except often with 2 lateroposterior depressions; color ranging from pale yellow to black. Ventral surface covered in dense, short pubescence except raised surfaces of meso- and metaventrites. Antennae 9-segmented, terminal 3 antennomeres forming compact club; bases visible dorsally, not hidden by lateral margin of head. Clypeus not separated from frons by a distinct suture, often truncate, extending anterior to eyes when viewed dorsally; mentum of variable shape, anterior margin subtruncate or emarginate, often impressed anteromedially. Maxillary palpi shorter than antennae, 2 nd palpomere swollen apically, length of apical segment equal to or slightly longer than penultimate; male maxilla bearing adhesion discs, though may be indistinct in smaller species. Elytra with distinct epipleura, at least in basal half. Middle portion of prosternum elevated to form a tectiform longitudinal keel, not differentiated laterally from antennal excavations. Median portion of mesoventrite elevated to form a median keel or plate; contacting metaventrite at a single point or separated from it by a narrow gap. Outer margin of profemora rounded; tarsal formula 5-5-5 with 1 st tarsomere always longer than 2 nd. Male genitalia of trilobed type, as in Fig. 7. Female genitalia as in Fig. 13.</p> <p>Notes on Identification. Along with members of the Staphylinidae, Cercyon is one of the more common beach-dwelling beetle taxa. As it is the only beach-dwelling hydrophilid genus in most regions, excluding Australia and New Zealand, it can be easily differentiated from other beach beetles by its evenly oval habitus, uninterrupted outline between pronotum and elytra, presence of a mesoventral tablet, maxillary adhesion discs, and clubbed antennae.</p> </div>	https://treatment.plazi.org/id/03FD136EB94BA93FFED5EEA4FCDDFD90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Suzumura, Alyssa Lee;Kobayashi, Norio;Ôhara, Masahiro	Suzumura, Alyssa Lee, Kobayashi, Norio, Ôhara, Masahiro (2019): Revision of the Beach-Dwelling Species of Cercyon Leach (Coleoptera: Hydrophilidae) of the West Coast of North America. The Coleopterists Bulletin 73 (3): 473-493, DOI: 10.1649/0010-065X-73.3.473, URL: http://dx.doi.org/10.1649/0010-065x-73.3.473
03FD136EB94BA938FD27EFFCFB3FF94B.text	03FD136EB94BA938FD27EFFCFB3FF94B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cercyon OF	<div><p>KEY TO THE SPECIES OF ADULT MARINE CERCYON OF WESTERN COASTAL NORTH AMERICA</p> <p>1. Dorsal surface of head, pronotum, and elytra completely without microsculpture; male with a patch of dense, felt-like setae covering mentum (Fig. 8C); size large, length over 3.5 mm; male genitalia as in Fig. 7D–F.............................................. C. dux Sharp</p> <p>1 ʹ. At least head with distinct, though sometimes fine microsculpture; mentum with at most sparse setae, without felt-like patch of setae; smaller, usually not exceeding 4.0 mm in length................................................ 2</p> <p>2. Pronotum with dense, distinct microsculpture, at least on lateral portions (Fig. 12D); posterior portion of elytra sparsely pubescent (Fig. 12G); very small, less than 2.8 mm in length; male genitalia as in Fig. 7M–O........................................ C. setulosus Sharp</p> <p>2 ʹ. Pronotum without apparent microsculpture except sometimes in a small area behind anterior angles (Figs. 5D, 10D); elytra lacking pubescence; larger, more than 2.8 mm in length................................................ 3</p> <p>3. Elytral striae deeply impressed, particularly at base where 1 st 2 often seem to connect (Fig. 9A); elytral intervals convex; mesoventral plate diamond-shaped (Fig. 9E, F); male genitalia as in Fig.7G–I................................................. C. fimbriatus Mannerheim</p> <p>3 ʹ. Elytral striae rather superficial, 1 st 2 striae not seemingly connected; elytral intervals flat; mesoventral tablet long and narrow (Figs. 5E, F, 10E, F)........................................... 4</p> <p>4. Pronotum piceous, elytra uniformly piceous to testaceous or fulvous (Fig. 1G), sometimes with darkened crescentic spot in apical 1/3; small area behind apical angles of pronotum with microsculpture (Fig. 10D); apices of parameres widened, as in Fig. 7J......................................... C. luniger Mannerheim</p> <p>4 ʹ. Pronotum and elytra piceous to testaceous, irregularly lightened at margins (Fig. 1A); pronotum completely without apparent microsculpture (Fig. 5D); apices of parameres narrow, as in Fig. 7A......................................... C. whuljensis Suzumura, new species</p></div> 	https://treatment.plazi.org/id/03FD136EB94BA938FD27EFFCFB3FF94B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Suzumura, Alyssa Lee;Kobayashi, Norio;Ôhara, Masahiro	Suzumura, Alyssa Lee, Kobayashi, Norio, Ôhara, Masahiro (2019): Revision of the Beach-Dwelling Species of Cercyon Leach (Coleoptera: Hydrophilidae) of the West Coast of North America. The Coleopterists Bulletin 73 (3): 473-493, DOI: 10.1649/0010-065X-73.3.473, URL: http://dx.doi.org/10.1649/0010-065x-73.3.473
03FD136EB94DA928FF0AEDB7FE5AFD75.text	03FD136EB94DA928FF0AEDB7FE5AFD75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cercyon (Cercyon) whuljensis Suzumura 2019	<div><p>Cercyon (Cercyon) whuljensis Suzumura, new species</p> <p>Zoobank.org/ urn:lsid:zoobank.org:act: 486BCAD2-AF82-46EF-AA92-838B183B1353 (Figs. 1A, B, 2A, 4C, 5, 6, 7A–C)</p> <p>Cercyon luniger: Smetana 1978: 150 (partim).</p> <p>Description. Body length = 2.1–4.2 mm, width = 1.7–2.9 mm; body oval, convex though dorsoventrally compressed (Figs. 1A, B, 5A), body outline continuous between pronotum and elytra. Surface shiny anteriorly, becoming matte, rugose on posterior half of elytra; head piceous; pronotum and elytra irregularly piceous to testaceous with lateral and posterior margins pale, variegated; base of elytra often with pale spots, elytral suture often dark. Ventrally piceous with white pubescence; femora testaceous or fulvous. Head covered with dense punctures separated by a little more than their diameter except a narrow space around epicranial suture; interspaces with dense, granulate microsculpture except along median line and posterolateral to epicranial sutures, clypeus anteriorly truncate with marginal stria, anterior margins rounded (Fig. 5B). Eyes separated by about 9X their width dorsally. Mentum trapezoidal (length:width index = 0.46); anterior margin carinate; surface slightly convex; covered in uniform, dense granulate microsculpture and setae on posterior half and around to anterolateral margins. Maxillary palpi a little longer than half width of head; second palpomere swollen apically, about 2X as long as wide; apical palpomere fusiform, a little longer than penultimate. Male with adhesion disks on maxillae, diameter a little over 1/5 width of mentum (Fig. 5C). Antenna about 0.85X width of head; pedicel about 1/5 length of scape; club compact, about 2X as long as wide. Pronotum widest at base, lateral margins evenly rounded; surface covered with dense punctures separated by about their diameter; very sparse, minute microsculpture only visible at high magnification (Fig. 5A, D). Elytra with sides regularly curved, widest at about the middle; with 10 relatively superficial striae, becoming less impressed posteriorly and laterally. Intervals flat, with dense punctures separated by about their diameter, interspaces with dense granulate microsculpture and sparse microrugae, though absent on anteromedial disk of elytra. Prosternum weakly tectiform, with or without complete carina medially, with grooves for reception of antennal club, pubescent, not reaching lateral margin. Mesoventral tablet extremely narrow, almost linear (length:width index = 7.3–13.0); surface flat, shiny, with fine punctures (Fig. 5E, F). Metaventrite with raised pentagonal plate, convex, shiny, punctate, punctures separated by about 2–3X their diameter, interspaces in anteromedial area sometimes with weak granulate microsculpture, projecting anteriorly between mesocoxae and contacting mesoventral tablet at a single point; lateral portions densely pubescent, lacking punctures or femoral lines (Fig. 5E). Femora punctate, punctures separated by 2–3X their diameter; interspaces with indistinct to dense granulate microsculpture. Tibia with dense granulate microsculpture. Aedeagus as in Fig. 7A–C; median lobe roundly narrowed to apex, apical third with short, dense setae, parameres narrow (genital ratio = 1.01).</p> <p>Type Material. Holotype male: CANADA: British Columbia, Big Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.5525&amp;materialsCitation.latitude=48.935" title="Search Plazi for locations around (long -125.5525/lat 48.935)">Ucluelet</a>, 48.93500°N, 125.55250°W, M. Ôhara, 2014.VII.1, BC14MO-025, pinned, with genitalia preserved in Canada balsam. Paratypes: Canada, British Columbia, Vancouver Isle: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.3&amp;materialsCitation.latitude=48.4981" title="Search Plazi for locations around (long -124.3/lat 48.4981)">Otter Point</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.3&amp;materialsCitation.latitude=48.4981" title="Search Plazi for locations around (long -124.3/lat 48.4981)">Sooke</a>, 48.39667°N, 123.86694°W, M. Ôhara, 2014.VI.26, BC14MO-007 (1 sex undetermined); Sombrio Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.3&amp;materialsCitation.latitude=48.4981" title="Search Plazi for locations around (long -124.3/lat 48.4981)">Juan de Fuca</a>, 48.49810°N, 124.30000°W, M. Ôhara, 2014.VI.27, BC14MO-010 (2 males); Little Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.54056&amp;materialsCitation.latitude=48.93389" title="Search Plazi for locations around (long -125.54056/lat 48.93389)">Ucluelet</a>, 48.93389°N, 125.54056°W, M. Ôhara, 2014.VII.1, BC14MO-023a (1 male); Big Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.5525&amp;materialsCitation.latitude=8.935" title="Search Plazi for locations around (long -125.5525/lat 8.935)">Ucluelet</a>, 8.93500°N, 125.55250°W, M. Ôhara, 2014.VII.1, BC14MO-025 (5 males, 2 females); “Bowser, B.C./VI.8: -55/W. J. Brown// C. luniger Mannerheim / Det. M. Hatch ” OSAC _ 0000186121 – OSAC _186126 (4 males, 1 sex undetermined); USA: Washington. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.78067&amp;materialsCitation.latitude=48.15557" title="Search Plazi for locations around (long -123.78067/lat 48.15557)">Whisky Creek</a> Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.78067&amp;materialsCitation.latitude=48.15557" title="Search Plazi for locations around (long -123.78067/lat 48.15557)">Port Angeles</a>, Clallam Co., 48.15557°N, 123.780673°W, A. Suzumura, 2018.VIII.22, WA18AS-008 (9 males, 9 females); Chito Beach Resort, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.17207&amp;materialsCitation.latitude=48.34528" title="Search Plazi for locations around (long -124.17207/lat 48.34528)">Sekiu</a>, Clallam Co., 48.305656°N, 124.428559°W, A. Suzumura, 2018.VIII.22, WA18AS-009 (6 sex undetermined); unnamed beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.17207&amp;materialsCitation.latitude=48.34528" title="Search Plazi for locations around (long -124.17207/lat 48.34528)">Sekiu</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.17207&amp;materialsCitation.latitude=48.34528" title="Search Plazi for locations around (long -124.17207/lat 48.34528)">Clallam Co.</a>, 48.345279°N, 124.17207°W, A. Suzumura, 2018.VIII.22, WA18AS-010 (4 males, 1 female); Hobuck Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.72963&amp;materialsCitation.latitude=48.85583" title="Search Plazi for locations around (long -122.72963/lat 48.85583)">Neah Bay</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.72963&amp;materialsCitation.latitude=48.85583" title="Search Plazi for locations around (long -122.72963/lat 48.85583)">Makah Reservation</a>, 48.336265°N, 124.663221°W, A. Suzumura, 2018.VIII.22, WA18AS-011 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.72963&amp;materialsCitation.latitude=48.85583" title="Search Plazi for locations around (long -122.72963/lat 48.85583)">Derelict Conveyor</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.72963&amp;materialsCitation.latitude=48.85583" title="Search Plazi for locations around (long -122.72963/lat 48.85583)">Ferndale</a>, Whatcom Co., 48.855832°N, 122.72963°W, A. Suzumura, 2018.VIII.28, WA18AS-015 (1 male, 1 female).</p> <p>Holotype deposited in BBM with six paratypes (BC14MO-007, 1 specimen; BC14MO-010, 1 specimen; BC14MO-023a, 1 specimen; BC14MO- 025, 3 specimens). Six paratypes will be deposited in MCZ (WA18 AS-008, 6 specimens) and five in OSAC (OSAC _0000186121– OSAC _0000186126, 5 specimens).The remaining paratypes will be deposited in SEHU (BC14 MO-010, 1 specimen; BC14MO-25, 3 specimens, WA18AS-008, 12 specimens; WA18AS- 010, 5 specimens; WA18AS-011, 1 specimen; WA18AS-015, 2 specimens).</p> <p>Distribution. British Columbia and Washington State.</p> <p>Etymology. “whulj” from the Lushootseed language means “salt water” and is the original name for the Salish Sea, a complex network of waterways extending from southwestern British Columbia through northwestern Washington State and the type locality for this species.</p> <p>Remarks. Cercyon whuljensis is similar to C. luniger and often found living with it (Smetana 1978) but differs in the vastly different shape of the male parameres, absence of a small area of microsculpture on the apical angles of the pronotum, and darker, more mottled coloration. Due to close external similarities, several specimens of C. whuljensis were previously identified by M. H. Hatch (OSAC_00000186121-00000186126) as C. luniger, but examination of the lectotype (GAC. 28090) and one paralectotype (MCZ- ENT00007930) of C. luniger revealed that the specimens Hatch identified are in fact a previously undescribed species. The genitalia illustrated as belonging to C. luniger in Smetana (1988) corresponds to C. whuljensis, whereas the genitalia of C. spathifer in Smetana (1978) corresponds to the true C. luniger.</p> <p>Cercyon (Cercyon) dux Sharp, 1873 (Figs. 1C, D, 2B, 4A, C, 7D–F, 8)</p> <p>Cercyon dux Sharp 1873: 65 [Japan, Kyushu]. Cercyon (Cyceron) dux: Shatrovskiy 1992: 366. Cercyon (Cercyon) dux: Shatrovskiy 1989: 281</p> <p>[southern Kurils?]; Hansen 1999: 278; Ohara and</p> <p>Jia 2006: 2.</p> <p>Redescription. Body length = 3.5–5.0 mm, width = 2.1–2.8 mm; body oval, outline slightly interrupted between pronotum and elytra, convex (Figs. 1C, D, 8A); shiny with dense punctation, no microsculpture dorsally. Color patterns as follows: 1) head piceous, pronotum and elytra piceous to testaceous, fulvous along lateral area, though often not reaching margin, often paler along elytral suture and striae (Fig. 1C, D); 2) head, pronotum, and scutellum piceous, lateral margins of pronotum often paler, elytra testaceous. Ventrally piceoustestaceous with white pubescence, legs testaceous. Head covered with dense punctures separated by a little less than their diameter except a narrow area around epicranial sutures; clypeus truncate with marginal striae (Fig. 8B). Eyes large, separated by about 6X their width dorsally. Mentum trapezoidal (length:width index = 1.79), anterior margin carinate, deeply impressed anteromedially, male mentum evenly covered with a dense patch of seate (Fig. 8C), female mentum punctate, interspaces filled with granulate microsculpture. Length of maxillary palpi about 0.5X width of head; 2 nd palpomere swollen apically, about 2.5X as long as wide; apical palpomere fusiform, about as long as penultimate. Male with large adhesion disks on maxillae, diameter about half width of mentum (Fig. 8C). Length of antenna a little less than width of head, pedicel about 1/3 length of scape, club compact, a little less than 2X as long as wide. Pronotum widest about 1/3 before base; lateral margins strongly convex, evenly rounded; surface covered with very dense punctures separated by a little less than their diameter. Elytra with sides regularly curved, widest at about middle; with 10 striae, deeply impressed posteriorly, 10 th stria superficial, pigmented but not impressed; intervals convex with dense punctures separated by less than their diameter (Fig. 8D); without pubescence. Pseudepipleura almost horizontal; epipleura densely pubescent. Prosternum tectiform, carinate medially; with welldefined grooves for reception of antennal club; pubescent. Mesoventral tablet narrowly triangular, widest posteriorly and narrowed anteriorly (length: width index = 5.3–8.0), surface shiny, densely punctate, flat though anterior 1/7 depressed (Fig. 8E, F). Metaventrite with raised pentagonal plate, convex, shiny, and densely punctate; projecting anteriorly between mesocoxae and almost contacting posterior margin of mesoventral plate, lateral portions densely pubescent, lacking punctures or femoral lines (Fig. 8E). Metepisterna rectangular, parallel, about 5X as long as wide. Femora densely punctate with punctures separated by about 2–3X their diameter, interspaces on meso- and metafemora with indistinct to dense granulate microsculpture. Tibiae with fine rugose microsculpture. Aedaegus as in Fig. 7D–F, median lobe wide, with opening at about 1/3 from apex, rounded and with dense setae from opening to tip; parameres wide, diagonally weakly tapered to diagonally truncate apices (genital ratio = 0.92).</p> <p>Specimens Examined. CANADA: British Columbia. Vancouver Isle: Cowichan Bay, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.94417&amp;materialsCitation.latitude=49.64917" title="Search Plazi for locations around (long -125.94417/lat 49.64917)">Dancan</a>, 48.740556°N, 123.941667°W, M. Ôhara, 2014.VI.28, BC14MO-012 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.94417&amp;materialsCitation.latitude=49.64917" title="Search Plazi for locations around (long -125.94417/lat 49.64917)">Connox Bay</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.94417&amp;materialsCitation.latitude=49.64917" title="Search Plazi for locations around (long -125.94417/lat 49.64917)">Royston</a>, 49.64917°N, 125.94417°W, M. Ôhara, 2014.VII.2, BC14MO-027 (1 male, 4 females); Arbutus Grove Provincial Park, Nanoose Bay, 49.26194°N, 124.18889°W, M. Ôhara, 2014.VII.3, BC14MO-030 (3 males, genitalia illustrated); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.48334&amp;materialsCitation.latitude=49.35664" title="Search Plazi for locations around (long -124.48334/lat 49.35664)">Qualicum Beach</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.48334&amp;materialsCitation.latitude=49.35664" title="Search Plazi for locations around (long -124.48334/lat 49.35664)">Nanaimo</a>, 49.35664°N, 124.48334°W, M. Ôhara, 2016.VIII.4, BC16MO- 032 (17 males, 6 females); Qualicum Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.11438&amp;materialsCitation.latitude=49.86327" title="Search Plazi for locations around (long -125.11438/lat 49.86327)">Nanaimo</a>, 49.86327°N, 125.11438°W, M. Ôhara, 2016.VIII.5, BC16MO-033 (1 female); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.23288&amp;materialsCitation.latitude=50.00677" title="Search Plazi for locations around (long -125.23288/lat 50.00677)">Campbell River</a>, 50.00677°N, 125.23288°W, M. Ôhara, 2016.VIII.7, BC16MO-040 (7 males, 6 females). USA: Washington. Neptune Beach, Lummi Reservation, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.71103&amp;materialsCitation.latitude=48.79467" title="Search Plazi for locations around (long -122.71103/lat 48.79467)">Ferndale</a>, 48.79467°N, 122.71103°W, B. Oxborrow, 2016.X.25, WA16AS-015 (4 males); Derelict Conveyor, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.72963&amp;materialsCitation.latitude=48.85583" title="Search Plazi for locations around (long -122.72963/lat 48.85583)">Ferndale</a>, Whatcom Co., 48.855832°N, 122.72963°W, A. Suzumura, 2018.VIII.26, WA18AS-015 (3 males, 3 females). JAPAN: Hokkaido. “ Tachimachi-misaki / Hakodate, Hokkaido / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.93158&amp;materialsCitation.latitude=44.624294" title="Search Plazi for locations around (long 142.93158/lat 44.624294)">Japan</a>, 8-IX-1995 / M. Ôhara [under seaweed]” (1 male: SEHU0000027966); “ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.93158&amp;materialsCitation.latitude=44.624294" title="Search Plazi for locations around (long 142.93158/lat 44.624294)">Tachimachi-misaki</a> / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.93158&amp;materialsCitation.latitude=44.624294" title="Search Plazi for locations around (long 142.93158/lat 44.624294)">Hakodate</a>, Hokkaido / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.93158&amp;materialsCitation.latitude=44.624294" title="Search Plazi for locations around (long 142.93158/lat 44.624294)">Japan</a>, 8-IX-1995 / M. Ôhara [under seaweed]” (1 male: SEHU0000027860); “ Hokkaido: JAPAN/ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.93158&amp;materialsCitation.latitude=44.624294" title="Search Plazi for locations around (long 142.93158/lat 44.624294)">Aikage</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.93158&amp;materialsCitation.latitude=44.624294" title="Search Plazi for locations around (long 142.93158/lat 44.624294)">Teuri-to</a>,/ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.93158&amp;materialsCitation.latitude=44.624294" title="Search Plazi for locations around (long 142.93158/lat 44.624294)">Off Haboro</a> / TE-08-MO-007/ 44°25’07”N, 141°19’16”E / 15.VI.2008, M. Ôhara / Under sea weed” (2 males: SEHU0000027838, SEHU0000027839); “ Hokkaido: JAPAN/ Tomisato, Okushiri-to/ Off Setana/OK-08-MO-015/ 42°04’18”N, 139°28’17”E / 12.VII.2008, M. Ôhara / under sea weed” (2 males: SEHU0000027901, SEHU0000027900); “Tokachi: Hokkaido: Japan/ Oshirabetsu, Hiroo/HK-09-MO-007/cobble beach/ 42°07’40”N, 143°19’03”E / 5.VI.2009, M. Ôhara / Under seaweed” (1 male: SEHU0000028026); “Nemuro: Hokkaido: JAPAN/ Shoji riv., Rausu,/ 44°07’09”N, 145°15’12”E / 13-IX-2009, M. Ôhara,/ H. Yamamoto and M. Furuta /under seaweeds (Laminaria)/[HK-09-MO097/CO]” (1 male: SEHU0000027994); Motoineppu, Oumu, Monbetsu, 44.624293°N, 142.931578°E, A. Suzumura leg., 2018.VI.26, HK18AS-027 (3 males). Honshu. “Japan: Shimane /Izumo-shi Takicho Kumura/Tamura beach/ 5.Oct.2008 / T. Hayama leg.” (1 male: SEHU0000028111). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.84278&amp;materialsCitation.latitude=26.492779" title="Search Plazi for locations around (long 127.84278/lat 26.492779)">Ryukyu Islands</a>. “Onna, Onnason/ Okinawa-ken, Japan/ 26°29’34”N, 127°50’34”E / [seashore], 25.vii.2015 / Takuma Yoshida leg.”</p> <p>Distribution. Palearctic: Russian Federation (Far East), Japan, Korea; Nearctic: British Columbia, northern Washington (Fig. 4A, C).</p> <p>Remarks. Cercyon dux is easily distinguished from other littoral Cercyon by its large size, lack of dorsal microsculpture, and presence of an oily, feltlike patch of setae covering the mentum in males. This is the first Nearctic record of this species, and it has so far only been collected from a few localities in the Strait of Georgia region of the Salish Sea (Fig. 4C). Only individuals of the second, darker coloration have been found in North American populations. The absence of records of this distinctively large species in the region prior to 2014 suggests a recent introduction. One possibility is that it may have been directly introduced from Asia via debris generated by the 2011 Tohoku Earthquake Tsunami Disaster, though more physiological and phylogenetic studies are needed to better support or refute this hypothesis.</p> <p>Cercyon (Cercyon) fimbriatus Mannerheim 1852 (Figs. 1E, F, 2C, 4B, 7G–I, 9)</p> <p>Cercyon fimbriatum Mannerheim 1852: 344 [USA, Alaska, Mt. Edgecumbe]; Smetana 1978: 105.</p> <p>Cercyon (Cyceron) fimbriatus: Shatrovskiy 1992: 326</p> <p>Cercyon (Cercyon) fimbriatus: Horn 1890: 293.</p> <p>Redescription. Body length = 2.8–3.5 mm, width = 1.6–2.1 mm; body oval, convex though dorsoventrally compressed (Figs. 1E–F, 9A), body outline interrupted between pronotum and elytra; glabrous with dense punctation. Surface shiny on head and pronotum, elytra matte, color patterns (Fig. 1E, F) as follows: 1) piceous with lateral margins and apex of elytra paler, fulvous; 2) head and disc of pronotum piceous, lateral margins of pronotum paler, fulvous, elytra yellow to light brown, opaque, often with dark crescentic spot on apical third, scutellum and elytral suture darkened. Ventrally piceous with white pubescence; femora fulvous to piceous; maxillary palpi and antennal club testaceous; scape, pedicel, and flagellum testaceous to fulvous. Head covered with punctures separated by about 2X their diameter except a narrow space around epicranial sutures; clypeus truncate, anterior margin emarginate, with marginal stria (Fig. 9B). Eyes separated by about 10X their width dorsally. Mentum trapezoidal, convex, widened to apical 2/5 and narrowed anteriorly, anterior margin truncate, carinate, depressed behind; with sparse punctures separated by about 3–5X their diameter, denser posteriorly; interspaces filled with dense, rugose microsculpture (length:width index = 0.44). Maxillary palpi about 0.6X width of head, 2 nd palpomere swollen apically, about 2X as long as wide, apical palpomere fusiform, slightly longer than penultimate. Male with adhesion disks on maxillae, their diameter a little over 1/3 width of mentum. Antenna about 0.70X width of head; pedicel about 0.27X as long as scape; club compact, about 2X as long as wide. Pronotum widest at base, lateral margins evenly rounded; surface covered with dense punctures separated by a little more than their diameter; without microsculpture. Elytra with sides regularly curved, widest at about half length; with 9 deeply impressed striae, 1 st 2 curved towards each other, often seemingly joined at base (Fig. 9D). Prosternum weakly tectiform, almost flat except for median carina, with well-defined grooves for reception of antennal club; pubescent, not reaching lateral margin. Mesoventral tablet variably shaped, fusiform, widest at about half length, acute anteriorly, rounded posteriorly (length:width index = 3.3–4.7) (Fig. 9E, F); surface flat, shiny, densely punctate. Metaventrite with raised pentagonal plate, convex, projecting anteriorly between mesocoxae and almost contacting posterior margin of mesoventral plate; shiny, densely punctate, punctures separated by about their diameter; lateral portions densely pubescent, lacking punctures or femoral lines. Femora punctate, punctures separated by about 2X their diameter; interspaces with dense granulate microsculpture. Tibiae with dense granulate microsculpture. Aedeagus as in Fig. 7G–I; median lobe with apical 1/5, tapered to a truncate apex, bearing short setae; parameres apically rounded with a membranous protrusion on inner apical margin (genital ratio = 0.71).</p> <p>Specimens Examined. Paratype: “Sitkha // MCZ- ENT00007932 ” (1 sex undetermined). CANADA: British Columbia. Vancouver Isle: Alberthead Regional Park, Metchosin, 48.39583°N, 123.48833°W, M. Ôhara, 2014.VI.26, BC14MO- 005 (1 male); French Beach, Juan de Fuca, 48.39583°N, 123.94528°W, M. Ôhara, 2014.VI.26, BC14MO-006 (1 sex undetermined); Otter Point, Sooke, 48.39667°N, 123.86694°W, M. Ôhara, 2014.VI.26, BC14MO-007 (2 sex undetermined); Wickaninnish Beach, Ucluelet, 49.01611°N, 125.67417°W, M. Ôhara, 2014.VI.29, BC14MO-015 (9 sex undetermined); Half Moon Bay, Ucluelet, 48.94444°N, 125.61278°W, M. Ôhara, 2014.VI.30, BC14MO-017a (3 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.47365&amp;materialsCitation.latitude=48.4753" title="Search Plazi for locations around (long -123.47365/lat 48.4753)">Florencia Bay</a>, Ucluelet, 49.98111°N, 125.61111°W, M. Ôhara, 2014.VI.30, BC14MO- 018 (4 sex undetermined); Long Beach, Ucluelet, 49.07028°N, 125.75583°W, M. Ôhara, 2014.VI.30, BC14MO-019 (1 sex undetermined); Esowista, Long Beach, Ucluelet, 49.06861°N, 125.78417°W, M. Ôhara, 2014.VI.30, BC14MO-020 (3 sex undetermined); Little Beach, Ucluelet, 48.93389°N, 125.54056°W, M. Ôhara, 2014.VII.1, BC14MO- 023a (2 males); Terrace Beach, Ucluelet, 48.92861°N, 125.54722°W, M. Ôhara, 2014.VII.1, BC14MO-024 (3 sex undetermined); Arbutus Grove Provincial Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.47365&amp;materialsCitation.latitude=48.4753" title="Search Plazi for locations around (long -123.47365/lat 48.4753)">Nanoose Bay</a>, 49.26194°N, 124.18889°W, M. Ôhara, 2014.VII.3, BC14MO- 030 (6 sex undetermined); Qualicum Beach, Nanaimo, 49.35664°N, 124.48334°W, M. Ôhara and N. Kobayashi, 2016.VIII.4, BC16MO-032 (1 sex undetermined); Qualicum Beach, Nanaimo, 49.86327°N, 125.11438°W, M. Ôhara and N. Kobayashi, 2016.VIII.5, BC16MO-033 (2 male); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.47365&amp;materialsCitation.latitude=48.4753" title="Search Plazi for locations around (long -123.47365/lat 48.4753)">Fort Rupert</a>, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.47365&amp;materialsCitation.latitude=48.4753" title="Search Plazi for locations around (long -123.47365/lat 48.4753)">Port Hadley</a>, 50.69515°N, 127.41035°W, M. Ôhara and N. Kobayashi, 2016.VIII.6, BC16MO-036 (1 sex undetermined); Sayward, 50.39783°N, 125.96224°W, M. Ôhara and N. Kobayashi, 2016.VIII.7, BC16MO- 039 (1 sex undetermined); Perimeter Park, Metchosin, Colwood, 48.47530°N, 123.47365°W, M. Ôhara, 2016.VIII.8, BC16MO-044 (3 sex undetermined). USA: Alaska. Unalaska, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Amaknak Island</a>, 53.88491°N, 166.55671°W, M. Ôhara, 2015.VIII.12-13, UN15MO-019 (5 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Land End</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Homer</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Kenai Peninsula</a>, 59.600849°N, 151.412078°W, M. Ôhara, 2015.VIII.17, AL15MO-022 (28 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Anchor Point</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Homer</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Kenai</a>, Alaska, 59.77291°N, 151.87000°W, M. Ôhara, 2015.VIII.18, AL15MO-023 (21 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Nash Road</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Seward Peninsula</a>, 60.08330°N, 149.35199°W, M. Ôhara, 2015.VIII.19, AL15MO-029 (55 sex undetermined). Washington. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Roosevelt Ocean Beach Access</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Copalis Crossing</a>, 47.17639°N, 124.19694°W, M. Ôhara, 2013.VI.21, WA13MO- 039 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Glenwood Creek</a>, Snohomish Co., 47.959751°N, 122.248971°W, A. Suzumura, 2016.VI.6, WA16AS-006 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Neptune Beach</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Lummi Reservation</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Ferndale</a>, 48.79467°N, 122.71103°W, B. Oxborrow, 2016.X.25, WA16AS-015 (8 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Howarth Park</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Snohomish Co.</a>, 47.96440°N, 122.24050°W, A. Suzumura, 2017.VI.5, WA17AS-001 (2 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Meadowdale</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Snohomish Co.</a>, 47.85948°N, 122.33526°W, A. Suzumura, 2017.VI.9, WA17AS- 004 (7 males); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">North of Battery Van Horne</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Whidbey Island</a>, 48.16159°N, 122.68286°W, A. Suzumura, 2017.VI.24, WA17AS-005 (2 males); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Port Williams County Park</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Sequim</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Clallam Co.</a>, 48.097421°N, 123.046779°W, A. Suzumura, 2018.VIII.14, WA18AS-004 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Dungeness Landing County Park</a> , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Sequim</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Clallam Co.</a>, 48.151984°N, 123.143518°W, A. Suzumura, 2018.VIII.21, WA18AS-005 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Cline Spit County Park</a> , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Sequim</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Clallam Co.</a>, 48.151984°N, 123.152187°W, A. Suzumura, 2018.VIII.21, WA18AS-006 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Port Angeles</a> waterfront, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Clallam Co.</a>, 48.119615°N, 123.428617°W, A. Suzumura, 2018.VIII.21, WA18AS-007 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Chito Beach Resort</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Sekiu</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Clallam Co.</a>, 48.305656°N, 124.428559°W, A. Suzumura, 2018.VIII.22, WA18AS-009 (1 sex undetermined); unnamed beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Sekiu</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Clallam Co.</a>, 48.345279°N, 124.172070°W, A. Suzumura, 2018.VIII.22, WA18AS-010(1 male); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Hobuck Beach</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Neah Bay</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Makah Reservation</a>, 48.336265°N, 124.663221°W, A. Suzumura, 2018.VIII.22, WA18AS-011 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Clallem Bay Spit</a> , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Clallam Bay</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Clallam Co.</a>, 48.255792°N, 124.26017°W, A. Suzumura, 2018.VIII.23, WA18AS-013 (1 sex undetermined); Washington <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Park Boat Landing</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Anacortes</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Skagit Co.</a>, 48.500156°N, 122.693686°W, A. Suzumura, 2018.VIII.26, WA18AS-016 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Libbey Beach Park</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Coupeville</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Whidbey Island</a>, 48.232156°N, 122.701029°W, A. Suzumura, 2018.VIII.26, WA18AS-017 (1 male). Oregon. OSAC _00000206873; OSAC _00000206879; OSAC _00000206882; OSAC _00000206894.</p> <p>Distribution: Pacific Coast of North America, Alaska through Mexico (Fig. 4B)</p> <p>Remarks: The most common species of beachdwelling Cercyon in the region, C. fimbriatus can be easily distinguished by the deeply impressed, often connecting bases of the first and second striae (Fig. 9D). Body size and coloration are highly variable within this species.</p> <p>Cercyon (Cercyon) luniger Mannerheim, 1853 (Figs. 1G, H, 2D, 4B, 7J–L, 10)</p> <p>Cercyon lunigerum Mannerheim 1853: 168 [type locality: Kodiak, AK].</p> <p>Heteryon luniger: Winters 1944: 94.</p> <p>Cercyon (Cercyon) spathifer: Smetana 1978: 149 [San Pedro, CA].</p> <p>Cercyon (Cyceron) luniger: Smetana 1988: 161; Shatrovskiy 1992: 362.</p> <p>Cercyon (Cyceron) spathifer: Smetana 1978: 150; Shatrovskiy 1992: 362.</p> <p>Redescription. Body length = 3.1–3.9 mm, width = 1.8–2.1 mm. Body oval, convex though dorsoventrally compressed (Figs. 1G, H, 10A); body outline continuous between pronotum and elytra. Surface shiny anteriorly, though becoming matte, rugose on posterior half of elytra. Head, pronotum, and scutellum piceous, lateral margins of pronotum often paler, testaceous; elytra fulvoustestaceous, often with crescentic oblique spot in apical third (Fig. 1G, H). Ventrally piceous with pale pubescence, legs and maxillary palpi testaceous to fulvous. Head covered with dense punctures separated by a little more than their diameter except a narrow space around epicranial suture; interspaces with dense, granulate microsculpture except along median line and posterolaterally to epicranial sutures; clypeus anteriorly truncate with marginal stria, anterior corners rounded (Fig. 10B). Eyes separated by about 9X their width dorsally. Mentum trapezoidal to almost rectangular (length:width index = 0.52), anterior margin carinate, depressed behind; surface with uniform, dense granulate microsculpture and long setae around lateral and posterior margins. Maxillary palpi a little longer than half width of head; 2 nd palpomere swollen apically, a little over 2X as long as wide; apical palpomere fusiform, slightly longer than penultimate. Male with adhesion discs on maxillae, diameter about 1/4 width of mentum (Fig. 10C). Antenna about 0.90X width of head; pedicel about 1/5 length of scape; club compact, about 2X as long as wide. Pronotum widest at base, lateral margins evenly rounded; surface covered with dense punctures separated by a little more than their diameter; without microsculpture except a small area behind anterolateral corners (Fig. 10D). Elytra with sides regularly curved; widest at about half length; with 10 relatively superficial striae, becoming less impressed posteriorly and laterally; 10 th stria almost completely unimpressed, only marked by dark pigmentation. Intervals flat with dense punctures separated by 1–2X their diameter; interspaces with sparse rugose microsculpture and dense granulate microsculpture. Prosternum weakly tectiform, with a weak or incomplete carina medially; with grooves for reception of antennal club; pubescent though not reaching lateral margin. Mesoventral tablet extremely narrow, almost linear (length:width index&gt;10.0), surface flat, shiny, with fine punctures (Fig. 10E, F). Metaventrite with raised pentagonal plate, convex, projecting anteriorly between mesocoxae and almost contacting posterior margin of mesoventral plate; shiny, sparsely punctate, punctures separated 2–5X their diameter; lateral portions densely pubescent, lacking punctures or femoral lines (Fig 10E). Femora with punctures separated by about 2–3X their diameter, interspaces with dense to indistinct granulate microsculpture. Tibiae apically rounded, with dense granulate microsculpture. Aedaegus as in Fig. 7J–L, median lobe roundly narrowed to apex; with short, indistinct setae; parameres extremely wide, apical 2/5 almost triangular, apical margin flat (genital ratio = 0.71).</p> <p>Specimens Examined. Lectotype: “Kadjak// Holmberg// Cercyon / lunigerum/ Mannerheim// Cercyon luniger / Mannh./ Smetana des 1974/ LECTOTYPE // Mus. Zool. H:fors/ Spec. typ. No 2379/ Cercyon lu-/ nigerum/ Mannh.// Mus.Zool. Helsinki/ Loan No./ CA 32// Photographed/ 2019/ Pekka Malinen// http://id.luomus.fi// GAC. 28090/ UNITED STATES Alaska/ Kodiak/ 57.47 N, 152.48 W / Holmberg leg.” (LUOMUS). Paralectotype: “Type 7930 // C. lunigerum / Kadjak. Mann. // MCZ-ENT00007930” (MCZ). CANADA: British Columbia, Vancouver Isle. Alberthead Regional Park, Metchosin, 48.39583°N, 123.48833°W, M. Ôhara, 2014.VI.26, BC14MO- 005 (1 sex undetermined); Otter Point, Sooke, 48.39667°N, 123.86694°W, M. Ôhara, 2014.VI.26, BC14MO-007 (1 male); Whiffin Spit Beach, Sooke, 48.35694°N, 123.86694°W, M. Ôhara, 2014.VI.27, BC14MO-008 (1 male); Sombrio Beach, Juan de Fuca, 48.4981°N, 124.30000°W, M. Ôhara, 2014.VI.27, BC14MO-010 (2 males); Nanaimo, Nanaimo, 49.17583°N, 123.94165°W, M. Ôhara, 2014.VI.28, BC14MO-013 (1 sex undetermined); Little Beach, Ucluelet, 48.93389°N, 125.54056°W, M. Ôhara, 2014.VII.1, BC14MO-023a (1 male); Big Beach, Ucluelet, 48.93500°N, 125.55250°W, M. Ôhara, 2014.VII.1, BC14MO-025 (6 males, 1 female); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.96224&amp;materialsCitation.latitude=50.39783" title="Search Plazi for locations around (long -125.96224/lat 50.39783)">Connox Bay</a>, Royston, 49.64917°N, 125.94417°W, M. Ôhara, 2014.VII.2, BC16MO- 027 (1 sex undetermined); Sayward, 50.39783°N, 125.96224°W, M. Ôhara and N. Kobayashi, 2016.VIII.7, BC16MO-039 (1 sex undetermined). USA: Washington. “Bowser, B.C./VI.8: -55/ W. J. Brown // C.luniger / det. M. Hatch ” OSAC _ 000184564, OSAC _000184565; just North of Battery Van Horne, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.17207&amp;materialsCitation.latitude=48.34528" title="Search Plazi for locations around (long -124.17207/lat 48.34528)">Whidbey Island</a>, 48.16159°N, 122.68286°W, A. Suzumura, 2017.VI.24, WA17AS- 005 (3 males, 1 female); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.17207&amp;materialsCitation.latitude=48.34528" title="Search Plazi for locations around (long -124.17207/lat 48.34528)">Whisky Creek Beach</a> , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.17207&amp;materialsCitation.latitude=48.34528" title="Search Plazi for locations around (long -124.17207/lat 48.34528)">Port Angeles</a>, Clallam Co., 48.15557°N, 123.780673°W, A. Suzumura, 2018.VIII.22, WA18AS-008 (13 sex undetermined); unnamed beach, Sekiu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.17207&amp;materialsCitation.latitude=48.34528" title="Search Plazi for locations around (long -124.17207/lat 48.34528)">Clallam Co.</a>, 48.345279°N, 124.172070°W, A. Suzumura, 2018.VIII.22, WA18AS-010 (4 sex undetermined).</p> <p>Distribution: Pacific Coast of North America, Alaska through Mexico (Fig. 4B).</p> <p>Remarks: Cercyon luniger is similar to C. whuljensis and often found living with it (Smetana 1978) but differs in the shape of the male parameres, the presence of a small area of microsculpture on the apical angles of the pronotum, and paler elytral coloration. Due to the close external similarities, specimens of C. whuljensis in OSAC were previously identified by Hatch (OSAC_0000186121 – OSAC_0000186126) and described within C. luniger in Smetana (1988), but examination of the genitalia revealed that what had previously been considered interspecific variation in coloration and microsculpture, corresponds, in fact, to two different species (Suzumura 2019). The genitalia illustrated as belonging to C. spathifer in Smetana (1978) correspond to this species, whereas the genitalia illustrated as belonging to C. luniger in Smetana (1988) correspond to C. whuljensis.</p> <p>Cercyon (Cercyon) setulosus Sharp, 1884 (Figs. 1I, J, 2E, 4A, 7M–O, 12, 13)</p> <p>Cercyon setulosus Sharp 1884: 48; Kuwayama 1967: 134 [Japan, Kuril Isles.: Kunashir].</p> <p>Cercyon (Cercyon) tolfino Hatch 1965: 41; Smetana 1978: 150; Shatrovskiy 1992: 367 (synonymy) [British Columbia, Washington].</p> <p>Cercyon (Cyrceon) setulosus: Shatrovskiy 1992: 367 [southern Kurils].</p> <p>Cercyon (Cercyon) setulosus: Hansen 1999: 289; Shatrovskiy 1989: 281 [southern Kurils]; Ohara and Jia 2006: 2 [Kurils].</p> <p>Redescription. Body length = 2.0– 2.8 mm, width = 1.0– 1.4 mm; body oblong, convex though dorsoventrally compressed (Figs. 1I, J, 12A); surface matte, predominately covered with microsculpture. Body outline interrupted between pronotum and elytra, piceous, lateral margins of pronotum and elytra, apical portion of elytra, and elytral suture often paler (Fig. 1I, J). Ventrally piceous with pale pubescence, legs and maxillary palpi testaceous. Eyes separated by about 10X their width. Head covered with fine punctures, separated by a little more than their diameter except around epicranial sutures, interspace with dense granulate microsculpture; clypeus anteriorly truncate with marginal striae, anterior corners rounded (Fig. 12B). Mentum trapezoidal, anterior margin emarginate and carinate, sparsely punctate with interspaces filled with microsculpture consisting of short, horizontal wrinkles (length:width index = 2.44). Maxillary palpi a little less than 0.5X width of head, 2 nd palpomere swollen apically, apical palpomere fusiform and a little longer than penultimate. Male with small, indistinct adhesion disks on maxillae, their diameter about 1/4 width of mentum (Fig. 12C). Antenna about 0.70X width of head, pedicel about ¼ length of scape, club compact, about 2X as long as wide. Pronotum widest at base, about 2X as wide as long, lateral margins evenly rounded, surface covered with dense punctures separated by a little more than their diameter, interspaces with granulate microsculpture (Fig. 12D), densest laterally and reduced towards center. Elytra with sides regularly curved, widest at about middle, with 10 deeply impressed striae, sometimes transversely indented parallel to margin; intervals flat, with fine punctures separated by 2–3X their diameter, interspaces with fine microrugae and sparse, indistinct, decumbent pubescence laterally from 6 th stria and in posterior third (Fig. 12G). Prosternum tectiform, carinate medially, with well-defined grooves for reception of antennal club, pubescent. Mesoventral plate fusiform though variably shaped, widest at about middle, acute and often constricted anteriorly (length:width index = 2.0–3.0) (Fig. 12E, F), surface flat, shiny, sparsely punctate. Metaventrite with raised pentagonal plate, convex, shiny and irregularly punctate, projecting anteriorly between mesocoxae and contacting mesoventral tablet at a single point, lateral portions densely pubescent, lacking punctures or femoral lines (Fig. 12E). Femora with punctures separated by about 3X their diameter, interspaces with dense granulate microsculpture. Tibiae with indistinct granulate microsculpture. Aedaegus as in Fig. 7M–O; median lobe evenly tapered to a point with long, sparse setae on apical fourth; parameres apically narrowed and diagonally truncate at tips (genital ratio = 0.92).</p> <p>Specimens Examined. Paratypes: “Bowser, B.C./VI.8 -55/ W. J. Brown // PARATYPE / Cercyon (s. str)/ tolfino Hatch /1960// OSAC _0000063017- 0000063021” (5 sex undetermined). CANADA: British Columbia, Vancouver Isle.: Alberthead Regional Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.48833&amp;materialsCitation.latitude=48.39583" title="Search Plazi for locations around (long -123.48833/lat 48.39583)">Metchosin</a>, 48.39583°N, 123.48833°W, M. Ôhara, 2014.VI.26, BC14MO- 005 (1 sex undetermined); French Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.94528&amp;materialsCitation.latitude=48.39583" title="Search Plazi for locations around (long -123.94528/lat 48.39583)">Juan de Fuca</a>, 48.39583°N, 123.94528°W, M. Ôhara, 2014.VI.26, BC14MO-006 (3 sex undetermined); Otter Point, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.86694&amp;materialsCitation.latitude=48.39667" title="Search Plazi for locations around (long -123.86694/lat 48.39667)">Sooke</a>, 48.39667°N, 123.86694°W, M. Ôhara, 2014.VI.26, BC14MO-007 (2 males, 2 females); Whiffin Spit Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.86694&amp;materialsCitation.latitude=48.35694" title="Search Plazi for locations around (long -123.86694/lat 48.35694)">Sooke</a>, 48.35694°N, 123.86694°W, M. Ôhara, 2014.VI.27, BC14MO- 008 (3 males); Sombrio Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.3&amp;materialsCitation.latitude=48.4981" title="Search Plazi for locations around (long -124.3/lat 48.4981)">Juan de Fuca</a>, 48.4981°N, 124.30000°W, M. Ôhara, 2014.VI.27, BC14MO-010 (9 sex undetermined); Brown’ s Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.55778&amp;materialsCitation.latitude=49.9375" title="Search Plazi for locations around (long -125.55778/lat 49.9375)">Ucluelet</a>, 49.93750°N, 125.55778°W, M. Ôhara, 2014.VI.29, BC14MO-016a (2 sex undetermined); Florencia Bay, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.61111&amp;materialsCitation.latitude=49.98111" title="Search Plazi for locations around (long -125.61111/lat 49.98111)">Ucluelet</a>, 49.98111°N, 125.61111°W, M. Ôhara, 2014.VI.30, BC14MO- 018 (1 sex undetermined); Chesterman Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.89694&amp;materialsCitation.latitude=49.1225" title="Search Plazi for locations around (long -125.89694/lat 49.1225)">Tolfino</a>, 49.12250°N, 125.89694°W, M. Ôhara, 2014.VI.30, BC14MO-022 (1 sex undetermined); Little Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.54056&amp;materialsCitation.latitude=48.93389" title="Search Plazi for locations around (long -125.54056/lat 48.93389)">Ucluelet</a>, 48.93389°N, 125.54056°W, M. Ôhara, 2014.VII.1, BC14MO-023a (1 male, 13 sex undetermined); Terrace Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.54722&amp;materialsCitation.latitude=48.92861" title="Search Plazi for locations around (long -125.54722/lat 48.92861)">Ucluelet</a>, 48.92861°N, 125.54722°W, M. Ôhara, 2014.VII.1, BC14MO-024 (20 sex undetermined); Big Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.5525&amp;materialsCitation.latitude=48.935" title="Search Plazi for locations around (long -125.5525/lat 48.935)">Ucluelet</a>, 48.93500°N, 125.55250°W, M. Ôhara, 2014.VII.1, BC14MO-025 (2 sex undetermined); Connox Bay, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.94417&amp;materialsCitation.latitude=49.64917" title="Search Plazi for locations around (long -125.94417/lat 49.64917)">Royston</a>, 49.64917°N, 125.94417°W, M. Ôhara, 2014.VII.2, BC14MO-027 (7 sex undetermined); Arbutus Grove Provincial Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.18889&amp;materialsCitation.latitude=49.26194" title="Search Plazi for locations around (long -124.18889/lat 49.26194)">Nanoose Bay</a>, 49.26194°N, 124.18889°W, M. Ôhara, 2014.VII.3, BC14MO-030 (4 males); Transfer Beach Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.18889&amp;materialsCitation.latitude=49.26194" title="Search Plazi for locations around (long -124.18889/lat 49.26194)">Ladysmith</a>, 49.26194°N, 124.18889°W, M. Ôhara, 2014.VII.3, BC14MO- 031 (32 sex undetermined); Qualicum Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.48334&amp;materialsCitation.latitude=49.35664" title="Search Plazi for locations around (long -124.48334/lat 49.35664)">Nanaimo</a>, 49.35664°N, 124.48334°W, M. Ôhara and N. Kobayashi, 2016.VIII.4, BC16MO-032 (8 males, 2 female, 69 sex undetermined); Fort Rupert, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-127.41035&amp;materialsCitation.latitude=50.69515" title="Search Plazi for locations around (long -127.41035/lat 50.69515)">Port Hadley</a>, 50.69515°N, 127.41035°W, M. Ôhara and N. Kobayashi, 2016.VIII.6, BC16MO- 036 (7 males); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.96224&amp;materialsCitation.latitude=50.39783" title="Search Plazi for locations around (long -125.96224/lat 50.39783)">Sayward</a>, 50.39783°N, 125.96224°W, M. Ôhara and N. Kobayashi, 2016.VIII.7, BC16MO-039 (1 sex undetermined); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.23288&amp;materialsCitation.latitude=50.00677" title="Search Plazi for locations around (long -125.23288/lat 50.00677)">Campbell River</a>, 50.00677°N, 125.23288°W, M. Ôhara, 2016.VIII.7, BC16MO-040 (7 males); Otter Point, Muir Creek Beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.86755&amp;materialsCitation.latitude=48.3797" title="Search Plazi for locations around (long -123.86755/lat 48.3797)">Sooke</a>, 48.37970°N, 123.86755°W, M. Ôhara and N. Kobayashi, 2016.VIII.8, BC16MO-042 (5 males); Whiffin Spit, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.86755&amp;materialsCitation.latitude=48.3797" title="Search Plazi for locations around (long -123.86755/lat 48.3797)">Sooke</a>, 48.37970°N, 123.86755°W, M. Ôhara and N. Kobayashi, 2016.VIII.8, BC16MO-043 (2 sex undetermined); “ Bowser, BC/VI.8 -55/ W.J. Brown ”. USA: Alaska. Whitter, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-148.71776&amp;materialsCitation.latitude=60.78064" title="Search Plazi for locations around (long -148.71776/lat 60.78064)">Kenai Peninsula</a>, 60.78064°N, 148.71776°W, M. Ôhara, 2015.VIII.17, AL15MO-021 (7 sex undetermined); Land End, Homer, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-151.41208&amp;materialsCitation.latitude=59.60085" title="Search Plazi for locations around (long -151.41208/lat 59.60085)">Kenai Peninsula</a>, 59.600849°N, 151.412078°W, M. Ôhara, 2015.VIII.17, AL15MO-022 (19 sex undetermined); Lowel Point, Seward, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-149.43613&amp;materialsCitation.latitude=60.07127" title="Search Plazi for locations around (long -149.43613/lat 60.07127)">Kenai Peninsula</a>, 60.07127°N, 149.43613°W, M. Ôhara, 2015.VIII.19, AL15MO- 028 (4 sex undetermined). Washington. Snohomish Co: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.24897&amp;materialsCitation.latitude=47.95975" title="Search Plazi for locations around (long -122.24897/lat 47.95975)">Glenwood Creek</a>, Snohomish Co., 47.959751°N, 122.248971°W, A. Suzumura, 2016.VI.6, WA16AS- 006 (3 males, 2 females, 1 sex undetermined); Chito Beach Resort, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.42856&amp;materialsCitation.latitude=48.305656" title="Search Plazi for locations around (long -124.42856/lat 48.305656)">Sekiu</a>, Clallam Co., 48.305656°N, 124.428559°W, A. Suzumura, 2018.VIII.22, WA18AS-009 (1 sex undetermined); unnamed beach, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.17207&amp;materialsCitation.latitude=48.34528" title="Search Plazi for locations around (long -124.17207/lat 48.34528)">Sekiu</a>, Clallam Co., 48.345279°N, 124.172070°W, A. Suzumura, 2018.VIII.22, WA18AS-010 (1 sex undetermined); Derelict Conveyor, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.72963&amp;materialsCitation.latitude=48.85583" title="Search Plazi for locations around (long -122.72963/lat 48.85583)">Ferndale</a>, Whatcom Co., 48.855832°N, 122.729630°W, A. Suzumura, 2018.VIII.26, WA18AS-015 (1 sex undetermined); Libbey Beach Park, Coupeville, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70103&amp;materialsCitation.latitude=48.232155" title="Search Plazi for locations around (long -122.70103/lat 48.232155)">Whidbey Island</a>, 48.232156°N, 122.701029°W, A. Suzumura, 2018.VIII.26, WA18AS-017 (1 sex undetermined). JAPAN: Hokkaido: “Daikoku-jima: Akkeshi: Hokkaido / 42°57’39”N, 144°52’11”E / 12.VII.2005, M. Ôhara”, DA05MO-005 (2 males: SEHU0000029423, SEHU0000029424); “ Hokkaido: Japan/ Yagishiri-to/ Off Haboro/ 44°26’02”N 141°25’41”E / 16.VI.2008, M. Ôhara / Under sea weed” YA08MO-010 (1 male: SEHU0000028193); “Oshima: Hokkaido: Japan/ Osatsube, Minami-kayabe/ 41°53’37”N, 141°00’42”E / 15.VII.2009, M. Ôhara /under seaweeds (Laminaria sp.)” HK09MO-041/SI (1 male: SEHU0000029596); “Soya: Hokkaido: JAPAN/ Higashiura, Wakkanai/ 45°25’09”N 142°02’02”E / 23.VII.2008, M. Ôhara/ under seaweeds (Laminaria) and eel grass (Zostera)” HK09MO-051/SI (1 male: SEHU0000029538; 2 females: SEHU0000029539, SEHU0000029540); Nosappu, Nemuro, 43.3855492°N,</p> <p>145.81350176°E, A. Suzumura, 2018.VI.17, HK 18AS-017 (2 males, 3 females).</p> <p>Distribution. Palearctic: Kuril Islands, Japan, Russia Federation; Nearctic: Alaska through Washington (Fig. 4A).</p> <p>Remarks. Cercyon setulosus is a very small, flat species characterized by the presence of dense microsculpture on most of the dorsal surface. Cercyon tolfino was placed in junior synonymy with C. setulosus by Shatrovskiy (1992), but the revision was either overlooked or not accepted as the two names have continued to be used. We are basing our understanding of this species on specimens examined by Shatrovskiy. This confirmed synonymy gives it a trans-Pacific distribution, ranging along the coastlines of the Northern Pacific Rim from northern Japan to Washington State.</p> </div>	https://treatment.plazi.org/id/03FD136EB94DA928FF0AEDB7FE5AFD75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Suzumura, Alyssa Lee;Kobayashi, Norio;Ôhara, Masahiro	Suzumura, Alyssa Lee, Kobayashi, Norio, Ôhara, Masahiro (2019): Revision of the Beach-Dwelling Species of Cercyon Leach (Coleoptera: Hydrophilidae) of the West Coast of North America. The Coleopterists Bulletin 73 (3): 473-493, DOI: 10.1649/0010-065X-73.3.473, URL: http://dx.doi.org/10.1649/0010-065x-73.3.473
