taxonID	type	description	language	source
03FA8790C84A5742B0B6FCD13A9E17C1.taxon	description	: protibia subapically strongly curved or angled and distinctly excavate on inner side (exception: C. immutatum); metaventrite flattened or shallowly impressed; aedeagus long and slender, 0.38 – 0.66 mm long; ventral process apically with more or less pronounced median extension (Figs 25 – 35); internal sac with weakly sclerotized structures, without distinct spines; parameres gradually tapering apicad and with long apical seta. Species of the C. perispinctum group are regionally or locally endemic in Northwest Turkey, South Turkey, Southeast Bulgaria, and the Caucasus region (Georgia, Azerbaijan).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8495740B0E8FF51380E1061.taxon	description	(Figs 1, 29 – 31, Map 1)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8495740B0E8FF51380E1061.taxon	materials_examined	Material examined: Turkey: Istanbul: 3 , 3 , Belgrad Ormanı, 41 ° 12 ' 52 " N, 28 ° 58 ' 17 " E, 130 m, 7 – 8. IV. 2010, leg. Brachat & Meybohm. Kocaeli: 1 , 2 , Pazarçayırı, 40 ° 37 ' 43 " N, 30 ° 03 ' 36 " E, 720 m, 5. V. 2012, leg. Brachat & Meybohm. Sakarya: 4 , 4 , 17 km N Hendek, 40 ° 52 ' 30 " N, 30 ° 46 ' 16 " E, 850 m, 3. V. 2012, leg. Brachat & Meybohm (cAss); 3 , 3 , 14 km N Hendek, 40 ° 52 ' 10 " N, 30 ° 44 ' 41 " E, 700 m, 3. V. 2012, leg. Brachat & Meybohm; 1 , 2  [1 teneral], 4 km S Dikmen, 40 ° 40 ' 21 " N, 30 ° 54 ' 22 " E, 1275 m, 4. V. 2012, leg. Brachat & Meybohm; 1 , 2 km W Dikmen, 40 ° 41 ' 57 " N, 30 ° 53 ' 19 " E, 700 m, 4. V. 2012, leg. Brachat & Meybohm. Düzce: 1 , 2  [1 teneral], 26 km S Düzce, 40 ° 40 ' 43 " N, 31 ° 08 ' 48 " E, 1200 m, 1. V. 2012, leg. Brachat & Meybohm; 1 , 2 , 23 km S Düzce, 40 ° 41 ' 21 " N, 31 ° 08 ' 18 " E, 1020 m, 1. V. 2012, leg. Brachat & Meybohm; 1 , 2 , 19 km S Düzce, 40 ° 41 ' 54 " N, 31 ° 09 ' 51 " E, 650 m, 1. V. 2012, leg. Brachat & Meybohm; 1 , 3 , Kaplandede Dağı, 40 ° 57 ' 04 " N, 31 ° 04 ' 03 " E, 1100 m, 2. V. 2012, leg. Brachat & Meybohm; 2 , 2 , Kaplandede Dağı, 40 ° 55 ' 24 " N, 31 ° 01 ' 53 " E, 710 m, 2. V. 2012, leg. Brachat & Meybohm. Bolu: 3 , 2 , 8 km NE Abant, 40 ° 38 ' 48 " N, 31 ° 21 ' 35 " E, 1010 m, 27. IV. 2012, leg. Brachat & Meybohm; 1 , S Abant, 40 ° 35 ' 05 " N, 31 ° 16 ' 13 " E, 1180 m, 27. IV. 2012, leg. Brachat & Meybohm; 3 , 2 , Bolu – Yığılca, 40 ° 50 ' 57 " N, 31 ° 38 ' 31 " E, 1060 m, 29. IV. 2012, leg. Brachat & Meybohm; 1 , Bolu – Yığılca, 40 ° 50 ' 55 " N, 31 ° 37 ' 17 " E, 1020 m, 29. IV. 2012, leg. Brachat & Meybohm; 2  [1 teneral], 2 , Bolu – Yığılca, 40 ° 50 ' 51 " N, 31 ° 36 ' 40 " E, 980 m, 29. IV. 2012, leg. Brachat & Meybohm; 3 , 2 , 11 km S Bolu, 40 ° 39 ' 52 " N, 31 ° 37 ' 50 " E, 1080 m, 30. IV. 2012, leg. Brachat & Meybohm; 1 , 2 , 10 km S Bolu, 40 ° 40 ' 03 " N, 31 ° 38 ' 06 " E, 1020 m, 30. IV. 2012, leg. Brachat & Meybohm; 1 , 7 km S Bolu, 40 ° 40 ' 10 " N, 31 ° 37 ' 58 " E, 950 m, oak and beech forest, sifted, 31. III. 2010, leg. Assing; 4 , 40 km W Mudurnu, 40 ° 35 ' 44 " N, 30 ' 57 ' 50 " E, 650 m, 23. IV. 2014, leg. Brachat & Meybohm; 1 , 3 , Mudurnu, Vakıfaktaş, 40 ° 25 ' 43 " N, 31 ° 13 ' 13 " E, 1180 – 1220 m, 23. IV. 2014, leg. Brachat & Meybohm; 6 , 8 , same data, but 24. IV. 2014; 1 , Kartalkaya, 40 ° 37 ' 03 " N, 31 ° 48 ' 32 " E, 1720 m, 25. IV. 2014, leg. Brachat & Meybohm; 2 , 2 , Kartalkaya, 40 ° 41 ' 57 " N, 31 ° 46 ' 07 " E, 1400 m, 25. IV. 2014, leg. Brachat & Meybohm; 1 , Bolu pass, 900 m, 20. VI. 1989, leg. Wolf. Zonguldak: 4 , 4 , Devrek, 41 ° 13 ' 20 " N, 31 ° 53 ' 06 " E, 590 m, 2. V. 2014, leg. Brachat & Meybohm; 8 , 3 , same data, but 3. V. 2014; 3 , Devrek, 41 ° 12 ' 27 " N, 31 ° 53 ' 59 " E, 550 m, 3. V. 2014, leg. Brachat & Meybohm; 1 , 1 , 15 km W Devrek, 41 ° 13 ' 45 " N, 31 ° 51 ' 45 " E, beech forest with rhododendron undergrowth, sifted, 26. III. 2010, leg. Assing; 2 , W Çaycuma, 41 ° 25 ' 53 " N, 31 ° 57 ' 51 " E, 570 m, 1. V. 2014, leg. Brachat & Meybohm. Karabük: 1 , Safranbolu, 41 ° 23 ' 31 " N, 32 ° 47 ' 16 " E, 950 m, 29. IV. 2014, leg. Brachat & Meybohm; 1 , 30 km S Devrek, 41 ° 03 ' 21 " N, 32 ° 03 ' 57 " E, 570 m, 4. V. 2014, leg. Brachat & Meybohm. All examined specimens are deposited in cAss. Material examined and identified by C. Besuchet: Type material: 3 , 2 : “ Asia Minor, Goek-Dagh., V. Bodemeyer ” (HNHM). Additional material: Turkey: Istanbul: 17 , 12 , Belgrad Ormanı, leg. Apfelbeck, Schubert (NHMW). Kocaeli: 1 , 3 , 2 exs., “ Goek- Dagh ” [40 ° 37 ' N, 29 ° 56 ' E], leg. v. Bodemeyer (various collections).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8495740B0E8FF51380E1061.taxon	discussion	Comment: Based on an unspecified number of syntypes from “ Anatolien: Goek-Dagh und Alem-Dagh ”, C. bodemeyeri was originally described as a distinct species (REITTER 1903). It was subsequently attributed to C. perispinctum KOLENATI, 1846 as a subspecies (VÍT & BESUCHET 2004). While the distribution of C. bodemeyeri is confined to Northwest Turkey, C. perispinctum is widespread and not uncommon in Georgia and has also been recorded from Azerbaijan. The previous record from Armenia (VÍT & BESUCHET 2004) is most likely based on Besuchet (unpublished notes), who erroneously attributed specimens labelled “ Armen. Geb. ” to Armenia rather than to Georgia (see ASSING & SCHÜLKE 2019). In any case, the distributions of both taxa are separated by a distance of nearly 800 km, thus casting doubt on the previous subspecific concept. A comparison of the aedeagi of C. bodemeyeri and C. perispinctum eventually revealed that they are similar, but not identical, and distinguished particularly by the shape of the apex of the median lobe in ventral view (C. bodemeyeri: ventral process more strongly constricted subapically and apico-laterally not distinctly angled). For comparison see the aedeagus of C. perispinctum in Figs 25 – 26. A third argument comes from the discovery of a third similar species of the same species group, which was recently collected in Southeast Bulgaria and which is described (as C. liguliferum) in an appendix at the end of this paper. In consequence, the above observations suggest that these taxa represent distinct, albeit closely related species rather than subspecies.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8495740B0E8FF51380E1061.taxon	diagnosis	Diagnosis: Body length 1.2 – 1.4 mm; length of pronotum 0.39 – 0.45 mm; width of pronotum 0.50 – 0.57 mm; length of antennae approximately 0.6 mm. Habitus as in Fig. 1. Colouration: body reddish to dark-brown with the elytra often somewhat darker than head and pronotum. Eyes composed of approximately ten ommatidia. Antenna with pronounced club formed by antennomeres IX – XI; antennomere VIII slightly smaller than antennomere VII and much smaller than antennomere IX; antennomere XII nearly twice as long as broad. Pronotum 1.20 – 1.25 times as broad as long, broadest in anterior half, and strongly convex in cross-section; lateral margins straight or indistinctly sinuate in posterior half. Elytra each anteriorly with a transverse impression with long and tomentose whitish pubescence; humeral carina nearly obsolete. : protibia subapically distinctly curved and somewhat flattened and excavate on inner side; aedeagus (Figs 29 – 31) 0.50 – 0.55 mm long; ventral process apically extending into a long and narrow median process in ventral view; internal sac with a pair of moderately sclerotized oblong structures; paramere bisinuate, far from reaching apex of ventral process, and apically with a long curved seta.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8495740B0E8FF51380E1061.taxon	distribution	Distribution and natural history: The known distribution of C. bodemeyeri is confined to Northwest Turkey from Istanbul province in the west to Karabük in the east (Map 1). The altitudes range from 130 to 1720 m, from the beech forest to the fir forest zone. Some specimens collected at the end of April and in the beginning of May are teneral.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8485740B0B6FCD13DAF1521.taxon	description	(Figs 2 – 3, 32, 46, Map 2)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8485740B0B6FCD13DAF1521.taxon	materials_examined	Type material examined: Holotype : “ N 39 ° 50 ' 08 E 28 ° 25 ' 31 (19), TR Bursa 16.4.2010, Devecikonagi 640 m, leg. Brachat & Meybohm / Holotypus  Cephennium fractum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 2 , 2 : same data as holotype (cAss); 1 , 2 : “ N 40 ° 06 ' 47 E 029 ° 04 ' 10 (21), TR Bursa 17.4.2010, Uludag 1430 m, leg. Brachat & Meybohm ” (cAss); 4 : “ N 40 ° 31 ' 40 E 029 ° 52 ' 10 (27), TR Bursa 20.4.2010, Samanli Daglari 740 m, leg. Brachat & Meybohm ” (cAss); 1 : “ N 40 ° 30 ' 41 E 029 ° 49 ' 50 (26), TR Bursa 19.4.2010, Samanli Daglari 720 m, leg. Brachat & Meybohm ” (cAss); 1 , 1 : “ TR – Balikesir (11), Kaz Daği-Gebiet, ca. 20 km NO Edremit, Kuruçam Tepe, 708 m, 13. IV. 2010 / N 39 ° 41 ' 39,2 E 27 ° 08 ' 31,7, leg. Meybohm & Brachat ” (cAss); 2 , 2 : “ N 40 ° 38 ' 47 E 30 ° 03 ' 25 (2), TR Kocaeli 22.4.2014, Pazarcayiri 880 m, Brachat & Meybohm ” (cAss); 1 , 1 : “ N 40 ° 37 ' 43 E 30 ° 03 ' 36, TR Kocaeli 25.4.2012, Pazarcayiri 720 m, Brachat & Meybohm (5) ” (cAss); 1 : “ N 40 ° 38 ' 47 E 30 ° 03 ' 25, TR Kocaeli 25.4.2012, Pazarcayiri 880 m, Brachat & Meybohm (6) ” (cAss); 1 , 2 : “ N 40 ° 39 ' 04 E 30 ° 03 ' 42, TR Kocaeli 24.4.2012, Kartepe 1030 m, Brachat & Meybohm (1) ” (cAss); 1 , 3 : “ N 40 ° 38 ' 54 E 30 ° 05 ' 32, TR Kocaeli 5.5.2012, Kartepe 1270 m, Brachat & Meybohm (35) ” (cAss); 1 , 2 : “ N 40 ° 39 ' 24 E 30 ° 04 ' 46, TR Kocaeli 24.4.2012, Kartepe 1140 m, Brachat & Meybohm (2) ” (cAss); 1 : “ N 39 ° 42 ' 45 E 27 ° 10 ' 23 (14), TR Canakkale, 14.4.2010, Kurucam Tepe 600 m, leg. Brachat & Meybohm ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8485740B0B6FCD13DAF1521.taxon	etymology	Etymology: The specific epithet is the past participle of the Latin verb frangere (to fold, to break) and alludes to the subapically distinctly curved male protibiae.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8485740B0B6FCD13DAF1521.taxon	description	Description: Body length 1.1 – 1.3 mm; length of pronotum 0.38 – 0.42 mm; width of pronotum 0.45 – 0.51 mm; length of antennae 0.55 – 0.60 mm. Habitus as in Figs 2 – 3. Colouration: body reddish to brown, often with the elytra somewhat darker than the head and pronotum. Eyes composed of 6 – 8 ommatidia. Antenna of similar shape as in C. bodemeyeri. Pronotum approximately 1.2 times as broad as long, broadest in anterior half, and moderately convex in crosssection; lateral margins straight in posterior half. Elytra noticeably broader than pronotum, anteriorly with a subcircular impression with tomentose whitish pubescence; humeral carina fine, approximately as long as the combined length of the basal three antennomeres. : protibia distinctly bent subapically and flattened in subapical portion, apically dilated; metaventrite extensively flattened, but not distinctly impressed; aedeagus (Figs 32, 46) long and slender, 0.60 – 0.66 mm long; ventral process constricted in the middle and apically with a moderately long apical extension in ventral view; internal sac with weakly sclerotized small structures; paramere long and straight, extending to apical fifth of ventral process, with a long and weakly curved apical seta. Comparative notes: This species is distinguished from the similar C. bodemeyeri by slightly shorter antennae, slightly smaller eyes, smaller and more circular anterior tomentose impressions on the elytra, subapically more sharply bent male protibiae, and a longer and more slen- der aedeagus with a differently shaped ventral process with a shorter apical extension, by straight parameres, and by internal structures of different shapes.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8485740B0B6FCD13DAF1521.taxon	distribution	Distribution and natural history: The distribution is confined to Bursa, Balıkesir, Kocaeli, and Çanakkale provinces, Northwest Anatolia (Map 2). The specimens were sifted from litter in beech and hornbeam forests, mostly with rhododendron undergrowth and often in dark stream valleys. The altitudes range from 600 to 1430 m. In one locality in Kocaeli, the species was collected together with C. bodemeyeri.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8485748B334F9F13BB11721.taxon	description	(Figs 4, 33, 47, Map 1)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8485748B334F9F13BB11721.taxon	materials_examined	Type material examined: Holotype : “ N 39 ° 56 ' 15 E 29 ° 15 ' 25 (24), TR Bursa 18.4.2010, Kocayayla 1200 m, leg. Brachat & Meybohm / Holotypus  Cephennium infractum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 1 , 4 : same data as holotype (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8485748B334F9F13BB11721.taxon	etymology	Etymology: The specific epithet is the past participle of the Latin verb infringere (to fold, to break) and alludes to the subapically distinctly curved male protibiae.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8485748B334F9F13BB11721.taxon	description	Description: Body length 1.2 – 1.3 mm. Habitus as in Fig. 4. Colouration: body reddish to brown, sometimes with the elytra somewhat darker than the head and pronotum. Eyes composed of 5 – 10 ommatidia. Other external characters as in C. fractum. : protibia distinctly bent subapically and excavated in subapical portion, apically dilated; metaventrite shallowly and extensively impressed; aedeagus (Figs 33, 47) long and slender, 0.55 – 0.58 mm long; ventral process not constricted, subapically abruptly narrowed, and apically with long median extension; internal sac with weakly sclerotized small structures; paramere long and straight, each with a long and curved apical seta. Comparative notes: This species is distinguished from the similar C. fractum especially by a shorter aedeagus with a ventral process of different shape and with a longer apical median extension. In addition, the male metaventrite is shallowly impressed and the colouration is on average paler.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8485748B334F9F13BB11721.taxon	distribution	Distribution and natural history: The type locality is situated in the south slopes of Uludağ, Bursa (Map 1). The specimens were sifted from litter in a grassy valley surrounded by pine forest at an altitude of 1200 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8405748B334FB513A761B21.taxon	description	: protibia subapically not, or only weakly curved, flattened to weakly excavate in apical third to apical half; metaventrite extensively and rather deeply impressed; aedeagus with ventral process more or less gradually narrowed apicad, apically of more or less triangular shape; internal sac with spine-shaped structures of speciesspecific shapes and arrangement; parameres relatively short, thin, and non-sinuate, apically (seta excluded) extending to apical fourth or third of ventral process, with short and indistinct apical seta. Species of the C. ericinum group are locally endemic in the region from Zonguldak province in the west to Ordu province in the east, Northwest and North Anatolia (Map 2).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8405748B0B6FBF13D021781.taxon	description	(Figs 5, 34 – 35, Map 1)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8405748B0B6FBF13D021781.taxon	materials_examined	Type material examined: Holotype : “ N 37 ° 26 ' 35 E 028 ° 18 ' 07 (4), Türkei Mugla Yatagan, Bozdogan 825 m 19.4.2006, l. Brachat & Meybohm / Holotypus  Cephennium immutatum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 1 : “ N 37 ° 28 ' 39 E 027 ° 28 ' 41 Türkei Mugla (1), Bafa See Südküste, l. J. Martens 22.3.2006 ” (cAss); 1 : “ N 38 ° 24 ' 19 E 027 ° 23 ' 31 (17), Türkei Izmir Nif Dagi, 970 m 24.4.2006, l. Brachat & Meybohm ” (cAss); 1 : “ TR – Adıyaman [22], 50 km NE Adıyaman, 8 km NE Narince, 870 m, 37 ° 54 ' 58 N, 38 ° 48 ' 45 E, 24. III. 2005, P. Wunderle ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8405748B0B6FBF13D021781.taxon	etymology	Etymology: The specific epithet (Latin, adjective: unchanged) alludes to the nearly unmodified male protibiae, one of the characters distinguishing this species from C. bodemeyeri.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8405748B0B6FBF13D021781.taxon	description	Description: Body length 1.20 – 1.25 mm; length of pronotum 0.4 mm; width of pronotum 0.47 – 0.50 mm; length of antennae approximately 0.55 mm. Habitus as in Fig. 5. Colouration: body dark-brown. Eyes composed of approximately ten ommatidia. Antenna with pronounced club formed by antennomeres IX – XI; antennomere VIII slightly smaller than antennomere VII and much smaller than antennomere IX; antennomere XII less than twice as long as broad. Pronotum 1.18 – 1.25 times as broad as long, broadest in anterior half, and moderately convex in cross-section; lateral margins straight in posterior half. Elytra each anteriorly with a subcircular to transversely oval impression with long and tomentose whitish pubescence; humeral carina fine, approximately as long as the combined length of the basal 2 – 3 antennomeres. : protibia nearly unmodified, very weakly curved subapically; aedeagus (Figs 34 – 35) approximately 0.38 mm long; ventral process apically with a short median extension in ventral view; internal sac with a pair of weakly sclerotized small structures and basally with a broad membranous structure; paramere weakly sinuate subapically, far from reaching apex of ventral process, and apically with a moderately long seta. Comparative notes: This species is distinguished from C. bodemeyeri by slightly smaller size, more slender elytra with less transverse basal impressions, nearly unmodified male protibiae, and a much smaller aedeagus of different shape.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8405748B0B6FBF13D021781.taxon	distribution	Distribution and natural history: The distribution extends from Izmir and Muğla in the West and Southwest Anatolia eastwards to Adıyaman (Map 1). The specimens were sifted on a rocky north slope with oak, near rocks, and in a stream valley with Platanus and Crataegus. The altitudes range from near sea-level to 970 m. The female paratype (collected in March) appears to be teneral.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85F5757B0E8FF513D5A1141.taxon	description	(Figs 6 – 7, 36 – 37, Map 2)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85F5757B0E8FF513D5A1141.taxon	materials_examined	Type material examined: Holotype : “ TR [22] – Ordu, 15 km S Ordu, S Kabaduz, 990 m, mixed forest, 40 ° 48 ' 59 N, 37 ° 54 ' 28 E, 30. VII. 2006, M. Schülke / Holotypus  Cephennium ericinum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 2  [one with aedeagus lost], 3 : same data as holotype (cAss, MNB).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85F5757B0E8FF513D5A1141.taxon	etymology	Etymology: The specific epithet is an adjective derived from the Latin noun ericius (hedgehog) and alludes to the spinose internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85F5757B0E8FF513D5A1141.taxon	description	Description: Rather large species; body length 1.6 – 1.8 mm; length of pronotum 0.47 – 0.53 mm; width of pronotum 0.60 – 0.67 mm; length of antennae approximately 0.7 mm. Habitus as in Fig. 6. Colouration: body dark-reddish to dark-brown. Eyes reduced to minute rudiments without ommatidia (Fig. 7). Pronotum approximately 1.25 times as broad as long, broadest in anterior half, and strongly convex in cross-section; lateral margins straight in posterior half. Elytra without tomentose impressions anteriorly; humeral carinae distinct, approximately as long as the combined length of the basal 3 – 4 antennomeres. : protibia subapically curved and somewhat excavate and flattened on inner side; aedeagus (Figs 36 – 37) 0.44 – 0.45 mm long; ventral process gradually narrowed apicad, of acutely triangular shape in ventral view; internal sac with long, weakly sclerotized, and apically spinose structures; parameres short and thin, not sinuate, apically extending approximately to middle of ventral process, and with rather short and thin apical seta. Comparative notes: Cephennium ericinum is readily distinguished from other congeners known from Turkey by larger body size alone. It is additionally characterized by strongly reduced eyes, the absence of tomentose anterior impressions on the elytra, and the morphology of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85F5757B0E8FF513D5A1141.taxon	distribution	Distribution and natural history: The type locality is situated to the south of Ordu, Ordu province, North Anatolia (Map 2). The specimens were sifted from leaf litter in a mixed forest with alder, spruce, bramble, and ivy at an altitude of 990 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85F5756B36EFD11383117E1.taxon	description	(Figs 8 – 9, 38 – 39, Map 2)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85F5756B36EFD11383117E1.taxon	materials_examined	Type material examined: Holotype : “ TR [11] – Ordu, 18 km NE Akkuş, 40 ° 56 ' 03 " [N], 37 ° 06 ' 47 " [E], 920 m, mixed deciduous forest, 15. VII. 2008, V. Assing / Holotypus  Cephennium varispinosum sp. n., det. V. Assing 2020 ” (cAss). Paratype : same data as holotype (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85F5756B36EFD11383117E1.taxon	etymology	Etymology: The specific epithet is an adjective composed of the adjective varius (various) and spinosum (spinose). It alludes to the presence of spines of various sizes and shapes in the internal sac of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85F5756B36EFD11383117E1.taxon	description	Description: Body length 1.4 – 1.5 mm; length of pronotum 0.45 mm; width of pronotum 0.54 mm; length of antennae approximately 0.6 mm. Habitus as in Fig. 9. Colouration: forebody reddish-brown; elytra reddish. Eyes reduced to small rudiments without ommatidia (Fig. 8). Antennomeres VIII – X strongly transverse. Pronotum rather large in relation to elytra and as broad as elytra, 1.2 times as broad as long, broadest in anterior half, and strongly convex in cross-section; lateral margins straight in posterior half. Elytra without tomentose impressions anteriorly; humeral carina distinct, approximately as long as the combined length of the basal 3 – 4 antennomeres. : protibia subapically not curved, but somewhat flattened on inner side; aedeagus (Figs 38 – 39) 0.38 mm long; ventral process gradually narrowed apicad in ventral view; internal sac with approximately eight weakly sclerotized and rather small spines of various sizes and shapes ventrally, and with a pair of larger and strongly curved spines dorsally; parameres very thin and indistinct, not sinuate, extending to apical third of ventral process. Comparative notes: Aside from its aedeagus, C. varispinosum is characterized among the Turkish Cephennium species by a rather large pronotum, strongly reduced eyes, and strongly transverse antennomeres VIII – X. It is additionally separated from the geographically close C. ericinum by much smaller size and weakly modified male protibiae.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85F5756B36EFD11383117E1.taxon	distribution	Distribution and natural history: The type locality is situated to the northeast of Akkuş, Ordu province, North Anatolia (Map 2). The specimens were sifted from leaf litter in a mixed forest with predominant Fagus at an altitude of 620 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85E5756B334FBB13AB31B21.taxon	materials_examined	Type material examined: Holotype : “ N 41 ° 28 ' 08 E 32 ° 35 ' 52 (21), TR Bartin 30.4.2014, Bahcedik 3 km s 510 m, Brachat & Meybohm / Holotypus  Cephennium spinigerum sp. n., det. V. Assing 2020 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85E5756B334FBB13AB31B21.taxon	etymology	Etymology: The specific epithet (Latin, adjective: spinose) alludes to the numerous spines in the internal sac of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85E5756B334FBB13AB31B21.taxon	description	Description: Body length 1.4 mm. Habitus as in Fig. 11. Colouration: body dark-reddish. Antennae with antennomeres VIII – X moderately transverse, other external characters as in C. varispinosum. : protibia subapically not curved, but somewhat flattened on inner side in apical portion; aedeagus (Figs 42 – 43) 0.38 mm long; ventral process apically of triangular shape in ventral view; internal sac with numerous rather stout spines; parameres thin, not sinuate, extending to apical third of ventral process. Comparative notes: This species is distinguished from the similar and geographically close C. victrix only by the shape and internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85E5756B334FBB13AB31B21.taxon	distribution	Distribution and natural history: The type locality is situated in Bartin province, Northwest Anatolia (Map 2). The holotype was sifted from litter near old trees in a beech forest with rhododendron undergrowth at an altitude of 510 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85E5756B0B6FAB13A8A10E1.taxon	description	(Figs 10, 40 – 41, Map 2)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85E5756B0B6FAB13A8A10E1.taxon	materials_examined	Type material examined: Holotype : “ TR [15] – Sinop, Çangal Dağı, Ordu, 30 km S Ayancık, 41 ° 48 ' 44 " N, 34 ° 39 ' 19 " E, 920 m, beech forest, 1. IV. 2009, V. Assing / Holotypus  Cephennium victrix sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 1 , 2 : same data as holotype (cAss); 3 , 6 : “ TR [16] – Sinop, Çangal Dağı, Ordu, 30 km S Ayancık, 41 ° 47 ' 50 " N, 34 ° 37 ' 29 " E, 360 m, sycamore forest, 1. IV. 2009, V. Assing ” (cAss); 1 : “ TR [20] – Sinop, 30 km SW Sinop, 41 ° 58 ' 03 " N, 34 ° 50 ' 12 " E, 65 m, mixed deciduous forest, 2. IV. 2009, V. Assing ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85E5756B0B6FAB13A8A10E1.taxon	etymology	Etymology: The specific epithet (Latin, adjective: winning) alludes to the shape of the median pair of internal structures of the aedeagus, which together form a “ V ” (as in victory).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85E5756B0B6FAB13A8A10E1.taxon	description	Description: Body length 1.3 – 1.4 mm. Habitus as in Fig. 10. Colouration: body reddish to reddish-brown with the elytra often slightly paler than head and pronotum. Antennae with antennomeres VIII – X moderately transverse, other external characters as in C. varispinosum. : protibia subapically not curved, but somewhat flattened on inner side in apical portion; aedeagus (Figs 40 – 41) 0.33 – 0.35 mm long; ventral process subapically abruptly narrowed and apically acute in ventral view; internal sac with a pair of lateral clusters of small weakly sclerotized spines and with a median pair of longer weakly sclerotized spines forming a “ V ”; parameres thin, not sinuate, extending to apical fourth of ventral process. Comparative notes: Cephennium victrix is reliably distinguished from the similar C. varispinosum especially by a smaller aedeagus with a ventral process and internal structures of different shapes and arrangement, respectively, additionally also by less transverse antennomeres VIII – X.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85E5756B0B6FAB13A8A10E1.taxon	distribution	Distribution and natural history: The specimens were collected in three localities in Sinop province, North Turkey, by sifting leaf litter in a beech forest, in a moist sycamore forest with rocks, and in a mixed deciduous forest with hornbeam, oak, and laurel at altitudes of 65 – 920 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5755B0E8FF5138061601.taxon	description	(Figs 12, 44 – 45, Map 2)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5755B0E8FF5138061601.taxon	materials_examined	Type material examined: Holotype : “ N 41 ° 25 ' 53 E 31 ° 57 ' 51 (24), TR Zonguldak 1.5.2014, Caycuma w 570 m, Brachat & Meybohm / Holotypus  Cephennium digitale sp. n., det. V. Assing 2020 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5755B0E8FF5138061601.taxon	etymology	Etymology: The specific epithet (Latin, adjective: of a / the finger) alludes to the finger-shaped spines in the internal sac of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5755B0E8FF5138061601.taxon	description	Description: Body length 1.4 mm. Habitus as in Fig. 12. Colouration: body dark-reddish. External characters as in C. spinigerum. : protibia subapically not curved, weakly excavate in apical portion; aedeagus (Figs 44 – 45) 0.39 mm long; apex of ventral process of narrowly triangular shape in ventral view; internal sac with several horn- or fingershaped structures of various sizes and shapes; parameres thin, not sinuate, extending to apical third of ventral process. Comparative notes: Cephennium digitale differs from the similar and geographically close C. spinigerum only by a more slender aedeagus with fewer internal spines of different shapes.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5755B0E8FF5138061601.taxon	distribution	Distribution and natural history: The type locality is situated in Zonguldak province, Northwest Anatolia (Map 2). The holotype was sifted from litter in a beech forest with rhododendron undergrowth at an altitude of 570 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5755B0E8FAD138F81B01.taxon	description	: protibia without distinct modifications; metaventrite extensively depressed or impressed; aedeagus (e. g., Figs 48 – 50) with bulbous capsule, basally subcircular in cross-section, approximately twice as long as broad; internal structures minute; parameres thin, apically extending slightly beyond apex of median lobe, with short and fine apical seta. The C. jonicum group includes several described and undescribed species distributed in the Balkans. In the study region, the C. jonicum group is represented by only a single species, C. reductum, which has been recorded from both the Taurus and the Pontic Mountains (South and North Anatolia) (Map 3).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5754B36EFF513DA11281.taxon	description	(Figs 13, 48 – 50, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5754B36EFF513DA11281.taxon	materials_examined	Type material: Holotype : “ TR Mersin (48), Güzeloluk – Erdemli, S Aydinlar 1350 m / (48) 7.5.2004, leg. Brachat & Meybohm, 36 ° 44 ' 59 N 34 ° 7 ' 48 E / Holotypus  Cephennium reductum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 1 : “ TR Mersin (46), Kirobasi – Güzeloluk, 14 km W Güz. 1430 m / 36 ° 45 ' 1 N 33 ° 57 ' 51 E (46) 8.5.2004, leg. Brachat & Meybohm ” (cAss); 1 , 1 : “ N 37 ° 52 ' E 031 ° 20 ', Türkei Umg. Beysehir, Gedikli 7 km s. 1000 m, Meybohm 14.5.2000 ” (cAss); 1 , 1 : “ N 36 ° 26 ' E 032 ° 53 ', Türkei nördl. Anamur, Ermenek Umg. 1450 m, Meybohm 20.5.2000 ” (cAss); 1 : “ N 36 ° 21 ' E 032 ° 56 ', Türkei Umg. Anamur, Abanoz 1240 m, Meybohm 19.5.2000 ” (cAss); 1 : “ N 36 ° 44 ' 01 E 33 ° 51 ' 16, Türkei Mersin (8), Kirobasi 1390 m, 1 km w. Kirobasi / Ku [l] turland am Bach, 19.4.2005 (8), leg. Brachat & Meybohm ” (cAss); 1 : “ TR Mersin (41), road Mut – Karaman, Sertavul Gec. 1570 m / 36 ° 55 ' 28 N 33 ° 16 ' 26 E (41), 5.5.2004, leg. Brachat & Meybohm ” (cAss); 1 : “ TR [36 a] – Kastamonu, 40 km NW Kastamonu, 41 ° 42 ' 05 " N, 33 ° 28 ' 17 " E, 1090 m, calcareous slope, 9. IV. 2009, V. Assing ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5754B36EFF513DA11281.taxon	etymology	Etymology: The specific epithet (Latin, adjective) alludes to the conspicuously small internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5754B36EFF513DA11281.taxon	description	Description: In external characters similar to species of the C. corruptum group (see below). Body length 1.0 – 1.1 mm. Habitus slender (Fig. 13). Eyes composed of approximately five defined ommatidia without pigmentation. Punctation of pronotum slightly to distinctly less fine than that of elytra. Elytra each with a small obliquely oval impression with whitish tomentose pubescence anteriorly. Hind wings completely reduced. : protibia unmodified; metaventrite extensively triangularly depressed (not impressed); aedeagus (Figs 48 – 50) 0.27 – 0.29 mm long, approximately twice as long as broad in ventral view; capsule bulbous, basally subcircular in cross-section; ventral process apically moderately broadly truncate and in the middle with small projection in ventral view; internal sac with two very small apico-median structures; paramere apically extending slightly beyond apex of median lobe, with short apical seta. Comparative notes: This species is distinguished from the externally similar representatives of the C. corruptum group by a more oblong aedeagus with a bulbous capsule (not strongly depressed dorso-ventrally), with a more slender and apically less broadly truncate ventral process, and with smaller internal structures of different shapes and arrangement, from the geographically close C. corruptum also by more distinct punctation of the pronotum. It differs from other described and undescribed species of the C. jonicum group, which are distributed in the Balkans, primarily by the shapes of the internal structures (especially the apical structure) of the aedeagus, from most of them also by a smaller aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85D5754B36EFF513DA11281.taxon	distribution	Distribution and natural history: Unlike the externally similar species of the C. corruptum group (see below), C. reductum is not locally endemic, but distributed in both the Pontic (Kastamonu) and the Taurus Mountains (Konya, Antalya, Mersin) (Map 3). The specimens were sifted from grass and moss in a calcareous slope and from litter of Crataegus and Berberis (partly near rocks). The altitudes range from 1000 to 1570 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85C5754B0B6FA913AB51541.taxon	description	: protibia without distinct modifications; metaventrite extensively and shallowly impressed, depressed, or unmodified; aedeagus strongly dorso-ventrally depressed and short, much less than twice as long as broad; ventral process short and broad, apically broadly truncate; internal structures weakly to moderately sclerotized, not conspicuously minute, apico-medially with a more or less transverse structure; parameres thin, apically extending to, nearly to, or beyond apex of median lobe, with short and fine apical seta. The species of the C. corruptum group can be assigned to two subgroups, one of them (C. unguicum subgroup) including five species (C. unguicum, C. deplectens, C. sicaferum, C. geminum, C. duplum) from Southwest Anatolia and characterized by the presence of pairs of large spines in the internal sac of the aedeagus. The C. corruptum subgroup, which comprises the remaining ten species, is distributed in central southern Anatolia, Lebanon, and Jordan and lacks such distinct spines. In the study region, the representatives of the C. corruptum group are locally endemic in South and Southwest Turkey (thirteen species), Lebanon (one), and Jordan (one) (Map 3).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85C5753B334F9513ACC1161.taxon	description	(Figs 14, 57, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85C5753B334F9513ACC1161.taxon	materials_examined	Type material: Holotype : “ N 37 ° 07 ' 41 E 34 ° 33 ' 03, Türkei Mersin (25 / 26), Camliyayla, 3 km w. Sebil 1060 – 1090 m / Streu an Felsen, 24.4.2005 (25 / 26), leg. Brachat & Meybohm / Holotypus  Cephennium corruptum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 8 , 1 : same data as holotype (cAss); 1 : “ TR – Mersin, ca. 30 km NNW Tarsus, 430 m, No. 7, 37 ° 08 ' 43 N, 34 ° 44 ' 29 E, Pinus, Q. ilex, Juglans, 26.12.2000, V. Assing ” (cAss); 1 : “ TR Umg. Mersin, Mersin – Arslanköy, 9 km sö Akdag, N 36 ° 55 ' E 34 ° 32 ' / Süd-Türkei, 6.5.2002, leg. Brachat & Meybohm ” (cAss); 1 : “ TR Mersin (33), road to Arslanköy, 5 km SE Aladag 700 m / 36 ° 54 ' 45 N 34 ° 31 ' 44 E (33), 2.5.2004, leg. Brachat & Meybohm ” (cAss); 1 : “ TR Mersin (34), road to Arslanköy, Aladag 830 m / 36 ° 56 ' 20 N 34 ° 30 ' 12 E (34), 2.5.2004, leg. Brachat & Meybohm ” (cAss); 2 : “ N 37 ° 08 ' 50 E 034 ° 42 ' 49, TR Mersin Daripinari 710 m, E Camliyayla 25.4.2011, Brachat & Meybohm (28) ” (cAss); 2 : “ TR Umg. Adana, Ciftehan Tal – Camlyayla [sic] 430 m, N 37 ° 8 ', E 34 ° 44 ' / Süd-Türkei, 5.5.2002, leg. Brachat & Meybohm ” (cAss); 2 : “ N 37 ° 29 ' 34 E 35 ° 23 ' 56, Türkei Adana (37), Karsanti 910 m (w Kozan), Hasandede Gec. / Hang mit Seidelbast, 28.4.2005 (37), leg. Brachat & Meybohm ” (cAss); 1 , 1 : “ N 37 ° 29 ' 52 E 35 ° 22 ' 48, Türkei Adana (38), Karsanti 1100 m (w Kozan), w Hasandede Gec. / Tannenwald, 28.4.2005 (38), leg. Brachat & Meybohm ” (cAss); 1 : “ N 37 ° 20 ' 33 E 034 ° 54 ' 41, TR Adana Belemedik, 710 m, 15.4.2011, Brachat & Meybohm (2) ” (cAss); 1 , 1 : “ N 37 ° 32 ' 30 E 034 ° 58 ' 55, TR Adana Kamisli E 5 km 1450 m, 24.4.2011, Brachat & Meybohm (30) ” (cAss); 2 , 1 : “ N 37 ° 33 ' 00 E 034 ° 58 ' 36, TR Adana Kamisli E 4 km 1385 m, 26.4.2011, Brachat & Meybohm (31) ” (cAss); 1 : “ TR Umg. Adana, südl. Pozanti, Tekir, 1200 – 1400 m, N 37 ° 18 ', E 34 ° 51 ' / Süd-Türkei, 4.5.2002, leg. Brachat & Meybohm ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85C5753B334F9513ACC1161.taxon	etymology	Etymology: The specific epithet is the past participle of the Latin verb corrumpere (to decay) and alludes to the fungus-shaped internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85C5753B334F9513ACC1161.taxon	description	Description: Body length 1.0 – 1.2 mm; length of pronotum 0.30 – 0.35 mm; width of pronotum 0.40 – 0.44 mm; length of antennae 0.45 – 0.50 mm. Habitus as in Fig. 14. Colouration: body yellowish-red. Eyes composed of approximately five ommatidia. Antenna with antennomeres VIII – X moderately transverse. Pronotum 1.20 – 1.25 times as broad as long, broadest in anterior half, and moderately convex in cross-section; lateral margins straight in posterior half; punctation as fine as that of elytra. Elytra noticeably broader than pronotum, anteriorly with small transversely oval impression with tomentose whitish pubescence; humeral carina pronounced, approximately as long as the combined length of the basal four antennomeres, inner margin of this carina sharply delimited. : protibia unmodified; metaventrite depressed or shallowly impressed; aedeagus (Fig. 57) short, broad, and apically truncate, approximately 0.25 mm long; internal sac with structures of distinctive shapes; paramere apically extending nearly to apex of median lobe, with short apical seta. Comparative notes: Among other species of this group, C. corruptum is reliably identified only based on the shape and the internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85C5753B334F9513ACC1161.taxon	distribution	Distribution and natural history: The distribution includes several localities in Mersin and Adana provinces, central South Anatolia (Map 3). The specimens were sifted in various forested (Pinus, Juglans, Quercus ilex, Cedrus, Juniperus, etc.) and unforested habitats (grassy slopes) or found under stones at altitudes of 430 – 1450 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85B5753B36EFD313A881421.taxon	description	(Figs 15, 52, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85B5753B36EFD313A881421.taxon	materials_examined	Type material: Holotype : “ N 37 ° 23 ' 45 E 036 ° 48 ' 05, TR Kahramanmaras, Uzunsögut 660 m (47), 30 km ssw K. Maras / Laubstreu, 4.5.2005, leg. Brachat & Meybohm / Holotypus  Cephennium marasicum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 1 , 1 : “ N 37 ° 26 ' 04 E 036 ° 44 ' 32 (8), TR Kahramanmaras, 25 km sw Maras, Yesilyöre 800 m, Brachat & Meybohm 23.4.2007 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85B5753B36EFD313A881421.taxon	etymology	Etymology: The specific epithet is an adjective derived from Maras, alluding to the fact that the type locality is situated in the Turkish province Kahramanmaraş.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85B5753B36EFD313A881421.taxon	description	Description: Habitus as in Fig. 15. Punctation of pronotum more distinct than that of elytra. Other external and male secondary sexual characters as in C. corruptum. : aedeagus (Fig. 52) short, broad, and apically truncate, 0.33 mm long; internal sac with structures of distinctive shape; paramere apically extending beyong apex of median lobe, with short apical seta. Comparative notes: Cephennium marasicum is distinguished from the externally similar and geographically close C. corruptum only by relatively more distinct punctation of the pronotum, as well as by the internal structures and longer parameres of the larger aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85B5753B36EFD313A881421.taxon	distribution	Distribution and natural history: This species is currently known from only two localities to the southwest of Kahramanmaraş, Kahramanmaraş province, central southern Anatolia (Map 3). The specimens were sifted from leaf litter near rocks at altitudes of 660 and 800 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85B5752B36EF8F13B4017A1.taxon	description	(Figs 16, 53, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85B5752B36EF8F13B4017A1.taxon	materials_examined	Type material: Holotype : “ N 37 ° 23 ' 10 E 36 ° 40 ' 51, TR Kahramanmaras, Doluca 1040 m (49), 40 km ssw K. Maras / Steine, Büsche, 4.5.2005 (49), leg. Brachat & Meybohm / Holotypus  Cephennium dolucanum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 1 : same data as holotype (cAss); 2 , 1 : “ N 37 ° 23 ' 10 E 36 ° 40 ' 51 (28), TR Kahramanmaras, W Doluca 1040 m, 40 km SSW Maras Brachat & Meybohm 30.4.2007 ” (cAss); 2 , 1 : same data, but “ (28 a) ... 2.5.2007 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85B5752B36EF8F13B4017A1.taxon	etymology	Etymology: The specific epithet is an adjective derived from Doluca, the populated place in whose environs the type locality is situated.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85B5752B36EF8F13B4017A1.taxon	description	Description: Habitus as in Fig. 16. External and male secondary sexual characters as in C. marasicum. : aedeagus (Fig. 53) short, broad, and apically truncate, 0.32 mm long; internal sac with structures of distinctive shape; paramere apically extending beyond apex of median lobe, with short apical seta. Comparative notes: Cephennium dolucanum is distinguished from the externally identical and geographically close C. marasicum only by the internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85B5752B36EF8F13B4017A1.taxon	distribution	Distribution and natural history: The type locality is situated near Doluca, some 40 km to the southwest of Kahramanmaraş, Kahramanmaraş province, central southern Anatolia (Map 3). The specimens were found under stones in the transition zone between arable land and a pine forest an altitude of 1040 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85A5752B334FF513A8B14E1.taxon	description	(Figs 17, 58, 70, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85A5752B334FF513A8B14E1.taxon	materials_examined	Type material: Holotype : “ N 37 ° 57 ' 20 E 36 ° 06 ' 22, Türkei Adana (35 / 36), Eyüplü 1560 m, s Saimbeyli (n Kozan) / Tannen, Polsterpfl., 27.4.2005 (35 / 36), Brachat & Meybohm / Holotypus  Cephennium fimbriatum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 5 , 4 : same data as holotype (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85A5752B334FF513A8B14E1.taxon	etymology	Etymology: The specific epithet is an adjective derived from the Latin noun fimbria (fringe) and alludes to the partly fimbriate internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85A5752B334FF513A8B14E1.taxon	description	Description: Habitus as in Fig. 17. Pronotum with slightly more distinct punctation than elytra. Other external and male secondary sexual characters as in C. marasicum and C. corruptum. : aedeagus (Fig. 58) short, broad, and apically truncate, approximately 0.27 mm long; internal sac with partly fimbriate structures of distinctive shape; paramere apically extending beyond apex of median lobe, with short apical seta. Comparative notes: Cephennium fimbriatum is distinguished from the externally similar C. marasicum and C. dolucanum by finer punctation of the pronotum and especially by the smaller aedeagus with internal structures of different shapes.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C85A5752B334FF513A8B14E1.taxon	distribution	Distribution and natural history: The type locality (Fig. 70) is situated to the north of Kozan, Adana province, central southern Anatolia (Map 3). The specimens were sifted from cushion plants and fir litter at an altitude of 1560 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595751B0E8F9F13DB11781.taxon	description	(Figs 19, 55, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595751B0E8F9F13DB11781.taxon	materials_examined	Type material: Holotype : “ N 37 ° 39 ' 14 E 36 ° 26 ' 27, TR Kahramanmaras, Andirin – Geben, 1500 m, 12,5 km von Andirin (45) / Felswand, Tannen, 3.5.2005 (45), leg. Brachat & Meybohm / Holotypus  Cephennium affimbriatum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 3 , 1 : same data as holotype (cAss); 2 : “ N 37 ° 33 ' 28 E 36 ° 25 ' 54, TR Kahramanmaras, Torlar 1110 m (41), 18 km sö Andirin / Streu, Steine (41), 30.4. / 2.5.2005 (45), leg. Brachat & Meybohm ” (cAss); 3 : “ N 37 ° 38 ' 19 E 36 ° 20 ' 51, TR Kahramanmaras, Andirin – Cokak 1150 m, 10 km n Andirin (41) / Carpinus, Quellen, 1. / 2.5.2005 (43), leg. Brachat & Meybohm ” (cAss); 1 : “ N 37 ° 39 ' 14 E 36 ° 26 ' 27, TR Kahramanmaras, Andirin 12,5 km NE, S Geben, 1500 m, Brachat & Meybohm, 27.4.2007 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595751B0E8F9F13DB11781.taxon	etymology	Etymology: The specific epithet (without a fringe) alludes to the absence of fimbriate structures in the internal sac of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595751B0E8F9F13DB11781.taxon	description	Description: Habitus as in Fig. 19. Punctation of pronotum not more distinct than that of elytra. Other external and male secondary sexual characters as in C. marasicum and C. corruptum. : aedeagus (Fig. 55) short, broad, and apically truncate, approximately 0.27 mm long; internal sac with non-fimbriate structures of distinctive shapes; paramere apically extending slightly beyond apex of median lobe, with short apical seta. Comparative notes: Cephennium affimbriatum is reliably distinguished from other species of the C. corruptum group only by the shape and internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595751B0E8F9F13DB11781.taxon	distribution	Distribution and natural history: The currently known distribution is confined to the environs of Andırın in the west of Kahramanmaraş province, central southern Anatolia (Map 3). The specimens were sifted from litter of fir, hornbeam, cedar, and other plants, partly between rocks, at altitudes of 1110 – 1500 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595751B0E8FF5138541521.taxon	description	(Figs 18, 54, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595751B0E8FF5138541521.taxon	materials_examined	Type material: Holotype : “ TR Karahmanmaras [sic] (18) Pass N Tekir S Göksun, 1400 – 1550 m / 37 ° 56 ' 56 N 36 ° 34 ' E (18) 26.4.2004, Brachat & Meybohm / Holotypus  Cephennium besucheti sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 2 , 3 : same data as holotype (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595751B0E8FF5138541521.taxon	etymology	Etymology: This species is dedicated to our late friend, Claude Besuchet, specialist of Pselaphinae and Scydmaeninae. He studied Cephennium, including the available type material of the described species, in the 1960 s, prepared superb drawings of the aedeagus of both described and numerous undescribed species, but unfortunately never published his results. He also participated in the field trip to Turkey in 2004.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595751B0E8FF5138541521.taxon	description	Description: Habitus as in Fig. 18. Pronotum with more distinct punctation than elytra. Other external and male secondary sexual characters as in C. marasicum and C. corruptum. : aedeagus (Fig. 54) short, broad, and apically truncate, approximately 0.33 mm long; internal sac with narrowly fimbriate structures of distinctive shape; paramere apically extending slightly beyond apex of median lobe, with short apical seta. Comparative notes: Cephennium besucheti is reliably distinguished from other species of the C. corruptum group only by the shape and internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595751B0E8FF5138541521.taxon	distribution	Distribution and natural history: The type locality is situated to the southeast of Göksun in the west of Kahramanmaraş province, central southern Anatolia (Map 3). The specimens were sifted from litter on slopes with cedar, fir, and cypress at an altitude between 1400 – 1550 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595750B36EFB51387C1241.taxon	description	(Figs 20, 56, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595750B36EFB51387C1241.taxon	materials_examined	Type material: Holotype : “ TR Gaziantep (17), Kartal Dag 1200 m, W Yamacoba / 37 ° 10 ' N E 37 ° 5 ' E (17) 25.4.2004, leg. Brachat & Meybohm / Holotypus  Cephennium kartalicum sp. n., det. V. Assing 2020 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595750B36EFB51387C1241.taxon	etymology	Etymology: The specific epithet is an adjective derived from Kartal, the name of the mountain range where the species was discovered.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595750B36EFB51387C1241.taxon	description	Description: Body length 1.0 mm. Habitus as in Fig. 20. Punctation of pronotum much more distinct than that of elytra. Other external and male secondary sexual characters as in C. marasicum and C. corruptum. : aedeagus (Fig. 56) short, broad, and apically truncate, 0.24 mm long; internal sac with non-fimbriate structures of distinctive shapes; paramere apically extending slightly beyond apex of median lobe, with short apical seta. Comparative notes: Cephennium kartalicum is reliably distinguished from other species of the C. corruptum group only by the shape and internal structures of the rather small aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8595750B36EFB51387C1241.taxon	distribution	Distribution and natural history: The type locality is situated in Kartal Dağı, Gaziantep province, central southern Anatolia (Map 3). The holotype was sifted at an alitude of 1200 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B334FE513C411561.taxon	description	(Figs 24, 67, 71)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B334FE513C411561.taxon	materials_examined	Type material: Holotype : “ N 32 ° 19 ' 32 E 35 ° 43 ' 26, Jordan Ajlun, Ajlun Castle 980 m (2), lg. Meybohm 21.2.2014 / Holotypus  Cephennium jordanicum sp. n., det. V. Assing 2020 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B334FE513C411561.taxon	etymology	Etymology: The specific epithet is an adjective derived from Jordan, where this species is at present the sole representative of the genus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B334FE513C411561.taxon	description	Description: Body length 1.1 mm. Habitus as in Fig. 24. Punctation of pronotum more distinct than that of elytra. Eyes small, composed of three or four weakly defined ommatidia without pigmentation. Other external and male secondary sexual characters similar to those of other species of the C. corruptum group. : aedeagus (Fig. 67) 0.24 mm long, broad, and apically truncate; internal sac with two weakly sclerotized apicomedian structures, the lower one fringed with minute spines; paramere apically extending slightly beyond apex of median lobe, with short apical seta. Comparative notes: Based on the similar shape and internal structures of the aedeagus, C. jordanicum is most closely allied to C. judaeum, from which it is distinguished by the more transverse apical and the more strongly angled subapical internal structure.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B334FE513C411561.taxon	distribution	Distribution and natural history: The type locality is situated near Ajlun, some 30 km to the south-southwest of Irbid, Northwest Jordan. The holotype was sifted in a pasture woodland at an altitude of 980 m (Fig. 71).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B0B6F9F13D841281.taxon	description	(Figs 68 – 69)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B0B6F9F13D841281.taxon	discussion	Comment: The original description is based on an unspecified number of syntypes, among them at least one male, from “ Libanon: Beirut ” (REITTER 1882). Claude Besuchet (unpublished notes) examined the following material: 1 . “ Beirut. type. coll. Croissandeau in Mus. Paris ”; 1 : “ Libanon. Sahlberg. Mus. Helsinki ”; 1 : “ Fleuve Lycus, Liban. Sahlberg. Mus Helsinki ”; 1 : “ Beirut. Appl. coll. mea [now probably in Muséum d'histoire naturelle, Genève] ”. Claude Besuchet´s drawing of the aedeagus (probably of the male in his collection) is illustrated in Figs 68 – 69. Based on this drawing, C. judaeum belongs to the C. corruptum group and is most closely allied to the following species. It has been reported only from Lebanon (SCHÜLKE & SMETANA 2015).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B0B6FE113BF41521.taxon	description	(Figs 21, 51, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B0B6FE113BF41521.taxon	materials_examined	Type material: Holotype : “ TR – Adıyaman [22], 50 km NE Adıyaman, 8 km NE Narince, 870 m, 37 ° 54 ' 58 N, 38 ° 48 ' 45 E, 24. III. 2005, P. Wunderle / Holotypus  Cephennium nemruticum sp. n., det. V. Assing 2020 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B0B6FE113BF41521.taxon	etymology	Etymology: The specific epithet is an adjective derived from Nemrut, the name of the mountain where the species was discovered.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B0B6FE113BF41521.taxon	description	Description: Body length 1.2 mm. Habitus as in Fig. 21. Punctation of pronotum only slightly more distinct than that of elytra. Eye rudiments very small, composed of approximately 3 weakly defined ommatidia. Other external and male secondary sexual characters as in C. marasicum and C. corruptum. : aedeagus (Fig. 51) short, broad, and apically truncate, approximately 0.33 mm long; internal sac with weakly sclerotized non-fimbriate structures of distinctive shapes; paramere apically extending to apex of median lobe, with short apical seta. Comparative notes: Cephennium nemruticum is reliably distinguished from other species of the C. corruptum group only by the shape and internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C8585750B0B6FE113BF41521.taxon	distribution	Distribution and natural history: The type locality is situated in Nemrut Dağı, Adıyaman province, central southern Anatolia (Map 3). The holotype was sifted from leaf litter in a rocky slope with oak at an alitude of 870 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C858575FB334F9713BB51401.taxon	description	(Figs 22, 59, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C858575FB334F9713BB51401.taxon	materials_examined	Type material: Holotype : “ N 37 ° 41 ' 26 E 027 ° 13 ' 43 (24), Türkei Aydin Dilek Dagi, s Güzelcamli 410 m 28.4.2006, l. Brachat & Meybohm / Holotypus  Cephennium unguicum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 3 , 2 : same data as holotype (cAss); 1 : “ N 37 ° 40 ' 19 E 027 ° 10 ' 41 (2), Türkei Aydin Dilek Dagi, s Kanyon 670 m 17.4.2006, l. Brachat & Meybohm ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C858575FB334F9713BB51401.taxon	etymology	Etymology: The specific epithet is an adjective derived from the Latin noun unguis (claw) and alludes to the conspicuous pair of claw-shaped structures in the internal sac of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C858575FB334F9713BB51401.taxon	description	Description: Body length 1.0 – 1.1 mm. Habitus as in Fig. 22. Punctation of pronotum more distinct than that of elytra. Eye rudiments minute, composed of three or four weakly defined ommatidia without pigmentation. Other external and male secondary sexual characters as in C. marasicum and C. corruptum, except as follows: : metaventrite not impressed, only indistinctly depressed in the middle at most; aedeagus (Fig. 59) small, broad, and apically truncate, 0.23 - 0.25 mm long; internal sac with a pair of large and strongly curved claw-shaped structures; paramere apically extending to apex of median lobe, with short apical seta. Comparative notes: Cephennium unguicum is distinguished from the species of the C. corruptum subgroup by the presence of claw-shaped spines in the internal sac of the aedeagus. For characters separating it from other species of the C. unguicum subgroup see the comparative notes in the following sections.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C858575FB334F9713BB51401.taxon	distribution	Distribution and natural history: The type locality is situated in Dilek Dağı (in the peninsula close to the Greek island Samos), Aydın province, Southwest Anatolia (Map 3). The specimens were collected at an altitude of 410 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C857575FB0E8F8D13C5B1B01.taxon	description	(Fig. 61, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C857575FB0E8F8D13C5B1B01.taxon	materials_examined	Type material: Holotype : “ N 37 ° 52 ' 59 E 027 ° 53 ' 54 (11), Türkei Aydin Pasayaylasi, 250 m 20.4.2006, l. Brachat & Meybohm / Holotypus  Cephennium sicaferum sp. n., det. V. Assing 2020 ” (cAss). Paratype : “ N 37 ° 56 ' 47 E 027 ° 53 ' 53 (8), Türkei Aydin Pasayaylasi, 1460 m 20.4.2006, l. Brachat & Meybohm ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C857575FB0E8F8D13C5B1B01.taxon	etymology	Etymology: The specific epithet is an adjective composed of the Latin noun sica (dagger) and the suffix - ferum (carrying). It alludes to the presence of a pair of daggershaped structures in addition to a pair of claw-shaped structures in the internal sac of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C857575FB0E8F8D13C5B1B01.taxon	description	Description: Body length 0.9 mm. Eye rudiments minute, composed of three distinct ommatidia without pigmentation. Other external and male secondary sexual characters as in C. unguicum. : aedeagus (Fig. 61) of similar shape as in C. unguicum, but smaller, 0.18 mm long; internal sac with a pair of large and strongly curved claw-shaped structures and additionally with a pair of dagger-shaped structures. Comparative notes: Cephennium sicaferum is distinguished from the geographically close C. unguicum by slightly smaller body size, eye rudiments composed of three distinct ommatidia, and by a smaller aedeagus with a pair of dagger-shaped structures in addition to a pair of smaller and differently shaped claw-shaped structures. The species differs from the externally similar C. deplectens (see below) only by the longer and more slender dagger-shaped structures and the differently shaped claw-shaped structures in the internal sac.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C857575FB0E8F8D13C5B1B01.taxon	distribution	Distribution and natural history: The type locality is situated near Paşayaylası, Aydın province, Southwest Anatolia (Map 3). The holotype was sifted from litter on a stream bank with Salix and Rubus at an altitude of 250 m, the paratype was found under stones or sifted near stones near a stand of walnut trees at an altude of 1460 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C856575EB0B6FF5138FE14A1.taxon	description	(Figs 23, 60, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C856575EB0B6FF5138FE14A1.taxon	materials_examined	Type material: Holotype : “ N 38 ° 24 ' 19 E 027 ° 23 ' 31 (17), Türkei Izmir Nif Dagi, 970 m 24.4.2006, l. Brachat & Meybohm / Holotypus  Cephennium deplectens sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 6 , 4 : same data as holotype (cAss); 1 : “ N 38 ° 24 ' 13 E 027 ° 23 ' 05 (18), Türkei Izmir Nif Dagi, 1080 m 24.4.2006, l. Brachat & Meybohm ” (cAss); 1 : “ N 38 ° 32 ' 51 E 027 ° 25 ' 14 (20), Türkei Manisa Sipil Dagi Milli Parki, 1250 m 25.4.2006, l. Brachat & Meybohm ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C856575EB0B6FF5138FE14A1.taxon	etymology	Etymology: The specific epithet is the present participle of the Latin verb deplecti (to claw) and alludes to the conspicuous pair of claw-shaped structures in the internal sac of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C856575EB0B6FF5138FE14A1.taxon	description	Description: Very small species; body length 0.9 – 1.0 mm. Habitus as in Fig. 23. Punctation of pronotum more distinct than that of elytra. Eye rudiments minute, composed of three or four weakly defined ommatidia without pigmentation. Other external and male secondary sexual characters as in other species of the C. unguicum subgroup, except as follows: : metaventrite not impressed, only indistinctly depressed in the middle at most; aedeagus (Fig. 60) very small, broad, and apically truncate, 0.18 mm long; internal sac with a pair of large and strongly curved claw-shaped structures, with additional pair of straight stout spines, and with a strongly transverse apicomedian structure; paramere apically extending to apex of median lobe, with short apical seta. Comparative notes: As can be inferred particularly from the similar morphology of the aedeagus (small size; internal sac with pair of claw-shaped structures) and the absence of a median impression on the male metaventrite, C. deplectens is evidently closely allied to C. unguicum, from which it is distinguished by smaller body size and particularly by a smaller aedeagus with internal structures of different shapes and arrangement, and to C. sicaferum, from which it differs only by the shapes of the internal structures of the aedeagus (see the comparative notes in the section on C. sicaferum above).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C856575EB0B6FF5138FE14A1.taxon	distribution	Distribution and natural history: The species was found in two geographically close mountains (Nif Dağı, Sipil Dağı) in Izmir and Manisa provinces, West Anatolia (Map 3). The specimens were sifted from litter near rocks at altitudes of 970 – 1250 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C856575DB334FF5138A11281.taxon	description	(Figs 62 – 64, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C856575DB334FF5138A11281.taxon	materials_examined	Type material: Holotype : “ TR – Antalya, Umgeb. Antalya, Termessos, 14.4.1984, leg. V. u. C. Brachat / Holotypus  Cephennium geminum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 3 , 3 exs.: same data as holotype (cAss); 2 , 8 exs.: “ N 36 ° 59 ' 23 E 30 ° 28 ' 6, TR Antalya Termessos, 880 m 9.3.1979, leg. Fülscher & Meybohm ” (cAss); 3 , 1 : “ Türkei Umg. Antalya, Termessos 550 m – 800 m 12.5.2000, leg. Brachat & Meybohm ” (cAss); 1 , 2 : “ N 36 ° 59 ' 48 E 30 ° 28 ' 35, TR Antalya Termessos, 700 m 21.4.2001, leg. Brachat & Meybohm ” (cAss); 1 : “ N 36 ° 43 ' E 30 ° 26 ', Türkei Umg. Antalya, südl. Hisar, 1120 m, Meybohm 23.4.2001 ” (cAss); 1 : “ Türkey Umg. Antalya, Strasse nach Saklikent, 1000 m Kiefernwald, Meybohm 11.5.2000 ” (cAss); 2 : “ N 36 ° 39 ' 16 E 30 ° 25 ' 27 (2), TR Antalya, Olimpos nw Ovacik 1255 m, Brachat & Meybohm 12.4.2008 ” (cAss); 1 : “ TR. – Antalya, No. 23, W Kemer, road to Ovacik, Quercus litter, 325 m, 36 ° 36 ' 18 N, 30 ° 28 ' 38 E, 2. IV. 2002, V. Assing ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C856575DB334FF5138A11281.taxon	etymology	Etymology: The specific epithet (Latin adjective: doubled) alludes to the presence of two pairs of claw-shaped structures in the internal sac of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C856575DB334FF5138A11281.taxon	description	Description: Body length 1.0 – 1.1 mm. Eye rudiments minute, composed of three or four weakly defined ommatidia without pigmentation. Other external and male secondary sexual characters as in the preceding species, except as follows: : metaventrite depressed in the middle; aedeagus (Fig. 62 – 64) approximately 0.23 mm long; internal sac with two pairs of large and strongly curved claw-shaped structures, with an additional pair of weakly curved spines, and with a transverse apico-median structure; paramere apically extending beyond apex of median lobe, with short apical seta. Intraspecific variation: The males from Termessos differ from those from other localities by slightly longer and more slender weakly curved structures (i. e., the most basal pair of internal structures). However, since slight variation was observed also in the shapes of the remaining structures and between specimens from all localities, and since no other evidence was found suggesting that these populations should represent distinct species, the observed differences are attributed to intraspecific variation. Comparative notes: This species is reliably distinguished from the preceding representatives of the C. unguicum subgroup by the presence of three pairs of large sclerotized spines in the internal sac of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C856575DB334FF5138A11281.taxon	distribution	Distribution and natural history: The known distribution of C. geminum is confined to the west of Antalya province, Southwest Anatolia (Map 3). The specimens were sifted from litter in oak and pine forests and in a north slope with cedar, cypress, and oak shrubs at altitudes of 325 – 1255 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575DB0E8FE513DAA1282.taxon	description	(Figs 65 – 66, Map 3)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575DB0E8FE513DAA1282.taxon	materials_examined	Type material: Holotype : “ N 36 ° 43 ' E 029 ° 14 ', Türkei Umg. Fethiye, Kadyanda 850 m, Meybohm 28.4.2001 / Holotypus  Cephennium duplum sp. n., det. V. Assing 2020 ” (cAss). Paratypes: 1 : “ N 36 ° 33 ' E 029 ° 07 ', T Umg. Fethiye, Ölüdeniz, 150 m, 27.4.2001, Meybohm / Brachat ” (cAss); 1 : “ N 36 ° 33 ' E 029 ° 07 ', Türkei Umg. Fethiye, Ölüdeniz, 150 m, Meybohm 27.4.2001 ” (cAss); 1 : “ N 36 ° 36 ' E 029 ° 06 ', Türkei Umg. Fethiye, Fethiye, 360 m, Meybohm 27.4.2001 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575DB0E8FE513DAA1282.taxon	etymology	Etymology: The specific epithet (Latin adjective: doubled) alludes to the presence of two pairs of claw-shaped structures in the internal sac of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575DB0E8FE513DAA1282.taxon	description	Description: Body length 0.9 – 1.0 mm. Eye rudiments minute, composed of three or four weakly defined ommatidia without pigmentation. Other external and male secondary sexual characters as in the preceding species. : aedeagus (Fig. 65 – 66) approximately 0.20 mm long; internal sac with two pairs of large and strongly curved claw-shaped structures, with an additional pair of weakly curved spines, and with a transverse apico-median structure; paramere apically extending beyond apex of median lobe, with short apical seta. Intraspecific variation: The internal structures of the two examined males are not completely identical (Figs 65 – 66). However, the differences are only slight and the two localities are close, so that the differently shaped internal structures are interpreted as intraspecific variation. Comparative notes: As can be inferred from the similar shapes and arrangement of the internal structures of the aedeagus, C. duplum is most closely allied to the geographically close C. geminum, from which it differs by slightly smaller body size and a smaller aedeagus with less massive and less strongly curved internal spines and with a differently shaped apical structure.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575DB0E8FE513DAA1282.taxon	distribution	Distribution and natural history: The specimens were found in three localities near Fethiye, Muğla province, Southwest Anatolia (Map 3). The altitudes range from 150 to 850 m. Unidentified and unnamed species The following material is represented exclusively by females, so that an adequate description is not possible. Based on external characters, all of them most likely belong to the C. corruptum group.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575DB36EFE513D7711A1.taxon	materials_examined	Material examined: 1 : “ N 39 ° 43 ' 29 E 27 ° 11 ' 01 (13), TR Canakkale 14.4.2010, Kurucam Tepe 430 m, leg. Brachat & Meybohm ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575DB36EFD713D061041.taxon	materials_examined	Material examined: 1 : “ N 41 ° 13 ' 20 E 31 ° 53 ' 6 (26,2), TR Zonguldak 3.5.2014, Devrek 590 m, leg. Brachat & Meybohm ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575DB36EFC113C3A1761.taxon	materials_examined	Material examined: 1 : “ TR Umg. Osmaniye, Nur Dagl, Strasse nach Zorkum, 550 – 900 m, N 37 ° 1 ' E 36 ° 16 ' / Süd- Türkei, 25.4.2002, Brachat & Meybohm ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575DB36EFB313D8D16A1.taxon	materials_examined	Material examined: 1 : “ N 37 ° 33 ' 32 E 036 ° 35 ' 20, TR Kahramanmaras, Baskonus Yaylasi 1450 m, 50 km von K. Maras (51) / Tannenwald, 5.5.2005 (51), leg. Brachat & Meybohm ” (cAss); 1 : “ N 37 ° 33 ' 56 E 036 ° 33 ' 38 (10), TR Kahramanmaras, Baskonus Yaylasi 1250 m, Brachat & Meybohm 24.4.2007 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575CB36EF8B13B5217A1.taxon	description	(Figs 27 – 28, Map 1)	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575CB36EF8B13B5217A1.taxon	materials_examined	Type material: Holotype : “ BG Strandzha Mts., Malko Tarnovo 300 m, N 41.98928 E 27.52467, l. Bekchiev 18.4.2010 / Holotypus  Cephennium liguliferum sp. n., det. V. Assing 2020 ” (cAss).	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575CB36EF8B13B5217A1.taxon	etymology	Etymology: The specific epithet (Latin, adjective: carrying a spoon) alludes to the spoon-shaped apex of the apical extension of the ventral process of the aedeagus in ventral view.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575CB36EF8B13B5217A1.taxon	description	Description: Body length 1.3 mm. External and male secondary sexual characters as in C. bodemeyeri and C. perispinctum: : aedeagus (Figs 27 – 28) 0.55 mm long, slender; ventral process in ventral view with nearly parallel lateral margins, not constricted, with marked subapical angles, and apically with a long median extension, this extension apically somewhat spoon-shaped; internal sac with a pair of long and rather stout structures; paramere basally stout, smoothly tapering apicad, strongly bisinuate, and apically with long curved seta. Comparative notes: Based on the similar external characters and particularly on the similar structure of the aedeagus, this species is undoubtedly closely allied to C. bodemeyeri and C. perispinctum. It is distinguished from both of them by the spoon-shaped apical extension of the ventral process and by much longer and more massive internal structures of the aedeagus.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
03FA8790C855575CB36EF8B13B5217A1.taxon	distribution	Distribution: The type locality is situated in Strandzha Mountains, Southeast Bulgaria, not far from the border with the Turkish province Kırklareli (Map 1). The holotype was collected at an altitude of 300 m.	en	Assing, Volker, Meybohm, Heinrich (2021): The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae). Beiträge Zur Entomologie = Contributions to Entomology 71 (1): 1-28, DOI: 10.21248/contrib.entomol.71.1.001-028
