taxonID	type	description	language	source
03FA87EAFFF4FF9D5988001EFC877774.taxon	materials_examined	Type species: Ozopemon laevis Strohmeyer 1911: 22	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF4FF9D5988001EFC877774.taxon	diagnosis	Diagnosis. The genus Dryocoetiops Schedl belongs in the tribe Dryocoetini (Wood 1986; Alonso-Zarazaga & Lyal 2009). Generic boundaries in the tribe are not always clear (Wood 1986), and Dryocoetiops is no exception. Many members of the tribe appear to have undergone mosaic evolution, and genera can be distinguished only by combinations of characters. Dryocoetiops is distinguished morphologically from other genera included in the tribe by the following combination of characters: 1) antennal funicle four-segmented (excluding the pedicel), club flattened or weakly obliquely truncate, sutures strongly displaced apically on posterior face, anterior face with basal segment confined to basal fifth or quarter, its apical margin convex to straight; and 2) eye emarginate, in a few species rather large, coarsely facetted and extending onto the frons; and 3) female frons without a brush of setae, sparsely setose, not aciculate; and 4) pronotum about as long as wide or wider than long, strongly declivous anteriorly, usually with rather coarse, rasp-like asperities, unequal in size and larger antero-laterally, the summit obtuse, humped, at or a little behind the middle, a weak to distinct transverse impression behind it, disc flat in lateral view, usually granulate-punctate; and 5) elytra with distinct striae and interstriae; and 6) elytral declivity steep, convex, interstriae granulate except in one species; and 7) protibiae with four socketed teeth (rarely five) in apical half, without further smaller teeth on external margin more basally.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF4FF9D5988001EFC877774.taxon	description	Description. Female. Antenna with 4 - segmented funicle (excluding the pedicel), club flattened or weakly obliquely truncate, without septum, sutures strongly displaced apically on posterior face, basal segment not extending beyond basal quarter on anterior face, its apical margin convex, sinuate or approximately straight. Eyes distinctly but not deeply emarginate anteriorly, either of normal size, finely faceted, or in a few species enlarged, more coarsely faceted, and extending onto frons. Frons punctate or granulate-punctate, usually with a median raised line, sometimes with a median tubercle. Pronotum strongly declivous anteriorly, rising to an obtuse hump-like elevation, approximately in the middle, with a weak to distinct transverse impression behind it. Anterior slope of pronotum with rasp-like asperities, often extending to anterior margin, usually increasing in size antero-laterally, the disc flat in lateral view, granulate-punctate. Elytra convex, striae and interstriae distinct, strial punctures with short, fine, recumbent setae, interstrial punctures with stouter, longer, erect setae, occasionally flattened on declivity. Elytral declivity steep, convex, striae 1, and less often striae 2 – 3 impressed, interstriae usually granulate, 1 – 3 raised and costate in two species. Procoxae usually narrowly separated, and prosternal process pointed, but contiguous and prosternal process blunt in a few species. Protibiae with four socketed teeth (rarely five), meso- and metatibiae usually with six teeth. Body length 1.9 – 4.0 mm. Male. Unknown. Presumed to be dwarfed and flightless as in Coccotrypes Eichhoff, 1878 a.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF4FF9D5988001EFC877774.taxon	distribution	Distribution. The genus is primarily palaeotropical in its distribution from India through South-East Asia and Indonesia to New Guinea and northern Australia. Two species (D. apatoides, D. moestus) occur as far north as Japan. Relationships. Early phylogenetic studies (Jordal et al. 2000, 2002) suggested that Dryocoetiops should be included within the genus Coccotrypes. The studies were based on D. coffeae and D. cf. eugeniae [sic]. Both D. coffeae and D. eugeniae are now considered to be synonyms of D. moestus (see below). Later phylogenetic studies (Jordal & Cognato 2012; Gohli et al. 2017) suggest that Dryocoetiops should rather be considered as a sister genus to Coccotrypes. It appears to have the same haplo-diploid breeding system as Coccotrypes (Jordal et al. 2002), but is morphologically distinct. Species of Dryocoetiops can be distinguished from Coccotrypes most easily by characters of the pronotum. In Dryocoetiops, 1) pronotum dome-shaped with a distinct summit near the middle, as wide as or wider than long, steeply sloping anteriorly, asperities often increasing in size antero-laterally, anterior margin broadly rounded, without a distinct row of asperities on the margin, slightly to distinctly impressed behind summit, disc flat in lateral view; and 2) frons without aciculation. In Coccotrypes, the pronotum is usually longer than wide, and if there is a distinct summit, it is often behind the middle. The anterior slope is less steep, and the asperities usually smaller, and not increasing in size antero-laterally. In some species, there is a row of larger asperities on the anterior margin only. The species of Dryocoetiops are myelophagous (Kirkendall et al. 2015) breeding in the pith of small branches and twigs; those of Coccotrypes breed in a variety of habitats, including fruits and seeds, phloem, leafstalks, and mangrove propagules (Browne 1961; Jordal et al. 2002), but almost never in the pith of twigs. Besides Coccotrypes, the genus is most similar morphologically to Taphrorychus (Schedl 1957), although it is not very closely related (Jordal & Cognato 2012; Gohli et al. 2017), and has a very different biology and breeding system (see below). However, the two genera have been confused in the past, and some species now placed in Dryocoetiops have previously been placed in Taphrorychus (see below). Hence, we provide distinguishing characters here. Dryocoetiops can be most easily distinguished morphologically as follows: 1) basal segment of labial palp enlarged, barrel-shaped; 2) frons without a brush of setae; and 3) pronotum with distinct summit and steep anterior slope; and 4) pronotal asperities coarse, increasing in size antero-laterally; and 5) elytra with distinct striae and interstriae; and 6) without a circular impression on the elytral declivity; and 7) protibiae without additional small denticles. In Taphrorychus, the basal segment of the labial palp is of normal size. The other characters can vary depending on the species and the sex of the individual, but the combination of all of them is never present.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF4FF9D5988001EFC877774.taxon	biology_ecology	Biology. The biology of Dryocoetiops has been investigated only in D. moestus (Browne 1961; Kalshoven 1958, both as Dryocoetes coffeae), but is probably similar in other species. The species are inbreeding, polyphagous twig-borers. Only the female disperses and constructs the nest. A longitudinal tunnel is constructed in the centre of the twig, usually extending both upwards and downwards from the entrance. Browne (1961) notes that the eggs are laid in clusters and are very variable in size. The larvae feed on the tissues surrounding the gallery, and eventually pupate in single file (Browne 1961). Browne (1961) also states that the species is monogamous with both parents playing a part in the care of the nest and the young brood. However, this part of his account of the biology is mistaken. As noted above, all known adults are female, and the male remains unknown. It is likely that Browne observed two females occupying the same gallery system. Dissections of females of D. moestus caught in ethanol traps show that the spermathecae contain sperm, indicating that the species is not parthenogenetic. By contrast, the morphologically similar species of Taphrorychus are outbreeding bark beetles, making gallery systems below the bark of angiosperm trees, particularly of the family Fagaceae. Both male and female are of the same size, and the sex ratio at emergence is approximately 1: 1. Both sexes are capable of flight, disperse, and are involved in gallery construction. The males are harem polygynous (Kirkendall et al. 2015), with more than one female associated with each male in most gallery systems.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF1FF9B598800E4FDB776EC.taxon	description	Body. Brown, 2.2 mm long, 2.44 times as long as wide. Head. Frons convex, moderately shining, sparsely finely punctured, the punctures with moderately long setae, fringe of dense setae on anterior margin of epistoma, a short elevated median line just above epistoma. Antenna with four-segmented funicle (excluding pedicle); first segment of antennal club extends one quarter of total length of club on anterior face, its apical margin convex apically. Eyes large, extending onto frons, coarsely faceted, upper part larger than lower part. Pronotum. 0.94 times longer than wide, widest about one-third of length from base, posterior angles broadly rounded, sides subparallel in basal half, broadly rounded anteriorly, anterior margin not tuberculate, summit at middle, a weak transverse depression just behind it; anterior slope with numerous asperities with rounded apex, larger and more elevated in antero-lateral areas, weaker and more strongly transverse towards the summit, each bearing on its posterior side a single stiff seta; disc shining, rather weakly, not very densely granulate, becoming rugulose at posterior angles, the granules with fairly short, stiff setae, scattered long setae on anterior and lateral margins. Scutellum. Shining, linguiform. Elytra. 1.56 times as long as wide, 1.75 times as long as pronotum, slightly wider than pronotum at base; sides parallel in basal 3 / 4, then evenly rounded to apex; striae not impressed, slightly wider than interstriae, strial punctures uniseriate and large, separated by about the width of one puncture, bearing minute, fine setae; interstrial punctures much smaller and more widely separated with thick, erect setae slightly flattened at the tips, becoming more scale-like toward declivity. Declivity with strial and interstrial punctures more closely placed; interstrial setae wedge shaped on interstriae 1 – 4. Metaventrite with a row of fairly closely spaced sclerolepidia along its dorsal margin. Legs. Anterior tibia narrowed toward apex with 4 socketed teeth in apical third. Procoxae narrowly separated.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF1FF9B598800E4FDB776EC.taxon	materials_examined	Type Material. Holotype: Female: Pen [insular] Malaysia, Negeri Sembilan, Pasoh Forest Reserve, P 93 - 2737, Tower-CS- 5, Ethanol trap, 27 Nov. – 18 Dec. 1993, coll. K. Maetô (NHML).	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF1FF9B598800E4FDB776EC.taxon	diagnosis	Diagnosis. This species is distinguished within the genus by: 1) the large eyes which extend onto the frons, with the upper part above the emargination larger than the lower part; 2) the interstrial setae which are short, flattened, and spatulate on interstriae 1 – 4 on the elytral declivity.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF1FF9B598800E4FDB776EC.taxon	etymology	Etymology. The specific name refers to the location (Pasoh Forest Reserve) where the species was found.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF1FF9B598800E4FDB776EC.taxon	distribution	Distribution. West Malaysia.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF0FF98598805C9FDE47774.taxon	description	Body. Piceous, 2.9 – 3.2 mm long, 2.31 – 2.33 times as long as wide. Head. Frons convex, moderately shining, sparsely, finely punctured, granulate punctate anteriorly, the punctures with moderately long setae, fringe of dense setae on anterior margin of epistoma, indistinct elevated median line. Antenna with 4 - segmented funicle (excluding pedicle); first segment of club extends one third of total length of club on anterior face, its apical margin transverse. Eyes of normal size, with upper part above emargination smaller than lower part, and not extending onto frons, finely faceted. Pronotum. 0.86 – 0.91 times longer than wide, widest close to base, posterior angles broadly rounded, sides subparallel in basal half, broadly rounded anteriorly, anterior margin not tuberculate, summit slightly behind middle, a very weak transverse impression just behind it; anterior slope with numerous, densely placed asperities of variable size, arranged more or less concentrically, larger and more elevated in antero-lateral areas, smaller towards summit, apex of asperities rounded; disc dull, densely granulate, the granules tending to be aligned in concentric rows, becoming rugulose at posterior angles; vestiture of moderately long, erect setae on anterior slope and margin, and at sides. Scutellum. Shining, linguiform. Elytra. 1.50 – 1.55 times as long as wide. 1.8 – 2.0 times as long as pronotum, slightly wider than pronotum at base, sides subparallel in basal 3 / 4, then angularly rounded to apex; striae impressed, wider than interstriae; strial punctures very coarse, biseriate on striae 1, except in basal third of elytra and apical part of declivity; striae 2 uniseriate at base and on posterior part of disc, but biseriate for a short distance in middle of disc and on declivity except close to apex, outer interstriae uniseriate; interstriae narrow, especially on declivity, where strongly raised with a single, regularly placed row of large, pointed granules extending to apex; disc of elytra almost glabrous apart from short setae near elytral margins, declivity with minute, fine setae in strial punctures, and a short, stouter, erect seta arising behind each granule on interstriae. Metaventrite with a row of fairly closely spaced sclerolepidia along its dorsal margin. Legs. Anterior tibia narrowed toward apex with 4 socketed teeth in apical third. Procoxae contiguous.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF0FF98598805C9FDE47774.taxon	materials_examined	Type material. Holotype: Female: BRUNEI, E 115.70 o, N 4.34 o, Kuala Belalong FSC, Dipterocarp forest, Dryobalanops beccarii, aerial FIT 4, 270 m alt., 7. vi. [19] 91, N. Mawdsley, NM 192, (BM (NH) 1991 – 173) (NHML). Paratypes: Female: 3 specimens: same data as holotype (1 in RABC); as holotype except: aerial FIT 6, 275 m alt., 23. v. [19] 91, NM 168 (1); as holotype except: aerial FIT 1 A, 260 m alt., 8. vi. [19] 91, NM 183 (1) (both in NHML).	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF0FF98598805C9FDE47774.taxon	diagnosis	Diagnosis. The species is unique in the genus in the very large strial punctures on the elytra, and the coarse, pointed granules on the elytral declivity. The punctures of striae 1 appear biseriate between the raised interstriae 1 and 2, except in the basal third of the elytra and the apical part of declivity; those of striae 2 are similarly biseriate on the upper part of the declivity. It resembles D. apatoides in the costate interstriae and strongly impressed striae 1 – 3 on the elytral declivity, but is easily distinguished from that species by its greater size and the unusual strial puncturation.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF0FF98598805C9FDE47774.taxon	etymology	Etymology. The specific name is a Latin adjective meaning rugged or rough, and refers to the coarse sculpture of the elytra.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF0FF98598805C9FDE47774.taxon	distribution	Distribution. Brunei Darussalam.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF3FF985988054DFE077353.taxon	distribution	Distribution. China (Yunnan), Japan, Taiwan, Vietnam. New records. CHINA, S – Yunnan, (Xishuangbanna), 28 km NW Jinghong, vic. An Ma Xi Zhan (NNNR), N 22 o 12, E 100 o 38, 700 m, forest, EKL, 05. iv. 2009 (L. Meng) (1). VIETNAM, Cao Bang Prov., 22 o 34.118 ’ N, 105 o 52.537 ’ E, 1048 m, ex FIT, 12 – 17. iv. 2014 (Cognato, Smith, Pham) (1)	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF3FF985988054DFE077353.taxon	biology_ecology	Biology. Unknown.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF3FF995988004CFAE17454.taxon	distribution	Distribution. Laos, Malaysia (E. and W.). Imported to Japan, but not established. New records. LAOS, C., Kham Mouan Prov., Ban Khoun Ngeun, 18 o 07 ′ N 104 o 29 E, c. 200 m, 24 - 29. iv. 2001 (Pacholátko) (1); Pen [insula] MALAYSIA, Negeri Sembilan, Pasoh Forest Reserve, Tower-B- 32, 11. iv. 1993 (K. Maetô) (1); [E. MALAYSIA: Sarawak], Kinabalu Park PHS, Lower montane, mixed dipterocarp [forest, fogging] Meliaceae sp., 19. iii. 1996 (A. Floren) (1).	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF3FF995988004CFAE17454.taxon	biology_ecology	Biology. The type series was taken from ‘ mersawa’, Anisoptera sp. (Dipterocarpaceae) (Schedl 1970).	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF2FF9959880756FE07717F.taxon	description	The species is very similar to D. nitidus but the eyes are larger and more coarsely faceted, and the sculpture of the elytra differs. On the declivity, interstriae 1 – 4 are irregularly biseriately punctured, not strictly uniseriate. Each puncture bears a long fine seta, so that D. inopinatus has a much more setose appearance than D. nitidus when viewed from above. Striae and interstriae 2 – 3 are distinctly outwardly curved on the declivity, converging again towards the apex, not almost straight as in other species of Dryocoetiops. The granules on the declivital interstriae are minute, far smaller than in D. nitidus.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF2FF9959880756FE07717F.taxon	distribution	Distribution. New Guinea.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF2FF9959880756FE07717F.taxon	biology_ecology	Biology. Unknown.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF2FF96598802A1FBCF7409.taxon	materials_examined	Taxonomy. This species has been the subject of some taxonomic confusion. It is superficially similar to some species of Xyleborini, but clearly has the characteristics of Dryocoetini not Xyleborini, and of Dryocoetiops and not Ozopemon (Wood 1986, contra Wood 1980). It is most closely similar to D. bicolor, and these two species are distinguished from other described species of Dryocoetiops by the broader, bicolored pronotum. Both species lack the sclerolepidia present in all other species. The anterior coxae of both species are contiguous, whereas in other species, they are usually slightly separated. The holotype was deposited in the Philippine Bureau of Science collection which was destroyed in World War II (Baltazar 2001). A paratype in Strohmeyer’s own collection (Strohmeyer, 2011) is also presumed to have been destroyed in World War II. We have examined photographs of the holotype of Ozopemon dipterocarpi (NMNH), a paratype of Xyleborus orbus (NHML) from the Philippines, and a paratype of P. loebli (NMW) from Sri Lanka, and further specimens in NHML and RAB from East and West Malaysia and Vietnam, some of which were directly compared to these paratypes. All represent a single species. In the absence of type material of O. laevis, we accept the synonymy given by the authors cited above and accepted by Wood and Bright (1992).	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF2FF96598802A1FBCF7409.taxon	distribution	Distribution. Indonesia (Sumatera, Kalimantan), Malaysia (E. and W.), Philippines, Sri Lanka, Vietnam. Imported to Japan in timber from Indonesia and the Philippines, but not established.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFF2FF96598802A1FBCF7409.taxon	biology_ecology	Biology. The species was recorded (as O. dipterocarpi) from Dipterocarpus grandiflorus (Dipterocarpaceae) (Hopkins 1915), and as O. laevis from Hopea odorata (Dipterocarpaceae) by Beeson (1961), and from Dipterocarpus sp. timber imported to Japan from Borneo, Sumatra and the Philippines (Ohno 1990; Ohno et al. 1986, 1987). It was recorded (as O. diversicolor) from ‘ lagan’ in the Simaloer islands off Sumatera (Eggers 1923). According to Schedl (1966), ‘ lagan’ refers to Premna odorata (Verbenaceae), but the name more usually refers to various species of Dipterocarpus, and the other host records suggest this is a more likely interpretation.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFDFF9459880100FD217408.taxon	distribution	Distribution. Australia, ‘ Borneo’, Brunei Darussalam, Cambodia, China (Sichuan, Yunnan), India (Bengal, Meghalaya), Indonesia (Java), Japan, Malaysia (E. and W.), Nepal, New Guinea, Singapore, Sri Lanka, Taiwan, Thailand, Timor, Vietnam. New records. BRUNEI, Temburong, nr. K. Belalong Field Stud. Centre, 4 ° 33 ′ N, 115 ° 09 ′ E, 250 m, ex liane, 2. ii. 1992 (R. A. Beaver) (2); as previous except: ex Nephelium sp., 14. ii. 1992 (1); as previous except: ex Shorea sp., 19. ii. 1992 (1); CAMBODIA, Siem Reap, Angkor Park, light trap, 4. xii. 2004 (Danny & Jump) (1); CHINA, Sichuan, Mt. Emei, 600 – 1050 m, 5 – 19. v. 1989 (Lad. Bocák) (1); S. Yunnan, (Xishuangbanna), 23 km NW Jinghong, vic. Na Ban (NNNR), 22 ° 09.49 ′ N 100 ° 38.92 ′, 730 m, second [ary] for [est], EKL, 30. x. 2008 (L. Meng) (1); Yunnan, Gaoligong Mts., 25 ° 22 ′ N 98 ° 49 ′ E, 1500 – 2500 m, 17 – 24. v. 1995 (Vit Kubáň) (1); INDIA, Meghalaya, 3 km E Tura, 25 ° 30 ′ N 90 ° 14 ′ E, 1150 m, 4. v. 1999 (Dombický & Pacholátko) (1); NEPAL, P. Seti, D. Bajheng, way Segu Bagar (29 ° 34 ’ 49 ’’ N 81 ° 13 ’ 44 ’’ E) to before Talkot (29 ° 36 ′ 17 ′′ N 81 ° 17 ′ 54 ′′ E, 1300 - 1400 m, 15. vi. 2009 (A. Kopetz) (1); Kathmandu V., Godavari, 1500 m, 29. iv. 1984 (B. Bhakta) (1); THAILAND, Chiang Mai, Doi Chiang Dao WS, Nature trail, 19 ° 24.278 ′ N, 98 ° 55.311 ′ E, 491 m, pan trap, 5 – 6. x. 2007 (Songkran & Apichart) (2); Chiang Mai, Doi Phahompok NP, Doi Phaluang, 20 ° 1.06 ′ N, 99 ° 9.581 ′ E, 1449 m, Malaise trap, 20 – 27. vii. 2007 (Wongchai P.) (1); Chiang Mai, Doi Pui, 18 ° 50 ′ 23 ′′ N, 98 ° 53 ′ 53 ′′ E, 1200 – 1300 m, EtOH trap, various dates from 11. vi. – 17. ix. 2014, 8. vii. – 11. xi. 2015, 25. v. 2016 (S. Sanguansub et al.) (22); as previous except: ex Mangifera indica, ix. 2014 (2); as previous except: ex Diospyros kaki, 27. vii. 2012 (R. A. Beaver) (4); Nakhon Sri Thammarat, Khao Luang N. P., EtOH trap, 1. ix. 2010 (W. Sittichaya) (1); TIMOR LESTE, Ermera, Mertutu, Railori, S 8.76809, E 125.41182, ex Persea americana shoot, 22. ii. 2019 (T. Popic) (3).	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFDFF9459880100FD217408.taxon	biology_ecology	Biology. The habits of the species have been briefly described above under the genus. Browne (1961) and Kalshoven (1958) (both as D. coffeae) list trees in thirteen different families as hosts, indicating the polyphagy of the species. To that list can be added, based on collections made by the authors: Diospyros kaki (Ebenaceae) in Thailand, and Gomphia serrata (Ochnaceae) in West Malaysia. The species has also been collected from the petiole of large fallen leaves of Campnosperma auriculata (Anacardiaceae), and of three species of Artocarpus (Moraceae) in West Malaysia (Beaver & Browne 1979 as D. coffeae), and from an unidentified liane in Brunei (see above). The diameter of the stems attacked varied from 0.25 – 1.2 cm. The gallery is sometimes started in an abandoned entrance gallery made by another species of beetle (Kalshoven 1958; R. A. Beaver, pers. obs.). Recorded brood sizes vary from 7 to 24 (Browne 1961; Kalshoven 1958). These authors also note an association with twig-boring xyleborine scolytines. Browne (1961) suggests that the species may be of economic importance, noting an occasion when stressed seedlings of Intsia palembanica (Leguminosae) were attacked and killed. However, attacks are normally secondary, following those of other species.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFFFF94598807B2FC4C71D3.taxon	materials_examined	Taxonomy. We have examined two syntypes from Java, and other specimens from W. Malaysia, Indonesia (Sumatera) and Papua New Guinea (NMW), and compared them with specimens in RAB from Brunei, China and E. Malaysia (Sabah). As pointed out by Wood and Bright (1992), Schedl’s (1979) citation of a holotype is invalid, because the species was described from several specimens without a holotype designation. The species is distinguished from D. moestus by its larger size and coarser sculpture (see key). On the basis of the material studied, there is a size gap between the two species. However, the geographical ranges of the two species overlap considerably, and it is possible further studies will show that the species should be added to the list of synonyms of D. moestus.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFFFF94598807B2FC4C71D3.taxon	distribution	Distribution. Brunei, China (Yunnan), Indonesia (Java, Sumatera, West Papua), Malaysia (E. and W.), Papua New Guinea. New records. BRUNEI, Temburong, nr. K. Belalong Field Stud. Centre, 4 ° 33 ′ N, 115 ° 09 ′ E, 300 m, 11. ii. 1992 (R. A. Beaver) (1); CHINA, Yunnan, P. Xishuanbanna, 20 km N Jinhong, Man Dian (NNNR), 22 ° 07 ′ 80 ′′ N 100 ° 40 ′ 08 ′′ E, 230 m, 8. vii. 2008 (A. Weigel) (2); MALAYSIA [East], Sabah, Sipitang, Mendolong, ex Macaranga petiole, 11. v. 1988 (S. Adebratt) (1).	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFFFF94598807B2FC4C71D3.taxon	biology_ecology	Biology. Recorded from Shorea acuminata (Dipterocarpaceae) and Coffea sp. (Rubiaceae) (Schedl 1942 a). The new record from Macaranga sp. (Euphorbiaceae) indicates that, like D. moestus, it can occasionally be found in the petiole of large fallen leaves. Breeding in this habitat has not been observed.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFFFF94598802CDFF79725D.taxon	distribution	Distribution. Malaysia (Sabah), Philippines. New Record. [MALAYSIA, Sabah], Kinabalu Park, Sorinsim, 21. ii. 1997 (A. Floren) (1)	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFFFF94598802CDFF79725D.taxon	biology_ecology	Biology. The specimen from Sabah was obtained by fogging the canopy of Melochia umbellata (Sterculiaceae).	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFFFF955988014FFA867613.taxon	distribution	Distribution. Indonesia (Java), Malaysia (W.), Thailand. The record of the species (as D. kepongi) from Taiwan in Beaver and Liu (2010) should be referred to D. moestus. New Record. THAILAND, Chumphon Pr., EtOH trap in durian plant [atio] n, 1. ii. 2010 (W. Sittichaya) (1).	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFFFF955988014FFA867613.taxon	biology_ecology	Biology. One specimen was collected by RAB from the petiole of a fallen leaf of Artocarpus lanceifolius (Moraceae) in Pulau Pinang, West Malaysia. Because of errors in identification in published papers, it is not certain which host records in the literature actually appertain to this species. Most probably refer to D. moestus.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFEFF95598805C0FE0773AE.taxon	description	Blandford (1895) was in some doubt as to the generic placement of the species, and noted resemblances to species such as Dryocoetes dinoderoides, now a synonym of Dryocoetiops moestus (see above). Presumably because of this statement, Wood and Bright (1992) transferred the species to Dryocoetiops although without giving any indication that this was a new combination. The species does not have the characteristics of Dryocoetiops. The pronotum is more elongate, is not steeply convex anteriorly and lacks a pronounced summit near the middle. The summit is close to the base. The elytral declivity is much less steep than in species of Dryocoetiops. The species belongs in the large genus, Coccotrypes, and is transferred here to that genus. The species has not been recorded again since its original description.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFEFF95598805C0FE0773AE.taxon	distribution	Distribution. Sri Lanka.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFEFF95598805C0FE0773AE.taxon	biology_ecology	Biology. Unknown.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFEFF925988010BFCF47774.taxon	distribution	Distribution. Thailand.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
03FA87EAFFFEFF925988010BFCF47774.taxon	biology_ecology	Biology. Collected only from the petioles of fallen leaves of Manglietia garrettii Craib (Magnoliaceae), incorrectly given as M. garelli in Beaver (1990). The beetles bored into the base of the petiole. The species appears to be monogamous. Immature stages were not found in the galleries, and it is possible that leafstalks are used primarily as a ‘ refuge’ in the absence of other suitable breeding material.	en	Beaver, Roger A., Smith, Sarah M., Sanguansub, Sunisa (2019): A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4712 (2): 236-250, DOI: 10.11646/zootaxa.4712.2.4
