taxonID	type	description	language	source
03FADC67C001FFB35F92FEBFFB010BD6.taxon	description	Redescription. Dorsal habitus (Figs. 1 A – C): Major male. Length 7.7 – 9.5 mm. Dorsal surface of the body almost glabrous, with only short dark brown setae on the clypeus, antero-lateral parts of pronotum and elytral striae. Color shining black, elytra slightly sericeous with a violet-blue sheen. Head and pronotum with a slight greenish sheen. Head: Clypeus transverse, concave, subsinuate at the apex. Central and posterior parts of clypeus with wide, scattered and confluent punctures. Genae wide, with the basal third weakly concave. Genal sutures fine. Clypeal carina feebly indicated in the center, smooth and evanescent on the sides; in dorsal view with a trapezoid shape, open posteriorly. Central part of the head distinctly convex. Frontal carina quite evenly elevated, with double anterior convexity, lying at the level of the eyes and completely effaced laterally. Surface of frons with fine punctures. Pronotum: With fine basal margin that is clearly visible medially. Lateral margin sub-linear between the anterior and the intermediate angle. Pronotal prominence pronounced and apically rounded, intermediate tubercles obsolete, posterior tubercles located near the middle of the pronotum. Pronotal disc with well-defined and irregular main punctures, with non-uniform distribution, and with more abundant, irregular, smaller intervening punctures, non-uniformly distributed. Finer microsculpturing not evident. Elytra: Striae wide, well impressed, with irregular and fine punctures. Elytral interstriae weakly convex, surface sculpturing composed of irregularly aligned small granules, and very fine setae-bearing punctures. Microsculpture quite conspicuous, uniformly reticulate. Legs: Protibia very feebly arched, slightly dilated towards the apex. External margin of protibia denticulate; the basal tooth prolonged toward the base as a smooth lamina. Apical spur curved medially. Pygidium: Punctation quite dense and confluent, forming irregular transversely arranged wrinkles. Female. Major female (Figs. 1 D, E). Length 7.2 – 9.1 mm. Similar to the male except for the clypeus being sub-trapezoidal, sinuate apically; clypeal carina distinctly developed; frontal carina strongly developed, protruded laterally as two vertical tubercles, flattened antero-posteriorly; pronotum with the posterior tubercles located more anteriorly than those of the male. Genitalia. Endophallite copulatrix (EC) (Fig. 2 A): Left lobe (LL) rounded, slightly widened. Inner lobe of the free margin of the left lobe (InL) short and rounded. Medial keel (MK) (or keel of basal margin of endophallite copulatrix) elongate and slightly curved. Right lobe (RL) much longer than the left lobe. Median keel of right lobe absent. Internal keel of right lobe (InK) forming a rounded and sinuate tooth. Apex of the right lobe (RLA) oval. Additional inferior endophallite (AIE) (Fig. 2 B): Subrectangular, relatively simple in appearance, without intricate folds, without development of the part closest to the margin where the lamellar spiny belt is inserted. Vagina (Fig. 2 C): Expanded portion of the vagina (VEP) developed, membranous, with semicircular apical margin. Ventral sclerotization of the vagina (VVS) with the centralsuperior region sinuate and the lateral-inferior regions wide and outwardly directed. Tubular tract of the infundibulum (ITT) elongate, narrow, straight or nearly so. Median tract of the infundibulum (IMT) weakly developed. Caudal tract of the infundibulum (ICT) well developed. Spermatheca (Sp) (Fig. 2 D): U-shaped, with the flex-de-sclerotized area elongated and narrowed, with a spermathecal gland inserted in the basal portion of the flex-de-sclerotized area. Spermathecal gland (SpG) well developed, smaller than spermatheca, with apex rounded and globose.	en	Rivera-Gasperín, Sara Lariza, Escobar-Hernández, Fernando, Favila, Mario E. (2021): An Update on the Feeding Preferences of the Dung Beetle Onthophagus moroni Zunino and Halffter (Coleoptera: Scarabaeidae) from the Cuetzalan Caves, Puebla, Mexico. The Coleopterists Bulletin 75 (1): 247-255, DOI: 10.1649/0010-065X-75.1.247, URL: http://dx.doi.org/10.1649/0010-065x-75.1.247
03FADC67C001FFB35F92FEBFFB010BD6.taxon	materials_examined	Type Locality. MEXICO: State of Puebla, Cuetzalan del Progreso: interior of the Tasalolpan cave. Material Examined. 21 ♂♂, 12 ♀♀: Verbatim label data: MEXICO: Estado de Puebla, Cuetzalan, Cueva de Tasalolpan, 18 - XI- 2018, 907 m., Fer. Escobar-Hdez and Rivera-Gasper´ın. leg., 20 ° 01 ′ 11.6 ′′ N 97 ° 32 ′ 31.1 ′′ W. Cloud forest. Collected directly under droppings of rodents, 70 m. inside the cave (Fig. 3 A). 6 ♂♂: Verbatim label data: MEXICO: Estado de Puebla, Cuetzalan, Gruta de Atepolihui, Cueva El Nido del Murciélago. 21 - XII- 2014, 1,030 m., Fer. Escobar-Hdez. leg., 20 ° 00 ′ 41.42 ′′ N 97 ° 32 ′ 32.44 ′′ W. Cloud forest. Collected directly in bat guano, 60 m. inside the cave. Paratype. 1 ♂ (Fig. 3 B): Verbatim label data: MEXICO: Edo. de Puebla, Cuetzalan. 11 - VIII- 1987, 1,520 m., G. Halffter, M. Zunino, C. Deloya, leg., Cueva de Tasalolpan. NOTE: This material, not yet registered in the collection of Dr. G. Halffter, was also examined; 1 ♂, 1 ♀: MEXICO: Puebla, Cuetzalan, C. Tasalolpan, North entrance, “ Zone D ”. 21 - XI- 1986 on rock, L. Arellano leg. 1 ♂: MEXICO: Puebla, Cuetzalan, C. Tasalolpan, “ Zone D ”. 19 - III- 1987, in guano, J. Monterrubio leg. 1 ♀: MEXICO: Puebla, Cuetzalan, C. Tasalolpan, North entrance, “ tunnel C ”. 21 - XI- 1992 in murcielaguina with soil, Montes de Oca leg. Affinities. There are five species within the Onthophagus brevifrons species complex: O. cavernicollis, O. subtropicus, O. brevifrons, O. cuevensis and O. moroni (Halffter et al. 2019). Taking into account the study of genital morphology within the O. brevifrons species complex, Zunino and Halffter (1988 b), consider O moroni as the sister species to O. cuevensis. Feeding. Onthophagus moroni is closely associated with the latrines of Peters’ climbing rat, Tylomys nudicaudus (Peters, 1866) (Rodentia: Cricetidae). A large number of live adults were observed interacting above and below the rat latrine; apparently, they feed and nest on accumulated rodent droppings of the rats (Ceballos 2014; Espinoza-Medinilla 2014; Morales and Vázquez 1987; Ram´ırez-Pulido et al. 2005) (Figs. 3 C – E).	en	Rivera-Gasperín, Sara Lariza, Escobar-Hernández, Fernando, Favila, Mario E. (2021): An Update on the Feeding Preferences of the Dung Beetle Onthophagus moroni Zunino and Halffter (Coleoptera: Scarabaeidae) from the Cuetzalan Caves, Puebla, Mexico. The Coleopterists Bulletin 75 (1): 247-255, DOI: 10.1649/0010-065X-75.1.247, URL: http://dx.doi.org/10.1649/0010-065x-75.1.247
