identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FA3948FFF7FFC2A5CFF9C243C5AC40.text	03FA3948FFF7FFC2A5CFF9C243C5AC40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades Herrich-Schaffer 1855	<div><p>Systematics of Abantiades</p><p>Molecular analyses</p><p>The Maximum-Likelihood phylogram (Fig. 1) reveals a well supported Abantiades species groups (compared to our six outgroup taxa). The “dark A. obscura group” comprises four clades, and is sister to Abantiades obscura sensu stricto (Fig. 1). Although this is only based on the single mitochondrial COI gene, it does delineate the four new species that are also defined by distinct morphological characters (see Taxonomy section below). We found close intraspecific sequence divergences for several instances where we obtained multiple individuals within species (Supplementary Table S2). For example, the greatest intraspecific sequence divergence varied between 0.3-2.2%, indicating variation that we would expect within species included here (Supplementary Table S2). These distances also correspond directly to those reported elsewhere for Lepidoptera that indicate a maximum of between 2-3% within species comparisons using the same COI gene (e.g. Hausmann et al. 2011; Grund et al. 2019; Beaver et al. 2020; but see Simonsen et al. 2019: 796−797 for some potential caveats).</p><p>FIGURE 1. Maximum Likelihood (ML) phylogram (using TrN) based on mitochondrial DNA (COI) dataset from 68 Abantiades sequences, including the four new Abantiades species. The tree with the highest log likelihood (-2,953.03) is shown with branch lengths measured in the number of substitutions per site. Bootstrap values from 1,000 pseudoreplicates greater than 50% are shown at each node. Sequences generated from this study have been combined with GenBank sequences only when the id’s could be confirmed morphologically, and a * indicates where we have corrected a GenBank id (see Table S1).</p><p>Sequence divergence comparisons between all Abantiades species compared to our outgroup taxa were between 8.2-12.0% (Supplementary Table S2). Examining between Abantiades species in the present study reveal interspecific sequence divergences of up to 9.9% (i.e. between A. centralia sp. nov. and A. labyrinthicus (Donovan, 1805) . However, we also observed several that are on the lower end of what has been reported for related groups (see above), for example between A. hutchinsoni sp. nov. and A. zonatriticum sp. nov. (1.2%), and between A. hutchinsoni sp. nov. and A. kayi sp. nov. (1.5%) (Supplementary Table S2). Although some levels of sequence divergence are low (less than 2%) our COI phylogram for Abantiades species (Fig. 1) indicates that these are well supported species (bootstrap values&gt; 76%), such divergence is corroborated by consistent morphological characters (see Taxonomy section below).</p></div>	https://treatment.plazi.org/id/03FA3948FFF7FFC2A5CFF9C243C5AC40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Four new tri-forked species of the Australian genus Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) from the “ dark obscura clade ”. Zootaxa 4801 (1): 115-141, DOI: 10.11646/zootaxa.4801.1.5
03FA3948FFF5FFC2A5CFFDE14743AE08.text	03FA3948FFF5FFC2A5CFFDE14743AE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades Herrich-Schaffer 1855	<div><p>Genus Abantiades Herrich-Schäffer, [1856]</p><p>Diagnosis</p><p>Simonsen (2018) produced a new diagnosis for the genus: “A patch of elongate scales at the forewing base and the ‘sensory tubercles’ on S2 are both unique and diagnostic for Abantiades .” He also noted that they have hepialine venation (Dumbleton 1966, Dugdale 1994), and mono-pectinate antennae that lack scales on the flagellum.</p></div>	https://treatment.plazi.org/id/03FA3948FFF5FFC2A5CFFDE14743AE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Four new tri-forked species of the Australian genus Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) from the “ dark obscura clade ”. Zootaxa 4801 (1): 115-141, DOI: 10.11646/zootaxa.4801.1.5
03FA3948FFF5FFC2A5CFFCA540F7AE99.text	03FA3948FFF5FFC2A5CFFCA540F7AE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades kayi Moore & Beaver & Velasco-Castrillón & Stevens 2020	<div><p>1. Abantiades kayi Moore &amp; Beaver sp. nov.</p><p>Figs 2 (a) and (b), 7(a), 8(a), 9(a), 10(a), 11(a) and (b).</p></div>	https://treatment.plazi.org/id/03FA3948FFF5FFC2A5CFFCA540F7AE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Four new tri-forked species of the Australian genus Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) from the “ dark obscura clade ”. Zootaxa 4801 (1): 115-141, DOI: 10.11646/zootaxa.4801.1.5
03FA3948FFF1FFC6A5CFF95E40FDABE0.text	03FA3948FFF1FFC6A5CFF95E40FDABE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades zonatriticum Moore & Beaver & Velasco-Castrillón & Stevens 2020	<div><p>2. Abantiades zonatriticum Moore &amp; Beaver sp. nov.</p><p>Figs 3 (a) and (b), 7(b), 8(b), 9(b), 10(b), 11(c) and (d).</p></div>	https://treatment.plazi.org/id/03FA3948FFF1FFC6A5CFF95E40FDABE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Four new tri-forked species of the Australian genus Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) from the “ dark obscura clade ”. Zootaxa 4801 (1): 115-141, DOI: 10.11646/zootaxa.4801.1.5
03FA3948FFFBFFCCA5CFFD6A43E0AFD4.text	03FA3948FFFBFFCCA5CFFD6A43E0AFD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades hutchinsoni Moore & Beaver & Velasco-Castrillón & Stevens 2020	<div><p>3. Abantiades hutchinsoni Moore &amp; Beaver sp. nov.</p><p>Figs 4, 7 (d), 8(d), 9(d), 10(d).</p></div>	https://treatment.plazi.org/id/03FA3948FFFBFFCCA5CFFD6A43E0AFD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Four new tri-forked species of the Australian genus Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) from the “ dark obscura clade ”. Zootaxa 4801 (1): 115-141, DOI: 10.11646/zootaxa.4801.1.5
03FA3948FFF9FFCEA5CFF85A40EAAAE4.text	03FA3948FFF9FFCEA5CFF85A40EAAAE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades centralia Moore & Beaver & Velasco-Castrillón & Stevens 2020	<div><p>4. Abantiades centralia Moore &amp; Beaver sp. nov.</p><p>Figs 5 (a) and (b), 7(e), 8(e), 9(e), 10(e), 11(e) and (f).</p></div>	https://treatment.plazi.org/id/03FA3948FFF9FFCEA5CFF85A40EAAAE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Four new tri-forked species of the Australian genus Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) from the “ dark obscura clade ”. Zootaxa 4801 (1): 115-141, DOI: 10.11646/zootaxa.4801.1.5
