taxonID	type	description	language	source
03FB87DBFF88A31119F9FD54FC48FB49.taxon	discussion	Comments. Acanthomegabunus sibiricus is the only species previously known from the Katunsky Biosphere Reserve. The species was recorded from the vicinity of Srednee Multinskoe Lake (erroneously named Srednee Kuktinskoe Lake) (Tchemeris 2015). The lake is actually located outside the reserve, close to its northern border where the reserve’s ranger station is situated, although this record requires confirmation because it is based on a single female. A closely related species, A. altaicus Tchemeris, 2015, may also occur there — its type locality lies close to the Katunsky Reserve (East Kazakhstan Area, Katon-Karagaiskii Districts). Habitat. Mountain moss-stony tundra, moss-pebble riverbanks, Abies forest, Anemone meadows, stony debris, dark coniferous forests, and aspen-fir forests (Tsurusaki et al. 2000 b; Trilikauskas 2016, 2017).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF88A31119F9FD54FC48FB49.taxon	distribution	Distribution. The mountains of South Siberia, Russia: the southern part of Krasnoyarsk Territory, Altai Territory, and Altai Republic, Kemerovo Area, Khakassia, and Tuva (Tchemeris 2015).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF88A31119F9FB3DFC84F91F.taxon	description	Figure 1 Farzalieva & Esyunin 2000: 193, figs 67 – 73. Tchemeris et al. 1999: 189, figs 1 – 7. Snegovaya & Cokendolpher 2021: 338, figs 28 – 29.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF88A31119F9FB3DFC84F91F.taxon	materials_examined	Material examined. RUSSIA: Altai Republic: Ust’ - Koksa District: 1 ♂ (ISEA O. 001.0250), Katunsky Biosphere Reserve, upper reaches of the Katun’ River, left bank of Zaichikha River, near mouth, [49 ° 37 ’ 08.3 ” N, 85 ° 43 ’ 03.8 ” E], 1302 m a. s. l., 15 – 26. VIII. 2020; 1 ♀ (ISEA O. 001.0251), Katunsky Biosphere Reserve, c. 4 km NE of the Zaichikha ranger station, [49 ° 36 ’ 25.6 ” N, 85 ° 44 ’ 13.4 ” E], 1511 m a. s. l., 16. VIII. 2020. Habitat. Dry valley meadow and synanthropic habitats (Tchemeris et al. 1999); scree, big stones near Salix, and grass thickets on small riverbanks (in Katunsky Biosphere Reserve).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF88A31119F9FB3DFC84F91F.taxon	distribution	Distribution. Finland to Korea (Tchemeris et al. 1999).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF88A31119F9FEB4FF50FDE3.taxon	discussion	Only four species of harvestmen, except for the five species listed below, Homolophus asiaticus Gritsenko, 1979, Opilio parietinus (De Geer, 1778), Homolophus pallens (Kulczyński, 1901), and Scleropilio insolens (Simon, 1895), are known to date from Russian part of Altai-Sayan Mountain region (Tchemeris et al. 1999; 2015; Tsurusaki et al. 2000 a).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF88A31719F9F8C8FE63FEC3.taxon	description	Figures 2 – 15 Gritsenko, 1979: 266, figs 7 – 13	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF88A31719F9F8C8FE63FEC3.taxon	materials_examined	Material examined. RUSSIA: Altai Republic: Ust’ - Koksa District: 1 ♂ (ISEA O. 001.0288), Katunsky Biosphere Reserve, NW shore of Taimenye Lake, [49 ° 49 ’ 35.9 ” N, 85 ° 48 ’ 25.5 ” E], 1745 – 1760 m a. s. l., scree, 8. VIII. 2019; 4 ♂, 1 ♀ (PCLT), Katunsky Biosphere Reserve, head of the Ozernaya River, shore of Taimenye Lake, near the ranger station, [49 ° 48 ’ 40.9 – 42.9 ” N, 85 ° 47 ’ 57.2 – 57.6 ” E], 1509 m a. s. l., 10. VIII. 2019; 2 ♂ (PCLT), upper reaches of the Katun’ River, mouth of Ozernaya River, [49 ° 40 ’ 16.7 ” N, 85 ° 41 ’ 49.6 ” E], 1268 m a. s. l., 14. VIII. 2019.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF88A31719F9F8C8FE63FEC3.taxon	discussion	Comments. The species was described from the Ivanovsky Mountain Range in north-eastern Kazakhstan (Gritsenko 1979) and has not been found since; this is the first record for the species from Russia. The genus Liropilio is morphologically similar to Zacheus C. L. Koch but differs by the presence of a tubercle with denticles on the basal segment of the pedipalps (Figs 3 – 6) and an alate part on the penis, apically (Figs 11 – 15). Palpal femur, trochanter, and tibia of this species slightly thickened latero-apically, densely covered with short, stout setae. Palpal femur bordered dorso-apically, with two rows of teeth ventro-retrolaterally, tarsus covered with setae (Figs 3 – 4, 7 – 10). Habitat. Scree, edges of mixed forests (Pinus silvestris, Abies sibirica, Betula pendula), stony riverbanks overgrown by moss and grass. The habitat of the type locality is unknown (Gritsenko 1979).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF88A31719F9F8C8FE63FEC3.taxon	distribution	Distribution. Eastern Kazakhstan (Ivanovsky Mountain Range, West Altai) and Altai Republic, Russia (Katunsky Mountain Range).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF8EA31619F9F8C7FD5FFDBF.taxon	description	Tchemeris et al. 1999: 194, figs 15 – 23.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF8EA31619F9F8C7FD5FFDBF.taxon	materials_examined	Material examined. RUSSIA: Altai Republic: Ust’ - Koksa District: 4 ♂, 2 ♀ (PCLT), Katunsky Biosphere Reserve, bank of Taimenye Lake near Soloukha River mouth, [49 ° 50 ’ 11.6 ” N, 85 ° 51 ’ 07.7 ” E], 1534 m a. s. l., 7. VIII. 2019; 1 ♂, 1 ♀ (PCLT), Katunsky Biosphere Reserve, head of the Ozernaya River, shore of Taimenye Lake, near the ranger station, [49 ° 48 ’ 40.9 – 42.9 ” N, 85 ° 47 ’ 57.2 – 57.6 ” E], 1509 m a. s. l., 10. VIII. 2019; 1 ♂, 3 ♀ (PCLT), Katunsky Biosphere Reserve vicinities of Taimenye Lake, middle reaches of Kharyuzovka River, [49 ° 48 ’ 53.6 ” N, 85 ° 47 ’ 42.0 ” E], 1652 – 1660 m a. s. l., 11. VIII. 2019; 6 ♀ (PCLT), SE shore of Taimenye Lake, [49 ° 48 ’ 08.2 ” N, 85 ° 48 ’ 00.5 ” E], 1551 m a. s. l., 7 – 13. VIII. 2019; 13 ♂, 5 ♀ (ISEA O. 001.0253), 11 ♂, 11 ♀ (PCLT), Katunsky Biosphere Reserve, upper reaches of the Katun’ River, left bank of Zaichikha River, near its mouth, [49 ° 37 ’ 08.3 ” N, 85 ° 43 ’ 03.8 ” E], 1302 m a. s. l., 15 – 26. VIII. 2020; 1 ♂ (PCLT), Katunsky Biosphere Reserve, 4 km NE of the Zaichikha ranger station, [49 ° 36 ’ 25.6 ” N, 85 ° 44 ’ 13.4 ” E], 1511 m a. s. l., 16.08.2020. Habitat. Mesophytic and moist meadows and clearings in birch and mixed forests, sloping mesophytic steppes, in sparse mountain larch forest, as well as agricultural areas (Tchemeris et al. 1999); blueberry-green moss and grass-green moss spruce forests and spruce woodland, Abies sibirica forest, subalpine grass meadow, thickets of sedge near lakes and streams (in Katunsky Biosphere Reserve).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF8EA31619F9F8C7FD5FFDBF.taxon	distribution	Distribution. Holarctic (Tchemeris et al. 1999).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF8CA31419F9F939FB93FF26.taxon	description	Tchemeris et al. 1999: 196, figs 24 – 31.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF8CA31419F9F939FB93FF26.taxon	materials_examined	Material examined. RUSSIA: Altai Republic: Ust’ - Koksa District: 1 ♂, 4 ♀ (ISEA O. 001.0254), 11 ♂, 20 ♀ (PCLT), Katunsky Biosphere Reserve, upper reaches of the Katun’ River, left bank of Zaichikha River near its mouth, [49 ° 37 ’ 08.3 ” N, 85 ° 43 ’ 03.8 ” E], 1302 m a. s. l., 15 – 26. VIII. 2020. Habitats. Mesophytic meadows and clearings in birch or mixed forests (Tchemeris et al. 1999); grass-green moss spruce forests (in Katunsky Biosphere Reserve).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF8CA31419F9F939FB93FF26.taxon	distribution	Distribution. Europe to Siberia (up to Yenisei River) (Tchemeris et al. 1999).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF8DA31419F9FE90FC07F87F.taxon	description	Tchemeris et al. 1999: 198, figs 49 – 58.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF8DA31419F9FE90FC07F87F.taxon	materials_examined	Material examined. RUSSIA: Altai Republic: Ust’ - Koksa District: 3 ♀ (PCLT), Katunsky Biosphere Reserve, head of the Ozernaya River, shore of Taimenye Lake, near the ranger station, [49 ° 48 ’ 40.9 – 42.9 ” N, 85 ° 47 ’ 57.2 – 57.6 ” E], 1509 m a. s. l., 10. VIII. 2019; 1 ♀ (PCLT), Katunsky Biosphere Reserve, vicinities of the Taimenye Lake, middle reaches of Kharyuzovka River, [49 ° 48 ’ 53.6 ” N, 85 ° 47 ’ 42.0 ” E], 1652 – 1660 m a. s. l., 11. VIII. 2019; 1 ♀ (PCLT), SE shore of Taimenye Lake, [49 ° 48 ’ 08.2 ” N, 85 ° 48 ’ 00.5 ” E], 1551 m a. s. l., 7 – 13. VIII. 2019; 2 ♀ (PCLT), Katunsky Biosphere Reserve, upper reaches of the Katun’ River, left bank of Zaichikha River near its mouth, [49 ° 37 ’ 08.3 ” N, 85 ° 43 ’ 03.8 ” E], 1302 m a. s. l., 15 – 26. VIII. 2020. Habitat. Moist and mesophytic deciduous and mixed forests and anthropogenic habitats (Tchemeris et al. 1999); blueberry-green moss and grass-green moss spruce forests (in Katunsky Biosphere Reserve).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF8DA31419F9FE90FC07F87F.taxon	distribution	Distribution. Holarctic (Tchemeris et al. 1999). MAP 2. Distribution of Sabacon sergeidedicatum and Sabacon zateevi sp. nov.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF82A31B19F9F8FCFBE7F879.taxon	discussion	To date, only two species of Sabaconidae, Sabacon crassipalpe (L. Koch, 1879) and S. sergeidedicatum Martens, 1989, are known from the Altai-Sayan Mountain region (Chemeris & Logunov 2001).	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF83A31C19F9FF4DFC09FB7F.taxon	description	Figures 18 – 21, 25 – 27, 31 – 33, 41 – 44, 50, 56, 64 – 65, 69 – 71, Map 2	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF83A31C19F9FF4DFC09FB7F.taxon	materials_examined	Type. Holotype ♂ (ISEA O. 001.0280), Russia, Altai Republic, Ust’ - Koksa District, upper reaches of Katun’ River, Katunsky Biosphere Reserve, Zaichikha River mouth, [49 ° 37 ’ 08.3 ” N, 85 ° 43 ’ 03.8 ” E], 1302 m a. s. l., 16 – 26. VIII. 2020, L. A. Trilikauskas. Paratypes. RUSSIA: Altai Republic: Ust’ - Koksa District: 1 ♀ (ISEA O. 001.0281), Katunsky Biosphere Reserve, head of Ozernaya River, vicinities of Taimenye Lake, middle reaches of Kharyuzovka River, [49 ° 48 ’ 53.6 ” N, 85 ° 47 ’ 42.0 ” E], 1652 – 1660 m a. s. l., 11. VIII. 2019, L. A. Trilikauskas; 1 ♂, 2 ♀ (MMUE G 7650.1), 1 ♀ (ISEA O. 001.0282), adjacent territory of Katunsky Biosphere Reserve, SE shore of Taimenye Lake, [49 ° 48 ’ 08.2 ” N, 85 ° 48 ’ 00.5 ” E], 1551 m a. s. l., 7 – 13. VIII. 2019, L. A. Trilikauskas; 1 ♂ (ISEA O. 001.0283), Katunsky Biosphere Reserve, upper reaches of Katun’ River, Katunsky State Nature Biosphere Reserve, Zaichikha River mouth, [49 ° 37 ’ 08.3 ” N, 85 ° 43 ’ 03.8 ” E], 1302 m a. s. l., 19. VIII. 2020, L. A. Trilikauskas; 1 ♂ (ISEA O. 001.0285), 1 ♀ (ISEA O. 001.0284), same but 17. VIII. 2020, L. A. Trilikauskas; 3 ♂ (ISEA O. 001.0286), 32 km SSW of Ust’ - Koksa, upper reaches of Petrushkina River, [50 ° 03 ’ N, 85 ° 21 - 22 ’ E], 1400 - 1500 m a. s. l., forest, 5 - 6. VI. 2005, R. Y. Dudko; 1 ♂ (ISEA O. 001.0287), 25 km SSW of Ust’ - Koksa, NE slope of Kabanukha Mt., [50 ° 05 ’ N, 85 ° 24 ’ E], 1850 - 1950 m a. s. l., forest, 28 - 29. V. 2005, R. Y. Dudko. Comparative material: Sabacon sergeidedicatum Martens, 1989 (Figs 16 – 17, 22 – 24, 28 – 30, 34 – 40, 45 – 49, 51 – 55, 57 – 63, 66 – 68, Map 2): RUSSIA: Novosibirsk Area: Bolotnoye District: 4 ♂, 2 ♀ (ISEA O. 001.0258) near Bolshaya Chyornaya, [55 ° 57 ’ 49 ” N, 84 ° 25 ’ 24 ” E], Pinus sibirica forest, pitfall traps, 16 – 31. V. 2015, L. A. Trilikauskas. Tomsk Area: Tomsk District: 9 ♂, 7 ♀ (ISEA O. 001.0274), Anikino vicinities, mixed forest, near spring, 20 – 30. VI. 2000 (A. N. Tchemeris); 4 ♂, 1 ♀ (ISEA O. 001.0275), same but 25. VI. 1994, S. Y. Rakov; 1 ♂ (ISEA O. 001.0276), same but 10. VI. 1994, S. Y. Rakov; 4 ♂, 1 ♀ (ISEA O. 001.0050), same but 21 – 23. VI. 1995, S. Y. Rakov. Altai Territory: Charyshskoe District: 1 ♂ (ISEA O. 001.0259), Bashchelak Mt. Range, ca. 30 km NWW of Sentelek, Zagrikha Mt., [51 ° 15 ’ N, 84 ° 11 ’ E], 1700 m a. s. l., sparse forest, 28. VI. 2000, G. N. Azarkina; 1 ♂ (ISEA O. 001.0260), 10 km S of Tigirek, upper reaches of Malyi Tigirek River, [ca. 51 ° 03 ’ N, 82 ° 59 ’ E], 1000 – 1400 m a. s. l., stone stream (kurum), 19 – 21. VI. 2000, A. N. Tchemeris; 1 ♂ (ISEA O. 001.0261), Malyi Tigirek River valley, [ca. 51 ° 06 ’ N, 83 ° 02 ’ E], 500 – 1000 m a. s. l., bank of river, on stone covered with moss, 16 – 18. VIII. 2000, R. Y. Dudko; 1 ♂, 2 ♀ (ISEA O. 001.0262), Korgon Mt. Range, Gorelyi Korgon River valley, [51 ° 00.01 ’ N, 83 ° 48.49 ’ E], 1. VII. 2011, R. Y. Dudko. Altai Republic: Choya District: 1 ♂ (ISEA O. 001.0257), near Uskuch Vil., [52 ° 02 ’ 07 ” N, 86 ° 54 ’ 30 ” E], detritus scree, 30. IV. – 02. V. 2011, L. A. Trilikauskas. Turochak District: 1 ♀ (ISEA O. 001.0002), ca. 4 km SW of Kebezen’, [51 ° 53 ’ 11.5 ” N, 87 ° 02 ’ 55.9 ” E], 450 m a. s. l., Pinus-Abies- birch forest, pitfall traps, 21. VI. – 6. VII. 2004, S. B. Ivanov; 3 ♂, 2 ♀ (ISEA O. 001.0001), ca. 2 km SEE of Kebezen’, [51 ° 54 ’ 06.1 ” N, 87 ° 07 ’ 54.0 ” E], 500 m a. s. l., Pinus forest, pitfall traps, 21. V. – 6. VI. 2004, S. B. Ivanov; 1 ♂ (ISEA O. 001.0263), ca. 2 km SW of Kebezen’, [51 ° 53 ’ 11.5 ” N, 87 ° 02 ’ 55.9 ” E], Abies - aspen forest, pitfall traps, 5 – 22. VIII. 2004, S. B. Ivanov; 2 ♂, 1 ♀ (ISEA O. 001.0026), ca. 4 km NNW of Kebezen’, [51 ° 57 ’ 00.8 ” N, 87 ° 04 ’ 37.3 ” E], Pinus - birch forest, pitfall traps, 7 – 19. VI. 2004, S. B. Ivanov; 1 ♀ (ISEA O. 001.0030), 2 ♂ (ISEA O. 001.0029), 4 ♂, 1 ♀ (ISEA O. 001.0018), near Obogo, Valley of Pyzha River, [51 ° 31.355 ’ N, 87 ° 17.737 ’ E], 980 m a. s. l., birch-aspen valley forest, pitfall traps, 20. VI. – 6. VII. 2003, 6 – 23. VII. 2003, S. B. Ivanov; 18 ♂, 9 ♀ (ISEA O. 001.0264), 2 ♂, 2 ♀ (ISEA O. 001.0051), same but Picea - birch valley forest, 17. V. – 26. VII. 2003, 1 – 20. VI. 2003, S. B. Ivanov; 1 ♂ (ISEA O. 001.0009), 1 ♂ (ISEA O. 001.0017), ca. 1.2 km E of Obogo, [51 ° 31.259 ’ N, 87 ° 18.458 ’ E], Abies-Pinus sibirica forest, logging area, 950 m a. s. l., pitfall traps, 1 – 20. VI. 2003, 6 – 23. VII. 2003, S. B. Ivanov; 1 ♀ (ISEA O. 001.0005), ca. 5.5 km SW of Obogo, [51 ° 30.048 ’ N, 87 ° 13.216 ’ E], Abies-Pinus sibirica forest, 860 m a. s. l., pitfall traps, 23. VI. – 14. VII. 2004, S. B. Ivanov; 6 ♂, 2 ♀ (ISEA O. 001.0265), ca. 8.8 NEE of Obogo, [51 ° 33.072 ’ N, 87 ° 24.678 ’ E], 1270 m a. s. l., Picea-Abies- Pinus sibirica forest, pitfall traps, 21. VI. – 7. VII. 2003, S. B. Ivanov; 2 ♂, 1 ♀ (ISEA O. 001.0024), ca. 15 km NE of Obogo, [51 ° 36.61 ’ N, 87 ° 26.395 ’ E], Betula rotundifolia tundra, 1870 m a. s. l., pitfall traps, 21. VI. – 8. VII. 2003, S. B. Ivanov; 1 ♂, 1 ♀ (ISEA O. 001.0031), head of Archa Mt., [51 ° 31.595 ’ N, 87 ° 26.316 ’ E], 1840 m a. s. l., sparse forest with meadows and Betula rotundifolia, pitfall traps, 19. V. – 2. VI. 2003, S. B. Ivanov; 2 ♀ (ISEA O. 001.0266), same but Pinus sibirica forest, [51 ° 32.865 ’ N, 87 ° 25.742 ’ E], 1600 m a. s. l., 16. V. – 2. VI. 2003, S. B. Ivanov; 3 ♂, 1 ♀ (ISEA O. 001.0028), ca. 14 km SW of Iogach, near Suuchak, [51 ° 41 ’ 21.6 ” N, 87 ° 07 ’ 13.6 ” E], birch-aspen forest, old fire site, pitfall traps, 11 – 22. VI. 2004, S. B. Ivanov. Shebalino District: 1 ♀ (ISEA O. 001.0267), Seminsky Mt. Range, near Kamlak, Kamyshlinskoe Plateau, Dezertirnaya cave, [ca. 51 ° 42 ’ N, 85 ° 34 ’ E], 25. VI. 2015, I. S. Turbanov. Ulagan District: 2 ♂, 1 ♀ (ISEA O. 001.0098), Altai Nature Reserve, right bank of Bayas River, [ca. 51 ° 20 ’ N, 87 ° 52 ’ E], Pinus sibirica forest, 20 – 23.08.2011, S. M. Ponomareva; 1 ♂ (ISEA O. 001.0105), Altai Nature Reserve, Kyga River basin, Malaya Kolyushta Mt., [ca. 51 ° 16 ’ N, 87 ° 58 ’ E], mountain stony tundra, 25 – 28. VI. 2011, S. M. Ponomareva; 1 ♂ (ISEA O. 001.0092), 2 km NW from Chiri, [ca. 51 ° 22 ’ N, 87 ° 49 ’ E], pine forest, 22. VI. – 6. VII. 2011, S. M. Ponomareva. RUSSIA: Kemerovo Area: 2 ♂, 2 ♀ (ISEA O. 001.0269), Mezhdurechensk vicinity, right bank of Usa River, 3 km from mouth, [ca. 53 ° 42 ’ N, 88 ° 02 ’ E], 23. IX. 1997, R. Y. Dudko, I. I. Lyubechanskii. Tashtagol District: 7 ♂, 9 ♀ (ISEA O. 001.0255), Shorsky National Park, near Verkhniy Taimet, Biyskaya Griva Mt. Range, [52 ° 26 ’ 18.2 ” N, 88 ° 17 ’ 42.5 ” E], Pinus sibirica forest, pitfall traps, 19. IX. – 11. X. 2016, L. A. Trilikauskas; 1 ♀ (ISEA O. 001.0161) same but 22. VIII. 2013, L. A. Trilikauskas; 5 ♂ 6 ♀ (ISEA O. 001.0152) same but 7 – 15. VI. 2013, L. A. Trilikauskas; 79 ♂, 64 ♀ (ISEA O. 001.0256), Shorsky National Park, near Verkhniy Taimet, Populus tremula- Abies sibirica forest (chernevaya taiga), [52 ° 26.309 ’ N, 88 ° 17.695 ’ E], 9 – 19. VI. 2016, L. A. Trilikauskas; 3 ♂ 3 ♀ (ISEA O. 001.0118) same but Pinus sibirica forest, 8 – 18. VI. 2012, L. A. Trilikauskas; 5 ♂, 5 ♀ (ISEA O. 001.0117), Shorsky National Park, Mrassu River valley, mouth of Kubansu River, [ca. 52 ° 28 ’ N, 88 ° 41 ’ E], dark coniferous forest, 22 – 26. VII. 2012, L. A. Trilikauskas; 2 ♂ 3 ♀ (ISEA O. 001.0052), 6 ♂, 4 ♀ (ISEA O. 001.0268), near “ Mednaya ”, [ca. 52 ° 52 ’ N, 88 ° 23 ’ E], Pinus sibirica-Abies forest, pitfall traps, 3 – 10. VI. 2010, L. A. Trilikauskas; 1 ♂, 1 ♀ (ISEA O. 001.0154), Sheregesh vicinity, subalpine Picea sibirica forest, [ca. 52 ° 57 ’ N, 88 ° 02 ’ E], 26. VII. – 1. VIII. 2013, L. A. Trilikauskas. Republic of Khakassia: Ust’ - Abakanskiy District: 1 ♀ (ISEA O. 001.0270), Kuznetskiy Alatau Mt. Range, 20 km NE of Balyksu, right tributary of Kainzas River, [ca. 53 ° 36 ’ N, 89 ° 20 ’ E], 1000 m a. s. l., forest, 22 – 23. V. 1997, R. Y. Dudko; 1 ♀ (ISEA O. 001.0271), Kuznetskiy Alatau, 7 – 20 km NE of Balyksu, Terensug River valley, [ca. 53 ° 30 ’ N, 89 ° 19 ’ E], 24. V. 1997, R. Y. Dudko. Tashtypskiy District: 3 ♂ (ISEA O. 001.0272), Abakanskiy Mt. Range system, Choochek Mt. Range, 20 km SSE of Mrassu, [ca. 52 ° 07 ’ N, 88 ° 33 ’ E], bold mountain (golets), 1600 – 1800 m a. s. l., 7 – 19. VII. 1999, D. E. Lomakin; 2 ♂ (ISEA O. 001.0273), Abakanskiy Mt. Range, ca. 20 km SSE of Mrassu, [ca. 52 ° 08 ’ N, 88 ° 38 ’ E], upper line of forest, ca. 1500 m a. s. l., 16 – 20. VII. 1999, D. E. Lomakin. KAZAKHSTAN: East Kazakhstan Province: 3 ♂ (ISEA O. 001.0279), Ivanovskiy Mt. Range, ca. 10 km S of Ridder [= Leninogorsk], [ca. 50 ° 15 ’ N, 83 ° 29 ’ E], Abies-Larix forest, 1700 m a. s. l., 30 – 31. V. 1996, R. Y. Dudko; 1 ♀ (ISEA O. 001.0277), Ivanovskiy Mt. Range, upper reaches of Gromotukha River, [ca. 50 ° 12 ’ N, 83 ° 45 ’ E – 50 ° 14 ’ N, 83 ° 47.5 ’ E], Larix forest, 1400 – 1800 m a. s. l., 3 – 5. VI. 1996, R. Y. Dudko; 1 ♀ (ISEA O. 001.0278), ca. 15 km NE of Ridder [= Leninogorsk], Belaya Uba River valley, Poperechnoe, [ca. 50 ° 26 ’ N, 83 ° 47 ’ E], 9. VI. 1996, R. Y. Dudko.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF83A31C19F9FF4DFC09FB7F.taxon	etymology	Etymology. The species is named in honour of Aleksandr V. Zateev, the director of the Katunsky Biosphere Reserve, who greatly helped the first author during fieldwork in the reserve.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF83A31C19F9FF4DFC09FB7F.taxon	diagnosis	Diagnosis. Sabacon zateevi sp. nov. is morphologically similar to S. sergeidedicatum, differs in shape of penis — subparallel lateral sides in S. sergeidedicatum and club-shaped in S. zateevi sp. nov. — cheliceral glands and palpal coxa. The main diagnostic characters to differentiate between the two species are given in Table 1.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF83A31C19F9FF4DFC09FB7F.taxon	description	Description. MALE (Holotype). Measurements. Body 3.10 long, 1.7 wide. Cephalothorax 2.00 long. Eye tubercle 0.25 wide. Clypeus (space between the ocularium and the front margin of the carapace) 0.08 long. Chelicera: basal segment 0.70 long; distal segment 0.57 long; forceps 0.35 long. Penis 1.85 long, 0.12 wide at its base. Body almost white. Carapace with a light brownish grey (in holotype) to dark grey (Fig. 19) trapezoidal spot; scutum and free tergites slightly tinged with grey (holotype) to dark brown stripe (in some paratypes; Figs 18 and 19, respectively). Ocularium black. Groups of short setae on both sides of the ocularium in the anterior part of the propeltidium. Scutum and free tergites with transverse rows of setae. Coxae of walking legs with short setae, with decreasing density from the first to the fourth leg. Scutum oval. Carapace smooth. Metapeltidium medially with a pair of sensory cones. Chelicerae. Whitish. Basal segment dorsally with cheliceral gland occupying half of its length. Cheliceral gland with numerous setae. Distal segment wide, almost flat retrolaterally, prolaterally with a projection situated in its apical half. Pedipalps grey. Palpal patella distally with ventral black teeth. Holotype has two teeth on one pedipalp and four on the other. Paratypes have three teeth on all palps. Palpal coxa with a small, hump-like projection. Legs grey. Cylindrical in cross-section with longitudinal rows of setae. FEMALE (Paratype, ISEA 001.8563). In general appearance, similar to the male but larger. Palpal patella without black teeth. Basal segment of chelicerae without cheliceral gland dorsally. Ovipositor as illustrated in Fig. 71. Habitat and biology. The species is occasionally collected in pitfall traps in subalpine spruce forests. It was hand-collected in these forests exclusively from logs of conifers. Some specimens were also found among stones and plant residues on an island in the floodplain of a small river and on pebbles at the confluence of streams running towards Taimenye Lake. Records of subadult and adult males in the spring and early juveniles in the autumn indicate that the species overwinters at an active stage. Thus, the life cycle of S. zateevi sp. nov. seems to be similar to that of S. sergeidedicatum (see Trilikauskas 2017). All habitats are located from 1300 to 1950 m a. s. l.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
03FB87DBFF83A31C19F9FF4DFC09FB7F.taxon	discussion	Comments. We re-studied material of S. sergeidedicatum from Chemeris & Logunov (2001) (except for the Holotype) as well as additional material from ISEA. Our investigation shows that figures 22 and 23 are actually macrographs of the right (Fig. 22) and left (Fig. 23) palps taken from retrolateral and prolateral views, respectively. Moreover, our study of many specimens from populations throughout the species’ range shows intraspecific variation in such important taxonomical characters as shape, size and proportions of penis (Figs 34 – 40), shape of male’s chelicerae (Figs 45 – 49, 51 – 55) and male’s palpal coxae (Figs 57 – 62). Only one exception among thousands of specimens was found: palpal coxae of one male from Shorsky National Park (Kemerovo Area, Russia) shows absence of arrowhead-shaped basal knob, which might be considered a developmental anomaly. The apical part of arrowhead-shaped basal knob seems to be lost in old specimens (see Figs 57 and 60) but the basal part always stays intact. Specimens from Kholzun Mt. Range differ from both S. sergeidedicatum and S. zateevi sp. nov. and most likely belong to a new underscribed species. Description of this species is beyond the aims of this paper and will be made in a separate paper. The Altai Mountains have a large degree of endemism in other invertebrate taxa, e. g. spiders (see Marusik et al. 2004; Marusik & Fomichev 2010; Marusik et al. 2014, 2019), beetles (see Dudko & Shilenkov 2001; Dudko 2006, Dudko 2011) etc. Finding a new species was not surprising to us. Moreover it is possible that more new species will be revealed especially from the poorly studied Mongolian part of Altai.	en	Trilikauskas, Laimonas A., Azarkina, Galina N. (2021): The harvestmen fauna (Arachnida: Opiliones) of the Katunsky Biosphere Reserve and adjacent territories (South Siberia, Russia), with a description of a new species of Sabacon (Sabaconidae) and notes on Sabacon sergeidedicatum Martens, 1989. Zootaxa 4990 (1): 117-133, DOI: 10.11646/zootaxa.4990.1.7
