identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FB87EAFF96FFE9FF1EFD1CFC60FF08.text	03FB87EAFF96FFE9FF1EFD1CFC60FF08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Willowsia cassagnaui	<div><p>Willowsia cassagnaui sp. nov.</p><p>Figs 1−15</p><p>Types. Holotypes: female on slide, Nepal: Machapuchare, alt. 2200 m, above Pokhara, forest of large trees ( Quercus semicarpifolia Sm. In Rees, Acer sp., rhododendrons, bamboos etc.), dry moss on a branch, 26.x.1976, P. Cassagnau leg. (# Nep. 76.51). Paratypes: 4 female and 4 young specimens on slides, same data as holotype. Two paratypes in NJAU and others in MNHN.</p><p>Description. Body length up to 1.82 mm.</p><p>Ground colour pale yellow in alcohol. Blue pigment forming irregular patterns on body. Ant. I −IV gradually becoming darker from anterior to posterior. A transverse band between the eye patches and a small lateral patch behind each eye patch. Lateral patches present from Th. II to Abd. VI. A pair of lighter small patches on median Th. III and Abd. I. Two transverse bands on posterior Abd. II and III. A pair of large median and small posterior patches on Abd. IV. Legs slightly pigmented. Furcula pale (Fig. 1). Scales on body pointed, striate heavily with basal ribs longer than distal ones (Fig. 2); scales on ventral manubrium obviously narrower than those on terga (Fig. 3). Scales absent on antennae, legs, ventral tube and dens.</p><p>Antenna twice as long as cephalic diagonal. Antennal segments ratio as I: II: III: IV = 1: 1.9: 1.8: 2.9. Ant. III organ not clearly seen. Antennal apical bulb bilobed. Labral margin with 4 rounded papillae. Labral intrusion slightly curved (Fig. 4). Tip of lateral process of labial palp nearly reaching apex of labial papilla (Fig. 5). Subapical chaeta of maxillary outer lobe slightly stronger than apical one; 3 hairs on sublobal plate (Fig. 6). Labial chaeta R 0.4−0.5 x the length of M (Fig. 7). Dorsal cephalic chaetotaxy with 6 antennal (An) and 5 sutural (S) mac (Fig. 8).</p><p>Th. II with 10 posterior mac; median mac absent; p4−6 and p5pi as mac. Th. III with 11 mac (Fig. 9). Thoracic S-chaetae not clearly seen. Metatrochanteral organ with about 14 smooth spiny chaetae; 10 in arms, V-shaped and 4 between arms (Fig. 10). Unguis with 4 inner teeth; basal inner pair with tip reaching 0.45 distance from base of ventral edge, median one at 0.71 and distal one 0.87 distance from base. Unguiculus lanceolate with outer edge slightly serrate. Tenent hair subequal to unguis in length (Fig. 11).</p><p>Abd. IV four times Abd. III in length along dorsal midline. Abd. I with 3 (m2−4) mac and 2 S-chaetae; ms internal to sens. Abd. II with 3 (a2, m3, m3e) central, 1 (m5) lateral mac and 2 sens. Abd. III with 2 (a2, m3) central, 3 (am6, pm6, p6) lateral mac, 1 ms and 2 sens. Abd. IV with 7 central mac and about 12 sens; homology of anterior three mac unclear; 3 lateral sens obviously shorter than others (Fig. 12). Accessory mic of bothriotrichal complexes on Abd. II −IV unmodified. Ventral tube anteriorly with about 14 ciliate chaetae on each side (Fig. 13), 2 of them larger than others; posteriorly with 11 chaetae, apical two smooth; each lateral flap with 7 smooth and 1 ciliate chaetae (Fig. 14). Distal smooth part of dens 1.9−2.5 x mucro in length (Fig. 15).</p><p>Etymology. The new species is named after P. Cassagnau, who collected the specimens.</p><p>Remarks. Willowsia cassagnaui sp. nov. is similar to W. ieti Yosii, 1971 in most features. It differs from it in colour pattern, lateral process of labial palp longer with tip nearly reaching apex of labial papilla, slightly curved labral intrusion, fewer chaetae on trochanteral organ (14 vs&gt;20), lanceolate unguiculus, presence of mac m2 on Abd. I, and absence of scales on Ant. I (Table 1). Lateral stripes, transverse band on Abd. II, and paired lighter small patches on median Th. III, Abd. I and Abd. IV are present in W. cassagnaui sp. nov., but are absent in W. ieti, which possesses a pair of small spots on Abd. II. The slightly curved labral intrusion of the new species is rarely seen in entomobryids. The shape of the unguiculus is also a characteristic separating them, as it is lanceolate in new species and truncate in W. ieti .</p><p>Type locality. Nepal: Khumbu Himal, Yaral near Pangpoche, alt. 4100 m.</p><p>Specimens examined. 4 females on slides, Nepal: Machapuchare, alt. 2740 m, level site with large trees (oaks, rhododendrons, bamboos), moss and lichen on a branch, 25.x.1976, P. Cassagnau leg. (# Nep. 76.40); 8 females on slides, Nepal: Gosaikund, alt. 3660 m, upper edge of the fir forest and juniper heath, moss and liverworts on a rock, moderately humid, 20.xi.1978 (# Nep. 78.59). Eight specimens on slides in NJAU and others in MNHN.</p><p>Additions to original description. Body scales heavily striate with pointed tips (Fig. 16); present on Ant. I, body and ventral side of manubrium; scales on manubrium narrower than those on body, some of them extremely elongate (Fig. 17). Tip of lateral process of labial palp not reaching apex of labial papilla (Fig. 18). Subapical chaeta of maxillary outer lobe stronger than apical one; 3 hairs on sublobal plate (Fig. 19). Labial chaeta R 0.7 x the length of M. Dorsal cephalic chaetotaxy as in Fig. 20. Metatrochanteral organ with more than 20 smooth spiny chaetae (Fig. 21). Ventral tube anteriorly with about 22 ciliate chaetae on each side, 2 of them much larger than others (Fig. 22). Manubrial plaque with about 6 ciliate chaetae; pseudopores not clearly seen. Distal smooth part of dens about twice mucro in length. Th. II with 10 posterior mac; median mac absent. Th. III with 11 mac (Fig. 23). Abd. I with 2 (m3−4) mac. Abd. II with 3 (a2, m3, m3e) central and 1 (m5) lateral mac. Abd. III with 2 central (a2, m3) and 3 (am6, pm6, p6) lateral mac (Fig.24). Abd. IV with 7−8 central and about 14 lateral mac; homology of anterior four central mac unclear, the second mac close to midline often as a mesochaeta (Fig. 25). Accessory mic of bothriotrichal complexes on Abd. II −IV unmodified or absent.</p><p>Remarks. The type specimens in Natural History Museum, Geneva, could not be studied as permission was not given to borrow it. The type locality (Yaral) is located between the two new sites for this species recorded here. The Gosaikund record is 100 km directly east of the type locality, Machapuchare about 75 km west of Yaral. Our specimens from these two new populations comply well with Yosii’s (1971) original description, particularly in colour and tergal chaetotaxy (Th. II‒Abd. III), so that they are designated as W. ieti Yosii here. We added further details of mouthpart, chaetae on dorsal and ventral head, ventral tube and total body chaetotaxy. For differences between W. ieti and W. cassagnaui sp. nov. see the remarks above.</p></div>	https://treatment.plazi.org/id/03FB87EAFF96FFE9FF1EFD1CFC60FF08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zhang, Feng	Zhang, Feng (2015): Some Willowsia from Nepal and Vietnam (Collembola: Entomobryidae) and description of one new species. Zootaxa 3905 (4): 489-499, DOI: 10.11646/zootaxa.3905.4.3
03FB87EAFF91FFE7FF1EFE8EFCE3FF2A.text	03FB87EAFF91FFE7FF1EFE8EFCE3FF2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Willowsia bartkei Stach 1965	<div><p>Willowsia bartkei Stach, 1965</p><p>Figs 26–32</p><p>Type locality. Vietnam: Cha-Pa.</p><p>Specimens examined. 7 females on slides and 14 in alcohol, all paratypes, Vietnam: Cha-Pa, alt. 1500 m, under stones and in moss, also under loose bark of trees and in litter in a forest, iii.1961, A. Bartke leg. All deposited in ISEA.</p><p>Redescription. Body length up to 2.3 mm.</p><p>Ground colour pale yellow in alcohol. Eye patches dark blue. Each antennal segment becoming gradually darker towards apex. Body dorsum with lateral stripes along mesothoracic margins, transverse bands on Abd. I posteriorly and Abd. II and III, and dark patches on posterior Abd. IV and V. Scattered pigment present on distal part of femora and tibiotarsi (Fig. 26). Scales hyaline, leaf-like with tip pointed and finely striate; scales absent on antennae, legs and furcula (Fig. 27).</p><p>Antenna 2.5 times as long as cephalic diagonal. Antennal segments ratio as I: II: III: IV = 1: 2: 2: 3. Ant. III organ with two rods. Antennal apical bulb unilobed. Labral margin with 4 conical papillae. Dorsal cephalic chaetotaxy with 6 sutural (S) mac; anterior area not clearly seen. Interocular area with 3 ciliate mic (Fig. 28).</p><p>Dorsal mac shown in Fig. 29. Th. II with 2 (m1, m2) medio-medial, 3 (m4, m 4i, m4p) medio-lateral and 15 posterior mac; p4 and p5 as mac. Th. III with 18 mac; m 5i and a 6i as mac. Metatrochanteral organ with about 30 smooth spiny chaetae. Unguis with 3 inner teeth, all minute; inner paired and distal ones 0.5 and 0.8 distance from base, respectively. Unguiculus lanceolate. Tenent hair subequal to unguis in length (Fig. 30).</p><p>Abd. IV 2.2 times as long as Abd. III along dorsal midline. Abd. I with 3 (m2−4) central mac. Abd. II with 3 (a2, m3, m3e) central and 1 (m5) lateral mac. Abd. III with 2 (a2, m3) central and 4 (am6, pm6, p6, m7) lateral mac. Abd. IV with 7 central mac. Accessory mic of bothriotrichal complexes on Abd. II −IV strongly modified or fan-shaped. Ventral tube anteriorly with 3 large ciliate chaetae and several ciliate chaetae on each side (Fig. 31); posterior face and lateral flap not clearly seen. Distal smooth part of dens 1.86−2.00 times mucro in length (Fig. 32).</p><p>Remarks. We provide here complete dorsal cephalic and body chaetotaxy based on paratypes of W. bartkei . Fresh material is needed for a fuller description. Willowsia bartkei is most similar to W. guangxiensis Shi &amp; Chen, 2004 from Guangxi Province, South China in having transverse bands on Abd. I to III, scale sculpture with short striations, abundant spiny chaetae (&gt;30) on the trochanteral organ, and in the chaetotaxy of Abd. III. It differs from the latter in having a posterior patch of dark pigment on Abd. IV, unilobed Ant. IV apical bulb, scale distribution (absent on antennae), lanceolate unguiculus, much longer distal smooth part of dens, 3+3 mac on Abd. I, 3+3 inner mac on Abd. II, and fewer inner mac (7+7) on Abd. IV.</p></div>	https://treatment.plazi.org/id/03FB87EAFF91FFE7FF1EFE8EFCE3FF2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zhang, Feng	Zhang, Feng (2015): Some Willowsia from Nepal and Vietnam (Collembola: Entomobryidae) and description of one new species. Zootaxa 3905 (4): 489-499, DOI: 10.11646/zootaxa.3905.4.3
03FB87EAFF9FFFE7FF1EFEE5FA77FA3E.text	03FB87EAFF9FFFE7FF1EFEE5FA77FA3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Willowsia pseudosocia Stach 1965	<div><p>Willowsia pseudosocia Stach, 1965</p><p>Figs 33−44</p><p>Type locality. Vietnam: Cha-Pa</p><p>Specimens examined. Holotype and paratype, both females, Vietnam: Cha-Pa, alt. 1500m, under a piece of bark lying on the ground, 20.iv.1961, A. Bartke leg. All deposited in ISEA.</p><p>Redescription. Body length up to 3.0 mm.</p><p>Ground colour pale in alcohol. Eye patches dark blue. A narrow stripe present on lateral margin from head to Abd. I. A pair of dark patches on Abd. V (Fig. 33). Scales coarsely striate and leaf-like with tip pointed; scales present on Ant. I but absent on legs (Fig. 34).</p><p>Ant. III and IV lost from examined type specimens. Labral papillae rounded. Lateral process of labial palp thin with tip not reaching apex of labial papilla (Fig. 35). Labial chaeta R 0.55 x the length of M (Fig. 36). Dorsal cephalic chaetotaxy with about 12 antennal (An), 4 median (M) and 5 sutural (S) mac. Interocular area with 3 ciliate mic (Fig. 37).</p><p>Th. II with 4 (m1, m 1i, m2, m 2i) medio-medial, 3 (m4, m 4i, m4p) medio-lateral and 26‒30 posterior mac. Th. III with 40‒44 mac (Fig. 38). Metatrochanteral organ with about 36 smooth spiny chaetae (Fig. 39). Unguis with 1 outer, 2 lateral and 3 inner teeth, all tiny; inner pair with tip reaching 0.50 distance from base, median one at 0.75 distance from base. Unguiculus lanceolate. Tenent hair shorter than unguis (Fig. 40).</p><p>Abd. IV 5.5 times Abd. III in length along dorsal midline. Dorsal mac shown in Fig. 41. Abd. I with 10 mac. Abd. II with 6 (a2, a3, m3, m3e, m3ea, m3ep) central and 1 (m5) lateral mac. Abd. III with 3 (a2, a3, m3) central and 4 (am6, pm6, p6, m7) lateral mac. Abd. IV with about 17 central and 22 lateral mac. Accessory mic of bothriotrichal complexes on Abd. II‒IV strongly modified with most being transformed into scales (Fig. 43). Ventral tube anteriorly with 3 large ciliate chaetae and many ciliate chaetae of different sizes on each side (Fig. 42); posterior and lateral flap not clearly seen. Distal smooth part of dens 0.5 times mucro in length (Fig. 44).</p><p>Remarks. We redescribed W. pseudosocia based on types and gave additional details of the dorsal cephalic and body chaetotaxy, interocular chaetae, labial chaetae, lateral process of labial palp, anterior face of ventral tube and trochanteral organ. This species is most similar to W. nivalis Yosii, 1971 from Nepal in colour (pale on dorsal body), scale sculpture, labral chaetae and labral papillae, claw structure, modified accessory mic of bothriotrichal complexes, 3+3 inner mac on Abd. III. It differs from W. nivalis in having lateral stripes on Th. II‒Abd. I, scales present on antennae, more numerous spine-like chaetae (&gt;40) on trochanteral organ and tergal mac obviously more abundant (such as 10+10 on Abd. I, 6+6 inner on Abd. II). Many undescribed Willowsia species from the Chinese region (Guangdong, Guangxi, Tibet Provinces) adjacent to Vietnam and Nepal have been discovered during recent expeditions. It is necessary to re-examine those known species for the further comparison and systematic revision.</p></div>	https://treatment.plazi.org/id/03FB87EAFF9FFFE7FF1EFEE5FA77FA3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zhang, Feng	Zhang, Feng (2015): Some Willowsia from Nepal and Vietnam (Collembola: Entomobryidae) and description of one new species. Zootaxa 3905 (4): 489-499, DOI: 10.11646/zootaxa.3905.4.3
