taxonID	type	description	language	source
03FB87F5FFA6FB4DFF08FE607D8E8F4C.taxon	description	urn: lsid: zoobank. org: act: D 1 F 98 CEE- 808 E- 4 DFB-A 291 - AE 3 E 6 A 357568	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA6FB4DFF08FE607D8E8F4C.taxon	type_taxon	Type species: Pericoma formosa Meunier, 1905, by present designation.	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA6FB4DFF08FE607D8E8F4C.taxon	diagnosis	Diagnosis. Only known from Eocene Baltic amber (34 – 48 Mya; Seyfullah et al. 2018). Antenna (Figs. 7; 17) with the following combination of character states: Scape elongate, at least three times longer than wide; male pedicel with basal half cylindrical, apical half globular; male flagellomere 1 fusiform to globular, always clearly differing in shape from remaining flagellomeres; flagellomeres 2 – 13 nodiform, symmetrical; flagellomere 14 acorn-shaped, with subapical digitiform process; flagellomeres 2 – 13 (unknown for f 14) with two ascoids, inserted on node opposite each other; ascoids (Figs. 10; 13) fan-like, each with 4 – 6 separate, digitiform branches in male, 3 branches in female. Gonocoxite (in dorsal view) with flat, quadrate extension basomedially (Figs. 4; 14; 18; 21).	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA6FB4DFF08FE607D8E8F4C.taxon	discussion	Comments. Characterizing Paleotelmatoscopus as a genus is somewhat precarious due to its similarity to several extant genera (e. g. Telmatoscopus Eaton, 1904; Seoda Enderlein, 1935; Jungiella Vaillant, 1972). Many diagnostic characters that are described for extant psychodine genera are of the internal male genitalia. Characters of the male genitalia, some of which were discussed by Kvifte (2014), are of paramount importance for separating genera and species; however, in fossil taxa this is limited to external characters such as the length and curvature of gonostyli and surstyli, or the number of tenacula. So far, these have proven to be less useful for separating species of Paleotelmatoscopus, as their male genitalia, on the exterior, are more uniform. The most reliable male genitalia character for distinguishing Paleotelmatoscopus from related genera is the dorsal part of the base of the gonocoxite. In Paleotelmatoscopus it is broad, quadrate and dorsoventrally compressed (Fig. 14), with gonocoxites clearly separate. In Telmatoscopus, it is narrowed to a point and the gonocoxites meet at the median line (Vaillant 1989, Fig. 2; Kvifte 2014, Fig. 1 C), while in Seoda it is rounded, with gonocoxites clearly separated (Ježek 1989, Fig. 13; Kvifte 2014, Fig. 2 C – D) and in Jungiella it is truncate, without an extension of the base (Omelková & Ježek 2017, Figs. 14; 38). In contrast to the variability of the gonocoxites, the gonostyli of Paleotelmatoscopus and related genera are remarkably similar. Curvature of the gonostyli, with a distinctive, angular bend at midlength, are most similar to Telmatoscopus advena Eaton, 1893 (Vaillant 1989, Figs. 2 – 3) and some species of Seoda (Ježek 1989; Figs. 27 – 28; 35; 41). Head capsules of Paleotelmatoscopus species observed in this study are typical of the form observed in related genera: rounded or slightly pyriform in frontal view; 4 facet rows in eye bridge, separated at median by distance of approximately two facet diameters; with corniculi in some species. The antennae appear to be the only character suite in the head capsule that can be used to separate genera. Most notably, flagellomeres in Paleotelmatoscopus are not strongly asymmetrical as in related genera, and their ascoids are palmate. Nonetheless, all specimens examined by us had incomplete antennae, missing one or more flagellomeres and ascoids; this prevented an accurate description of flagellomere 14 with regard to whether it bears ascoids. Diagnostic characters given above for the genus are based on a survey of all specimens available to us; however, this will require revision based on the study of additional material. Wings of Paleotelmatoscopus are also similar to related genera, except some species of the fossil genus (e. g. P. formosa) have a posterior wing margin that is comparatively more convex, with a CuA cell that is expanded posteriorly. Paleotelmatoscopus can be readily distinguished from Telmatoscopus because species of the former have the apex of R 5 terminating posterior to the wing apex, while the latter have the apex of R 5 terminating at the wing apex; however, separation of the new genus from other related genera requires greater scrutiny. Female Paleotelmatoscopus do not appear to have distinctive genus level characters. Association of males and females thus far is mainly based on specimens that are syninclusions.	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA7FB4AFF08FB197F338F04.taxon	diagnosis	Diagnosis. Species known only from Eocene Baltic amber. Male with the following combination of character states: head with corniculi inserted at postocciput (Figs. 2; 5; 8; 10; 12); antennal scape with prominent, bulbous mesial lobe at apex (Figs. 2; 5 – 8; 12); flagellomere 1 fusiform, with node broader than remaining flagellomeres; posterior wing margin convex between Cu and A 1, changing abruptly to straight between A 1 and alula (Figs. 1; 5; 9); gonopods at point of articulation between gonocoxite and gonostyli conspicuously wider than all preceding abdominal segments (Figs. 3; 4; 9).	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA7FB4AFF08FB197F338F04.taxon	description	Description. Male. Outline of head pyriform in frontal view, narrowing from vertex to postocciput; simple, mushroom-shaped cornicula inserted on either side of postocciput; eyebridge comprised of four complete facet rows, separated by two facet diameters at median; interocular suture arched toward vertex, not distinctly U-shaped; frons, vertex, occiput covered with setae alveoli; frons conspicuous, bulging between antennae; clypeus with ventral margin straight; labrum inconspicuous, triangular in frontal view; labellum bulbous, equal in width or narrow compared to clypeus; palp with segments cylindrical, nondescript; apical palp segment elongate, approximately twice the length of preceding segment; scapus elongate, more than three times longer than wide, with prominent mesial lobe at apex; mesial surface of scape covered in dark, spatulate setae; pedicel with basal half cylindrical, apical half globular, narrow compared to node of flagellomere 1; flagellomere 1 broad-fusiform, symmetrical, with node broader than that of remaining flagellomeres; flagellomeres 2 – 13 nodiform, with nodes and internodes narrowing progressively from base to apex of antenna; apical flagellomere with node spherical, apiculus digitiform, setose. Thorax with lateral sclerites poorly visible in most specimens; expandable lobes and other remarkable features ab-sent; legs with femora and tibiae increasing in length from foreleg to hindleg; wing with C approximately equal in thickness compared to longitudinal veins; Sc extending to base of R 2 + 3; radial fork placed slightly apical to medial fork, both placed slightly basal to apex of Cu. Terminalia with hypandrium band-like, without increase in length at median, not extending beyond anterior margins of gonocoxites; gonopods at point of articulation between gonocoxite and gonostyli conspicuously wider than all preceding abdominal segments; gonocoxite reniform in dorsal view; gonostylus acuminate, progressively tapered from base to apex, with slight bend at midlength, longer than gonocoxite; parameres membranous, fused into sheath, completely surrounding aedeagus; epandrium narrowing from base to apex in dorsoventral view, emarginate posteriorly, with paired aperture ventrally; surstyli elongate, curved dorsally, slightly inflated at base, with cluster of more than 10 tenacula inserted dorsoapically; hypoproct conspicuous, triangular in dorsoventral view, narrowing from base to apex. Female. Head without corniculi; eye bridge comprised of four complete facet rows; scape cylindrical, approximately 2.5 times longer than wide, pedicel globular, flagellomere 1 of similar shape to flagellomeres 2 – 13; ascoids palmate, with three branches.	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA7FB4AFF08FB197F338F04.taxon	materials_examined	Type material. A male specimen of Pericoma formosa Meunier, 1905 numbered Z 2681 (GZG. BST 06169) is here designated as the lectotype. Other material examined. See Table 1 and the corresponding folio in Morphobank: http: // morphobank. org / permalink /? F 895	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA7FB4AFF08FB197F338F04.taxon	discussion	Comments. The lectotype male shows all diagnostic characters for this species, which is why this specimen was chosen, despite the fact that the genitalia is not clearly visible. Considering the diagnostic characters listed above, an examination of the original series of specimens as listed by Meunier (1905) shows that “ Pericoma ” formosa included at least two species. Some of these specimens are identified here as Paleotelmatoscopus formosa, while others are identified as a new species, described below. P. formosa has a significantly broader wing and longer scape than P. madrizi, and a wider head capsule in at least the male, but probably the female as well (see measurements in Table 1). Unfortunately, there are no female specimens of P. madrizi in the material studied by us, of which the head capsule could be accurately measured for comparison.	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA0FB44FF08FB517B4B89C0.taxon	description	urn: lsid: zoobank. org: act: 3137 EB 10 - 98 D 9 - 4940 - A 214 - 6677 AA 4 BE 228	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA0FB44FF08FB517B4B89C0.taxon	diagnosis	Diagnosis. Species known only from Eocene Baltic amber. Male with the following combination of character states: head capsule with postocciput lacking corniculi (Figs. 13; 15; 19); antennal scape without prominent mesial lobe at apex (Figs. 13; 15; 17; 21); flagellomeres approximately equal in width from base to apex; anterior and posterior wing margin evenly convex (Figs. 13; 15; 17; 21).	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA0FB44FF08FB517B4B89C0.taxon	description	Description. Male. Outline of head round in frontal view, vertex broadly rounded, narrowing abruptly at postocciput; corniculi absent from postocciput; eyebridge comprised of four facet rows, separated by two facet diameters at median, with ventral-most row interrupted by base of scape; interocular suture arched toward vertex, broadly U-shaped; frons, vertex, occiput covered with setae alveoli; frons inconspicuous, bulging slightly between antennae; clypeus with ventral margin convex; labrum inconspicuous, triangular in frontal view; labellum somewhat laterally compressed, narrow compared to clypeus; palp with segments cylindrical, nondescript; apical palp segment elongate, approximately 2.5 times the length of preceding segment; scapus elongate, approximately 2.5 times longer than wide, without mesial lobe at apex; mesial surface of scape covered in dark, spatulate setae; pedicel with basal third cylindrical, apical 2 / 3 globular, broad compared to node of flagellomere 1; flagellomere 1 fusiform, symmetrical, with node approximately equal in width to those of remaining flagellomeres; flagellomeres 2 – 13 nodiform, with nodes approximately equal in width, internodes shorter at base and apex of antenna; apical flagellomere with node oblong, with minute subapical projection, apiculus digitiform, setose. Thorax with lateral sclerites poorly visible in most specimens; expandable lobes and other remarkable features absent; midleg with femur longer than that of fore- or hindleg, tibia shorter than that of hindleg; wing with C slightly greater in thickness compared to longitudinal veins; Sc not reaching base of R 2 + 3; radial fork placed slightly apical to medial fork. Terminalia with hypandrium band-like, without increase in length at median, not extending beyond anterior margins of gonocoxites; gonopods at point of articulation between gonocoxite and gonostyli approximately equal in width to preceding abdominal segments; gonocoxite with apical 2 / 3 cylindrical in dorsal and lateral view; gonostylus longer than gonocoxite, acuminate, with pronounced bend at midlength, apical half conspicuously narrowed in comparison to base; parameres membranous, fused into sheath, partially surrounding aedeagus; epandrium wider than long, narrowing from base to apex in dorsoventral view, emarginate posteriorly, with paired aperture ventrally; surstyli elongate, curved dorsally, tapered from base to apex, with cluster of fewer than 10 tenacula inserted dorsoapically; hypoproct conspicuous, tongue-shaped in dorsoventral view, narrowing from base to apex. Female. Head with eye bridge comprised of four facet rows, with ventral-most row incomplete; scape conical, widening from base to apex, at least twice as long as wide, pedicel globular, flagellomere 1 of similar shape to flagellomeres 2 – 13; ascoids palmate, with three branches.	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA0FB44FF08FB517B4B89C0.taxon	etymology	Etymology. The specific epithet is a patronym in honor of Isaí Madriz, our friend and colleague who expertly illustrated the male genitalia, revised the habitus illustration and provided many hours of guidance during the process of capturing and processing photomicrographs for this work.	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA0FB44FF08FB517B4B89C0.taxon	materials_examined	Type material. Holotype: male, CCHH 1800 - 3, deposited in SDEI. For a list of paratypes, please see Table 1 and the corresponding folio in Morphobank: http: // morphobank. org / permalink /? F 896 Other material examined. See Table 1 and the corresponding folio in Morphobank.	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFA0FB44FF08FB517B4B89C0.taxon	discussion	Comments. Some specimens of P. formosa examined by Meunier (1905, p. 244) were described as “ Var. ”. This is based on the flagellomeres being roughly the same width rather than being progressively more narrow from the base to apex of the antenna. These specimens are regarded here as Paleotelmatoscopus madrizi (Table 1). Male and female specimens of P. madrizi have a shorter scape and smaller wing (i. e. shorter and more narrow), and males have a more narrow head capsule in comparison to P. formosa. This may be true for females as well, but measurement was not possible for this character. In the line illustration of the holotype male (Fig. 13), ascoids are absent from some flagellomeres; this is to indicate that, on this particular specimen, we did not see ascoids inserted on those flagellomeres. Presumably they were lost prior to, or during preservation of the specimen in the amber.	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFAEFB42FF08FC947D188928.taxon	diagnosis	Diagnosis. Species known only from Eocene Baltic amber. Male with the following combination of character states: head capsule with antennal scape and pedicel globular, flagellomeres fusiform (Fig. 25); wing profile ovate, with bluntly pointed apex (Figs. 25 – 26; 28).	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFAEFB42FF08FC947D188928.taxon	description	Description. Male. Eyebridge comprised of three facet rows; labellum bulbous, conspicuous; palp with segments cylindrical, nondescript; scapus globular, slightly larger than pedicel; pedicel globular; flagellomeres uniform in shape, fusiform, twice as long as wide; ascoids not visible in specimen; Thorax without remarkable characters; legs approximately equal in length, with proportions of femora, tibiae and tarsi of fore-, mid- and hindleg roughly equal; wing with C nearly twice as thick compared to longitudinal veins; radial fork placed slightly apical to medial fork. Terminalia with gonostylus straight, slightly inflated at base, evenly tapered from inflated part to apex; surstyli moderately elongate, slightly curved dorsally, inflated at base, without noticeable tapering from base to apex; cluster of more than 10 tenacula inserted dorsoapically.	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFAEFB42FF08FC947D188928.taxon	materials_examined	Type material. A male specimen originally examined by Meunier (1905) numbered Z 2726 (GZG. BST- 06177) is here designated as lectotype. Other material examined. See Table 1 and the corresponding folio in Morphobank: http: // morphobank. org / permalink /? F 897. An additional female specimen, probably congeneric with P. speciosa, is placed in a separate folio in Morphobank: http: // morphobank. org / permalink /? F 907	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
03FB87F5FFAEFB42FF08FC947D188928.taxon	discussion	Comments. Generic classification of this species is currently inadequate, as it is not Pericoma, but also can not be confidently assigned to another genus or described as new. Few specimens are available for study and the lectotype male shows only a lateral view of the genitalia. Unfortunately, a second male specimen (MZ 17325) with ovate wings and fusiform antennal flagellomeres agreeing to P. speciosa can only be tenusouly identified as such. The eye bridge is obscured and genitalia is visible in partial dorsoventral view. Additional specimens of this type are needed to clarify the genus and species-level taxonomy.	en	Curler, Gregory R., Skibińska, Kornelia (2021): Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species. Zootaxa 4927 (4): 505-524, DOI: 10.11646/zootaxa.4927.4.2
