taxonID	type	description	language	source
03FBE052FFF0FF9FFF38FCEA7129FC76.taxon	materials_examined	Type locality. Small spring, Budha Valley N of Yangpachem, Tibet South Central. Type material studied. Holotype male (NMB): “ TIBET South Central, Budha Valley N of Yangpachem, about 30 ° 10 ' 38 N 90 ° 30 ' 18 E, 20. VI. 2007, 5100 m ” [white printed label]. “ Holotype Agabus joachimschmidti sp. n. Brancucci & Hendrich des. 2008 ” [red printed label]. Paratypes: 9 specimens with same data as holotype (3 males and 4 females NMB, 1 male and 1 female CLH). All specimens are provided with a red printed paratype label.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF0FF9FFF38FCEA7129FC76.taxon	description	Description. Measurements: Holotype: total length = 6.20 mm, total length-head = 5.60 mm, total width = 3.20 mm. Paratypes (male and female do not differ in size): total length = 5.60 – 6.10 mm (5.85 mm, n = 7), total length-head = 5.20 – 5.50 mm (5.35 mm, n = 7), total width = 2.90 – 3.20 mm (3.08 mm, n = 4). Habitus: Body short-oval, distinctly convex, black on head and pronotum and dark brown on elytra (Fig. 1). Head: Black, shining, labrum testaceous. Antennae testaceous at base, becoming darker brown on distal part from the 4 th joint on. Reticulation consisting of medium-sized and well-impressed polygonal meshes, irregular in size, with a small puncture on the inner side of some meshes. Row of punctures alongside eyes and clypeal grooves well-impressed. A few punctures beside eyes well-impressed but not confluent. Antennae short, joints almost globose, the 5 th only 1.2 times as long as broad. Pronotum: Black, shining. Reticulation consisting of polygonal meshes, irregular in size and often incomplete particularly on disc; some meshes with 1 very minute puncture, seldom more, on their inner surfaces. Latero-basal part with few deep wrinkles. Anterior row of punctures complete; punctures very large, confluent and in groups, leaving small spaces free. Posterior row broadly interrupted at middle; punctures large, irregularly distributed, confluent at sides. Lateral margin distinctly bordered with punctures alongside the grooves. Sides of pronotum rather straight, forming a distinct pronoto-elytral angle, base of pronotum narrower than base of elytra (Fig. 1). Elytra: Brownish-black with a distinct bronze lustre. Epipleura brown. Reticulation consisting of wellimpressed polygonal meshes, with one minute puncture on the inner side of some meshes. Sutural row of punctures restricted to apical two-thirds and consisting of some medium-sized well-spaced punctures. Discal, sublateral and lateral rows not reaching base; punctures well-spaced, here and there in groups. Punctures present in interspaces between discal and sublateral rows. Epipleura broad at base, strongly narrowed by first sternite and evenly but slightly tapered as far as apical part. Venter: Dark ferrugineous-brown, legs testaceous, femora dark brown. Prosternal process lanceolate, very narrow and strongly convex. Metacoxal line complete. Metafemora with a short row of setae, slightly transversely reticulated. Metatibiae with two rows of punctures, one along outer margin and consisting of 7 – 9 larger punctures each with a strong but short seta, other along inner margin, shortened distally with 4 – 5 large punctures; ground-surface distinctly shining. Sexual dimorphism: Male with protarsi and mesotarsi distinctly dilated, with numerous small rounded pads. Protarsal and mesotarsal claws almost as long as protarsomere 5 and mesotarsomere 5, respectively. Anal sternite microreticulate on anterior third, somewhat wrinkled and punctured on posterior half. Posterior margin distinctly flattened, finely bordered. Median lobe of aedeagus, in lateral view, regularly curved, bidentate in apical part (Fig. 3); in dorsal view, suddenly tapered and prolonged into a sharp point at apex. Basal part of parameres short but narrow (Fig. 5). Female similar to male. Anal sternite distinctly reticulate on posterior half, with coarse and irregularly distributed punctures on both sides of middle. Protarsal and mesotarsal claws only 1.4 and 1.3 times shorter than protarsomere 5 and mesotarsomere 5.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF0FF9FFF38FCEA7129FC76.taxon	etymology	Derivatio nominis. Dedicated to Joachim Schmidt (Admannshagen, Germany) a well-known specialist of Himalayan ground beetles. Differential diagnosis. This species clearly belongs to the confinis - group as delimited by Nilsson (1990, 2001), mainly characterized by a bifid median lobe and a typical habitus of the species. It is related to A. tibetanus, but can be easily distinguished by its smaller size. The total length is 5.85 mm as the mean between all paratypes, whereas it is 6.8 mm in A. tibetanus. The sides of the pronotum are rather straight and not regularly rounded, forming a distinct pronoto-elytral angle, the base of the pronotum being narrower than base of elytra (Fig. 1); in A. tibetanus, the base of pronotum and the base of elytra are approximately of the same width (Fig 2). The posterior enlarged part of the metacoxal processes is provided laterally just with a few larger punctures, whereas it is distinctly bordered in A. tibetanus. The sutural rows of the elytra are incomplete and just marked by a few larger and sparse punctures, but are complete and very distinct in A. tibetanus according to the re-description by Nilsson (1990). Finally, the anal sternite is less roughly puncturate and wrinkled. The median lobe of aedeagus does not allow a clear identification to be made, but the subapical tooth is more elongate (Figs 3 & 4).	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF0FF9FFF38FCEA7129FC76.taxon	description	Habitat. These specimens were collected at an altitude of 5100 m, which may be an altitude record for dytiscids. The habitat was a flat area of debris with water trickling over it, with the source exposed to the south. This appears to be a temporary habitat as the ice-free period at this altitude seems to be quite short.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF0FF9FFF38FCEA7129FC76.taxon	distribution	Distribution. Only known from the type locality but probably more widespread in south central Tibet.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF2FF98FF38FB42714FFE8E.taxon	materials_examined	Material studied. “ Bhutan, Thimpu Prov., Druk Path, marsh near Janye Tsoh, 4100 m, 10.12.2005, A. Bloch ” (3 exs., NMW; 1 ex., NMB). “ China, SE Qinghai road Toramarkog-Nangqén, pass 20 km N NANGQÉN, alpine meadows, 4500 m, Rejsek leg., 25.6.1995 ” (2 exs., CLH).	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF2FF98FF38FB42714FFE8E.taxon	discussion	Remarks. Agabus longissimus Régimbart was described in 1899 on the basis of two male specimens collected in 1897 at “ Su-Tchuen ” by local people. One of these was found in the collections of the Muséum National d’Histoire Naturelle, Paris, and was designated as lectotype (Brancucci 1983). The second specimen mentioned in the original description is not present in the Paris Museum collections. In the same paper, Brancucci (1983) recorded this species from Gnatong (Sikkim) on the basis of a small series of specimens that were also found in the Paris Museum. Wu (1937) recorded this species from Mongolia, and Stastný (1996) reported it from oligotrophic pools in the subalpine and alpine zones of Sichuan and Qinghai. Later, Pederzani (2002) mentioned a further locality in Sichuan, SW of Kanding, leg. L. & R. Businský, 2001, at an altitude between 3200 and 4600 m. The material recorded here and collected a century after the first known specimens shows that this species is apparently widespread but remains very rarely found. It occurs only at very high altitudes, between 3200 and 4600 m, as indicated on the labels of the known specimens. This species undoubtedly belongs to the guttatus - group, characterized by a medial gap in the anterior row of punctures on pronotum (Nilsson 1990). In its form it is very similar to A. biguttatus (Olivier) but is somewhat narrower. One striking characteristic is the coarse upper surface of the females, which seems to be constant and not accidental as it is in many other Agabus - species. For further details, refer to the original description (Régimbart 1899) and to Brancucci (1983).	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF2FF98FF38FB42714FFE8E.taxon	distribution	Distribution. China (Sichuan, Quinghai), Bhutan and India (Sikkim), between 3200 – 4600 m.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF5FF98FF38FE807100FCF8.taxon	materials_examined	Type material studied. Paratype No. 8 (male): “ China Yünnan prov.; ca. 4000 m SE ZHONGDIAN; 99 ° 52 / 27 ° 48 flat valley, slowly flowing stream ca. 2 m wide; sandy-gravely 11. - 12.10.1999, leg. Stastný ” (CLH). Paratype No. 43 (female): “ China Yünnan prov. 30 km SE ZHONGDIAN 99 ° 52 / 27 ° 48; ca. 3900 m a. s. l. ground water pools, sandy-gravel 11. - 12.10.1999, leg. Stastný ” (CLH). Additional material studied. “ China, N-Yunnan, Diqing Tibet Aut. Pref., Deqin Co., Baima Shan, pass 15 km Se Deqin, 4330 m, 28 ° 22.68 N 99.00. 57 E, meadows, Azalea Juniperus leaf litter, grass, moss, sifted, 12. VI. 2005, M. Schülke [C 2005 - 10] ” (1 male, 2 females, CLH).	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF5FF98FF38FE807100FCF8.taxon	discussion	Remarks. This is the first record of the species since its description in 2003. It was only known from the type locality around Zhongdian, in the northern part of Yünnan.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF5FF98FF38FE807100FCF8.taxon	distribution	Distribution. China, northern part of Yünnan Province. At altitudes around 4000 m above sea level.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF5FF98FF38FC0D77C2FA33.taxon	materials_examined	Material studied. “ V. tja. Toso-nora 13.300 ' II. 95 Rob. I Kozl, Paralectotype, Agabus tibetanus Zaitzev, 1908, des. A. Nilsson 1988 ” (1 male, ZISP); “ China, Qinghai prov., Qingshuihe, 1. - 5.7.1992, 4200 m ”, “ Agabus tibetanus Zaitz. Det. Anders Nilsson 93 ” (1 male, CLH).	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF5FF98FF38FC0D77C2FA33.taxon	discussion	Remarks. A very rare species, known only from a very few specimens apart from the type series. The type locality is Toso-noor (Toson-Hu) in the Province Tsinghai (Qinghai) of China, at an elevation of about 4000 m. We were able to study one paralectotype from ZISP. Contrary to the redescription by Nilsson (1990), the male protarsal claw is definitely not longer than protarsomere 5.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF5FF98FF38FC0D77C2FA33.taxon	distribution	Distribution. China (Tibet) at an elevation of around 4000 m.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF5FF9AFF38F9C275BEFE2E.taxon	materials_examined	Material studied. 12 males, 3 females: “ Tibet South Central, Budha Valley N of Yangpachem, about 30 ° 10 ' 38 N 90 ° 30 ' 18 E, 20. VI. 2007, 4750 m ” (12 males, 3 females, NMB, NMW, CLH).	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF5FF9AFF38F9C275BEFE2E.taxon	discussion	Remarks. Just a year after the description of this remarkable species, based on two specimens collected by the Russian entomologist Gorbatchev at Zhangmu, southern Tibet near the Nepalese border, a good series has been found in Tibet near Lhasa. This new record also shows that this species is more widespread in Tibet, and the longer series collected by another non-waterbeetle-specialist suggests that it is also not very rare, at least locally. Female. Similar to male. Anal sternite microreticulate, distinctly wrinkled longitudinally on posterior half and broadly rounded. In the male, the latter is somewhat flattened on the middle.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
03FBE052FFF5FF9AFF38F9C275BEFE2E.taxon	distribution	Distribution. Tibet between 4000 m and 4750 m. Habitat. All the specimens were collected among gravel on the banks of a small brook of melting and source water at 4750 m (Figs 6 & 7). Fifteen specimens were taken, 12 males and 3 females, bringing the sex ratio for this collection to 4: 1. The unusual sex ratio is confirmed by the fact that the two specimens of the type-series were also males.	en	Brancucci, Michel, Hendrich, Lars (2008): 5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae). Zootaxa 1825: 51-58, DOI: 10.5281/zenodo.183097
