taxonID	type	description	language	source
03FB3D2FFFB5FFC6F088FD08E789FDC2.taxon	description	Fig. 1 A – E	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB5FFC6F088FD08E789FDC2.taxon	materials_examined	Type material examined. Holotype ♂ “ LAOS, 1. – 9. ν. 1999, Louangphrabang 20 ° 33 - 4 ’ N 102 ° 14 ’ E Ban Song Cha (5 km W), 1200 m, Vit Kubáň leg. / 1372 Sericini Asia spec. ” (ZFMK). Paratypes: 11 ♂♂, 2 ♀♀ “ LAOS, 1. – 9. ν. 1999, Louangphrabang 20 ° 33 - 4 ’ N 102 ° 14 ’ E Ban Song Cha (5 km W), 1200 m, Vit Kubán leg. ” (ZFMK), 1 ♂ “ NW THAILAND MAE HONG SON 28.4. – 3.5.1992 leg. P. Pacholatko / 1380 Sericini Asia spec. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB5FFC6F088FD08E789FDC2.taxon	description	Description of holotype. Length: 8.9 mm; length of elytra: 6.5 mm; maximum width: 5.6 mm. Dorsal surface dark brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, slightly longer than remaining antennomeres combined. Eyes moderately small, ratio of diameter / interocular width: 0.59. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures and a very few short setae on odd intervals; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.69. Pygidium moderately convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, without longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a small, convex extension at basal third and glabrous. Metatibia moderately short and wide, widest at middle; ratio width / length: 1 / 2.9; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere almost as long as following two tarsomeres combined, a quarter of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 1 A – C. Habitus: Fig. 1 D, E. Variation. Length: 8.9 – 10.3 mm; length of elytra: 6.5 – 7.1 mm; maximum width: 5.6 – 5.7 mm. Female: antennal club in female composed of three antennomeres, distinctly shorter than the remaining antennomeres combined.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB5FFC6F088FD08E789FDC2.taxon	diagnosis	Diagnosis. Tetraserica similis Ahrens, Pacholátko & Pham, sp. nov. differs from the very similar T. pingjiangensis Liu, Fabrizi, Bai, Yang & Ahrens, 2014 by the mesally extremely widened left paramere which has a large triangular extension on its mesodorsal margin (lateral view).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB5FFC6F088FD08E789FDC2.taxon	etymology	Etymology. The name of the new species is derived from the Latin adjective similis (similar), with reference to its strong resemblance with T. pingjiangensis.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB6FFC0F088FD3FE727FE52.taxon	description	Fig. 1 F – J	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB6FFC0F088FD3FE727FE52.taxon	materials_examined	Type material examined. Holotype ♂ “ LAOS-C; KHAM MOUAN pr Ban Khoun Ngeun; ~ 200 m; 18 ° 07 ° N 104 ° 29 ′ E; Pacholátko leg.; 24. – 29. iv. 2001 / 1377 Sericini Asia spec. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB6FFC0F088FD3FE727FE52.taxon	description	Description of holotype. Length: 10.4 mm; length of elytra: 7.4 mm; maximum width: 6.0 mm. Dorsal surface dark brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, as long as remaining antennomeres combined. Eyes small, ratio of diameter / interocular width: 0.56. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures and a very few short setae on odd intervals; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.71. Pygidium weakly convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, without longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, without particular extension at basal third and glabrous. Metatibia moderately short and wide, widest at middle; ratio width / length: 1 / 2.9; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere almost as long as following two tarsomeres combined, a quarter of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 1 H – J. Habitus: Fig. 1 F, G. Female unknown.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB6FFC0F088FD3FE727FE52.taxon	diagnosis	Diagnosis. Tetraserica elargata Ahrens, Pacholátko & Pham, sp. nov. differs from the very similar T. pingjiangensis Liu, Fabrizi, Bai, Yang & Ahrens, 2014 by the apically extremely widened left paramere. The left paramere has a large, blunt, tooth-like extension on its mesodorsal margin shortly before the apex of the paramere (lateral view). Furthermore, in T. elargata Ahrens, Pacholátko & Pham, sp. nov. the blunt extension on the posterior margin of metafemur is less distinct than in T. pingjiangensis.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB6FFC0F088FD3FE727FE52.taxon	etymology	Etymology. The name of the new species is derived from the Latin adjective elargatus (widened), with reference to its apically widened left paramere.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB0FFC2F088FE44E11AFE77.taxon	description	Fig. 2 A – F	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB0FFC2F088FE44E11AFE77.taxon	materials_examined	Type material examined. Holotype ♂ “ S LAOS, Champasak, Ban Itou, 800 m, 10. – 18. iv. 1999, 15 ° 10,4 ’ N, 106 ° 05,8 ’ E, M. Strba leg. / 1376 Sericini Asia spec. ” (ZFMK). Paratypes: 2 ♂♂ “ S LAOS, Champasak, Ban Itou, 800 m, 10. – 18. iv. 1999, 15 ° 10,4 ’ N, 106 ° 05,8 ’ E, E. Jendek & O. Šausa leg. / 1376 Sericini Asia spec. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB0FFC2F088FE44E11AFE77.taxon	description	Description of holotype. Length: 8.3 mm; length of elytra: 6.0 mm; maximum width: 5.2 mm. Dorsal surface dark reddish brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, 1.2 times as long as remaining antennomeres combined. Eyes moderately small, ratio of diameter / interocular width: 0.6. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures and a very few short setae on odd intervals; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.58. Pygidium moderately convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, without longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, straight and glabrous. Metatibia moderately short and wide, widest at middle; ratio width / length: 1 / 3.0; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere as long as following two tarsomeres combined, a third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 2 A – D. Habitus: Fig. 2 E, F. Female unknown. Variation. Length: 8.0 – 8.8 mm; length of elytra: 5.6 – 6.1 mm; maximum width: 4.8 – 5.2 mm. Surface in paratypes darker brown.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB0FFC2F088FE44E11AFE77.taxon	diagnosis	Diagnosis. Tetraserica strbai Ahrens, Pacholátko & Pham, sp. nov. differs distinctly from T. jakli Fabrizi, Dalstein & Ahrens, 2019 by the shape of parameres: the left paramere is not cleft before the apex and has at the middle of its dorsal margin a blunt triangular tooth (lateral view); the mesoapical phallobasal apophysis is more strongly curved at apex having the tip less strongly pointed; the ventral branch of the right paramere has its preapical ventrolateral tooth almost at apex (instead at apical third of basal branch as in T. jakli), and its dorsal branch is shorter and not produced towards the middle (dorsal view).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB0FFC2F088FE44E11AFE77.taxon	etymology	Etymology. The new species is names after one of its collectors, M. Strba.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB2FFC3F088FE60E5CAFEBF.taxon	description	Fig. 2 G – L	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB2FFC3F088FE60E5CAFEBF.taxon	materials_examined	Type material examined. Holotype ♂ “ LAO, Phongsaly prov., 21 ° 41 ’ - 2 ’ N 102 ° 6 ′ - 8 ’ E, 28. v. – 20. vi. 2003, ~ 1500 m PHONGSALY env., P. PACHOLATKO Igt. / 1382 Sericini Asia spec. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB2FFC3F088FE60E5CAFEBF.taxon	description	Description of holotype. Length: 8.0 mm; length of elytra: 6.2 mm; maximum width: 4.9 mm. Body more elongate, dorsal surface dark brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, 1.1 times as long as remaining antennomeres combined. Eyes large, ratio of diameter / interocular width: 0.95. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures, with very minute setae in punctures; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.38. Pygidium strongly convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, without longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, straight and glabrous. Metatibia moderately long and narrow, widest at middle; ratio width / length: 1 / 3.6; basal group of dorsal spines of metatibia shortly before middle, apical one at three quarters of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere as long as following two tarsomeres combined, a third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 2 I – L. Habitus: Fig. 2 G, H. Female unknown.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB2FFC3F088FE60E5CAFEBF.taxon	diagnosis	Diagnosis. Tetraserica misofi Ahrens, Pacholátko & Pham, sp. nov. is in shape of aedeagus most similar to T. jakli Fabrizi, Dalstein & Ahrens, 2019 and T. strbai Ahrens, Pacholátko & Pham, sp. nov. It differs from both by the larger eyes of male and convex pygidium as well as by the shape of aedeagus: the left paramere being composed by a short dorsal and a long ventral branch, the straight, much longer and simply pointed mesoapical phallobasal apophysis, as well as by the much longer basal lobe of the robust right paramere which is strongly developed mesally and which almost exceeds the left side of the aedeagus.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB2FFC3F088FE60E5CAFEBF.taxon	etymology	Etymology. The name species is names after Bernhard Misof, director of the LIB / Museum Koenig, who helped immensely to provide funds for the purchase of the Sericinae collection of Petr Pacholátko by the Museum Koenig, Bonn.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB3FFCCF088FE28E40CFEE3.taxon	description	Fig. 3 A – F	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB3FFCCF088FE28E40CFEE3.taxon	materials_examined	Type material examined. Holotype ♂ “ VS 24 - 0066 / VIETNAM: Quang Tri Pro., Bac Huong Hoa NR-UV 1 - behind guest house, 11. v. 2024, P. V. Pham leg. ” (IBHV). Paratype: 1 ♂ “ VS 24 - 0185 / VIETNAM: Quang Tri Pro., Bac Huong Hoa NR, UV 6 - up to the hill, 10. v. 2024, P. V. Pham leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB3FFCCF088FE28E40CFEE3.taxon	description	Description of holotype. Length: 9.0 mm; length of elytra: 7.4 mm; maximum width: 6.2 mm. Body oval, dorsal surface dark reddish brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, 1.2 times as long as remaining antennomeres combined. Eyes moderately large, ratio of diameter / interocular width: 0.62. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures, on odd intervals moderately concentrated along striae, with very minute setae in punctures; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.45. Pygidium moderately convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, with numerous longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, straight and glabrous. Metatibia moderately short and wide, widest at middle; ratio width / length: 1 / 2.4; basal group of dorsal spines of metatibia at first third, apical one at three quarters of metatibial length; ventrodistal margin strongly truncated, ventral margin one quarter of its length shorter than dorsal one, ventral terminal spine very long, as long as metatibia, and strongly curved. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere as long as following two tarsomeres combined, a quarter of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 3 A – D. Habitus: Fig. 3 E, F. Female unknown. Variation. Length: 9.0 – 10.6 mm; length of elytra: 7.4 – 8.1 mm; maximum width: 6.2 – 6.3 mm.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB3FFCCF088FE28E40CFEE3.taxon	diagnosis	Diagnosis. The species is most similar to T. bachmaensis Fabrizi, Dalstein & Ahrens, 2019, both species share the extremely elongate ventral metatibial spine and the ventroapically truncate metatibia (Fig. 3 E). Tetraserica quangtri Ahrens, Pacholátko & Pham, sp. nov. differs mainly by the shape of right paramere of which the dorsal lobe is strongly curved and of which the apex is ending well distant from that of the ventral lobe (lateral view); the ventral love somewhat longer and strongly curved in apical half rather than being almost straight as in T. bachmaensis.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB3FFCCF088FE28E40CFEE3.taxon	etymology	Etymology. The name of the new species is derived from its occurrence in Quang Tri Province (noun in apposition).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBDFFCDF088FF2CE40CF847.taxon	description	Fig. 3 G – L	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBDFFCDF088FF2CE40CF847.taxon	materials_examined	Type material examined. Holotype ♂ “ VS 24 - 0644 / Vietnam: Son La Pro., Thuan Chau Dist., Copia NP., UV 3, 8. vi. 2024, PV Pham leg. ” (IBHV). Paratypes: 1 ♂ “ VS 24 - 0720 / Vietnam: Son La Pro., Thuan Chau Dist., Copia NP., UV 2, 7. vi. 2024, PV Pham leg. ” (IBHV), 1 ♂ “ VS 24 - 0802 a / Vietnam: Son La Pro., Thuan Chau Dist., Copia NP., UV 1, 6. vi. 2024, PV Pham leg. ” (ZFMK), 1 ♂ “ VS 24 - 0802 b / Vietnam: Son La Pro., Thuan Chau Dist., Copia NP., UV 1, 6. vi. 2024, PV Pham leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBDFFCDF088FF2CE40CF847.taxon	description	Description of holotype. Length: 11.3 mm; length of elytra: 8.1 mm; maximum width: 6.8 mm. Dorsal surface dark brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, as long as remaining antennomeres combined. Eyes moderately small, ratio of diameter / interocular width: 0.65. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.67. Pygidium weakly convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, with numerous longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny, anterior margin acute, ventral posterior margin smooth and only weakly widened in apical half, with a sharp, sharply pointed tooth at basal third, which is a quarter as long as maximum width of metafemur, dorsal posterior margin smooth and glabrous. Metatibia moderately short and wide, widest at middle; ratio width / length: 1 / 3.3; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere little shorter than following two tarsomeres combined, a third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 3 I – L. Habitus: Fig. 3 G, H. Female unknown. Variation. Length: 9.8 – 11.3 mm; length of elytra: 6.6 – 8.1 mm; maximum width: 5.6 – 6.8 mm.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBDFFCDF088FF2CE40CF847.taxon	diagnosis	Diagnosis. Tetraserica sonla Ahrens, Pacholátko & Pham, sp. nov. differs from all other species that share a sharp, tooth-like spine at the ventral posterior margin of metafemur the median lamina of phallobase which is reduced in size and much shorter in size than width of phallobase (in lateral view), being usually not visible. Furthermore, the new species differs by the shape of parameres: the left paramere is composed by a single lobe, while lobes of the right parameres are short and almost of the same length.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBDFFCDF088FF2CE40CF847.taxon	etymology	Etymology. The name of the new species is derived from its occurrence in Son La Province (noun in apposition).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBEFFCEF088FF2CE1EEF8D7.taxon	description	Fig. 4 A – G	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBEFFCEF088FF2CE1EEF8D7.taxon	materials_examined	Type material examined. Holotype ♂ “ THAI-NE, Loei pro PHU KRADUNG N. 16 ° 52 ’ N 101 ° 49 ’ E, 1000 m, 16. – 18. v. 1999, D. Hauck leg. ” (ZFMK). Paratypes: 1 ♂ “ THAI-NE, Loei pro PHU KRADUNG N. 16 ° 52 ’ N 101 ° 49 ’ E, 1000 m, 16. – 18. v. 1999, D. Hauck leg. ” (ZFMK), 1 ♂ “ THAI, NE, Loei prov., Phu Kradung N. P., 1000 m, 16 ° 52 ’ N, 101 ° 49 ’ E, 16. – 17. v. 1999, M. Riha leg. / 1378 Sericini Asia spec. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBEFFCEF088FF2CE1EEF8D7.taxon	description	Description of holotype. Length: 8.8 mm; length of elytra: 6.5 mm; maximum width: 5.5 mm. Body oval, dorsal surface dark brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, as long as remaining antennomeres combined. Eyes moderately large, ratio of diameter / interocular width: 0.67. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures, with very minute setae in punctures; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.64. Pygidium moderately convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, with a few single, long setae at apex. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, straight and glabrous. Metatibia short and moderately wide, widest at middle; ratio width / length: 1 / 3.2; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere as long as following two tarsomeres combined, a third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 4 A – E. Habitus: Fig. 4 F, G. Female unknown. Variation. Length: 8.4 – 8.8 mm; length of elytra: 5.9 – 6.5 mm; maximum width: 4.9 – 5.5 mm.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBEFFCEF088FF2CE1EEF8D7.taxon	diagnosis	Diagnosis. Tetraserica lucieae Ahrens, Pacholátko & Pham, sp. nov. is in shape of male genitalia similar to T. chiangdaoensis Kobayashi, 2017. The new species differs by having an uncleft, short left paramere and a more robust dorsal lobe of the right paramere, which is narrowed only shortly before the apex (and not as in T. chiangdaoensis being narrowed already at middle).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBEFFCEF088FF2CE1EEF8D7.taxon	etymology	Etymology. The new species is names after Petr’s friend, Lucie (noun in genitive singular case).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB8FFC8F088FF2CE666F823.taxon	description	Fig. 4 H – L	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB8FFC8F088FF2CE666F823.taxon	materials_examined	Type material examined. Holotype ♂ “ VS 1047 / VS 1047 Vietnam Kon Tum Pro., Kon Plong Dist., near Giraffe tree 15. v. 2023 14 ° 43 ’ 50 ” N, 108 ° 21 ’ 41 ’’ E P. V. Pham & G. Sabatinelli Tetraserica spnVi 10 / Asia Sericini spec. 1362 ” (IBHV). Paratypes: 1 ♂ “ VS 1053 / VS 1053 Vietnam Kon Tum Pro., Kon Plong Dist., near Giraffe tree 15. v. 2023 14 ° 43 ’ 50 ” N, 108 ° 21 ’ 41 ’’ E P. V. Pham & G. Sabatinelli Tetraserica spnVi 10 ” (ZFMK), 1 ♂ “ VS 1068 / VS 1068 Vietnam Kon Tum Pro., Kon Plong Dist., Thach Nham Forest Stat. 3 13. v. 2023 14 ° 42 ’ 13 ” N, 108 ° 21 ’ 21 ’’ E P. V. Pham & G. Sabatinelli Tetraserica spnVi 9 ” (IBHV).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB8FFC8F088FF2CE666F823.taxon	description	Description of the holotype. Length: 11.0 mm; length of elytra: 8.4 mm; maximum width: 6.8 mm. Dorsal surface dark brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, as long as remaining antennomeres combined. Eyes moderately small, ratio of diameter / interocular width: 0.65. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures and a very few short setae on odd intervals; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.65. Pygidium strongly convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, with a few longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny or dull, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a small, sharp tooth at the middle and few short setae basally. Metatibia moderately short and wide, widest at middle; ratio width / length: 1 / 2.9; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere almost as long as following two tarsomeres combined, a third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 4 J – L. Habitus: Fig. 4 H, I. Female unknown. Variation. Length: 10.5 – 11.2 mm; length of elytra: 8.2 – 8.4 mm; maximum width: 6.7 – 6.8 mm.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB8FFC8F088FF2CE666F823.taxon	diagnosis	Diagnosis. Tetraserica guidosabatinellii Ahrens, Pacholátko & Pham, sp. nov. is in shape of aedeagus most similar to Tetraserica miniatula (Moser, 1915) and T. giulianae Fabrizi, Dalstein & Ahrens, 2019. The new species differs by the much shorter right paramere, of which both branches are of the same length. Furthermore, T. guidosabatinellii Ahrens, Pacholátko & Pham, sp. nov. has the dorsal lobe of the left paramere strongly and abruptly double bent at apex, the latter is slightly shorter than the ventral lobe of the left paramere.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB8FFC8F088FF2CE666F823.taxon	etymology	Etymology. The new species is named after our colleague and chafer specialist, who was one of the collectors of this new species, Guido Sabatinelli (noun in genitive singular).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB9FFC9F088FF2CE072F823.taxon	description	Fig. 5 A – F	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB9FFC9F088FF2CE072F823.taxon	materials_examined	Type material examined. Holotype ♂ “ VS 1051 / VS 1051 Vietnam Kon Tum Pro., Kon Plong Dist., near Giraffe tree 15. v. 2023 14 ° 43 ’ 50 ” N, 108 ° 21 ’ 41 ’’ E P. V. Pham & G. Sabatinelli Tetraserica spnVi 10 / Asia Sericini spec. 1364 ” (IBHV). Paratype: 1 ♂ “ VS 1048 / VS 1048 Vietnam Kon Tum Pro., Kon Plong Dist., near Giraffe tree 15. v. 2023 14 ° 43 ’ 50 ” N, 108 ° 21 ’ 41 ’’ E P. V. Pham & G. Sabatinelli Tetraserica spnVi 10 ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB9FFC9F088FF2CE072F823.taxon	description	Description of the holotype. Length: 10.8 mm; length of elytra: 8.2 mm; maximum width: 6.1 mm. Dorsal surface dark reddish brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, as long as remaining antennomeres combined. Eyes small, ratio of diameter / interocular width: 0.52. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals weakly convex, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures and a very few short setae on odd intervals; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.74. Pygidium moderately convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, with a few longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny or dull, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, with a small, blunt tooth at basal third, posterior margin smooth dorsally, with a few short setae basally. Metatibia short and wide, widest at middle; ratio width / length: 1 / 2.8; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere almost as long as following two tarsomeres combined, a third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 5 A – D. Habitus: Fig. 5 E, F. Female unknown. Variation. Length: 10.8 – 11.3 mm; length of elytra: 8.2 – 8.3 mm; maximum width: 6.1 – 6.9 mm.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB9FFC9F088FF2CE072F823.taxon	diagnosis	Diagnosis. Tetraserica konplong Ahrens, Pacholátko & Pham, sp. nov. is in shape of aedeagus quite similar to T. yongbelar Ahrens, 2023. The new species differs by the straight left parameres (dorsal view) the in basal half much narrower mesoventral apical process of phallobase (lateral view), and the distinctly longer right paramere, of which the portion distal of its bend is twice as long as the portion basal of it. Furthermore, T. konplong Ahrens, Pacholátko & Pham, sp. nov. has a posterior margin with a small, blunt tooth at basal third, which is lacking in T. yongbelar. From the also similar Tetraserica tanahrataensis Fabrizi, Dalstein & Ahrens, 2019, the new species differs by the much longer right paramere, the more robust mesoapical apophysis of phallobase, and the almost straight, deeply split left paramere.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFB9FFC9F088FF2CE072F823.taxon	etymology	Etymology. The new species is named after its occurrence in the Kon Plong district (noun in apposition).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBAFFCBF088F8A8E096F91F.taxon	description	Fig. 5 G – K	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBAFFCBF088F8A8E096F91F.taxon	materials_examined	Type material examined. Holotype ♂ “ VS 1009 / VS 1009 Vietnam Quang Nam Pro., Nam Tra My Dist., Ngoc Linh Gingsen sta. 18. v. 2023 14 ° 59 ’ 10 ” N, 108 ° 03 ’ 08 ’’ E P. V. Pham & G. Sabatinelli Tetraserica spnVi 5 / Asia Sericini spec. 1363 ” (IBHV).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBAFFCBF088F8A8E096F91F.taxon	description	Description of the holotype. Length: 11.0 mm; length of elytra: 8.5 mm; maximum width: 6.5 mm. Dorsal surface dark brown and glabrous, labroclypeus, pronotal margins and ventral surface reddish brown, femora and antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, 1.2 times as long as remaining antennomeres combined. Eyes moderately small, ratio of diameter / interocular width: 0.65. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins densely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures and a very few short setae on odd intervals; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.49. Pygidium moderately convex and dull, densely punctate, without smooth midline, almost glabrous, with a few longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny or dull, anterior margin acute, posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally. Posterior margin of metafemur generally straight or slightly convex. Metatibia short and wide, widest at middle; ratio width / length: 1 / 2.5; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; apex of metatibia strongly truncated ventrally, ventral terminal spine very long and s-shaped, situated subapically at two thirds of metatibial length on ventral margin, ending at the level of distal margin of metatarsomere 1; distal margin extremely deeply truncated ending at the ventral margin at posterior third of metatibial ventral margin. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere slightly shorter than following two tarsomeres combined, one third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 5 I – K. Habitus: Fig. 5 G, H. Female unknown.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBAFFCBF088F8A8E096F91F.taxon	diagnosis	Diagnosis. Tetraserica quangnam Ahrens, Pacholátko & Pham, sp. nov. is in shape of metatibia similar to Tetraserica bartolozzii Ahrens, 2023. The new species, however, differs by the less strongly truncated metatibia of which the ventral terminal spine is situated not at middle of the ventral margin but at posterior third of metatibial ventral margin; furthermore, the shape of parameres differs distinctly between the both species: The right paramere is in the new species strongly hooked at apex, while the left parameres is curved evenly ventrally instead of being reflexed dorsally (as in T. bartolozzii).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBAFFCBF088F8A8E096F91F.taxon	etymology	Etymology. The new species is named after its occurrence in the Quang Nam province (noun in apposition).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBBFFD5F088F893E6B7F9AE.taxon	description	Fig. 6 A – E	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBBFFD5F088F893E6B7F9AE.taxon	materials_examined	Type material examined. Holotype ♂ “ LAOS, Bolikhamxal pr 18 ° 16 ′ N 103 ° 11 E, 70 km NEE Vientian 27. – 30. iv. 1997, 150 m Vit Kuban leg. / 1374 Sericini Asia spec. ” (ZFMK). Paratype: 1 ♂ “ LAOS, Bolikhamxal pr 18 ° 16 ′ N 103 ° 11 E, 70 km NEE Vientian 27. – 30. iv. 1997, 150 m Vit Kuban leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBBFFD5F088F893E6B7F9AE.taxon	description	Description of holotype. Length: 8.4 mm; length of elytra: 6.1 mm; maximum width: 5.2 mm. Dorsal surface dark brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, 1.1 times as long as remaining antennomeres combined. Eyes moderately small, ratio of diameter / interocular width: 0.63. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.66. Pygidium moderately convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, without longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny, anterior margin acute, ventral posterior margin smooth and only weakly widened in apical half, with a sharp, sharply pointed tooth at basal third, which is a quarter as long as maximum width of metafemur, dorsal posterior margin smooth and glabrous. Metatibia moderately short and wide, widest at middle; ratio width / length: 1 / 3.1; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere little shorter than following two tarsomeres combined, a third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 6 A – C. Habitus: Fig. 6 D, E. Female unknown. Variation. Length: 8.1 – 8.4 mm; length of elytra: 6.1 – 6.2 mm; maximum width: 5.1 – 5.2 mm.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBBFFD5F088F893E6B7F9AE.taxon	diagnosis	Diagnosis. This new species differs from the similar T. spinicrus (Frey, 1972) by the shorter ventral lobe of the left paramere, consequently its dorsal and ventral lobe as well as left and right paramere being of the same length; the secondary (more mesal) dorsal lobe of the right paramere is distinctly larger than in T. spinicrus.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFBBFFD5F088F893E6B7F9AE.taxon	etymology	Etymology. The name of the new species is derived from the combined Greek prefix pseudo - (false) and the species name spinicrus, with reference to its similarity to T. spinicrus.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA5FFD7F088F918E4D2FF07.taxon	description	Fig. 6 F – K	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA5FFD7F088F918E4D2FF07.taxon	materials_examined	Type material examined. Holotype: ♂ “ Laos, 21 ° 09 ’ N, 101 ° 19 ’ E, Louangnamtha pr. Namtha-Muang Sing, 5 – 31. v. 1997, 900 – 1200 m Vit. Kubáň leg. / 1384 Sericini Asia spec. ” (ZFMK). Paratypes: 4 ♂♂ “ LAOS north, 5 – 11. V. 1997, 20 km NW Louang Namtha, N 21 ° 09.2, E 101 ° 18.7, alt. 900 ± 100 m, E. Jendek & O. Šauša leg. ” (ZFMK), 7 ♂♂ “ Laos, 21 ° 09 ’ N, 101 ° 19 ’ E, Louangnamtha pr. Namtha-Muang Sing, 5 – 31. v. 1997, 900 – 1200 m Vit. Kubáň leg. ” (ZFMK), 2 ♂♂ “ LAOS north, 5 – 11. V. 1997, 20 km NW Louang Namtha, N 21 ° 09.2, E 101 ° 18.7, alt. 900 ± 100 m, M. Strba & R. Hergovits leg. ” (ZFMK), 1 ♂ “ Laos 1963 Umgeb. Pak Lay / CF ” (CF), 1 ♂ “ X-DA 4778 labcode: VD 098, LAOS, Stupa GH, 5 km W Muang Sing, 750 m, 21.1482 N 101.1711 E, 9. v. – 2. vi. 2011, M. Murzin, O. Shulga leg. Tetraserica spLA _ V 57 / X-DA 4778 ” (ZFMK), 1 ♂ “ X-DA 4793 labcode: VD 099, LAOS, Stupa GH, 5 km W Muang Sing, 750 m, 21.1482 N 101.1711 E, 9. v. – 2. vi. 2011, M. Murzin, O. Shulga leg. Tetraserica 132 / sp LA _ V 57 / sp-LA-V 57 / X-DA 4793 ” (ZFMK), 1 ♂ “ Laos, 21 ° 09 ’ N, 101 ° 19 ’ E, Louangnamtha pr. Namtha-Muang Sing, 5. – 31. v. 1997, 900 – 1200 m Vit. Kubáň leg / coll. P. Pacholátko / LS 55 ” (ZFMK), 1 ♂ “ LAOS north, 5. – 11. V. 1997, 20 km NW Louang Namtha, N 21 ° 09.2, E 101 ° 18.7, alt. 900 ± 100 m, E. Jendek & O. Šauša leg. / Coll. Pacholátko / 183 Sericini Asia spec. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA5FFD7F088F918E4D2FF07.taxon	description	Description of holotype. Length: 8.2 mm; length of elytra: 6.2 mm; maximum width: 5.2 mm. Dorsal surface dark brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, 1.1 times as long as remaining antennomeres combined. Eyes moderately small, ratio of diameter / interocular width: 0.69. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.62. Pygidium strongly convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, with a few long setae at apex. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny, anterior margin acute, ventral posterior margin smooth and only weakly widened in apical half, with a sharp, sharply pointed tooth at basal third, which is a quarter as long as maximum width of metafemur, dorsal posterior margin smooth and glabrous. Metatibia moderately short and wide, widest at middle; ratio width / length: 1 / 2.9; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere as long as following two tarsomeres combined, a quarter of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 6 H – K. Habitus: Fig. 6 F, G. Female unknown. Variation. Length: 7.5 – 8.5 mm; length of elytra: 6.1 – 6.5 mm; maximum width: 4.9 – 5.8 mm.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA5FFD7F088F918E4D2FF07.taxon	diagnosis	Diagnosis. This new species differs from the similar T. pseudospinicrus Ahrens, Pacholátko & Pham, sp. nov. by the longer ventral lobe of the left paramere, its dorsal lobe is mesally strongly widened and the teethed dorsal margin is ending abruptly at middle by a deep concave emargination of the dorsal margin; the dorsal lobe of the right paramere is simple (not divided in two smaller lobes) and distinctly larger than in T. spinicrus and T. pseudospinicrus Ahrens, Pacholátko & Pham, sp. nov., being almost half as long as the ventral lobe.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA5FFD7F088F918E4D2FF07.taxon	etymology	Etymology. The name of the new species is derived from the combined Greek prefix para- (close by) and the species name spinicrus, with reference to its similarity to T. spinicrus.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA5FFD7F088F918E4D2FF07.taxon	discussion	Remark. This new species includes most of the northern and western Laos specimens previously referred to T. spinicrus (Frey) (Fabrizi et al. 2019). The latter species is actually known only from its type locality in southern Laos. The status of only one specimen from western Laos (1 ♂ “ Haut Mekong Vien Poukha 3. V. 1918 R. V. de Salvazza ” (NHMUK )), cited by Fabrizi et al. (2019) as T. spinicrus, remains uncertain, since we could not reexamine it for this study.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA7FFD0F088FEF0E1FEFF4F.taxon	description	Fig. 7 A – F	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA7FFD0F088FEF0E1FEFF4F.taxon	materials_examined	Type material examined. Holotype ♂ “ LAOS-NE; HUA PHAN prov 25 km SE Vieng Xai (by road) BAN KANGPABONG env.; 20 ° 19 ′ N 104 ° 25 ′ E; D. Hauck leg.; 14. – 18. v. 2001 / 1370 Sericini Asia spec. ” (ZFMK). Paratypes: 3 ♂♂ “ LAOS-NE; HUA PHAN prov 25 km SE Vieng Xai (by road) BAN KANGPABONG env.; 20 ° 19 ′ N 104 ° 25 ′ E; D. Hauck leg.; 14. – 18. v. 2001 / 1370 Sericini Asia spec. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA7FFD0F088FEF0E1FEFF4F.taxon	description	Description of the holotype. Length: 8.4 mm; length of elytra: 6.4 mm; maximum width: 5.8 mm. Dorsal surface dark brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, 1.2 times as long as remaining antennomeres combined. Eyes large, ratio of diameter / interocular width: 0.76. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures and a very few short setae on odd intervals; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.51. Pygidium strongly convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, without longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny or dull, anterior margin acute, posterior margin smooth ventrally and straight, only weakly widened in apical half, posterior margin smooth dorsally, glabrous basally. Metatibia moderately short and wide, widest at middle; ratio width / length: 1 / 3.2; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere almost as long as following two tarsomeres combined, a third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 7 A – D. Habitus: Fig. 7 E, F. Female unknown. Variation. Length: 8.1 – 9.0 mm; length of elytra: 6.0 – 6.5 mm; maximum width: 5.1 – 5.8 mm.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA7FFD0F088FEF0E1FEFF4F.taxon	diagnosis	Diagnosis. Tetraserica haucki Ahrens, Pacholátko & Pham, sp. nov. differs from T. damaidiensis Liu, Fabrizi, Bai, Yang & Ahrens, 2014 by the distinctly wider (dorsal view) right paramere (lateral view), of which the apex is mesally more strongly narrowly produced (dorsal view); the right paramere is in T. haucki at middle and apex twice as wide as in T. damaidiensis and its sharp basal dorsal tooth is in the new species situated slightly more basally.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA7FFD0F088FEF0E1FEFF4F.taxon	etymology	Etymology. The species is named after its collector, D. Hauck (noun in genitive case singular).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA0FFD0F088FEB8E064F823.taxon	description	Fig. 7 G – L	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA0FFD0F088FEB8E064F823.taxon	materials_examined	Type material examined. Holotype ♂ “ LAOS, 1. – 16. v. 1999 Louangphrabang pr 20 ° 33 - 4 ′ N 102 ° 14 ′ E Ban Song Cha (5 km W), 1200 m, Vit Kubáň leg. / 1371 Sericini Asia spec. ” (ZFMK). Paratype: 1 ♂ “ LAO, Phongsaly prov., 21 ° 41 ’ - 2 ’ N 102 ° 6 ′ - 8 ’ E, 28. v. – 20. vi. 2003, ~ 1500 m PHONGSALY env., P. PACHOLATKO Igt. / 1381 Sericini Asia spec. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA0FFD0F088FEB8E064F823.taxon	description	Description of the holotype. Length: 8.4 mm; length of elytra: 6.0 mm; maximum width: 5.0 mm. Dorsal surface dark brown and glabrous, frons and pronotum with weak greenish shine, labroclypeus, ventral surface and legs yellowish, partly reddish brown, antenna yellow. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye twice as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, 1.2 times as long as remaining antennomeres combined. Eyes moderately large, ratio of diameter / interocular width: 0.7. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and convex, widest shortly before base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals flat, with fine and almost evenly dense punctures weakly concentrated along striae, with very minute setae in punctures; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.51. Pygidium moderately convex in apical half and dull, densely punctate, without smooth midline, almost glabrous, without longer setae along apical margin. Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny or dull, anterior margin acute, posterior margin smooth ventrally and straight, only weakly widened in apical half, posterior margin smooth dorsally, glabrous basally. Metatibia moderately short and wide, widest at middle; ratio width / length: 1 / 3.0; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; distal margin not truncated, terminal spines of normal length. Tarsomeres dorsally smooth, with fine, dense setae ventrally on distal half, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere almost as long as following two tarsomeres combined, a quarter of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 7 I – L. Habitus: Fig. 7 G, H. Female unknown. Variation. Length: 8.4 – 8.6 mm; length of elytra: 6.0 – 6.2 mm; maximum width: 5.0 – 5.2 mm.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA0FFD0F088FEB8E064F823.taxon	diagnosis	Diagnosis. Tetraserica schnelli Ahrens, Pacholátko & Pham, sp. nov. differs from T. semidamaidiensis Fabrizi, Dalstein & Ahrens, 2019 by the entirely straight left paramere (lateral view), and the longer and less strongly curved right paramere (lateral view) of which the preapical lateral tooth is less distinct and its mesal lateral tooth is directed distally instead of basally (dorsal view) (as in T. semidamaidiensis).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA0FFD0F088FEB8E064F823.taxon	etymology	Etymology. The species is dedicated to Michael Schnell (Museum Koenig Bonn) who helped with the construction of light traps which are in use in our Vietnam biodiversity project (noun in genitive case singular).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFA2FFD2F088FF2CE745FE56.taxon	discussion	Remark. The name is here emended, since its etymology is based on the species name of T. damaidiensis Liu, Fabrizi, Bai, Yang & Ahrens, 2014 which was in the respective text of etymology passage (Fabrizi et al. 2019: 139) mistakenly misspelled (damadiensis). In the etymology of T. damaidiensis it was clearly stated that the latter name was based on its type locality, Damaidi.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFAAFFDAF088FE06E126FD99.taxon	materials_examined	Material examined. 24 ♂♂ “ LAOS, 1. - 16. v. 1999 Louangphrabang pr. 20 ° 33 - 4 ’ N 102 ° 14 ’ E, Ban Song Cha (5 km W), 1200 m, Vit Kuban leg. ” (ZFMK), 1 ♂ “ LAOS, 24. - 30. iv. 1999 Louangphrabang pr. 20 ° 33 - 4 ’ N 102 ° 14 ’ E, Ban Song Cha (5 km W), 1200 m, Vit Kuban leg. ” (ZFMK), 2 ♂♂ “ LAO, Phongsaly prov., 21 ° 41 ’ - 2 ′ N 102 ° 6 ’ - 8 ’ E, 28. v. – 20. vi. 2003, ~ 1500 m, PHONGSALY env., P. PACHOLATKO Igt. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFAAFFDAF088FCBAE46BFC79.taxon	materials_examined	Material examined. 8 ♂♂ “ NW-THAILAND. 2. - 6.5. Chiang Mai distr. 1996 Dai-Pui vill; 18 ° 49 ′, 98 ° 54 ’ J. Horák lgt.: 1600 m ” (ZFMK), 5 ♂♂ “ Thai, Chiang Mai prov., 18 ° 49 ’ N, 98 ° 54 ’ E, 1600 m, DOI PUI mt., 2. – 6. v., Vít Kubáň leg. 1996 ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFAAFFDAF088FD2EE025FD2D.taxon	materials_examined	Material examined. 1 ♂ “ BURMA; SW SHAN State: Taunggyi; 1. – 18. vi. 1997; J. Kaláb leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFAAFFDAF088FC0EE66FFBC5.taxon	materials_examined	Material examined. 10 ♂♂ “ LAOS, 21 ° 09 ′ N 101 ° 19 ’ E Louangnamtha pr. Namtha -> Muang Sing 5. – 31. v. 1997, 900 – 1200 m Vit Kubán leg ” (ZFMK), 4 ♂♂ “ LAOS north, 5. – 11. v. 1997, 20 km NW Louang Namtha N 21 ° 09.2 ’, E 101 ° 18.7 ’, alt. 900 ± 100 m E. Jendek & O. Šauša leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFAAFFDAF088FBD2E084FB59.taxon	materials_examined	Material examined. 1 ♂ “ THAI: 29. IV. 1993, PAI City; Pacholatko & Dembicky leg. / coll. Pacholátko ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFAAFFDAF088FB6EE49FFAC9.taxon	materials_examined	Material examined. 3 ♂♂ “ Thai-N 1. – 15. v. 1998, Chiang Mai prov., 19 ° 19 ’ N, 98 ° 50 ’ E, SAN PAKIA, 1400 m, Vít Kubáň leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFAAFFDAF088FADEE7BAF93D.taxon	materials_examined	Material examined. 2 ♂♂ “ LAOS-C; KHAM MOUAN pr. Ban Khoun Ngeun; ~ 200 m; 18 ° 07 ′ N 104 ° 29 ′ E; Pacholátko leg.; 24. – 29. iv. 2001 ” (ZFMK), 6 ♂♂ “ LAOS, 1. – 18. v. 2001. Bolikhamxai prov., 18 ° 21 ’ N 105 ° 08 ’ E, Ban Nape (8 km NE), ~ 600 m, Vit Kubáň leg. ” (ZFMK), 5 ♂♂ “ LAOS centr., Bolikhamsai prov. BAN NAPE-Kaew Nua Pass 18.4. – 1.5.1998, alt. 600 ± 100 m, N 18 ° 22.3, E 105 ° 09.1 (GPS), M. Strba & R. Hergovits leg. ” (ZFMK), 16 ♂♂ “ LAOS centr., Bolikhamsai prov. BAN NAPE-Kaew Nua Pass, 18.4. – 1.5.1998, alt. 600 ± 100 m, N 18 ° 22.3, E 105 ° 09.1 (GPS), E. Jendek & O. Šauša leg. ” (ZFMK), 2 ♂♂ “ LAOS centr., Khammouan prov. NAKAI env., 4. – 8.5.1998, Route No 8, alt. 560 ± 20 m N 17 ° 42.8, E 105 ° 08.9 (GPS), E. Jendek & O. Sauša leg. ” (ZFMK), 2 ♂♂ “ LAOS centr., Khammouan prov. NAKAI env., 4. – 8.5.1998, Route No 8, alt. 560 ± 20 m N 17 ° 42.8, E 105 ° 08.9 (GPS), M. Strba & R. Hergovits leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFAAFFDAF088F94AE715F8AD.taxon	materials_examined	Material examined. 1 ♂ “ S Vietnam, 1. – 5.5.1994 Nam Cat Tien Nat. park P. Pacholátko & L. Dembicky ́ leg. / coll. P. Pacholátko / VS 118 ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFAAFFDBF088F83AE043FF2B.taxon	materials_examined	Material examined. 3 ♂♂ “ LAOS, Hua Phan prov., 2004 20 ° 13 ’ N 103 ° 59 ′ E, 6. – 18. v. Ban Saluei, Phu Phan Mt. env. Petr Kresl leg .. alt. 1500 m ” (ZFMK), 1 ♂ “ LAOS-NE; HUA PHAN prov. BAN SALUEI; Phu Phan Mt., 20 ° 15 ′ N 104 ° 02 ′ E; 1500 – 2000 m; D. Hauck leg.; 26. iv. – 11. v. 2001 ” (ZFMK), 1 ♂ “ LAOS-NE: HUA PHAN prov.: BAN SALUEI; Phu Phan Mt. 20 ° 15 ′ N 104 ° 02 ′ E; 1500 – 2000 m; J. Bezděk leg.; 26. iv. – 11. v. 2001 ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFABFFDBF088FF46E4AEFE35.taxon	materials_examined	Material examined. 8 ♂♂ “ N VIETNAM, 21,27 N 105,39 E 70 km NW Hanoi, Tam Dao 9. – 19. v. 1996, 900 – 1200 m Pacholátko & Dembický / VS 82 ” (ZFMK), 1 ♂ “ N VIETNAM, 21,35 N 106,30 E 52 km SW of Lang Son, 27. iv. – 6. v. 1996, 370 m Pacholátko & Dembický leg. ” (ZFMK), 4 ♂♂ “ N VIET NAM (Tonkin) pr. Vinh Phu 1990 TAM DAO 17. – 21. V. P. Pacholátko leg. ” (ZFMK), 4 ♂♂ “ N VIETNAM (Tonkin) pr. Vinh Phu 1990 TAM DAO 6. – 9. V. P. Pacholátko leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFABFFDBF088FE4FE124FDA8.taxon	materials_examined	Material examined. 1 ♂ “ CHINA: Guangxi, Fangchenggang, Naliang, Tansan Vill. AEdz " 21.666031 ° N, 107.734480 ° E Shiwandashan Mount Range 325 m 2023 V. 15 native leg. ” (SCAU).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFABFFDBF088FD3FE5EEFD18.taxon	materials_examined	Material examined. 1 ♂ “ THAILAND bor. 1997 FANG, THA TON 26. V Lgt. M. Snížek / 1379 Sericini Asia spec. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFABFFDBF088FC73E5EEFB8C.taxon	materials_examined	Material examined. 1 ♂ “ NW THAILAND MAE HONG SON 28.4. – 3.5.1992 leg. P. Pacholatko ” (ZFMK), 1 ♂ “ THAI-N, Nan prov., 19 ° 13 ’ N 101 ° 7 ’ E, Doi Phukha N. P. Headq. 22. – 26. iv. 1999, ca 1500 m D. Hauck leg. ” (ZFMK), 3 ♂♂ “ THAILAND, Nan prov., Doi Phuka Nat. Park, 28. iv. – 12. v. 2002 P. Průdek & M. Oboril leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFABFFDBF088FCAFE482FC64.taxon	materials_examined	Material examined. 3 ♂♂ “ Thai, Chiang Mai prov., 18 ° 49 ’ N, 98 ° 54 ’ E, 1600 m, DOI PUI mt., 2 – 6. V., Vít Kubáň leg. 1996 ” (CPPB), 3 ♂♂ “ NW-THAILAND. 2. - 6.5. Chiang Mai distr. 1996 Doi-Pui vill; 18 ° 49 ’, 98 ° 54 ’ J. Horák lgt.; 1600 m ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFABFFDBF088FB1BE5EEFA00.taxon	materials_examined	Material examined. 8 ♂♂ “ NW Thailand 9. - 16. v. MAE HONG SON 199 Ban Huai Po 1600 m leg. P. Pacholatko ” (ZFMK), 2 ♂♂ “ THAI-N, 1 – 15. v. 199, Chiang Mai prov. SAN PAKIA vill., 19.19 N 98.50 E, 1400 m, Vít Kubáň leg. ” (ZFMK), 1 ♂ “ THAI-N, 1. – 19.5.1998 Chiang Mai prov., BAN SAN PAKIA Bednařik leg., 1400 m ” (ZFMK), 1 ♂ “ THAI, Mae Hong Son prov., 19 ° 27 ’ N 98 ° 20 ’ E, 1500 m. SOPPONG, 7. – 12. v. Vit Kubán leg., 1996 ” (ZFMK), 1 ♂ “ THAI 28 / 6.1993 14.01 N 99.31 E KANCHANABURI 50 m Vit Kuban leg. ” (ZFMK), 1 ♂ “ THAILAND, 1000 – 1600 m CHIANG MAI, 26. vi. – 9. v. 1996, 18 ° 49 ’ N 98 ° 54 ’ E, DOI PUI mt. lgt. S. Becvar ” (ZFMK), 1 ♂ “ THAI, N, Mae Hong Son prov., SE of Soppong, 1500 m, 19 ° 27 ’ N, 98 ° 20 ’ E. 23 – 27. ν. 1999, D. Hauck leg. ” (ZFMK), 1 ♂ “ THAILAND, 1000 – 1600 m CHIANG MAI, 26. iv. – 9. v. 1996, 18 ° 49 ’ N 98 ° 54 ’ E, DOI PUI mt. Igt. S. Becvar ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFABFFDBF088F997E618F928.taxon	materials_examined	Material examined. 2 ♂♂ “ N VIET NAM pr. Vinh Phu TAM DAO (Tonkin) 1990 17. – 21. v. P. Pacholátko leg. ” (ZFMK), 2 ♂♂ “ LAOS-NE: HUA PHAN prov.: BAN SALUEI; Phu Phan Mt. 20 ° 15 ′ N 104 ° 02 ′ E; 1500 – 2000 m; J. Bezděk leg.; 26. iv. – 11. v. 2001 ” (ZFMK), 1 ♂ “ LAOS, Hua Phan prov., 2004 20 ° 13 ’ N 103 ° 59 ′ E, 6. – 18., Ban Saluei, Phu Phan Mt. env Petr Kresl leg., alt. 1500 m ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFABFFDBF088F8BFE6B4F898.taxon	materials_examined	Material examined. 11 ♂♂ “ S-Vietnam, 14.10 N 108.30 E 40 Km NW of An Khe Buon Luoi, 620 – 750 m 28.3 – 12.4.1995 Pacholátko & Dembicky ́ leg. / (coll. Pacholátko) ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFABFFE4F088F82FE777FF4F.taxon	materials_examined	Material examined. 1 ♂ “ MYANMAR Shan State ca. 35 km N Aungban 20 ° 55.20 ′ N 96 ° 33.60 E 31.5. – 8.6.2002, ca. 1320 m at light, leg. Schillhammer (81 C) ” (NHMW).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFFABFFE4F088F82FE777FF4F.taxon	discussion	Remarks. First record for Myanmar.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF94FFE4F088FF62E42AFE11.taxon	materials_examined	Material examined. 6 ♂♂ “ LAOS Bolikhamxai pr. 18 ° 16 ’ N, 103 ° 11 ’ E 70 Km NEE Vientiane 27. – 30. iv. 1997, 150 m, Vít Kubáň leg. / coll. P Pacholátko ” (ZFMK), 6 ♂♂, 1 ♀ “ LAOS central, Saisombun zone PHOU KHAO KHOUAY Nat. Park TAD LEUK 18 ° 23 ′ N, 103 ° 04 ’ E 15. – 21. v. 2001, alt. 150 – 200 m E. Jendek & O. Šauša leg. ” (ZFMK), 4 ♂♂, 1 ♀ “ LAOS centr., Bolikhamsai prov., BAN NAPE-Kaew Nua Pass 18.4. – 1.5.1998, alt. 600 ± 100 m N 18 ° 22.3, E 105 ° 09.1 (GPS), E. Jendek & O. Šauša leg. / Coll. P. Pacholátko Invt. No. ” (ZFMK), 1 ♂ “ LAOS centr., Khammouan prov. NAKAI env., 17 ° 43 ’ N, 105 ° 09 ’ E 22. v. – 8. vi. 2001, alt. 500 – 600 m. E. Jendek & O. Šauša leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF94FFE4F088FDACE752FD60.taxon	materials_examined	Material examined. 9 ♂♂, 5 ♀♀ “ THAILAND, Nan prov. Doi Phuka Nat. Park, 28. iv. – 12. v. 2002 P. Průdek & M. Oboril leg. ” (ZFMK), 1 ♂ “ LAOS, 1. – 16. v. 1999 Louangphrabang pr. 20 ° 33 - 4 ’ N 102 ° 14 ’ E Ban Song Cha (5 km W), 1200 m, Vit Kuban leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF94FFE4F088FD7DE70EFCD6.taxon	materials_examined	Material examined. 1 ♂ “ LAOS, 21 ° 09 ’ N 101 ° 19 E, Louangnamtha pr. Namtha-> Muang Sing, 5 – 31. v. 1997, 90 – 1200 m Vit Kubán leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF94FFE4F088FCEAE694FA55.taxon	materials_examined	Material examined. 4 ♂♂ “ THAI-N, 1. - 19.5.198, Chiang Mai prov. BAN SAN PAKIA Bednařík leg., 1400 m ” (ZFMK), 4 ♂♂ “ THAI, N, Mae Hong So prov., SE of Soppong, 1000 m, 19 ° 27 ′ N 98 ° 20 ’ Ε. 23. – 27. v. 1999. M. Riha leg. ” (ZFMK), 12 ♂♂ “ THAI, N, Mae Hong Son prov., SE of Soppong, 1500 m, 19 ° 27 ’ N, 98 ° 20 ’ E. 23. – 27. ν. 1999, D. Hauck leg. (ZFMK), 7 ♂♂ “ THAI 28 / 6.1993 14.01 N 99.31 E KANCHANABURI 50 m Vít Kubáň leg. ” (ZFMK), 3 ♂♂ “ THAI, Chiang Mai prov. 18 ° 49 ′ N 98 ° 54 ′ E, 1600 m DOI PUI mt., 2. – 6. V. Vit Kubán leg., 1996 ” (ZFMK), 6 ♂♂ “ THAI, Mae Hong Son prov. 19 ° 27 ’ N 98 ° 20 ′ E, 1500 m, SOPPONG, 7. – 12. V. Vit Kubán leg., 1996 ” (ZFMK), 7 ♂♂ “ THAI, 2. – 6. v. 1996 Chiang Mai prov 18 ° 49 N 98 ° 54 ’ E, DOI PUI mt., 1600 m, Vit Kubán leg. ” (ZFMK), 7 ♂♂ “ THAI-N, 1. – 15. v. 1998, Chiang Mai prov., SAN PΑΚΙΑ vill., 19.19 N 98.50 E, 1400 m, Vit Kubán leg. ” (ZFMK), 5 ♂♂ “ NW-THAILAND. 2. - 6.5 Chiang Mai distr. 1996 Doi-Pui vill; 18 ° 49 ’, 98 ° 54 ’ J. Horák Igt.; 1600 m ” (ZFMK), 6 ♂♂ “ THAI 2 – 3 / 6 1995 18.49 N 98.54 E DOI PUI 1400 m Vít Kubáň leg. ” (ZFMK), 1 ♂ “ NW THAILAND, 25. iv. – 7. v. Chiang Mai prov., BAN SAN PAKIA 1996, Sv. Bílý leg., 1700 m ” (ZFMK), 7 ♂♂, 4 ♀♀ “ THAI 28 – 31 / 5 1995 19.27 N 98.20 E SOPPONG 1500 m Vit Kubáň leg. ” (ZFMK), 9 ♂♂ “ NW Thailand 9. – 16. v. MAE HONG SON 1991 Ban Huai Po 1600 m leg. P. Pacholatko ” (ZFMK), 15 ♂♂ “ THAI 1 – 8. v. 1993 SOPPONG PAI 1800 m Pacholatko & Dembicky leg. ” (ZFMK), 1 ♂ “ THAI-N. 1. – 19. v. 1998 Chiang Mai prov., BAN SANPAKIA, 1400 m Ivo Martinŭ leg. ” (ZFMK), 1 ♂ “ THAILAND, 1000 – 1600 m CHIANG MAI, 26. iv. – 9. v. 1996, 18 ° 49 ’ N 98 ° 54 ’ E, DOI PUI mt. lgt. S. Becvar ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF94FFE4F088FA68E5EEF9C8.taxon	materials_examined	Material examined. 4 ♂♂ “ NW Thailand 9. - 16. V MAE HONG SON 199 Ban Huai Po 1600 m leg. P. Pacholatko ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF94FFE4F088F9E5E6CDF91A.taxon	materials_examined	Material examined. 1 ♂ “ NW THAI 30. iv- 4.7.199 MAE HONG SON BAN HUSI PO 1600 – 2000 m J. HORÁK LEG. ” (ZFMK), 2 ♂♂ “ NW THAILAND 24. – 27.4. 1991 CHOM THONG LEG. PACHOLÁTKO ” (ZFMK), 1 ♂ “ THAILAND bor. 1997 FANG, THA TON 26. V Lgt. M. Snížek ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF94FFE4F088F8B6E703F891.taxon	materials_examined	Material examined. 3 ♂♂ “ LAOS, 1. – 16. v. 1999 Louangphrabang pr 20 ° 33 - 4 N 102 ° 14 ′ E Ban Song Cha (5 km W), 1200 m, Vit Kubáň leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF94FFE4F088F82CE5EEF804.taxon	materials_examined	Material examined. 1 ♂ “ Thai 21 – 26. V. 1993, Namuang Pacholátko & Dembicky leg. / coll. P. Pacholátko / TS 92 ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF95FFE5F088FF2CE17BFF07.taxon	materials_examined	Material examined. 5 ♂♂ “ Laos centr., Khammouan prov. NAKAI env., 4 – 8.5.1998, Route No 8 alt. 560 ± 20 m, N 17 ° 42.8 ’, E 105 ° 08.9 (GPS), E. Jendek & O. Šauša leg. / Coll. P Pacholátko ” (CPPB).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF95FFE5F088FE9CE4CDFE77.taxon	materials_examined	Material examined. 1 ♂ “ Laos Bolikhamxai pr., 18 ° 16 ’ N, 103 ° 11 ’ E, 70 km NEE Vientiane, 27. – 30. iv. 1997, 150 m, Vít Kubáň leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF95FFE5F088FE0CE67DFDE7.taxon	materials_examined	Material examined. 2 ♂♂ “ LAOS-NE. HUA PHAN prον. 25 km SE Vieng Xai (by road); BAN KANGPABONG env. 20 ° 19 ′ N 104 ° 25 ′ E; D. Hauck leg.; 14. – 18. v. 2001 ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF95FFE5F088FDFCE177FCEB.taxon	materials_examined	Material examined. 5 ♂♂, 9 ♀♀ “ S LAOS, Attapu, Nong Lom (lake), 18. – 30. iv. 1999 800 m, 15 ° 02 N, 106 ° 35 ’ E, E Jendek & O Šauša leg. / Coll. P Pacholátko ” (ZFMK), 2 ♂♂ “ S LAOS, Champasak, Ban Itou, 800 m, 10. – 18. iv. 1999 15 ° 10,4 ’ N, 106 ° 05,8 ’ E, E. Jendek & O. Šauša leg. ” (ZFMK), 3 ♂♂, 3 ♀♀ “ S LAOS, Champasak, Ban Itou, 800 m, 10. – 18. iv. 1999, 15 ° 10,4 ’ N, 106 ° 05,8 ’ E, M. Strba leg. ” (ZFMK), 4 ♂♂, 8 ♀♀ “ S LAOS, Attapu, Nong Lom (lake), 18. – 30. iv. 1999 800 m, 15 ° 02 N, 106 ° 35 ’ E, M. Strba leg. / Coll. P Pacholátko ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF95FFE5F088FCF8E746FC37.taxon	materials_examined	Material examined. 2 ♂♂ “ N Vietnam, (Tonkin) pr. Vinh Phu 1990 Tam Dao, 6. – 9. v. P. Pacholátko leg. / coll. P. Pacholátko / VS 80 ” (ZFMK), 1 ♂ “ N VIETNAM, 21,35 N 106,30 E 52 km SW of Lang Son, 27. iv. – 6. v. 1996, 370 m Pacholátko & Dembický leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF95FFE5F088FC4CE02EFB5C.taxon	materials_examined	Material examined. 1 ♂ “ LAO, Phongsaly prov., 21 ° 41 ’ - 2 ′ N 102 ° 6 ’ - 8 ’ E, 28. v. – 20. vi. 2003, ~ 1500 m, PHONGSALY env., P. PACHOLATKO Igt. / 1375 Sericini Asia spec. ” (ZFMK), 5 ♂♂ “ LAO, Phongsaly prov., 21 ° 41 ’ - 2 ′ N 102 ° 6 ’ - 8 ’ E, 28. v. – 20. vi. 2003, ~ 1500 m, PHONGSALY env., P. PACHOLATKO Igt. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF95FFE5F088FC4CE02EFB5C.taxon	discussion	Remarks. This species was originally described from Vietnam. It is for first time recorded from Laos.	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF95FFE5F088FB74E060FA87.taxon	materials_examined	Material examined. 4 ♂♂ “ LAOS-NE: HUA PHAN prov.: BAN SALUEI; Phu Phan Mt. 20 ° 15 ′ N 104 ° 02 ′ E; 1500 – 2000 m; J. Bezděk leg.; 26. iv. – 11. v. 2001 ” (ZFMK), 1 ♂ “ LAOS-NE: HUA PHAN prov.: BAN SALUEI; Phu Phan Mt. 20 ° 15 ′ N 104 ° 02 ′ E; 1500 – 2000 m; D. Hauck leg.; 26. iv. – 11. v. 2001 ” (ZFMK), 1 ♂ “ LAOS, Hua Phan prov., 2004 20 ° 13 ′ N 103 ° 59 ′ E, 6. – 18. v. Ban Saluei, Phu Phan Mt. env J. Bezděk leg., 1300 – 2000 m ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF95FFE6F088FA1CE124FE2F.taxon	materials_examined	Material examined. Thailand. 4 ♂♂ “ NE THAI NAN distr. BAN PHA KHAP 15. – 20. v. 1992 leg. P. Pacholatko ” (ZFMK), 2 ♂♂ “ THAI 11 – 15. v. 1993 NAN-PHA KHAB Pacholatko & & Dembicky leg. ” (ZFMK), 2 ♂♂ “ THAILAND bor- 1997 FANG, THA ΤΟΝ 26. v. Lgt. K. Snížek ” (ZFMK), 1 ♂ “ THAI, NE, Loei prov., Phu Kradung N. P., 1000 m, 16 ° 52 ’ N, 101 ° 49 ′ E, 16. – 17. v. 1999, M. Riha leg. ” (ZFMK), 7 ♂♂, 1 ♀ “ THAI-NE, Loei prov., PHU KRADUNG N. P. 16 ° 52 ’ N 101 ° 49 ° E. 16. – 18 v. 1999. D. Hauck leg. 1000 m ” (ZFMK), 1 ♂, 2 ♀♀ “ NW THAILAND 24. – 27.4. 1991 CHOM THONG LEG. PACHOLÁTKO ” (ZFMK), 7 ♂♂ “ THAI 21. – 26. v. 1993 NAMUANG Pacholatko & Dembicky leg. ” (ZFMK). Vietnam. 12 ♂♂ “ VIETNAM N SaPa- 1530 m 25. v. – 9. vi. 1991 Strnad Jan lgt. ” (ZFMK), 2 ♂♂ “ N VIETNAM Cuc Phuong Nat. Park 21. – 22. v. 1996 Pacholátko & Dembický leg. ” (ZFMK), 2 ♂♂, 3 ♀♀ “ N VIET NAM (Tonkin) HA NOI (city) 4. – 5. V. 1990 P. Pacholátko leg. ” (ZFMK), 1 ♂ “ N VIETNAM, 21,27 N 105,39 E 70 km NW Hanoi, Tam Dao 9. – 19. v. 1996, 900 – 1200 m Pacholátko & Dembický ” (ZFMK), 1 ♂ “ VIETNAM SA-PA 26.5.1991 Dr. Křeček lgt. ” (ZFMK), 7 ♂♂, 13 ♀♀ “ VIETNAM N. TONKIN CUC-PHUONG NAT. PARK 2. – 12. v. 1991 E JENDEK LEG. ” (ZFMK), 1 ♂ “ NW Vietnam, 6.1990 Hoang Lien Son dist. Sa Pa, 1200 m Duong Tat Tu leg. ” (ZFMK). Laos. 26 ♂♂ “ LAOS, 21 ° 09 ’ N 101 ° 19 E, Louangnamtha pr. Namtha-> Muang Sing, 5. – 31. v. 1997, 90 – 1200 m Vit Kubán leg. ” (ZFMK), 3 ♂♂ “ LAOS north, 13. – 24. V. 1997 15 km NW Louang Namtha N 21 ° 07.5, E 101 ° 21.0, alt. 750 ± 100 m E. Jendek & O. Šauša leg. ” (ZFMK), 8 ♂♂ “ LAOS north, 24. – 30. V. 1997, 20 km NW Louang Nam ha N 21 ° 09.2 ’, E 101 ° 18.7 », alt. 900 ± 100 m E. Jendek & O. Sauša leg. ” (ZFMK), 2 ♂♂ “ LAOS, 31. v. 1997 Louangnamtha pr. 21 ° 00 ’ N 101 ° 25 ′ E LOUANG NAMTHA, Vít Kubáň leg., 600 m ” (ZFMK), 1 ♂ “ LAO, Phongsaly prov., 21 ° 41 ’ - 2 ’ N 102 ° 6 ′ - 8 ’ E, 28. v. – 20. vi. 2003, ~ 1500 m PHONGSALY env., P. PACHOLATKO Igt. ” (ZFMK), 1 ♂ “ LAOS north, 5. – 11. v. 1997 20 km NW Louang Namtha N 21 ° 09.2, E 101 ° 18.7. alt. 900 ± 100 m. M. Strba & R. Hergovits leg. ” (ZFMK), 1 ♂ “ LAOS-NE; HUA PHAN prov., 25 km SE Vieng Xai (by road) BAN KANGPABONG env., 20 ° 19 ’ N 104 ° 25 ′ E; D. Hauck leg.: 14. – 18. v. 2001 ” (ZFMK), 1 ♂ “ LAOS, 1. – 16. v. 1999, Louangphrabang pr. 20 ° 33 - 4 ’ N 102 ° 14 ′ E Ban Song Cha (5 km W), 1200 m, Vit Kubán leg. ” (ZFMK). China. 1 ♂ “ CHINA: Guangxi, Fangchenggang, Naliang, Tansan Vill. AEdz " 21.666031 ° N, 107.734480 ° E Shiwandashan Mount Range 325 m 2023 V. 12 native leg. ” (SCAU).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF96FFE6F088FE44E5EEFD9F.taxon	materials_examined	Material examined. 6 ♂♂ “ NW Thailand, Mae Hong Son, 9. – 16. v. 1991 Ban Huai Po 1600 m leg. P. Pacholátko ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF96FFE6F088FD34E05BFCC7.taxon	materials_examined	Material examined. 1 ♂ “ E Thailand, 5. - 13.5. Chanthaburi Dist. Khao Soi Dao, 1998 J. Rolčík Igt. ” (ZFMK), 2 ♂♂ “ NW THAILAND MAE HONG SON 29. – 30.4.1992 J. Horak leg. ” (ZFMK), 5 ♂♂ “ NW Thailand 9. – 16. V MAE HONG SON 199 Ban Huai Po 1600 m leg. P. Pacholatko ” (ZFMK), 5 ♂♂, 4 ♀♀ “ THAI, 9. – 14. v. 1991 CHIANG DAO 350 m 19 ° 22 ′ N 98 ° 57 ′ E Vít Kubáň leg. / Thailand 91 Thanon Thong Chai, Král & V. Kubán ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF96FFE6F088FCDCE483FC37.taxon	materials_examined	Material examined. 1 ♂ “ China, NE Guangdong prov., Qifeng Shan, 900 – 1200 m 23 ° 51 ’ N 115 ° 21 ’ E Jatua leg., 18. VI. 2015 ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF96FFE6F088FC4CE07EFB83.taxon	materials_examined	Material examined. 8 ♂♂ “ N VIETNAM, 21,27 N 105,39 E 70 km NW Hanoi, Tam Dao 9. – 19. v. 1996, 900 – 1200 m Pacholátko & Dembický / VS 21 ” (ZFMK), 2 ♂♂ “ N-Vietnam, (Tonkin) Tamdao 12. – 24.5.1989 PACHOLÁTKO leg. / VS 21 ” (ZFMK), 1 ♂ “ N. VIETNAM / Tonkin / TAMDAO / pr. Vinhphu 2. – 11.6.1985 Vit. Kubán leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF96FFE6F088FB10E42AFAF3.taxon	materials_examined	Material examined. 1 ♂ “ THAILAND, Nan prov., Doi Phuka Nat. Park, 28. iv. – 12. v. 2002 P. Průdek & M. Oboril leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF96FFE6F088FA80E5EDF9F7.taxon	materials_examined	Material examined. 2 ♂♂ “ S Vietnam, 1. – 5.5.1994 Nam Cat Tien Nat. park P. Pacholátko & L. Dembicky ́ leg. / coll. P. Pacholátko / VS 66 ” (ZFMK), 1 ♂ “ NW Thailand 9. – 16. v. 1991 Mae Hong Son Ban Huai Po, 1600 m leg. P. Pacholatko ” (ZFMK), 1 ♂ “ E Thailand, 5. - 13. Chanthaburi Dist. Khao Soi Dao, 1996 J. Rolèik igt. ” (ZFMK), 2 ♂♂ “ S-Vietnam, Bin Phuoc Pr, Bu Gia Map NP, 16. – 20.4 2024, 12 ° 11 ’ 38 ” N, 107 ° 12 ’ 21 ” E, 450 – 600 m, A. Skale ” (CASG).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF96FFE6F088F98CE1B1F943.taxon	materials_examined	Material examined. 3 ♂♂ “ THAI-N, 1 – 15. v. 1998, Chiang Mai prov., SAN PAKIA vill., 19.19 N 98.50 E, 1400 m, Vit Kubáň leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF96FFE6F088F98CE1B1F943.taxon	discussion	Remark. The publication year of this species was updated by Ahrens & Bezděk (2024).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF96FFE6F088F950E5EEF847.taxon	materials_examined	Material examined. 1 ♂ “ LAOS, 1. – 18. v. 2001, Bolikhamxai prov., 18 ° 21 ’ N 105 ° 08 ′ E, Ban Nape (8 km NE), ~ 600 m, Vit Kubáň leg. ” (ZFMK). Tetraserica semiruiliensis Fabrizi, Dalstein & Ahrens, 2019 Material examined. 3 ♂♂, 3 ♀♀ “ E Thailand, 5. - 13.5. Chanthaburi Dist. Khao Soi Dao, 1998 M. Rolčik lgt. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF97FFE7F088FF2CE18BFEE3.taxon	materials_examined	Material examined. 2 ♂♂ “ THAI 24. – 29. iv. 1993, DOI SUTHEP Pacholátko & Dembicky leg / coll. P. Pacholátko ” (ZFMK), 1 ♂ “ THAI 1. – 8. v. 1993 SOPPONG PAI 1800 m Pacholatko & Dembicky leg. ” (ZFMK), 2 ♂♂ “ NW Thailand 9. – 16. v. Mae Hong Son 1991, Ban Huai Po, 1600 m, leg. P. Pacholátko ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF97FFE7F088FEF0E49AFCC7.taxon	materials_examined	Material examined. Laos. 1 ♂ “ LAOS-C: KHAM MOUAN Ban Khoun Ngeun; ~ 200 m 18 ° 07 ′ N 104 ° 29 ’ E: Pacholátko leg.: 24. – 29. ιν. 2001 ” (ZFMK), 1 ♂ “ S LAOS, Champasak, Ban Itou, 800 m, 10. – 18. iv. 1999 15 ° 10,4 ’ N, 106 ° 05,8 ’ E, E. Jendek & O. Šauša leg. ” (ZFMK), 5 ♂♂, 4 ♀♀ “ LAOS centr., Khammouan pov. NAKAI env., 17 ° 43 ′ N, 105 ° 09 ’ E 22. v. – 8. vi. 2001, alt. 500 – 600 m, E. Jendek & O. Šauša leg. ” (ZFMK), 8 ♂♂ “ LAOS centr., Bolikhamsai prov. BAN NAPE-Kaew Nua Pass 18.4. – 1.5.1998, alt. 600 ± 100 m, N 18 ° 22.3, E 105 ° 09.1 (GPS), M. Strba & R. Hergovits leg. ” (ZFMK), 16 ♂♂ “ LAOS centr., Bolikhamsai prov. BAN NAPE-Kaew Nua Pass, 18.4. – 1.5.1998, alt. 600 ± 100 m, N 18 ° 22.3, E 105 ° 09.1 (GPS), E. Jendek & O. Šauša leg. ” (ZFMK), 9 ♂♂ “ LAOS centr., Khammouan prov. NAKAI env., 4. – 8.5.1998, Route No 8, alt. 560 ± 20 m N 17 ° 42.8, E 105 ° 08.9 (GPS), E. Jendek & O. Sauša leg. ” (ZFMK), 1 ♂ “ LAOS centr., Khammouan prov. NAKAI env., 4. – 8.5.1998, Route No 8, alt. 560 ± 20 m N 17 ° 42.8, E 105 ° 08.9 (GPS), M. Strba & R. Hergovits leg. ” (ZFMK). Vietnam. 3 ♂♂ “ N VIETNAM, 21,27 N 105,39 E 70 km NW Hanoi, Tam Dao 9. – 19. v. 1996, 900 – 1200 m Pacholátko & Dembický / VS 82 ” (ZFMK). Thailand. 1 ♂ “ THAILAND, Nan prov., Ban Huay Kon env., 27. v. – 10. vi. 2002 P. Průdek & M. Oboril leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF97FFE7F088FCDCE74DFBEF.taxon	materials_examined	Material examined. 2 ♂♂ “ THAI, N, Mae Hong Son prov., SE of Soppong, 1500 m. 20 19 ° 27 ’ N, 98 ° 20 ’ E 23 – 27. ν. 1999, D. Hauck leg. ” (ZFMK), 1 ♂ “ THAI 28 – 31 / 5.1995 19.27 N 98.20 E SOPPONG 1500 m Vít Kubáň leg. / Coll. P. Pacholátko / TS 49 ” (ZFMK), 1 ♂ “ THAI, Mae Hong Son prov. 19 ° 27 ’ N 98 ° 20 ’ E, 1500 m SOPPONG, 7. – 12. v. Vit Kubán leg., 1996 ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF97FFE7F088FB84E5EEFB5F.taxon	materials_examined	Material examined. 6 ♂♂ “ NW Thailand, 9. – 16. v. MAE HONG SON. 1991 Ban Huai Po 1600 m leg. P. Pacholátko ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF97FFE7F088FB74E70BFACF.taxon	materials_examined	Material examined. 1 ♂ “ MALAYSIA, Pahang state Cameron Highlands TANAH RATA, 1400 – 1650 m 4. – 11.4.2005, Martinů leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF97FFE7F088FAE4E7F4FA3F.taxon	materials_examined	Material examined. 1 ♂ “ LAOS north, 5. – 11. v. 1997, 20 km NW Louang Namtha, N 21 ° 09.2 E 101 ° 18.7, alt. 900 ± 100 m, M. Štrba & R. Hergovits leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF97FFE7F088FA54E17BF9AF.taxon	materials_examined	Material examined. 1 ♂ “ E Thailand, 5. - 13.5. Chanthaburi Dist. Khao Soi Dao, 1998 M. Knížek lgt. ” (ZFMK), 1 ♂ “ E Thailand, 5. - 13.5. Chanthaburi Dist. Khao Soi Dao, 1998 J. Rolčik lgt. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF97FFE7F088F9C4E5EEF91F.taxon	materials_examined	Material examined. 2 ♂♂ “ THAI, 9. – 13. iv. 1991 THIMONGHTA 350 m 15 02 ’ N, 98 35 ’ E, P. Pacholátko leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF97FFE7F088F8B4E178F86B.taxon	materials_examined	Material examined. 2 ♂♂ “ Laos Bolikhamxai pr., 18 ° 16 ’ N, 103 ° 11 ’ E, 70 km NEE Vientiane 27. – 30. iv. 1997, 150 m, Vít Kubáň leg. / coll. P. Pacholátko ” (ZFMK), 1 ♂ “ LAOS centr., 27. iv. – 1. v. 1.1997, 70 km NE Vientiane, BAN PHABAT env., 150 m, N 18 ° 16.1, E 103 ° 10.9, M. Štrba & R. Hergovits leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF97FFE0F088F878E7B6FF2B.taxon	materials_examined	Material examined. 1 ♂ “ China, S. Jiangxi, Nat. Res. Jiulian Shan, 750 m 24 ° 32 ’ N 114 ° 28 ’ E Jatua leg., 27. v. – 20. vi. 2015 / Asia Sericini spec. 1350 ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF90FFE0F088FED4E1ECFDE7.taxon	materials_examined	Material examined. 12 ♂♂, 2 ♀♀ “ S Vietnam, 12.03 N 108.27 E 12 km N of Dalat-Lang Bian, 1580 – 1750 m, 17. – 21. iv. 1999, Pacholátko & Dembický leg. / coll. P. Pacholátko / VS 72 ” (ZFMK), 4 ♂♂ “ S-Vietnam, Bin Phuoc Pr, Bu Gia Map NP, 16. – 20.4 2024, 12 ° 11 ’ 38 ” N, 107 ° 12 ’ 21 ” E, 450 – 600 m, A. Skale ” (CASG), 1 ♂ “ S-Vietnam, Lam Dong Pr. Lac Duong Dis., Da Nhim town, 22. – 28. 4. 2024 leg. A. Skale, LF, 1400 m ” (CASG).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF90FFE0F088FEB8E049FEBF.taxon	materials_examined	Material examined. 2 ♂♂ “ THAI 24. – 29. iv. 1993 DOI SUTHEP Pacholatko & & Dembicky leg. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
03FB3D2FFF90FFE0F088FDFCE746FD57.taxon	materials_examined	Material examined. 1 ♂, 2 ♀♀ “ China, SE Yunnan, Xichou-E env. 1400 – 1700 m, 13. - 18.5.95 23 ° 22 - 26 ’ N / 104 ° 41 - 49 ’ E L. + R. BUSINSKY lgt. ” (ZFMK).	en	Pham, Phu V., Pacholátko, Petr, Ahrens, Dirk (2025): New species and records of Tetraserica Ahrens, 2004 from Indochina (Coleoptera: Scarabaeidae: Sericinae). Zootaxa 5691 (2): 193-230, DOI: 10.11646/zootaxa.5691.2.2, URL: https://doi.org/10.11646/zootaxa.5691.2.2
