taxonID	type	description	language	source
03F887D8FFBFFFD911759C6E1FBCFD32.taxon	materials_examined	Material examined PAN 17 - 1204 - PAN 01, Satpura National Park, Bariaam, Panchmari, Madhya Pradesh, Lat / Lon: 22.501, 78.4559, elevation: 1005.85 m; PAN 17 - 1209 - PAN 06, Satpura National Park, Bee Fall, Panchmari, Madhya Pradesh, Lat / Lon: 22.4664, 78.4412, elevation: 1014.08 m; PAN 17 - 1211 - PAN 08, PAN 17 - 1213 - PAN 010, Satpura National Park, Pandav Caves, Panchmari, Madhya Pradesh, Lat / Lon: 22.4702, 78.4444, elevation: 650.756 m, collected by Ashutosh Vishwakarma, collected on 1 st of October 2017; MNP 17 - 945 - 36 A 7, Keibul Lamjao National Park, Bishnupur, Manipur, Lat / Lon: 24.5475, 93.8521, elevation: 748.903 m, collected on 3 rd of October 2017; MNP 17 - 968 - 37 A 7, MNP 17 - 972 - 37 A 11, MNP 17 - 973 - 37 A 12, Keibul Lamjao National Park, Bishnupur, Manipur, Lat / Lon: 24.4953, 93.8319, elevation: 729.09 m, collected on 4 th of October 2017; MNP 17 - 985 - 39 A 2, MNP 17 - 1005 - 39 A 23, Churachandpur, Manipur, Lat / Lon: 24.3728, 93.7097, collected on 6 th of October 2017; MNP 17 - 1015 - 40 A 9, close to forest office, Churachandpur, Manipur, Lat / Lon: 24.3339, 93.7294, collected on 6 th of October 2017; MNP 17 - 1145 - 46 A 9, Heingang, close to Imphal River, Manipur, Lat / Lon: 24.8359, 93.95 elevation: 815.655 m, collected on 12 th of October 2017, collected by Shweta Yadav (Fig. 2).	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFBFFFD911759C6E1FBCFD32.taxon	description	Description Length 109 – 156 mm, diameter 5 – 5.5 mm, total number of segments 89 – 128. Colour dark pigmented, dark brown or greyish brown on dorsal side and paler at ventral side in preserved specimen. Prostomium rudimentary. Setae perichaetine, starts from segment ii. First dorsal pore in xii / xiii. Clitellum annular, extends from xiv – xvi, setae unrecognizable, dorsal pores absent. Spermathecal pore four pairs in an intersegmental furrow of v / vi, vi / vii, vii / viii, and viii / ix minute, superficial, more than 1 / 3 C apart. Female pore single median, encircled with circular whitish line on segment xiv. Male pores minute, superficial in segment xviii, each in a relatively circular area (usually slightly depressed) encircled by U-shaped ridge that opens mesially. Each male pore is 13 – 15 setae apart. Genital marking absent. Septa, 6 / 7 – 7 / 8 (and occasionally 5 / 6) substantially thickened, 10 / 11 – 11 / 12 less so. Septa, 8 / 9 – 9 / 10 absent. Intestine originate in segment xv. Lateral intestinal caeca, simple, in xxvii and extends up to segment xx. Lamelliform typhlosole. Holandric. Seminal vesicles present in xi, xii. Testis sac paired, vertical or sometimes unpaired, horseshoe-shaped including the hearts of x-xi and seminal vesicles of xi. Prostates in xviii, extends from xvi-xxii, duct muscular and two to three times looped. Spermathecal ampullae sac-like, with slender stalk diverticulum that originates from the spermathecal duct near parietes, longer than main axis and a variously looped wider portion entally.	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB9FFD912CD9A871A10FDD6.taxon	materials_examined	Material examined MNP 17 - 1051 - 42 A 10, MNP 17 - 1053 - 42 A 12, Yangoupokpi-Lokchao Wildlife Sanctuary, Manipur, Lat / Lon: 24.2542, 94.2992, elevation: 418.8 m, collected on 8 th of October 2017; MNP 17 - 1063 - 43 A 3, MNP 17 - 1068 - 43 A 8, Yangoupokpi-Lokchao Wildlife Sanctuary, Kwatha Range, Manipur, Lat / Lon: 24.3238, 94.2923, collected on 9 th of October 2017, collected by Shweta Yadav (Fig. 3).	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB9FFD912CD9A871A10FDD6.taxon	description	Description Length 97 – 110 mm, diameter 4 – 5 mm. Total number of segments 82 – 110. Body colour on the anterior segment dorsally brownish purple or chestnut, later paler, also pale ventrally. Prostomium open epilobic. First dorsal pore begins in intersegmental region of xii / xiii. The setae very small, numerous, arranged without any break on dorsal and ventral side, approximately 60 – 65 on segment ix. The clitellum covers three segments, i. e., xiv – xvi, without dorsal pore and setae. The male pores are present in the setal zone of segment xviii, approximately one-third of the circumference of the body apart each other with 9 – 11 (N = 6) setae intervening, placed on broad little prominent circular porophores which covers the whole segment. The female pores sometimes paired very close to each other, while in some appears as a single whitish oval patch on segment xiv. The spermathecal pores are four pairs, in the intersegmental furrow of v / vi – viii / ix, ventrolateral, approximately one-third of the circumference apart. Genital markings single median, very prominent, often transversely oval glandular cushions, variously places across two or more of xix / xx – xxvi / xxvii successively. The distance between two genital markings is very small, only restricted to the setal zone of the intermediate segments. Septa are as per the original description of Stephenson (1929). Oesophagus with a single gizzard between septa viii – x. The lateral intestinal caeca simple, originated in segment xxvii and extends up to segment xxiii. The last pair of heart in segment xiii. Male organs holandric, testis sac in segment x and xi, two pairs of seminal vesicles in segment xi and xii, prostates large, racemose, in segment xviii, extends from xvi – xxii, densely fissured surface, duct forming a loop extending backward. Prostatic duct opens directly, without the intermediary of copulatory bursae. The spermathecal ampulla large ovoidal sac (like original description of Stephenson, 1929), without sharp delimitation of spermathecal duct. Unidiverticulate, diverticulum originates at the extreme ectal end of the duct, tubular, with club shaped dilation at its ental end, variously looped, or arranged in somewhat zig-zag manner.	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB9FFD911759AFB1A35F8E0.taxon	materials_examined	Material examined MNP 17 - 1087 - 44 A 13, Yangoupokpi-Lokchao Wildlife Sanctuary, Tengnoupal, Manipur, Lat / Lon: 24.3831, 94.1492, collected on 10 th of October 2017, collected by Shweta Yadav. PAN 17 - 1220 (identified using morphoanatomical characterization, it was collected with Metaphire houlleti and subsequently preserved in sample tube PAN 17 - 1217 - PAN 014; the specimen was not retained for molecular investigations), Satpura National Park, Sahastradhara Waterfalls, Panchmari, Madhya Pradesh, Lat / Lon: 22.5834, 78.5041, elevation: 446.842 m, collected on 1 st of October 2017, collected by Ashutosh Vishwakarma (Fig. 4).	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB9FFD911759AFB1A35F8E0.taxon	description	Description Length approx. 83 mm, diameter 2.9 mm. Total number of segments approx. 105. Prostomium open epilobic. First dorsal pore begins from intersegmental region of xi / xii. Clitellum annular covering three segments, i. e., xiv – xvi, without setae. Female pore single, median on segment xiv, bordered with small, white circular patch. Male pores in segment xviii superficial on oval porophores, approximately one-fourth circumference apart each other with 13 setae intervening between male pores. The spermathecal pores are four pairs in intersegmental region of v / vi / vii / viii / ix. Genital markings small, circular, paired, presetal in segment vii, viii, and ix, also three small circular genital markings present near male pores, curbed within concentric circular folds. Oesophagus with a single gizzard between septa viii – x and two pairs of intramural calciferous glands in segment xii and xiii. A pair of lateral intestinal caeca originates in segment xxvii and extends up to segment xx. Last pair of heart in segment xiii. Male organ holandric, testis sac in x and xi, two pairs of seminal vesicles in segment xi, xii, prostate absent, duct present in segment xviii. Four pairs of spermathecae present in segment vi – ix, ampulla oval sac, with slender straight stalked diverticulum having oval seminar chamber at its ental end. Stalked glands associated with external genital markings.	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB8FFDB12CD9D9A1FECFA88.taxon	materials_examined	Material examined MNP 17 - 1107 - 45 A 4, MNP 17 - 1111 - 45 A 8, MNP 17 - 1122 - 45 A 19, MNP 17 - 1123 - 45 A 20, MNP 17 - 1130 - 45 A 28, Shiroi National Park, Shirui Kashong (Peak), Manipur, Lat / Lon: 25.1302, 94.4202, elevation: 2106.19 m, collected on 11 th of October 2017, collected by Shweta Yadav (Fig. 5).	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB8FFDB12CD9D9A1FECFA88.taxon	description	Description Length 56 – 77 mm, diameter 3 – 4 mm. Total number of segments 78 – 92. Body colour brownish yellow dorsally, paler on ventral side. Prostomium open epilobic. First dorsal pore starts from intersegmental furrow of x / xi. Clitellum covers three segments, i. e., xiv – xvi, setae partially visible. The female pore, single, median on segment xiv in small circular whitish patch. Male pores in the setal zone of segment xviii, superficial, on a small porophores, bounded by concentric ridges. Three pairs of spermathecal pores in intersegmental furrow of v / vi / vii / viii. Genital markings present on segment xviii near male pores, post-setal 2 – 3 small discs with fine circular swollen outline. A single spherical gizzard in segment viii. The lateral intestinal caeca simple with lobed margins, originates in segment xxvii and extends up to segment xxv. Last pair of heart in segment xiii. Intestine begins from segment xv. Male organs holandric, testis sacs in x and xi and paired seminal vesicles in segment xi and xii. Prostate racemose well developed large in segment xviii, extends xvi – xxii with long muscular ducts and at the ectal end of prostatic duct present a genital marking gland. Three pairs of spermathecae in segment vi – viii, large saccular ampullae, duct slender, equal to or smaller than ampulla, from the base of the duct arises a thin tubular diverticulum, each with a small, a little iridescent terminal bulb.	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFBBFFDB12CD9D1E1B14F9DC.taxon	materials_examined	Material examined MNP 17 - 1008 - 39 A 28, Churachandpur, Manipur, Lat / Lon: 24.3728, 93.7097, MNP 17 - 1024 - 40 A 20, Churachandpur, close to forest office, Manipur, Lat / Lon: 24.3339, 93.7294, collected on 6 th of October 2017, collected by Shweta Yadav (Fig. 6).	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFBBFFDB12CD9D1E1B14F9DC.taxon	description	Description Length approximately 90 mm, diameter 4 mm. Total number of segments 79 – 83. Prostomium open epilobic. First dorsal pore begins from segment x / xi. Clitellum covers three segment xiv – xvi. Single, median female pore on segment xiv. Male pores paired in the setal zone of segment xviii, superficial on small, circular porophores, intervened with 18 setae between two male pores. Two pairs of spermathecal pores present in the intersegmental furrow of v / vi and vi / vii. Genital markings in post-clitellar region in segment xviii, two instantly median to individual male pore, one pre-setal and one post-setal, encircled by several concentric furrows. Additionally, presetal genital markings on xviii and xix in both specimens, numbers variable, mid-ventral or median to male pores. Also, genital markings present on pre-clitellar regions single and / or paired small discs paired on vii median to spermathecal pore, unpaired on vii, mid-ventral, pre-setal. A single gizzard in segment x. The lateral intestinal caeca, simple with serrated margins originates in segment xvii and extends up to segment xv. Last pair of heart in segment xiii. Male organs holandric, testis sac in segment x and xi, two pairs of seminal vesicles in segment xi and xii. Prostate racemose in segment xviii, extends up to xxii, duct long, muscular. Intestine starts from segment xiv. Two pairs of spermathecae in segment vi and vii, ampulla ovoid, duct longer than ampulla, from the base of the duct arises a thin tubular, slightly iridescent club shaped diverticulum.	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFBBFFD711759EE21C87FD32.taxon	materials_examined	Material examined MNP 17 - 1152 - 47 A 1, MNP 17 - 1153 - 47 A 2, MNP 17 - 1165 - 47 A 15, MNP 17 - 1172 - 47 A 22, Lamdeng Makhaleikai forest range, Manipur, Lat / Lon: 24.8376, 93.874, elevation: 844.306 m, collected on 12 th of October 2017, collected by Shweta Yadav (Fig. 7).	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFBBFFD711759EE21C87FD32.taxon	description	Description Length 77 – 96 mm, diameter 4 – 5 mm. The total number of segments 93 – 110. Prostomium open epilobic. Setae very small, equally placed, without any break on dorsal and ventral side. The first dorsal pore in intersegmental furrow of xii / xiii. Clitellum annular, covers three segment xiv – xvi, dorsal pore absent. The female pore single, median on segment xiv. Male pores minute, each at centre of a small disc on segment xx, in copulatory pouch with transversely slit like equatorial aperture each distantly situated 1 / 6 th of the body circumference. There are 15 – 18 (N = 4) setae between male pores. In one specimen, two spermathecal pores observed which were placed on intersegmental furrow of vi / vii on left side and vii / viii on right side, while in rest of the three specimen spermathecal pores absent. Genital markings small, paired, disclike, each with apparent central pore and in a parietal invagination with transversely slit like equatorial aperture resembling male pores but slightly smaller than male pores, on post-clitellar regions, in the setal zone, placed slightly further apart than male pores usually in segment xviii, xix, xxi, xxii, sometimes in xvii paired and / or unpaired. Approximately, 20 setae present between two genital markings. A single, large gizzard in segment viii. The lateral intestinal caeca simple, originates in segment xvii and extends up to segment xiv. Last pair of heart in segment xiii. Male organs holandric, six pairs of testes present from segment v – xiii. Prostate large, extends from segment xviii to xxiii, duct muscular, long and looped. Spermathecae only in one specimen in intersegmental region of vi / vii, on left side, duct slender and approximately equal to the vaguely demarcated ampulla, diverticulum with short and slender stalk, twisted middle region in which lumen is gradually widened, and a clubshaped seminal chamber. On the other hand, small, immature spermathecae on right side in segment vii / vii was observed.	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB7FFD712CD9A871DB2FC9E.taxon	materials_examined	Material examined MNP 17 - 1006 - 39 A 26, Churachandpur, Manipur, Lat / Lon: 24.3728, 93.7097, collected on 6 th of October 2017, collected by Shweta Yadav (Fig. 8).	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB7FFD712CD9A871DB2FC9E.taxon	description	Description Length approximately 100 mm, diameter 4.6 mm. The total number of segments 97. Prostomium epilobic. First dorsal pore begins from segment xii / xiii. Clitellum annular covers three segments, i. e., xiv – xvi, setae and dorsal pores absent. Female pore single, median on segment xiv visible in small, intact, white circular patch. Male pores in the setal zone of segment xviii, inside a copulatory pouch, bounded by tumescent lip, opens with transverse slit- like aperture, intervening 13 setae between the male pores. Three pairs of spermathecal pores in the ventrolateral side of intersegmental furrow of segment v / vi / vii / viii, opens with transverse slit-like aperture. Genital markings absent. Oesophagus with a single gizzard in segment ix. The lateral intestinal caeca manicate, originates in segment xxvii and extends up to xxiv. Last pair of heart in segment xiii. Male organs holandric, testis sacs in x and xi, two pairs of seminal vesicles in xi and xii. Prostate racemose, deeply lobuled, in xviii, extends from xvi – xxii, duct uniformly thickened, muscular, U-shaped looped. Spermathecae paired in segment vi – viii, oval-shaped ampulla, duct short approximately half the length of ampulla, at the base of duct attached a diverticulum, strongly waved with 4 – 5 small seminal chambers.	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB7FFD611759B231BF2FECF.taxon	materials_examined	Material examined MNP 17 - 1011 - 40 A 3, MNP 17 - 1019 - 40 A 15, MNP 17 - 1025 - 40 A 21, Churachandpur, close to forest office, Manipur; Lat / Lon: 24.3339, 93.7294; collected on 6 th of October 2017; collected by Shweta Yadav; PAN 17 - 1207 - PAN 04, Satpura National Park, Bee Fall, Panchmari, Lat / Lon: 22.4664, 78.4412, elevation: 1014.08 m, PAN 17 - 1210 - PAN 07, Satpura National Park, Boat Club Lake, Panchmari, Lat / Lon: 22.4539, 78.416, elevation: 957.998 m, PAN 17 - 1219 - PAN 016, PAN 17 - 1217 - PAN 014 Satpura National Park, Sahastradhara Waterfalls, Panchmari, Lat / Lon: 22.5834, 78.5041, elevation: 446.842 m, collected on 1 st of October 2017, collected by Ashutosh Vishwakarma; MP 17 - 1235 - 1 SGO, MP 17 - 1239 - 5 SGO, Dr Hari Singh Gour University, Sagar, Lat / Lon: 23.8298, 78.7763, elevation: 559.924 m, collected on 26 th October of 2017, collected by Ashutosh Vishwakarma (Fig. 9).	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB7FFD611759B231BF2FECF.taxon	description	Description Length 75 – 90 mm, diameter 3.5 – 3.7 mm. The total number of segments 82 – 93. First dorsal pore begins from intersegmental furrow of ix / x. Clitellum annular, covers three segments, i. e., xiv – xvi. Female pore, single, median on segment xiv, in setal zone, bounded by semicircular patch. Male pores in the setal zone of segment xviii, inside a copulatory pouch, opens with a C-shaped aperture. Each male pore placed 0.30 × apart with 9 – 12 setae intervening between them. Three pairs of spermathecal pores in intersegmental region of vi / vii / viii / ix, sometimes deep slits. Genital marking absent or present; when present, in pre-clitellar region, within the intersegmental furrow, next to each spermathecal pore. Oesophagus with large, single gizzard in segment viii / ix. Lateral intestinal caeca simple, originates in segment xxvii, extends up to xxv. Male organs holandric, testis sac in segment x and xi, two pairs of seminal vesicles in segment xi and xii. Prostate racemose, in xviii, extends from xvi – xxii, duct U-shaped. Spermathecae three pairs in segment vii – ix, ampulla oval, with relatively larger swollen spermathecal duct, diverticulum enlarged with looped end. Genital marking gland single stalked, attached to spermathecal ducts.	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB6FFD1117599D11C45F91E.taxon	materials_examined	Material examined MNP 17 - 1137 - 46 A 1, MNP 17 - 1138 - 46 A 2, MNP 17 - 1139 - 46 A 3, Heingang, paddy field, close to Imphal River, Manipur, Lat / Lon: 24.8359, 93.95, elevation: 815.655 m, collected on 12 th of October 2017, collected by Shweta Yadav; UP 18 - 1275 - 4 A 6, paddy field, village Aaripur, Ghazipur, Uttar Pradesh, Lat / Lon: 25.6941, 83.6187, elevation: 53 m, collected on 14 th of October 2017; UP 18 - 1303 - 13 A 1, UP 18 - 1304 - 13 A 2, UP 18 - 1305 - 13 A 3, UP 18 - 1306 - 13 A 4, UP 18 - 1307 - 13 A 5, Jungle Kaudiya, Gorakhpur, Uttar Pradesh, Lat / Lon: 26.8777, 83.3055, elevation: 104 m, collected on 12 th of October 2017; UP 18 - 1310 - 14 A 1, UP 18 - 1311 - 14 A 2, Jungle Dhushad, Haidergunj, Gorakhpur, Uttar Pradesh, Lat / Lon: 26.7915, 83.4214, elevation: 52 m, collected on 12 th of October 2017, collected by Nalini Tiwari (Fig. 10).	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB6FFD1117599D11C45F91E.taxon	description	Description Length 75 – 111 mm, diameter 5 mm. The total number of segments 82 – 110. First dorsal pore starts from xii / xiii. Clitellum annular, covering three segments, i. e., xiv – xvi. Female pore on xiv single, median, presetal or just adjacent to setal line. Male pores in the setal line of segment xviii, on small disc in crescentic slits of copulatory pouches with about 17 – 22 setae between the male pores. Spermathecal pore four pairs in the intersegmental furrow of v / vi / vii / viii / ix. Genital markings paired, on xvii and xix, in setal line, slightly internal to the male pores. Oesophagus with single gizzard in ix / x. Lateral intestinal caeca simple, starts from xxvii, extends up to xxiv. Male organs holandric, testis sacs in x and xi, two pairs of seminal vesicles in segment xi and xii. Prostate racemose, in xviii, extending through xv – xxi, ducts long and looped. Four pairs of spermathecae in vi-ix, each with a short, stout duct and oval shaped ampulla; diverticulum stalk short and slender. Genital marking glands present in xvii and xix, corresponding to external genital markings, sessile.	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB1FFD012CD9EA01C22FE68.taxon	materials_examined	Material examined MP 17 - 1231 - 1 A 3, Union Carbide Factory, Bhopal, Madhya Pradesh, Lat / Lon: 23.2806, 76.4102, elevation: 362.716 m, collected on 11 th of September 2017; MP 17 - 1232 - 2 A 2, Javik Krishi Kendra, Patidar Krishi Farm, Fanda Block, Bhopal, Lat / Lon: 23.5669, 77.4584, elevation: 509.937 m, collected on 12 th of September 2017, collected by Ashutosh Vishwakarma; MNP 17 - 966 - 37 A 5, MNP 17 - 967 - 37 A 6, MNP 17 - 1147 - 46 A 11; Keibul Lamjao National Park, Bishnupur, Manipur, Lat / Lon: 24.4953, 93.8319, elevation: 729.09 m; Heingang, paddy fields, close to Imphal River, Manipur; Lat / Lon: 24.8359, 93.95, elevation: 815.655 m; collected by Shweta Yadav, collected on 4 th and 12 th of October 2017 (Fig. 11).	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
03F887D8FFB1FFD012CD9EA01C22FE68.taxon	description	Description Length 80 – 246 mm, diameter 4 – 5 mm. Total number of segments 110 – 156. Prostomium pro-epilobic. First dorsal pore starts from xii / xiii. In specimen collected from Manipur, setae a and b enlarged from segment v – xiii, later on setae a enlarged up to anal end. Clitellum annular, covers three segments xiv – xvi. Female pore single median on xiv. Male pore on xviii, in setal line, on small disc intervened with 10 – 12 setae. Spermathecal pores usually absent, only observed in one specimen, three pairs of spermathecal pores in each intersegmental furrows of v / vi / vii. Genital marking large, raised, smooth disks, paired, presetal on xix, xx, xxi, xxii. Oesophagus with single gizzard between septa viii – x. Lateral intestinal caeca absent. Last pair of heart in xiii. Intestine begins in xv. Holandric male organs, testis sac in x and xi, seminal vesicles in xi and xii. Ovaries in xiii. Prostate in xviii extended upto xvi – xx, with looped muscular duct, wider on ectal end. Spermathecae usually absent, when present, polythecal, in vi and vii, with a duct shorter than ampulla, stalked diverticulum attached to the base of spermathecal duct, with ovoidal end. Genital marking glands sessile.	en	Tiwari, Nalini, James, Samuel W., Yadav, Shweta (2024): Pheretimoid earthworms of India: new insight from molecular systematics and phylogenetic relationship with new records. Organisms Diversity & Evolution 24 (3): 313-333, DOI: 10.1007/s13127-024-00646-x, URL: https://doi.org/10.1007/s13127-024-00646-x
