identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F887E1FFDC883551D9FBCCF63E1371.text	03F887E1FFDC883551D9FBCCF63E1371.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spodochlamys keitai KobaYashi and Ratcliffe 2023	<div><p>Spodochlamys keitai Kobayashi and Ratcliffe, new species</p><p>zoobank.org/ urn:lsid:zoobank.org:act: 98807D14-6B85-4432-BA8A-1E3D4CA3EDE6 (Figs. 1–2)</p><p>Type Material. Holotype labeled “ COLOMBIA / Boyacá, / Otanche, / III-2021 ” and with our red holotype label (Fig. 2F) . Three paratypes with the same data and each with our red paratype label . Holotype deposited at the University of Nebraska State Museum (UNSM), Lincoln, Nebraska, USA. One paratype was deposited at the Natural History Museum (NHMUK), London, England, and two paratypes in the Keita Maruyama personal collection (KMC), Tokyo, Japan.</p><p>Description of Holotype. Male (Figs. 1A, 2A–F). Length 27.0 mm; width across humeri 10.1 mm. Color: Head, pronotum, elytra, and tibiae reddish brown with shiny, metallic copper reflections; scutellar shield shiny metallic green (Fig. 1A); pygidium, venter and femora shiny reddish brown. Elytra lacking waxy bloom. Head: Head completely, densely rugulopunctate. Clypeal apex (dorsal view) broadly semicircular, rounded (Fig. 1A), weakly reflexed; surface (frontal view) with upper and lower margins subparallel. Labrum at middle 3 times wider than venter of clypeal apex (Fig. 2B), apex broadly, weakly subacute; surface with minute, sparse, setigerous punctures, setae short and pale. Mentum slightly tumescent with faint longitudinal depression from apex to middle of disc (Fig. 2B). Interocular width equals 3.8 transverse eye diameters. Antenna with club subequal in length to entire stem. Pronotum: Form evenly convex in lateral view. Surface completely, densely, coarsely rugopunctate, becoming more so toward slightly explanate lateral margins. Lateral margin evenly rounded, weakly crenulate (Fig. 1A). Basal margin with complete bead. Length at middle 1.6 times longer than frons. Scutellar shield: Surface densely rugulopunctate, glabrous. Elytra: Surface completely, densely, coarsely rugopunctate, rugae becoming larger and transverse laterally. Striae completely absent. Epipleuron (ventral view; Fig. 2A) slightly concave, broad at base and gradually tapering apically, glabrous. Apex at suture nearly right angled. Apical umbone well developed, conical (Figs. 1A, 2E). Pygidium: Surface shiny, with minute, sparse, shallow punctures, glabrous. Form in lateral view strongly convex.Apical bead slender, slightly wider at middle. Venter: Mesoventrite not produced at apex. Meso- and metaventrite with long, dense, pale setae (Fig. 2A). Abdominal sternites with small, dense punctures. Legs: All tibiae coarsely rugopunctate. Protibia not elongate or curved, lacking ventroapical modification, inner margin straight (Fig. 1A); external margin tridentate, basal tooth slightly removed from apical teeth. Protarsal inner claw entire at apex; meso- and metatarsal inner claws minutely split at apices (Fig. 1A). Protarsomere 5 with weak median tooth. Profemur not elongated, lacking ventromedial projection. Procoxa not ventrally produced and enlarged. Metafemur with posterior margin “normal”, not weakly produced posteriorly, not laminate. Meso- and metatibial apices truncate, upper angle acute (Fig. 2e). Metatibial apical spurs simple, acute (Fig. 2A). Parameres: Form symmetrical, long, slender (Figs. 2C–D).</p><p>Variation. The three male paratypes (Figs. 1B–D) do not differ from the holotype. Length: 27.8–31.4 mm.</p><p>Etymology. The specific epithet is named in honor of Keita Maruyama who provided the specimens for this study.</p><p>Distribution. Spodochlamys keitai is known only from Otanche in Boyacá Department, Western Boyacá Province, in northeastern Colombia. Otanche is at an elevation of 1,050 m and surrounded by tropical rain forest. The specimens were purchased by Mr. Keita Maruyama from a local collector in Colombia.</p><p>Temporal Distribution. March (4).</p><p>Diagnosis. Spodochlamys keitai is immediately distinguished by its remarkably long and slender parameres (Figs. 2C–D) that are unlike any other in the genus. It is further differentiated from other species of Spodochlamys based on the characters in the key. The female remains unknown and so cannot be characterized.</p><p>We have not included the four species described by Soula (2010) ( Spodochlamys peruvianus Soula, Spodochlamys lydiae Soula, Spodochlamys curoei Soula, and Spodochlamys jamesonae Soula) in our key because Soula’s typically brief descriptions and images do not include some diagnostic characters, including the form of the labrum and whether all the claws are split or not; those are essential characters for our key. Based on available information, these species would go to couplet 9 in our key. The form of the parameres in Soula’s paper might distinguish those species, although S. curoei and S. jamesonae seem remarkably close to each other and Spodochlamys neitai Jameson and Ratcliffe, and S. peruvianus seems close to Spodochlamys feyeri Ohaus s ensu Jameson and Ratcliffe (2011), not Soula (2010).</p><p>In gestalt, S. keitai resembles S. curoei but differs in the following respects: 1) the surface adjacent to the elytral suture is more coarsely rugose and not finely punctate; 2) slender, hair-like setae on the parameres are completely absent; and 3) the parameres are more elongated and bi-arcuate.</p><p>Natural History. Nothing is known of the life history of S. keitai .</p></div>	https://treatment.plazi.org/id/03F887E1FFDC883551D9FBCCF63E1371	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kobayashi, Kazuo;Ratcliffe, Brett C.	Kobayashi, Kazuo, Ratcliffe, Brett C. (2023): A New Species of Spodochlamys Burmeister, 1855 (Coleoptera: Scarabaeidae: Rutelinae: Anatistini) from Colombia and a Key and Checklist of Spodochlamys Species. The Coleopterists Bulletin 77 (1): 120-126, DOI: 10.1649/0010-065X-77.1.120, URL: http://dx.doi.org/10.1649/0010-065x-77.1.120
03F887E1FFDB88365188FC9AF5E31124.text	03F887E1FFDB88365188FC9AF5E31124.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spodochlamys Burmeister 1855	<div><p>KEY TO SPECIES OF MALE SPODOCHLAMYS</p><p>Note: Males of Spodochlamys letiranti Jameson and Ratcliffe are not known. The four Soula species ( S. peruvianus, S. lydiae, S. curoei, S. jamesonae) are not included in the key for lack of known characters. See Diagnosis in text.</p><p>1. Head and pronotum brown, black, or cupreous, occasionally with metallic green reflections (with magnification and illumination). Elytral epipleuron (ventral view) lacking short, dense setae. Elytral surface lacking minute setae .... 2</p><p>1′. Head and pronotum greenish in color (with magnification and illumination). Elytral epipleuron (ventral view) with short, dense setae. Elytral surface with minute setae ..... 13</p><p>2. Pronotum distinctly vaulted in lateral view. Protibia greatly elongate and bowed. Elytra with waxy bloom ............. S. iheringi Ohaus</p><p>2′. Pronotum regularly convex in lateral view. Protibia simple, not elongate and not bowed. Elytra with or without bloom........................ 3</p><p>3. Clypeal apex strongly narrowed to a recurved, acute, narrowly bidentate tooth. Elytra with waxy bloom.............. S. caesarea Burmeister</p><p>3′. Clypeal apex rounded, lacking strong attenuation. Elytra without waxy bloom ................ 4</p><p>4. Pygidial disc protuberant in lateral view ..... 5</p><p>4′. Pygidial disc evenly convex in lateral view, not noticeably produced posteriorly.................... 9</p><p>5. Inner claws (all legs) simple. Apical umbone of elytra rounded, lacking spinose projection ................................ S. flavofemorata Ohaus</p><p>5′. Inner claw of meso- and metathoracic legs split, prothoracic leg with claw weakly split or simple. Apical umbone of elytra with spinose projection or tubercle ......................................... 6</p><p>6. Posterior margin of metafemur not narrowly laminate, not produced posteriorly ............... 7</p><p>6′. Posterior margin of metafemur narrowly laminate and produced posteriorly ............... 8</p><p>7. Inner metatibial spur strongly hooked. ........... ...... S. ancistrospina Jameson and Ratcliffe</p><p>7′. Inner metatibial spur not hooked ................... .............................................. S. soulai Curoe</p><p>8. Apical umbone of elytra with well developed, spinose projection ................. S. feyeri Ohaus</p><p>8′. Apical umbone of elytra tuberculate .............. ................... S. neitai Jameson and Ratcliffe</p><p>9. Apex of labrum acutely angulate ................... ............................................ S. gigas Murray</p><p>9′. Apex of labrum obtusely angulate .............. 10</p><p>10. Inner claw split at apex on all legs ................. ........................................... S. cupreola Bates</p><p>10′. Inner claw of prothoracic leg simple; claw split or not on meso- and metathoracic legs. ............................................................ 11</p><p>11. Elytral color dark brown, opaque ................... .......... S. marahuaca Jameson and Ratcliffe</p><p>11′. Elytral color reddish brown, shiny or with waxy bloom .......................................................... 12</p><p>12. Elytra with waxy bloom.................................. ................ S. neblina Jameson and Ratcliffe</p><p>12′. Elytra shiny, with weak metallic copper reflection ................ S. keitai Kobayashi and Ratcliffe, new species</p><p>13. Pronotum greatly vaulted in lateral view; in dorsal view, basolateral margins greatly explanate. Anterior margin of profemur with well developed, forward-projecting spine near apex............................................................. 14</p><p>13′. Pronotum regularly convex in lateral view; in dorsal view, basolateral margins not greatly explanate.Anterior margin of profemur lacking forward-projecting spine near apex ............ 15</p><p>14. Anterior margin of elytra with weak, external tooth. Mesotibia at apex acutely produced on external edge. Metatibia not greatly flattened in lateral view ........ S. mirabilis Waterhouse</p><p>14′. Anterior margin of elytra without external tooth. Mesotibia at apex quadrate, not produced on external edge. Metatibia greatly flattened in lateral view .......................... S. latipes Arrow</p><p>15. Pronotum on lateral margin near middle rounded, lacking sharp angulation. Pronotal surface rugopunctate ................... S. nazareti Arnaud</p><p>15′. Pronotum on lateral margin near middle with sharp angle. Pronotal surface densely punctate....................................................... 16</p><p>16. Inner margin of protibia weakly curved. Mentum with median, longitudinal, parallel-sided depression that extends from apex to base........... S. curvibrachialis Ohaus</p><p>16′. Inner margin of protibia straight, not weakly curved. Mentum with median depression that extends from apex to mid-disc ....................... ........................................... S. poultoni Shipp</p></div>	https://treatment.plazi.org/id/03F887E1FFDB88365188FC9AF5E31124	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kobayashi, Kazuo;Ratcliffe, Brett C.	Kobayashi, Kazuo, Ratcliffe, Brett C. (2023): A New Species of Spodochlamys Burmeister, 1855 (Coleoptera: Scarabaeidae: Rutelinae: Anatistini) from Colombia and a Key and Checklist of Spodochlamys Species. The Coleopterists Bulletin 77 (1): 120-126, DOI: 10.1649/0010-065X-77.1.120, URL: http://dx.doi.org/10.1649/0010-065x-77.1.120
