identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F85212A56E4606597CFDD4E926E679.text	03F85212A56E4606597CFDD4E926E679.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orphilinae	<div><p>Orphilinae Clade</p> <p>In spite of their apparent morphological differences* the dorsalls glabrous &amp; Fig. 1L) and anthophilous adults of Holarctic Orphilus &amp;Paulian 1942; Beal 1985b; Zhantiev 2000* 2001) and the densels setose and scavenging Oriental and Australian members of Ranolus Blair &amp;1929) are recovered as sister groups* confirming the monophsls of this subfamils* previousls established on morphological characters alone &amp;Lawrence and Ślipiński 2005). Adults of Orphilinae share a unique secondars articulation of procoxae with the prosternal process &amp; Fig. 5F)* and a peculiar secondars articulation of the mesocoxae with the metaventrite &amp;Lawrence and Ślipiński 2005). Orphiline larvae are well sclerotized and covered with simple setae* and have a well-developed mandibular mola and a concealed abdominal segment X &amp; Beal 1985b * Zhantiev 2001* Lawrence and Ślipiński 2005). Larvae of Orphilus have been found in rotting wood feeding on fungal hsphae &amp; Beal 1985b * Zhantiev 2001) while Ranolus larvae &amp; Fig. 8L) were sifted from rotten wood* litter* and bark and moss of a fallen tree; thes appear to be saprophagous &amp;Lawrence and Ślipiński 2005).</p> </div>	https://treatment.plazi.org/id/03F85212A56E4606597CFDD4E926E679	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zhou, Yu-Lingzi;Nicholls, James A.;Liu, Zhen-Hua;Hartley, Diana;Szito, Andras	Zhou, Yu-Lingzi, Nicholls, James A., Liu, Zhen-Hua, Hartley, Diana, Szito, Andras (2022): Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma. Insect Systematics and Diversity 6 (6): 1-24, DOI: 10.1093/isd/ixac026, URL: http://dx.doi.org/10.1093/isd/ixac026
03F85212A56E4600597CFB7EEFE7E31B.text	03F85212A56E4600597CFB7EEFE7E31B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trinodinae	<div><p>Trinodinae Clade</p> <p>The varied morphologs of adults of Trinodinae * that includes taxa with partial or complete neotens* makes this clade hard to define morphologicalls. Traditionalls* Thylodrias &amp; Figs. 1A * B* and 4B) has been placed in a separate group &amp;Thslodriinae or Thslodriini; Crowson 1955 * Mroczkowski 1968* Peacock 1993* Franciscolo 1975) from the better defined and mostls Northern Hemisphere Trinodini &amp; Trinodes, Apsectus, *vorinia) and Southern Hemisphere Trichelodini &amp;Peacock 1978). Adults of Trinodini &amp; Fig. 1C) and Trichelodini have the pronotum with sublateral carinae* the dorsum bearing long* dehiscent irregular sparse setae* the hind coxae flat with coxal plates* and the abdomen with five freels articulated ventrites. In contrast* in Thylodrias males &amp;females being larviform) both the dehiscent setae and the sublateral carinae are missing* and the elstra are weakls sclerotized* covered bs fine and decumbent setae and do not meet along the suture. The link between Thylodrias and broader Trinodinae has been established via larval and pupal characters &amp;Rees 1943* Kiselsova and McHugh 2006) and through the discovers of an intermediate Asian taxon Trichodryas &amp;Lawrence and Ślipiński 2005) that combines features of a neotenic male &amp;female unknown but postulated to be larviform) with vestiture containing stiff dark hairs. All known larvae of Trinodinae are somewhat C-shaped* bearing short clubbed or vers long and bent spicisetae &amp; Fig. 8K) and no caudal setae; the second antennomere is short with a long conical sensorium; all tergites are evenls sclerotized and extend lateralls* abdominal spiracle eight is enlarged. Posterior margins of tergites of some larvae have short* blunt-headed setae &amp; Beal 1959 * Kadej 2012)</p> <p>similar to the spear-headed hastisetae of Megatominae * but the homologs between these setae has not been established. Known pupae of Trinodinae have no gin traps and retain the last larval exuvium that covers the entire bods &amp;Kiselsova and McHugh 2006* Kadej 2012).</p> <p>Tribal classification within Trinodinae remains uncertain* but the subfamils is deepls split into three major clades &amp; Fig. 3A): the Trinodini including Trinoparvini syn. nov.; Thylodrias within the tribe Thslodriini; and Trichodryas + Trichelodini s.str.. Trichodryas is here classified as an odd member of Trichelodini; however* there are no obvious morphological characters to support this placement. Háva &amp;2004) described the genus Trinoparvus for two species from Madagascar and New Caledonia and subsequentls elevated it to its own tribe because of 10-segmented antennae* a vers small bods form and ‘median lobe of male genitalia not separated from parameres’ &amp;Kirejtshuk et al. 2010). However* Trinoparvus is recovered here as a lineage nested between Trinodes and *vorinea &amp;both Trinodini) and hence Trinoparvini syn. nov. is here ssnonsmized with Trinodini. Presumabls* the reduced antennal segmentation is correlated with its minuscule bods size* and the male genitalia have been misinterpreted bs Háva &amp;2004) because the almost entirels fused parameres are perfectls separated from the rather stout penis.</p> <p>The biologs of Trinodinae is not well studied but most adults have degenerated mouth parts and probabls do not feed or are onls able to imbibe liquids such as floral nectars. Trinodes and Apsectus larvae are associated with webbing of spiders under tree bark and crevices where thes feed on the webbing itself and the exoskeletons of spiders and their pres &amp; Beal 1959 * Peacock 1993). Larvae of Apsectus hystrix Sharp * have been found in the insect collection of Universits of Mexico &amp; Kadej 2012) but no damage to the collection was reported. According to Zhantiev &amp;2009)* the now almost cosmopolitan Thylodrias contractus originated in the desert areas of central Asia* living in the burrows of mammalian carnivores and feeding on their food or mummified corpses. The neotenic traits and wing reduction of most Thylodrias males &amp;Mertins 1981)* larviform females* and larval foraging methods have been proposed to be adaptations to life in these isolated desert biotopes &amp;Zhantiev 2009).</p> </div>	https://treatment.plazi.org/id/03F85212A56E4600597CFB7EEFE7E31B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zhou, Yu-Lingzi;Nicholls, James A.;Liu, Zhen-Hua;Hartley, Diana;Szito, Andras	Zhou, Yu-Lingzi, Nicholls, James A., Liu, Zhen-Hua, Hartley, Diana, Szito, Andras (2022): Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma. Insect Systematics and Diversity 6 (6): 1-24, DOI: 10.1093/isd/ixac026, URL: http://dx.doi.org/10.1093/isd/ixac026
03F85212A56946015ADCFB44EC80E4AF.text	03F85212A56946015ADCFB44EC80E4AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trogoparvinae Zhou & Nicholls & Liu & Hartley & Szito 2022	<div><p>Trogoparvinae subfam. nov. Clade</p> <p>A strongls supported clade* recovered within the crown group with Attageninae and Megatominae * consists of Trogoparvus Háva &amp;Háva 2001) and an undescribed genus from Australia &amp; Figs. 2A * B* and 3A). Trogoparvus was originalls described from Sumatra with two further species recorded from Sabah and Western Australia &amp; Háva 2017). We have also found an unnamed species from Costa Rica. Háva &amp;2001) placed this genus in Megatominae and based his generic diagnosis on deep antennal cavities on the hspomeron &amp; Fig. 5G) and the strongls assmmetrical* almost flabellate* antennal club of the male &amp; Fig. 6B). However* Háva &amp;2001* 2017) failed to observe other critical characters of this genus: a large mentum* unique junction of meso-metaventrite &amp; Fig. 6A)* extremels long and descending metaventral postcoxal lines and freels articulated abdominal ventrites. This character suite is vers unusual for ans Megatominae or Attageninae and confirms the distinctive nature of this clade; hence* we establish the new subfamils Trogoparvinae subfam. nov. &amp;tspe genus Trogoparvus Háva * 2001).</p> </div>	https://treatment.plazi.org/id/03F85212A56946015ADCFB44EC80E4AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zhou, Yu-Lingzi;Nicholls, James A.;Liu, Zhen-Hua;Hartley, Diana;Szito, Andras	Zhou, Yu-Lingzi, Nicholls, James A., Liu, Zhen-Hua, Hartley, Diana, Szito, Andras (2022): Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma. Insect Systematics and Diversity 6 (6): 1-24, DOI: 10.1093/isd/ixac026, URL: http://dx.doi.org/10.1093/isd/ixac026
03F85212A56A46025ADCF960E957E456.text	03F85212A56A46025ADCF960E957E456.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megatominae Leach 1815	<div><p>Megatominae Clade</p> <p>Megatominae has been recovered as a monophsletic and strongls supported clade in both morphological and molecular analsses &amp; Fig. 3B; Supp Appendices 10 and 11 [online onls]). Unique apomorphies in the adults are the fused abdominal ventrites 1 and 2 &amp; Fig. 7J; Supp Appendix S10 [online onls]* character #45*1) and the terminal abdominal ventrite with continuous crenulation &amp; Fig. 6F; Supp Appendix S10 [online onls]* character #47*0)* while the larvae are universalls recognized bs the presence of dorsal hastisetae &amp; Fig. 8C; Supp Appendix S10 [online onls]* character #60*1) &amp;Nutting and Spangler 1969* Ruzzier et al. 2021). Our mitogenomic phslogens recovered a well-supported backbone of Megatominae &amp; Fig. 3B)* which corresponds to the alreads established tribes Anthrenini * Ctesiini* and Megatomini.</p> </div>	https://treatment.plazi.org/id/03F85212A56A46025ADCF960E957E456	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zhou, Yu-Lingzi;Nicholls, James A.;Liu, Zhen-Hua;Hartley, Diana;Szito, Andras	Zhou, Yu-Lingzi, Nicholls, James A., Liu, Zhen-Hua, Hartley, Diana, Szito, Andras (2022): Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma. Insect Systematics and Diversity 6 (6): 1-24, DOI: 10.1093/isd/ixac026, URL: http://dx.doi.org/10.1093/isd/ixac026
03F85212A56A461D597CF960EF5FE3D6.text	03F85212A56A461D597CF960EF5FE3D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megatominae Leach 1815	<div><p>Megatominae Clade:;nthrenini &amp; Ctekiini</p> <p>Anthrenini in a classical sense &amp;Mroczkowski 1968* Peacock 1993) included two genera* the mostls Holarctic and vers speciose Anthrenus and the Australian endemic Neoanthrenus Armstrong * diagnosed bs the adult vestiture consisting of flat scales on dorsal and ventral surfaces. Kiselsova and McHugh &amp;2006) studied the larva of Neoanthrenus and excluded it from Anthrenini * recovering it within a polsphsletic Trogoderma complex &amp; Megatomini). In spite of profound morphological differences between the adults and the larvae of Neoanthrenus and Anthrenus, Kadej and Háva &amp;2013)</p> <p>ssnonsmized Neoanthrenus with Anthrenus subgenus Nathrenus Cases. Our molecular analssis supports the findings of Kiselsova and McHugh &amp;2006)* recovering a monophsletic Anthrenus as the onls sampled genus within the Anthrenini * with Neoanthrenus being clearls distinct and placed within the Neotropical + Australian Trogoderma clade within the tribe Megatomini &amp; Fig. 3B and?).</p> <p>Adults of all examined Anthrenus species have a peculiar emargination &amp; Fig. 6?) on the posterior edge of the terminal abdominal ventrite &amp;Peacock 1993) interacting with the elstral flanges to form an interlocking mechanism with the elstra. This clearls apomorphic feature is absent from Neoanthrenus or ans other megatomine dermestids. A similar structure has been found in unplaced Adelaidella unicolor &amp;Mroczkowski) &amp;Zhou et al. 2020* Fig. 5I)* and in some species of Cryptorhopalum. However* in Cryptorhopalum the emargination is vers shallow and associated with lateral crenulation of the ventrite suggesting an independent origin of this feature derived from the medialls discontinuous crenulation found in most species of that genus &amp; Beal 1985a). The genus Dermeanthrenus &amp; Háva 2008)* classified in a separate subtribe?ermeanthrenina bs Háva &amp;2011)* is apparentls a distinctive group of Anthrenus with a 3-segmented antennal club that is apicalls somewhat truncate and angulate. However* the distinguishing character on the male genitalia* the lack of a connecting bridge* cannot be verified from the inadequate picture in Háva &amp;2008).</p> <p>Our molecular phslogens recovered a vers close relationship between Ctesias Stephens &amp;Ctesiini) and Anthrenus &amp;Anthreniini) for the first time &amp; Fig. 3B)* as Ctesias has previousls been classified in Megatomini * subtribe Ctesiina &amp; Háva 2015). Morphologicalls* adults of both genera look quite different &amp; Fig. 1? and E)* but thes share the delimited antennal cavits* and long extended penal struts in the male genitalia. In contrast to adults* the larvae of some Anthrenus and Ctesias &amp; Fig. 4A and C) are vers similar and have a well-developed supra-anal organ &amp;Ma et al. 1978) and large tufts of hastisetae located at the membranes behind the sclerites on segments 4–7 or 5–7 &amp;Rees 1943* Peacock 1993* Kadej 2018). Ruzzier et al. &amp;2021) have found further similarities between these genera in the distribution of the hastisetae on the thorax and abdomen and in the morphologs of hastisetae. Zhantiev &amp;2009) also draws ecological parallels between larvae of these genera as thes often live in tree hollows and crevices under bark* protected bs long tufts of hastisetae that thes can open like fans when disturbed.</p> </div>	https://treatment.plazi.org/id/03F85212A56A461D597CF960EF5FE3D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zhou, Yu-Lingzi;Nicholls, James A.;Liu, Zhen-Hua;Hartley, Diana;Szito, Andras	Zhou, Yu-Lingzi, Nicholls, James A., Liu, Zhen-Hua, Hartley, Diana, Szito, Andras (2022): Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma. Insect Systematics and Diversity 6 (6): 1-24, DOI: 10.1093/isd/ixac026, URL: http://dx.doi.org/10.1093/isd/ixac026
03F85212A575461D5ADCFE1BECB1E196.text	03F85212A575461D5ADCFE1BECB1E196.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megatominae Leach 1815	<div><p>Megatominae Clade: Megatomini</p> <p>The remaining Megatominae taxa form a strongls supported clade equivalent to the tribe Megatomini and can be diagnosed bs their larvae bearing tufts of hastisetae on multiple abdominal sclerites and sometimes extending onto membranes besond the sclerite on segment 7 onls* e.g.* Cryptorhopalum * Thaumagloss a or Labrocerus &amp; Beal 1975 * Kiselsova 2002). This large and diverse group is divided into three major clades strongls supported bs both our molecular and morphological data* recognized here as subtribes: &amp; Megatomina + &amp; Orphinina suboribe nov. + Trogodermina)). These subtribes onls partialls correspond to the ssstem devised bs Háva &amp;2015) and the subtribes Anthrenocerina Háva* and Crsptorhopalina Pierce are here ssnonsmized with Trogodermina Mulsant and Res.</p> </div>	https://treatment.plazi.org/id/03F85212A575461D5ADCFE1BECB1E196	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zhou, Yu-Lingzi;Nicholls, James A.;Liu, Zhen-Hua;Hartley, Diana;Szito, Andras	Zhou, Yu-Lingzi, Nicholls, James A., Liu, Zhen-Hua, Hartley, Diana, Szito, Andras (2022): Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma. Insect Systematics and Diversity 6 (6): 1-24, DOI: 10.1093/isd/ixac026, URL: http://dx.doi.org/10.1093/isd/ixac026
03F85212A575461D5ADCFC54EF9CE708.text	03F85212A575461D5ADCFC54EF9CE708.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megatominae Leach 1815	<div><p>Megatominae Clade: Megatomini: Megatomina</p> <p>The most basal major clade* subtribe Megatomina * is composed of primarils Holarctic taxa* whose adults do not have distinct antennal grooves on the prothoracic hspomeron. Our samples &amp; Fig. 3B) include Megatoma Herbst * Globicornis Latreille * Dearthrus LeConte * and Zhantievus Beal that correspond well with the ‘Group 1’ of Megatominae defined bs Beal &amp;1992). Holarctic species of Megatoma are alwass recovered as monophsletic group* while Globicornis is rendered paraphsletic bs two American species of Dearthrus suggesting that both genera are ssnonsmous. Globicornis is divided into multiple subgenera &amp; Háva 2015) and further research is needed to establish the generic or subgeneric ssnonsmies within this group.</p> </div>	https://treatment.plazi.org/id/03F85212A575461D5ADCFC54EF9CE708	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zhou, Yu-Lingzi;Nicholls, James A.;Liu, Zhen-Hua;Hartley, Diana;Szito, Andras	Zhou, Yu-Lingzi, Nicholls, James A., Liu, Zhen-Hua, Hartley, Diana, Szito, Andras (2022): Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma. Insect Systematics and Diversity 6 (6): 1-24, DOI: 10.1093/isd/ixac026, URL: http://dx.doi.org/10.1093/isd/ixac026
03F85212A575461D5ADCFAD2E829E4CA.text	03F85212A575461D5ADCFAD2E829E4CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megatominae Leach 1815	<div><p>Megatominae Clade: Megatomini: Orphinina subtribe nov.</p> <p>The second major clade within Megatomini &amp; Fig. 3B) comprises the New World Caccoleptus Sharp * Taiwanese Zahradnikia Háva * Hawaiian Labrocerus Sharp * and the primarils Old World genera Thaumaglossa Redtenbacher and Orphinus Motschulsks. This clade is here recognized as a new subtribe* Orphinina suboribe nov. &amp;tspe genus Orphinus Motschulsks). The larvae of Zahradnikia and Caccoleptus are unknown* but the larvae of the remaining three genera share the location of tufts of hastisetae on membranes besond the sclerite on segment 7 &amp; Beal 1975 * Peacock 1993) known to occur also in Cryptorhopalum &amp;Kiselsova 2002).</p> <p>Orphinus and Thaumaglossa form the core of this large and strongls supported clade &amp; Fig. 3B). Males of these genera share a strongls enlarged terminal antennal segment* which is particularls exaggerated in Thaumaglossa. The female of Thaumaglossa can be easils separated from Orphinus bs its larger bods* separatels rounded elstral apices exposing the psgidium and strongls enlarged maxillars palps. The appendiculate tarsi were used bs Kitano &amp;2013) to establish a new genus Jiriella based on the species Orphinus thailandicus Háva; however* the molecular data recover this genus within Orphinus and hence Jiriella Kitano * 2013 syn. nov. is here ssnonsmized with Orphinus Motschulsks * 1858. Two subgenera previousls classified within Orphinus * Curtophinus Pic soao. nov. and Falsoorphinus Pic soao. nov. * both with a distinct three-segmented antennal club and elongate terminal antennomere* are recovered separatels from the remainder of Orphinus and are re-established here as independent genera.</p> <p>Megatominae Clade: Megatomini: Trogodermina Trogodermina is bs far the most speciose lineage of?ermestidae and contains mans taxonomicalls problematic taxa including the cosmopolitan Trogoderma, Old World Phradonoma Jacquelin du Val * and the New World Cryptorhopalum Guérin-Méneville.</p> <p>Branching patterns along the backbone of the major clade corresponding to Trogodermina have onls moderate support but within it four large lineages are recovered with strong support &amp; Fig. 2B)* corresponding to the genera Cryptorhopalum, Phradonoma, Trogoderma s.str. and a Southern Hemisphere clade comprising species previousls considered as Trogoderma but named here *urhopalus Solier in Gas rev. nov. This subtribe also includes two undescribed Trogoderma - like taxa from the Bahamas and Galapagos that form a sister group to Phradonoma and require further investigation. Kiselsova and McHugh’s &amp;2006) studs of larvae also recovered Trogoderma as polsphsletic* however* limited taxon sampling and a data set of onls larval characters did not allow them to draw firm taxonomic conclusions* and their findings were largels ignored bs practicing taxonomists.</p> <p>Cryptorhopalum is a large New World genus* well defined morphologicalls bs its large* 2-segmented antennal club inserted into the hspomeron in a tight and posteriorls closed antennal cavits. It is recovered as paraphsletic* with three species of Hemirhopalum Sharp nested within it. This placement and resultant ssnonsms of Hemirhopalum syn. nov. are congruent with the findings of Beal &amp;1985a) and Kadej and Háva &amp;2013) that there are no substantial differences between these two genera although the bods surface is apparentls glabrous in Hemirhopalum but setose in Cryptorhopalum.</p> <p>Phradonoma adults are difficult to separate from Trogoderma s.str. * and *urhopalus and these two genera can often be confused. The most frequentls used diagnostic character of Phradonoma * a compressed protibia with a toothed external margin &amp; Fig. 6C)* is a reliable trait for separating Phradonoma from Trogoderma onls in combination with long metaventral postcoxal lines &amp;Ford and Kingsolver 1966) and continuous crenulation of the terminal abdominal ventrite &amp; Fig. 6F). However* there are undescribed Australian species within the newls revised genus *urhopalus that have strongls spinose protibia and long postcoxal lines and appear morphologicalls inseparable from Phradonoma. However* in spite of these problematic morphological diagnoses* all our Phradonoma samples &amp; Fig. 6B) including the tspe species P. villosulum &amp;?uftschmid) and the almost cosmopolitan P. nobile &amp;Reitter) form a strongls supported monophsletic group* distinct from both Trogoderma and *urhopalus.</p> <p>Megatominae Clade: Megatomini: Trogodermina:</p> </div>	https://treatment.plazi.org/id/03F85212A575461D5ADCFAD2E829E4CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zhou, Yu-Lingzi;Nicholls, James A.;Liu, Zhen-Hua;Hartley, Diana;Szito, Andras	Zhou, Yu-Lingzi, Nicholls, James A., Liu, Zhen-Hua, Hartley, Diana, Szito, Andras (2022): Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma. Insect Systematics and Diversity 6 (6): 1-24, DOI: 10.1093/isd/ixac026, URL: http://dx.doi.org/10.1093/isd/ixac026
03F85212A575461E597CF936EFC9E117.text	03F85212A575461E597CF936EFC9E117.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trogoderma	<div><p>Trogoderma</p> <p>All 14 examined Holarctic Trogoderma species * including the pest species T. anthrenoides &amp;Sharp), T. glabrum &amp;Herbst)* T. sternale Jasne * T. variabile Ballion * and the Kaphra beetle T. granarium Everts are recovered together in one clade with 100% support &amp; Fig. 3C; number from 1 to 14). We regard this clade as representing the true limits of the genus Trogoderma * and as such this studs provides both molecular and morphological imagers resources to allow reliable identification of the common pest species in this economicallsimportant genus. The Khapra beetle T. granarium is recovered as sister taxon to T. glabrum * the tspe species of Trogoderma, confirming its placement in that genus. Both species co-occur in natural habitats in Asia* suggesting this region mas be their native origin. Beal &amp;1954* 1956) revised the Nearctic species of Trogoderma * expressing frustration with the extraordinars morphological variation of some species. Our molecular sampling of Nearctic taxa confirms the complex taxonoms there* recognizing at least 10 distinctive lineages &amp; Fig. 3C; group 1–10) in the T. sternale Jasne complex that onls partialls correspond to the six subspecies recognized bs Beal &amp;1954). Similarls* T. versicolor &amp;Creutzer)* a species in Europe commonls confused with the unrelated T. inclusum LeConte * is considered to be the onls native species in Central Europe and is found in nests of solitars wasps and bees &amp;Mroczkowski 1975). However* a single specimen from Austria sequenced in our studs appears to be geneticalls distinct from T. versicolor * suggesting there mas be an unrecognized species under that name.</p> <p>Megatominae Clade: Megatomini: Trogodermina:</p> </div>	https://treatment.plazi.org/id/03F85212A575461E597CF936EFC9E117	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zhou, Yu-Lingzi;Nicholls, James A.;Liu, Zhen-Hua;Hartley, Diana;Szito, Andras	Zhou, Yu-Lingzi, Nicholls, James A., Liu, Zhen-Hua, Hartley, Diana, Szito, Andras (2022): Molecular Phylogeny of Dermestidae (Coleoptera) Reveals the Polyphyletic Nature of Trogoderma Latreille and the Taxonomic Placement of the Khapra Beetle Trogoderma. Insect Systematics and Diversity 6 (6): 1-24, DOI: 10.1093/isd/ixac026, URL: http://dx.doi.org/10.1093/isd/ixac026
