identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F987C08A7301733AEE8F34FB4448E2.text	03F987C08A7301733AEE8F34FB4448E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidraus Simon 1900	<div><p>Amphidraus Simon, 1900</p><p>Amphidraus Simon, 1900: 60 (type species: Amphidraus auriga Simon, 1900, original designation).</p></div>	https://treatment.plazi.org/id/03F987C08A7301733AEE8F34FB4448E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Salgado, Alexandre;Ruiz, Gustavo R. S.	Salgado, Alexandre, Ruiz, Gustavo R. S. (2019): Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini). Zootaxa 4563 (3): 451-481, DOI: 10.11646/zootaxa.4563.3.3
03F987C08A7301703AEE8FD5FB9C4F9A.text	03F987C08A7301703AEE8FD5FB9C4F9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidraus cornipalpis Salgado & Ruiz 2019	<div><p>Amphidraus cornipalpis sp. nov.</p><p>Figs 1–6, 14</p><p>Type material. Holotype: Ƌ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.55578&amp;materialsCitation.latitude=-9.175389" title="Search Plazi for locations around (long -43.55578/lat -9.175389)">Parque Nacional da Serra</a> das Confusões, [09°10'31.4"S, 43°33'20.8"W], Guaribas, Piauí, Brazil, 15.XII.10, L.S. Carvalho et al. leg. (MPEG 35014). Paratypes: 1 ♀, same data as holotype [MPEG 35015]; 1♂ and 1♀ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-35.96853&amp;materialsCitation.latitude=-8.285723" title="Search Plazi for locations around (long -35.96853/lat -8.285723)">Caruaru</a>, [08°17'08.6"S, 35°58'06.7"W], Pernambuco, Brazil, 2009–2010, H.P. Amorim leg. [IBSP 160615 (♂), 160616 (♀)] .</p><p>Etymology. The specific epithet is a compound adjective in Latin (cornus + palpus) and refers to the shape of distal process on the embolic disc (Fig. 3A).</p><p>Diagnosis. Males of Amphidraus cornipalpis sp. nov. share similarities with those of Amphidraus loxodontillus Salgado &amp; Ruiz, 2017, both with an embolic disc with one poorly-sclerotized, distal lamella that covers part of dPED ventral view (white arrows in Figs 3 A–B). However, males of A. cornipalpis sp. nov. can be distinguished from those of A. loxodontillus by having a longer and acute distal process on embolic disc (dPED) with the tip curved distally (shaped like a cow horn) (Figs 2C, 3A, 5A, 5D, 6 D–E), whereas in A. loxodontillus the dPED is short and rounded (Fig. 3B; see also Salgado &amp; Ruiz 2017: fig. 26C). In addition, A. cornipalpis sp. nov. has a reduced RTA (as large as RvTA in A. loxodontillus) and a small ITA in the male palp (Figs 5B, 5E), which is absent in A. loxodontillus (see Salgado &amp; Ruiz 2017: figs 26B, 26D). Also, males of A. cornipalpis sp. nov. differ from those of all other species of Amphidraus by having a tubercle placed on proximal dorso-retrolateral portion of the cymbium (black arrows in Figs 2 C–F, 5A–B, 5E, 6C) (present also in Yacuitella nana Galiano, 1999). Females of A. cornipalpis sp. nov. are similar to those of Y. nana by the arrangement of copulatory ducts and spermathecae in the epigyne, but differ from that species by having more separate copulatory openings, longer copulatory ducts and no tubercle on the epigynal plate (Fig. 5F), which is medially placed in Y. nana (see Galiano 1999: figs 12–14).</p><p>Description. Male holotype (MPEG 35014). Total length: 3.33. Carapace 1.78 long, 1.23 wide and 0.89 high. Ocular quadrangle 0.83 long. Anterior eye row 1.25 wide, posterior 1.13 wide. Legs 4312. Length of leg: I 2.65 (0.83 + 1.00 + 0.82); II 2.54 (0.81 + 0.93 + 0.80); III 3.12 (1.01 + 1.09 + 1.02); IV 3.40 (0.97 + 1.19 + 1.24). Palp (Figs 2 C–F, 3A, 5A–E, 6A–E): femur with large and low bump on median-ventral portion (Fig. 5C); patella with no modifications; tibia with a prolateral rounded expansion and a ventro-distal depression (Figs 2 D–F, 5A, 5E, 6A); large RvTA with acute tip (Figs 2D, 5B, 6C); short, poorly-sclerotized ITA retrolaterally projected (Figs 2F, 5E, 6C); cymbium oval (Figs 2F, 5E), with conductor (cc) wider than long with sinuous proximal border (Figs 2E, 6 A–B); tegulum with slender lobe curved to prolateral side (Figs 2C, 3A, 5A, 5D); embolus shaft (e) with tip strongly curved to retrolateral side (Figs 2C, 3A, 5A); embolic filament (ef) longer than embolus shaft (Figs 3A, 5A, 5D). Color in alcohol (Figs 1, 2 A–B): cephalic area black; thoracic area light brown with a black procurved arc; chelicera light brown; palp light brown, with black mark ventrally on femur and with dark brown cymbium; mouthparts and sternum light brown; abdomen dorsally and ventrally light brown variegated with dark brown; legs light brown with a distal black ring on metatarsi III and two black rings on metatarsi IV, one proximal and another distal.</p><p>Female paratype (MPEG 35015). Total length: 3.77. Carapace 1.66 long, 1.15 wide and 0.81 high. Ocular quadrangle 0.80 long. Anterior eye row 1.13 wide, posterior 1.00 wide. Legs 4312. Length of leg: I 2.54 (0.83 + 1.00 + 0.71); II 2.25 (0.73 + 0.88 + 0.64); III 3.08 (1.05 + 1.05 + 0.98); IV 3.55 (1.08 + 1.23 + 1.24). Epigyne/vulva (Figs 4C, 5F): epigynal plate with posterior border slightly bilobed; copulatory openings separated by approximately two diameters of copulatory ducts; long and convoluted copulatory ducts, forming a pair of rounded masses with spermathecae; oval spermathecae diagonally projected; fertilization ducts anteriorly placed and laterally projected. Color in alcohol (Figs 4 A–B): cephalic area black; thoracic area dorso-laterally dark brown and centrally light brown; abdomen dorsally and laterally variegated with brown; ventrally pale; pale legs with black spots, being more intense on metatarsi III and IV (same pattern as in male).</p><p>Distribution. Known from the states of Pernambuco and Piauí (Brazil) (Fig. 14).</p></div>	https://treatment.plazi.org/id/03F987C08A7301703AEE8FD5FB9C4F9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Salgado, Alexandre;Ruiz, Gustavo R. S.	Salgado, Alexandre, Ruiz, Gustavo R. S. (2019): Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini). Zootaxa 4563 (3): 451-481, DOI: 10.11646/zootaxa.4563.3.3
03F987C08A7101743AEE8D0CFBCF4D6A.text	03F987C08A7101743AEE8D0CFBCF4D6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidraus boomerang Salgado & Ruiz 2019	<div><p>Amphidraus boomerang sp. nov.</p><p>Figs 7–8, 14</p><p>Type material. Holotype: Ƌ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.587276&amp;materialsCitation.latitude=-4.7509446" title="Search Plazi for locations around (long -42.587276/lat -4.7509446)">Fazenda Nazareth</a>, [4°45'03.4"S; 42°35'14.2"W], José de Freitas, Piauí, Brazil, 31.I.2004, V.O. Costa leg. (MPEG 9881).</p><p>Etymology. The specific epithet is a noun in apposition and refers to embolic disc, boomerang-shaped (Figs 7C, 8A).</p><p>Diagnosis. A. boomerang sp. nov. differs from all Amphidraus species by having the embolic disc boomerangshaped, due its expanded retrolateral border (Figs 7C, 8A). The retrolateral triangular lamella on the embolus shaft of A. boomerang sp. nov. is also present in A. argentinensis Galiano, 1997, near the emergence point of the embolic filament (see Galiano 1997: figs 9–10). A. boomerang sp. nov. can be distinguished from A. argentinensis by shape of the RvTA, large and with a ventral lobe in A. argentinensis (Galiano 1997: fig. 11) and reduced in A. boomerang sp. nov. (Fig. 8B).</p><p>Description. Male holotype (MPEG 9881). Total length: 3.08. Carapace 1.62 long, 1.07 wide and 0.85 high. Ocular quadrangle 0.83 long. Anterior eye row 1.17 wide, posterior 0.98 wide. Legs 4312. Length of leg: I 2.65 (0.82 + 1.01 + 0.82); II 2.49 (0.80 + 0.91 + 0.78); III 3.15 (0.99 + 1.05 + 1.11); IV 3.47 (1.02 + 1.21 + 1.24). Palp (Figs 7 C–F, 8A–D): femur and patella with no modifications; short RvTA with two apical teeth (Figs 7D, 8B); straight RTA with the same length of tibia and with membranous external border (Figs 7D, 7F, 8B, 8D); cymbium oval, with proximal dorso-prolateral portion prominent (Figs 7F, 8D); tegulum with thick and straight lobe (Figs 7C, 8A); embolic disc with a process with acute tip (dPED) placed on the median distal portion and projected to prolateral side, with a ventral process (vPED) ventrally projected and with a proximal process (pPED) projected to proximal side, hidden under the tegulum (Figs 7 C–E, 8A, 8C); embolus shaft (e) with tip curved to retrolateral side and with two processes: a triangular lamella placed at the middle of retrolateral border and a rounded basal process (Figs 7C, 8A); embolic filament (ef) approximately 1/2 the length of embolus shaft (Figs 7C, 8A). Color in alcohol (Figs 7 A–B): cephalic area black; thoracic area brown with light brown longitudinal stripe; abdomen dorsally variegated with light brown and with longitudinal stripe of white setae; ventrally pale; pale legs.</p><p>Female. Unknown.</p><p>Distribution. Known only from the type locality (state of Piauí) (Fig. 14).</p></div>	https://treatment.plazi.org/id/03F987C08A7101743AEE8D0CFBCF4D6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Salgado, Alexandre;Ruiz, Gustavo R. S.	Salgado, Alexandre, Ruiz, Gustavo R. S. (2019): Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini). Zootaxa 4563 (3): 451-481, DOI: 10.11646/zootaxa.4563.3.3
03F987C08A74017A3AEE8E9EFEC24BA9.text	03F987C08A74017A3AEE8E9EFEC24BA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidraus sacrificatus Salgado & Ruiz 2019	<div><p>Amphidraus sacrificatus sp. nov.</p><p>Figs 9–10, 14</p><p>Type material. Holotype: ♂ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.408&amp;materialsCitation.latitude=-7.9898615" title="Search Plazi for locations around (long -49.408/lat -7.9898615)">Conceição do Araguaia</a>, 07°59’23.5”S, 49°24’28.8”W, Pará, Brazil, 16.I.2018, R.P.S Almeida. &amp; F.V. Arruda leg. (MPEG 35027). Paratypes: 2♂, same data as holotype (MPEG 34602, IBSP 221863) .</p><p>Etymology. The specific epithet an adjective and refers to the heart-shaped membrane linked to proximal process on embolic disc (Fig. 10B), in allusion to human sacrificial rituals practiced by some ancient cultures.</p><p>Diagnosis. Differs from all Amphidraus species in having a blade-shaped embolus shaft (Figs 9F, 10 A–B) and a proximal process on embolic disc (pPED) linked to a heart-shaped membrane [Figs 9 C–D, 9F (white arrow), 10A–B, 10D]. In addition, the cymbial conductor (cc) is “Y-shaped” and faces retrolaterally (Figs 9E, 10C) (cc faces ventrally in all the other described species; see Figs 13A, 17A).</p><p>Description. Male holotype (MPEG 35027). Total length: 4.45. Carapace 2.13 long, 1.46 wide, 1.03 high. Ocular quadrangle 1.10 long. Anterior eye row 1.48 wide, posterior 1.19 wide. Legs 4312. Length of legs: I 3.79 (1.22 + 1.43 + 1.14); II 3.62 (1.19 + 1.28 + 1.15); III 4.46 (1.40 + 1.45 + 1.61); IV 4.75 (1.39 + 1.61 + 1.75). Palp (Figs 9 C–F, 10A–E): femur and patella with no modifications; RvTA with two short branches, the more ventral branch is reduced and rounded, and the more dorsal branch is larger, with acute tip (Figs 9D, 10D); RTA with wide base, shorter than the tibia and with acute tip (Figs 9D, 10 D–E); cymbium oval (Fig. 10E); tegulum with stout lobe curved to prolateral side (Figs 9C, 9F, 10A); embolic disc with three processes very close together on its distal portion: two short and rounded distal process (dPED) and one short and triangular ventral process (vPED) (Figs 9D, 9F, 10 A–B, 10D); embolic filament (ef) very short, approximately 1/5 the length of embolus shaft (Figs 9F, 10 A–B). Color in alcohol (Figs 9 A–B): cephalic area black with yellow setae near the eyes; thoracic area laterally black and centrally dark brown with yellow setae on median-posterior portion; abdomen dorsally variegated with black and dark brown, and with a longitudinal stripe of white setae; ventrally variegated with brown; legs I almost black and legs II–IV dark brown; tarsi pale in all legs.</p><p>Female. Unknown.</p><p>Note. The RvTA was interpreted herein as an apophysis with two branches (Figs 9D, 10D). However, in other Amphidraus species, such as A. araripe sp. nov., these branches are not fused and therefore interpreted as two distinct apophyses: one RvTA and one ITA (Figs 11F, 13B). The processes placed proximally on embolic disc and herein named as the pPED in A. sacrificatus sp. nov. and A. boomerang sp. nov. (Figs 7C, 8A) are possibly homologous. However, in A. sacrificatus sp. nov. it is linked to a heart-shaped membrane [Figs 9C, 9F (white arrow), 10A–B]. The homologies of all these structures across the genus need to be tested in future studies.</p><p>Distribution. Known only from the type locality (state of Pará, Brazil) (Fig. 14).</p><p>Natural History. The specimens were collected from an ecotonal zone between the Cerrado and the Amazon rainforest.</p></div>	https://treatment.plazi.org/id/03F987C08A74017A3AEE8E9EFEC24BA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Salgado, Alexandre;Ruiz, Gustavo R. S.	Salgado, Alexandre, Ruiz, Gustavo R. S. (2019): Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini). Zootaxa 4563 (3): 451-481, DOI: 10.11646/zootaxa.4563.3.3
03F987C08A7B017C3AEE8DA0FBA94A95.text	03F987C08A7B017C3AEE8DA0FBA94A95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidraus araripe Salgado & Ruiz 2019	<div><p>Amphidraus araripe sp. nov.</p><p>Figs 11–14</p><p>Type material. Holotype: Ƌ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.447002&amp;materialsCitation.latitude=-7.3110003" title="Search Plazi for locations around (long -39.447002/lat -7.3110003)">Floresta Nacional do Araripe-Apodi</a>, 7°18'39.6"S, 39°26'49.2"W, Crato, Ceará, Brazil, 06.II.2011, L.S. Carvalho et al. leg. (MPEG 35011); Paratypes: 1 Ƌ and 1♀ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.899612&amp;materialsCitation.latitude=-3.8387222" title="Search Plazi for locations around (long -40.899612/lat -3.8387222)">Parque Nacional de Ubajara</a>, Trilha da Samambaia, 3°50'19.4"S, 40°53'58.6"W, Ubajara, Ceará, Brazil, 14.V.2013, L.S. Carvalho et al. leg. (UFMG 14887) .</p><p>Etymology. The specific epithet is a noun in apposition taken from the locality of the holotype.</p><p>Diagnosis. Among all species of Amphidraus, A. araripe sp. nov. is most similar to A. pulvinus Salgado &amp; Ruiz, 2017 by having the tegulum with small retrolateral spermophore loop (RSPL) (Figs 11D, 13B; see Salgado &amp; Ruiz 2017: fig. 8C) and the same tibial apophysis pattern: the retroventral tibial apophysis (RvTA) and intercalary tibial apophysis (ITA) are short and blade-shaped (the ITA with an internal basal projection; black arrow in Fig. 11E; see also Salgado &amp; Ruiz 2017: fig. 8D) and the RTA is linked to an inflated membrane (Figs 11 E–G, 13B–C; see Salgado &amp; Ruiz: figs 8B, 8D). These species also have an embolic disc with a distal process projected to prolateral side and a ventral process projected to retrolateral side (Figs 11C, 13A; see Salgado &amp; Ruiz 2017: fig. 8C). However, A. araripe sp. nov. can be distinguished by having the RvTA and ITA more separated from each other (Figs 11F, 13B), whereas these are almost fused in A. pulvinus (see Salgado &amp; Ruiz 2017: fig. 8D). In addition, A. araripe sp. nov. has a vPED placed near the distal processes (Figs 11C, 13A), whereas in A. pulvinus the vPED is placed on the prolateral border of embolic disc (see Salgado &amp; Ruiz 2017: fig. 8C). The female of A. pulvinus is unknown, but among the Amphidraus species currently known, the epigyne of A. araripe sp. nov. is most similar to that of A. manaura sp. nov., both with glands near copulatory openings and with epigynal plate with border slightly bilobed (Figs 12C, 13D, 16D, 17D). However, the female of A. araripe sp. nov. differs by having a longer copulatory duct (with one loop ventrally projected) and by having the less expanded border of epigynal plate (Figs 13D, 17D).</p><p>Description. Male holotype (MPEG 35011). Total length: 2.73. Carapace 1.32 long, 0.88 wide and 0.65 high. Ocular quadrangle 0.54 long. Anterior eye row 0.93 wide, posterior 0.81 wide. Legs 4312. Length of leg: I 2.09 (0.63 + 0.77 + 0.69); II 2.04 (0.62 + 0.70 + 0.72); III 2.50 (0.75 + 0.82 + 0.93); IV 2.78 (0.79 + 0.94 + 1.05). Palp (Figs 11 C–G, 13A–C): femur and patella without modifications; cymbium oval (Fig. 13C); tegulum with thick lobe curved to prolateral side (Figs 11C, 13A); embolus shaft (e) strongly arched to prolateral side, with tip retrolaterally projected (Fig. 13A); embolic filament (ef) short, with nearly half the length of embolus shaft (Fig. 13A). Color in alcohol (Figs 11 A–B): cephalic area black; thoracic area brown with pale longitudinal stripe; abdomen dorsally variegated with brown, and with a pale longitudinal stripe bearing white setae; ventrally pale with posterior portion dark; legs pale.</p><p>Description. Female paratype (UFMG 14887). Total length: 2.95. Carapace 1.40 long, 0.97 wide and 0.67 high. Ocular quadrangle 0.68 long. Anterior eye row 1.02 wide, posterior 0.90 wide. Legs 4312. Length of leg: I 2.18 (0.71 + 0.82 + 0.65); II 2.06 (0.66 + 0.76 + 0.64); III 2.50 (0.79 + 0.86 + 0.85); IV 2.88 (0.81+ 0.99 + 1.08). Epigyne/vulva (Figs 12 C–D, 13D: copulatory openings separated by about two diameters of copulatory duct; spermathecae kidney-shaped; fertilization ducts anteriorly placed and laterally projected. Color in alcohol (Figs 12 A–B): cephalic area black; thoracic area dark brown with a longitudinal light brown stripe; abdomen dorsally black with longitudinal interrupted pale stripe; ventrally pale; femora pale with black spots; other leg segments light brown with black spots.</p><p>Distribution. Known only from the type locality (state of Ceará, Brazil) (Fig. 14).</p></div>	https://treatment.plazi.org/id/03F987C08A7B017C3AEE8DA0FBA94A95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Salgado, Alexandre;Ruiz, Gustavo R. S.	Salgado, Alexandre, Ruiz, Gustavo R. S. (2019): Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini). Zootaxa 4563 (3): 451-481, DOI: 10.11646/zootaxa.4563.3.3
03F987C08A7C01633AEE8C24FB5E4DFE.text	03F987C08A7C01633AEE8C24FB5E4DFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidraus manauara Salgado & Ruiz 2019	<div><p>Amphidraus manauara sp. nov.</p><p>Figs 15–17, 19 A–B, 19E, 20</p><p>Type material. Holotype: ♂ from Fazenda Experimental da UFAM, 02°38'55.8"S, 60°03'09.4"W, Manaus, Amazonas, Brazil, unknown date, E. P. Franken &amp; A.C.K Silva leg. (MPEG 35025). Paratype: 1♀, same data as holotype (MPEG 35026) .</p><p>Additional material examined. BRAZIL: Amazonas: 1♀ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.05261&amp;materialsCitation.latitude=-2.6488333" title="Search Plazi for locations around (long -60.05261/lat -2.6488333)">Fazenda Experimental</a> da UFAM, 02°38'55.8"S, 60°03'09.4"W, Manaus, unknown date, E.P. Franken &amp; A.C.K Silva leg. (MPEG 34601) .</p><p>Etymology. The specific epithet (to be treated as a noun in apposition) means a person who lives in Manaus (Brazil).</p><p>Diagnosis. Among all species of Amphidraus, the male of A. manauara sp. nov. is most similar to that of A. caxiuanan Salgado &amp; Ruiz, 2017 by having the embolic disc with a long, prolaterally projected distal process (dPED) with rounded tip, and a short, ventrally projected ventral process (vPED) (Figs 15C, 17A, 18B); also by having the tibial apophyses with similar pattern (RvTA short and rounded and the RTA is curved on its basal portion and points to apex of cymbium; Figs 15D, 17C, 19G); and by having the cymbium with the proximal, dorso-prolateral portion prominent and one cavity placed on proximal, dorso-retrolateral portion (Figs 17B, 19D). However, the male of A. manauara sp. nov. can be distinguished by having the embolus shaft and embolic filament shorter in proportion to embolic disc (Figs 19A, 19C), by having shorter proximal, dorso-prolateral projection of cymbium (Figs 19B, 19D) and shorter RTA (Figs 19 E–G). The female of A. caxiuanan is unknown, but among Amphidraus females currently known, A. manaura sp nov. is most similar to A. complexus Zhang &amp; Maddison, 2012 by having short copulatory ducts, and the copulatory openings far apart from the posterior border of the epigyne (Figs 16D, 17D); see Zhang &amp; Maddison 2012: figs 5–6). However, the female of A. manauara sp. nov. can be distinguished by having glands associated with the copulatory openings, and longer copulatory ducts (Figs 16 C–D, 17D).</p><p>Description. Male holotype (MPEG 35025). Total length: 2.59. Carapace 1.37 long, 0.90 wide, 0.60 high. Ocular quadrangle 0.67 long. Anterior eye row 0.98 wide, posterior 0.84 wide. Legs 4312. Length of legs: I 2.03 (0.66 + 0.74 + 0.63); II 1.89 (0.60 + 0.67 + 0.62); III 2.29 (0.71 + 0.77 + 0.81); IV 2.61 (0.78 + 0.86 + 0.97). Palp (Figs 15 C–E, 17A–C, 19A–B, 19E): femur and patella with no modifications; cymbium oval (Figs 15E, 17B); tegulum with stout and straight lobe (Figs 15C, 17A); embolus shaft (e) with no modifications; embolic filament short, approximately 1/3 the length of embolus shaft (Fig. 17A). Color in alcohol (Figs 15 A–B): cephalic area black; thoracic area brown with pale longitudinal stripe; abdomen dorsally with two longitudinal brown lateral stripes and one longitudinal central pale stripe intersected by smaller pale stripes; ventrally pale; legs pale.</p><p>Description. Female paratype (MPEG 35026). Total length: 2.75. Carapace 1.34 long, 0.87 wide, 0.62 high. Ocular quadrangle 0.59 long. Anterior eye row 0.93 wide, posterior 0.85 wide. Legs 4321. Length of legs: I 1.94 (0.62 + 0.73 + 0.59); II 1.84 (0.59 + 0.65 + 0.60); III 2.27 (0.70 + 0.76 + 0.81); IV 2.60 (0.74 + 0.86 + 1.00). Epigyne (Figs 16 C–D, 17D): epigynal plate with a large, posterior dorsal pocket; copulatory openings separeted by approximately two diameters of the copulatory duct; short copulatory ducts connected to anterior, outer portion of spermathecae; rounded spermathecae with narrower terminal portion; fertilization ducts anteriorly placed and laterally projected; Color in alcohol (Figs 16 A–B): carapace as in male; abdomen dorsally brown with longitudinal pale stripe intersected by three pale stripes on posterior region; ventrally pale.</p><p>Distribution. Known only from the type locality (state of Amazonas, Brazil) (Fig. 20).</p></div>	https://treatment.plazi.org/id/03F987C08A7C01633AEE8C24FB5E4DFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Salgado, Alexandre;Ruiz, Gustavo R. S.	Salgado, Alexandre, Ruiz, Gustavo R. S. (2019): Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini). Zootaxa 4563 (3): 451-481, DOI: 10.11646/zootaxa.4563.3.3
03F987C08A6301633AEE8AD1FB7E4AFB.text	03F987C08A6301633AEE8AD1FB7E4AFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidraus caxiuanan Salgado & Ruiz 2017	<div><p>Amphidraus caxiuanan Salgado &amp; Ruiz, 2017</p><p>Figs 18 A–H, 19C–D, 19F–G, 20</p><p>Amphidraus caxiuanan Salgado &amp; Ruiz, 2017: 408, figs 5A–B, 6A–D (Holotype ♂ from Melgaço, [01°42'24"S, 51°27'34.3"W], Pará, Brazil, 14.IV.2006, J.A.P. Barreiros leg., deposited in MPEG 29052); World Spider Catalog 2018.</p><p>Additional material examined. BRAZIL: Pará: 1♂ from Floresta Estadual do Trombetas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.434444&amp;materialsCitation.latitude=-1.1028056" title="Search Plazi for locations around (long -56.434444/lat -1.1028056)">Rio Acapu</a> (along about 25 km on the North margin), [01°06'10.10"S, 56°26'04.00"W], Oriximiná, 10–18.X.2014, M.B. Aguiar-Neto et al. leg. (MPEG 35013).</p><p>Revised diagnosis. Amphidraus caxiuanan Salgado &amp; Ruiz, 2017 and A. manauara sp. nov. have the embolic disc with one long, prolaterally projected, distal process (dPED) with rounded tip, and one short, ventrally projected ventral process (vPED) (Figs 17A, 19C); also, the same pattern of tibial apophyses: the RvTA is short and rounded and the RTA emerges from retrodorsal portion of tibia, being curved and projected to the apex of palp (Figs 19 E–G); proximal dorso-prolateral portion of cymbium is prominent in both species, and there is a proximal retro-dorsal cavity on cymbium (Figs 17B, 19B, 19D). However, Amphidraus caxiuanan can be distinguished by having a longer proximal dorso-prolateral projection of cymbium (Figs 19B, 19D) and longer RTA (Figs 19 E–G). Moreover, the embolus shaft and the embolic filament are longer in proportion to the embolic disc in A. caxiuanan (Figs 19A, 19C). The female of A. caxiuanan is still unknown.</p><p>Description. See Salgado &amp; Ruiz (2017).</p><p>Variation. Based on comparisons between the type specimen of A. caxiuanan from Melgaço, Pará, Brazil, and this new male from Oriximiná, Pará, Brazil (about 550 km from the type locality; Fig. 20), we observed several similarities, including in tibial apophyses and especially in the embolic disc, which made us refrain from describing a new species from Oriximiná. That male is herein recognized as being a variant form of A. caxiuanan . Hence, palpal variation within the species is summarized as follows (Figs 18 B–D, 18F–H, 19C–D, 19F–G): the RTA has a slight (Figs 18G, 19F) or accentuated curvature (Figs 18C, 19G); the dorso-prolateral, proximal cymbial projection varies in length (Figs 18D, 18H); RTA length is also variable (Figs 18 C–D, 18G–H, 19F–G). However, the proportions of these structures in the male from Oriximiná are intermediate when comparing the holotype of A. caxiuanan and the holotype of A. manauara (whose type locality is about 450 km away from Oriximiná; Fig. 20), which could indicate a cline across the Amazon River. Nevertheless, the embolic disc of the male palp of A. manauara is different enough to convince us to describe it as a new species (see diagnoses above).</p><p>Distribution. Known only from the state of Pará, Brazil (Fig. 20). It is worth mentioning that, if the specimens from Oriximiná and Melgaço are indeed conspecific, this small species of jumping spider occurs in preserved forests on both margins of the old Amazon River, which is between two and eight kilometers wide in that portion of South America. This provides evidence for the high ability of dispersal present in this species.</p></div>	https://treatment.plazi.org/id/03F987C08A6301633AEE8AD1FB7E4AFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Salgado, Alexandre;Ruiz, Gustavo R. S.	Salgado, Alexandre, Ruiz, Gustavo R. S. (2019): Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini). Zootaxa 4563 (3): 451-481, DOI: 10.11646/zootaxa.4563.3.3
03F987C08A6301673AEE8DD2FEC24E36.text	03F987C08A6301673AEE8DD2FEC24E36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidraus shenlong Salgado & Ruiz 2019	<div><p>Amphidraus shenlong sp. nov.</p><p>Figs 21, 22, 26 E–F, 26H, 28B, 30</p><p>Type material. Holotype: ♂ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.37419&amp;materialsCitation.latitude=-7.9982505" title="Search Plazi for locations around (long -49.37419/lat -7.9982505)">Conceição do Araguaia</a>, 07°59’53.7”S, 49°22’27.1”W, Pará, Brazil, 16.I.2018, R.P.S Almeida. &amp; F.V. Arruda leg. (MPEG 35020). Paratypes: 2♂, same data as holotype (MPEG 35021, IBSP 221864) .</p><p>Additional material examined. BRAZIL: Pará: 4♂ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.37419&amp;materialsCitation.latitude=-7.9982505" title="Search Plazi for locations around (long -49.37419/lat -7.9982505)">Conceição do Araguaia</a>, 07°59’53.7”S, 49°22’27.1”W, 16.I.2018, R.P.S Almeida. &amp; F.V. Arruda leg. (MPEG 32794, 32807).</p><p>Etymology. The specific epithet (noun in apposition) is the name of a dragon of Chinese mythology and refers to the big projection of the embolic disc, which recalls a dragon’s tail (white arrow in Fig. 21D).</p><p>Diagnosis. A. shenlong sp. nov. is similar to A. draconicaudatus Salgado &amp; Ruiz, 2017 by having an elongate and stout projection on the retrolateral border of the embolic disc that extends towards the retrolateral tegulum border and bears several branches (Figs 21 C–D, 26A, 27A), and by having the RvTA ventrally projected with projections on its border (Figs 21F, 22B, 26G). However, A. shenlong sp. nov. can be distinguished by having the RvTA with four projections (three with acute tip and one with serrated tip) and the RTA serrated tip reaching the same height of the RvTA tip along palp axis (Figs 21F, 22B, 26H), whereas in A. draconicaudatus the RvTA has a single serrated projection and the RTA is reduced to spikes (Figs 26G, 27 C–D).</p><p>Description. Male holotype (MPEG 35020). Total length: 2.68. Carapace 1.31 long, 0.89 wide, 0.69 high. Ocular quadrangle 0.71 long. Anterior eye row 1.01 wide, posterior 0.86 wide. Legs 3412. Length of legs: I 1.82 (0.61 + 0.62 + 0.59); II 1.68 (0.59 + 0.57 + 0.52); III 2.42 (0.83 + 0.84 + 0.75); IV 2.29 (0.69 + 0.75 + 0.85). Chelicera with pocket on outer surface (between dorsum and retrolateral face) (Fig. 28B). Palp (Figs 21 C–F, 22A– B): femur with median-ventral region slight prominent (Fig. 21F); cymbium oval; cymbial conductor leaf-shaped (Fig. 22A); tegulum with narrow lobe curved to retrolateral side (Figs 21E, 22A); embolic disc with three processes very closer together: two rounded distal processes (dPED) almost fused and one short triangular ventral process (vPED) (Figs 21E, 22A). Embolic filament about as long as embolus shaft (Figs 21C, 21E). Color in alcohol (Figs 21 A–B): cephalic area black; thoracic area yellow; abdomen dorsally variegated with yellow and brown; ventrally yellow; legs yellow.</p><p>Female. Unknown.</p><p>Distribution. Known only from the type locality (state of Pará, Brazil) (Fig. 30).</p><p>Natural History. The specimens were collected from ecotonal zone between the Cerrado and the Amazon rainforest.</p></div>	https://treatment.plazi.org/id/03F987C08A6301673AEE8DD2FEC24E36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Salgado, Alexandre;Ruiz, Gustavo R. S.	Salgado, Alexandre, Ruiz, Gustavo R. S. (2019): Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini). Zootaxa 4563 (3): 451-481, DOI: 10.11646/zootaxa.4563.3.3
03F987C08A6701653AEE8D66FAAA4F4E.text	03F987C08A6701653AEE8D66FAAA4F4E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidraus draconitupan Salgado & Ruiz 2019	<div><p>Amphidraus draconitupan sp. nov.</p><p>Figs 23 A–G, 24A–D, 25A–D, 30</p><p>Type material. Holotype: ♂ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.502193&amp;materialsCitation.latitude=-22.095667" title="Search Plazi for locations around (long -50.502193/lat -22.095667)">Fazenda Palma</a>, Varpa, 22°05'44.4"S, 50°30'07.9"W, Tupã, São Paulo, Brazil, XII.2016, G.R.S Ruiz leg. (MPEG 35022). Paratypes: 1♂ from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.96&amp;materialsCitation.latitude=-21.726084" title="Search Plazi for locations around (long -44.96/lat -21.726084)">Cachoeira Eubiose</a>, 21°43'33.9"S, 44°57'36.0"W, São Thomé das Letras, Minas Gerais, Brazil, XII.2017, A. Salgado leg. (MPEG 35016) and 2♀, same data as holotype (MPEG 35017, 35028) .</p><p>Additional material examined. BRAZIL: Mato Grosso do Sul: 1♂, Brasilândia, Usina Hidrelétrica Sérgio Motta [21°16’22.7”S, 51°55’52.5”W], 2000, Equipe IBSP leg. (IBSP 30911); Minas Gerais: 1♀, São Thomé das Letras, Vale das Borboletas, 21°43'24.4"S, 45°00'24.4"W, XII.2017, A. Salgado leg. (MPEG 35023); 1♂ and 2♀ from São Thomé das Letras, Cachoeira Eubiose, 21°43'33.9"S, 44°57'36.0"W, XII.2017, A. Salgado leg. (MPEG 35024); São Paulo: 9♀, Tupã, Fazenda Palma, Varpa, 22°05'44.4"S, 50°30'07.9"W, XII.2016, G.R.S Ruiz leg. (MPEG 35029–35031) .</p><p>Etymology. The specific epithet is a compound name that refers to the large projection of embolic disc (homologous to projection with dragon's tail shape of A. draconicaudatus) and the type locality (Tupã). The final - n is added in order to mimic phonetics of the name of the locality.</p><p>Diagnosis. The male of Amphidraus draconitupan sp. nov. is most similar to those of A. draconicaudatus Salgado &amp; Ruiz, 2017 and A. shenlong sp. nov. by having an elongate and stout projection on the retrolateral border of the embolic disc that extends towards the retrolateral tegulum border (Figs 22A, 25A, 26A). However, the male of A. draconitupan sp. nov. differs from the males of these two species by having the superior border of its elongate projection of embolic disc with a rounded projection followed by several reduced spikes (Figs 23D, 25A), whereas in A. draconicaudatus and A. shenlong sp. nov. the rounded projection is absent and the elongate projections have several branches (Figs 26 A–F). The epigyne of A. draconitupan sp. nov. is similar to that of A. draconicaudatus, both with spermathecae kidney-shaped, with glands near to copulatory openings and with a narrow dorsal pocket placed posteriorly on epigynal plate (Figs 25D, 27E) (the female of A. shenlong sp. nov. is still unknown). However, the female of A. draconitupan sp. nov. can be distinguished from that of A. draconicaudatus by having shorter copulatory ducts connected to spermathecae on its outer border (Fig. 25D), whereas they connect to the inner posterior border in A. draconicaudatus (see Salgado &amp; Ruiz 2017: fig. 24F).</p><p>Description. Male holotype (MPEG 35022). Total length: 2.87. Carapace 1.79 long, 1.18 wide and 0.95 high. Ocular quadrangle 0.82 long. Anterior eye row 1.33 wide, posterior 1.14 wide. Legs 3412. Length of leg: I 2.93 (0.92 + 1.14 + 0.87); II 2.61 (0.86 + 0.93 + 0.82); III 3.42 (1.18 + 1.16 + 1.08); IV 3.39 (1.09 + 1.10 + 1.20). Chelicerae with pockets, as in A. shenlong sp. nov. (Fig. 28C). Palp (Figs 23 D–G, 25A–C): femur with medianproventral region prominent (Fig. 25B); RvTA ventrally projected with one single projection pointed to apex of palp (Figs 23G, 25C); cymbium oval; cymbial conductor leaf-shaped (Fig. 25A); tegular lobe dorsally curved (Fig. 23F); embolic disc with membranous prolateral portion, with two short distal processes (dPED) almost fused, and one triangular ventral process (vPED) distally placed (Figs 23D, 25A); straight embolus shaft, with its basal portion covered by the stout projection of embolic disc (Figs 23D, 25A); embolic filament with nearly half the length of embolus shaft (Fig. 25A). Color in alcohol (Figs 23 A–C): carapace black; abdomen black, dorsally variegated with brown; ventrally black; legs generally black; legs III and IV with pale metatarsi and tarsi.</p><p>Female paratype (MPEG 35028). Total length: 3.07. Carapace 1.63 long, 1.08 wide and 0.75 high. Ocular quadrangle 0.87 long. Anterior eye row 1.17 wide, 1.00 posterior wide. Legs 4312. Length of leg: I 2.39 (0.79 + 0.88 + 0.72); II 2.32 (0.76 + 0.86 + 0.70); III 3.12 (1.04 + 1.10 + 0.98); IV 3.32 (0.98 + 1.11 + 1.23). Epigyne/vulva (Figs 24D, 25D): epigynal plate with lateral depressions. Color in alcohol (Fig. 24 A–B): cephalic area black; thoracic area dark brown; abdomen dorsally variegated with black and brown; ventrally pale; legs pale with black spots. Compare this with color of live specimen in Fig. 24C.</p><p>Distribution. Known from the states of Mato Grosso do Sul, Minas Gerais and São Paulo (Brazil) (Fig. 30).</p><p>Natural History. The specimens were collected in leaf-litter and tree trunk, close to waterfalls and lakes.</p></div>	https://treatment.plazi.org/id/03F987C08A6701653AEE8D66FAAA4F4E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Salgado, Alexandre;Ruiz, Gustavo R. S.	Salgado, Alexandre, Ruiz, Gustavo R. S. (2019): Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini). Zootaxa 4563 (3): 451-481, DOI: 10.11646/zootaxa.4563.3.3
03F987C08A6501683AEE8C24FC3B4D36.text	03F987C08A6501683AEE8C24FC3B4D36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidraus draconicaudatus Salgado & Ruiz 2017	<div><p>Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017</p><p>Figs 26 A–D, 26G, 27A–E, 28A, 29A–B, 30</p><p>Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017: 427, figs 22–24 (Holotype ♂ from Pium, 09°58'38.7"S, 50°02'05.0"W, Tocantins, Brazil, 11–26.V.2016, Serrão et al. leg., deposided in MPEG 32689; 1♂ and 2♀ paratypes, same data as holotype, deposited in IBSP 211843–211844, MPEG 32690); World Spider Catalog 2018.</p><p>Additional material examined. BRAZIL: Tocantins: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.95789&amp;materialsCitation.latitude=-9.304334" title="Search Plazi for locations around (long -49.95789/lat -9.304334)">Caseara</a>, 9°18'15.6"S, 49°57'28.4"W: 2♂, 11.V.2016, A.B. Bonaldo et al. leg. (MPEG 35018); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.03472&amp;materialsCitation.latitude=-9.977416" title="Search Plazi for locations around (long -50.03472/lat -9.977416)">Pium</a>, 09°58'38.7"S, 50°02'05.0"W: 1♂, 19.V.2016, A.B. Bonaldo et al. leg. (MPEG 35019) .</p><p>Revised diagnosis. A. draconicaudatus is similar to A. shenlong sp. nov. by having an elongate and stout projection on the retrolateral border of the embolic disc that extends towards the retrolateral tegulum border and bears several branches (Figs 21D, 26A, 26C, 27A); also, by having the RvTA pointing ventrally, with projections on its border (Figs 21F, 22B, 26G). However, the male of A. draconicaudatus can be distinguished by having the RvTA with a single serrated projection, and a RTA reduced to spikes (Figs 26G, 27 B–D), whereas in A. shenlong sp. nov. the RvTA has four projections (three with acute tip and one with serrated tip), and the RTA serrated tip reaches the same height of the RvTA tip along palp axis (Figs 21F, 22B, 26H). The female of A. shenlong sp. nov. is unknown, but among the currently known females of Amphidraus, A. draconicaudatus sp. nov. is most similar to those of A. draconitupan sp. nov., both with spermathecae kidney-shaped, with glands near copulatory openings and with a narrow dorsal pocket placed posteriorly on epigynal plate (Figs 24D, 25D, 27E). However, the female of A. draconicaudatus sp. nov. can be distinguished from that of A. draconitupan sp. nov. by having longer copulatory ducts connected to the inner posterior border of spermathecae (see Salgado &amp; Ruiz 2017: fig. 24F), whereas they connect to the outer border of the spermathecae in A. draconitupan (Fig. 25D).</p><p>Description. See Salgado &amp; Ruiz (2017).</p><p>Variation. Among the males examined, we observed the following variation in palpal structures (Figs 26 A–D, 26G): the superior border of the branched projection of embolic disc can be smooth (Figs 26 A–B) or occupied by large serial spikes (similar to a comb) (Figs 26 C–D). We decided to refrain from describing this variation as a new species and all these forms are considered conspecific. Also, the number and shape of branches on the RvTA is variable (Fig. 26G; see Salgado &amp; Ruiz 2017: figs 23B, 24B, 24D).</p><p>Note. Amphidraus draconicaudatus, A. shenlong sp. nov. and A. draconitupan sp. nov. clearly compose a particular group within the genus. The males in this group have one stout projection that emerges from the retrolateral border of the embolic disc and extends towards to retrolateral border of tegulum (Figs 22A, 25A, 26A, 27A). Moreover, males also have pockets on outer faces of chelicerae (Figs 28 A–C, 29A–B), of unknown use.</p><p>Distribution. Known only from the state of Tocantins (Brazil) (Fig. 30).</p></div>	https://treatment.plazi.org/id/03F987C08A6501683AEE8C24FC3B4D36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Salgado, Alexandre;Ruiz, Gustavo R. S.	Salgado, Alexandre, Ruiz, Gustavo R. S. (2019): Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini). Zootaxa 4563 (3): 451-481, DOI: 10.11646/zootaxa.4563.3.3
