taxonID	type	description	language	source
03F9B062FFC6FFE6FF47FE1E5CF176E7.taxon	materials_examined	Type species. Ceroderma asperata Raffray (subsequent monotypy by Raffray, 1890 b: 199).	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC6FFE6FF47FE1E5CF176E7.taxon	diagnosis	Diagnosis. Head triangular; frontal rostrum low; lacking frontal fovea; antennomeres XI elongate and conical. Pronotum transversely triangular, lacking median antebasal fovea, with punctiform lateral antebasal foveae; with lateral and antebasal spines. Elytra with three small basal foveae, lacking discal striae. Abdomen with inner and outer marginal carinae of tergite IV forming broad triangular ridge at lateral margins; tergite IV longest, not much longer than tergite V, lacking discal carinae.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC6FFE6FF47FE1E5CF176E7.taxon	description	Redescription. Body flat; length about 2.6. Head triangular, with frontal rostrum low, antennal tubercles moderately prominent; lacking frontal fovea; vertex largely and broadly concave; vertexal foveae small, connected by shallow and broad triangular sulcus (foveae and sulcus absent in male due to presence of sexual feature); postocular margins roundly expanded laterally; with eleven antennomeres, clubs formed by apical three antennomeres, antennomeres XI elongate and conical; ocular-mandibular carinae well-defined; eyes oval; maxillary palpi with palpomeres III small and triangular, IV gradually narrowed in basal half; small gular foveae close. Pronotum transversely triangular; lateral antebasal foveae punctiform, lacking median antebasal foveae; with small lateral and antebasal spines; disc barely convex; basolateral foveae absent; paranotal carinae distinct; lateral procoxal foveae barely indicated. Elytra with three basal foveae, lacking discal striae, sutural and marginal striae complete. Thorax with small median and lateral mesoventral foveae; lateral mesocoxal foveae present; with lateral metaventral foveae; posterior margin narrowly notched medially. Legs with tarsomeres II and III subequal in length. All abdominal segments lacking mediobasal impression and foveae; tergite IV with lateral margins produced as thick triangular ridge edged by inner and outer marginal carinae, ridges extending entire tergal length, lacking discal carinae; tergites VI – VII with ridges much thinner; tergite VII with indistinct lateral tubercles. Sternite IV with median carina; sternites V – VII with short triangular basolateral ridges. Male with vertex, postocular margins and protibiae modified. Aedeagus with paramere fused to median lobe to from elongate ventral lobe; lacking obvious dorsal lobe. Comparative notes. Placed close to Hingstoniella Jeannel from Sikkim and Tibet, and Cerochusa gen. n. from Hainan described below by sharing a flat and broad general form, the transversely triangular pronotum, and the lack of a median antebasal fovea on the pronotum. Ceroderma can be separated from Hingstoniella by having the head with postocular margins broadly expanded laterally, the pronotum with small but distinct lateral and antebasal spines, and each elytron with three basal foveae. Hingstoniella has the head with postocular margins roundly narrowed basally, the pronotum lacking any spines, and each elytron with single large basal fovea. Ceroderma differs from Cerochusa by the lack of discal carinae on tergite IV and the lack of basomedian and basolateral foveae of sternite IV. In contrast, Cerochusa has short, close discal carinae on tergite IV, and sternite IV has distinct mediobasal and two pairs of basolateral foveae.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC6FFE4FF47F8895DA27704.taxon	materials_examined	Other material examined (NSMT). 2 3, 1 Ƥ, labeled ‘ Ulu Gombak, Selangor / (termite nest) / [MALAY- SIA] / 27. vii. 2003, M. Maruyama // Host: Coptotermes curvi - / gnathus Holmgren, 1913 / M. Maruyama det., 2003 // FAUNA MALAYSIA / Staphylinidae, Pselaphinae / Ceroderma asperata Raffray / S. Nomura det., 2003 ’.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC6FFE4FF47F8895DA27704.taxon	diagnosis	Diagnosis. Head triangular; vertex largely concave; postocular margins roundly expanded laterally; triangular pronotum broadly incised at basal half; elytra evenly constricted at basal half, with three basal foveae; tergite IV slightly longer than V.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC6FFE4FF47F8895DA27704.taxon	description	Redescription. Male. BL 2.66. Body (Figs. 1, 19) reddish brown, maxillary palpi and tarsi lighter; head, pronotum and elytra covered with dense setae, microtuberculate where setae arise. Head (Figs. 3, 19) much wider than long, HL 0.48, HW 0.81; vertex with tufted setae at anterior and posterior borders of large transverse median concavity, vertexal foveae and sulcus overlapped by concavity; postocular margins largely angularly expanded laterally, with dense long lateral setae; eyes well-developed, each composed of about 60 facets; antennomeres II – VIII (Fig. 4) of similar width, clubs formed by enlarged antennomeres IX – XI. Pronotum much wider than long, PL 0.55, PW 0.81; with shallow longitudinal median and lateral sulci; short discal carinae at basal half; lateral spines slightly larger than antebasal spines. Elytra wider than long, EL 0.82, EW 1.03; evenly constricted at basal half. Metaventrite concave along midline. Legs with femora thickened; protibiae (Fig. 5) angularly expanded mesally at apical half; mesotibiae (Fig. 5) with tiny membranous apical projection; metatrochanters (Fig. 7) with large elongate ventral projection. Abdomen wider than long, AL 0.81, AW 1.02; tergite IV with thick triangular lateral ridges; tergites V – VI with ridges much thinner; tergite VII (Fig. 8) with small lateral tubercles sharp. Sternite IV twice length of V at midline, with short median carina; sternites V – VII with short triangular basolateral ridges. Aedeagus (Figs. 9 – 11) well-sclerotized, length 0.37; broad basal bulb with large foramen, median lobe expanded apically. Female. Similar to male in general. Measurements: BL 2.62, HL 0.47, HW 0.74, PL 0.55, PW 0.78, EL 0.78, EW 1.04, AL 0.82, AW 0.98. Vertex lacking setose concavity, widely separated foveae connected by shallow triangular sulcus in large impressed area; postocular margins less expanded than in male, lacking long lateral setae. Each eye composed of about 55 facets. Protibiae lacking mesal expansion at apical half; mesotibiae lacking apical tubercle.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC6FFE4FF47F8895DA27704.taxon	distribution	Distribution. Malaysia: Penang, Selangor. Host termite. Coptotermes curvignathus Holmgren (det. M. Maruyama).	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC3FFE2FF47FA515C44778E.taxon	materials_examined	Type species. Cerochusa cilioceps, new species	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC3FFE2FF47FA515C44778E.taxon	diagnosis	Diagnosis. Head nearly triangular; lacking frontal rostrum; lacking frontal fovea; antennomeres XI robust and conical. Pronotum transversely triangular, deeply incised at level of punctiform lateral antebasal foveae; lacking median antebasal fovea; with large lateral and antebasal spines. Elytra with two basal foveae, discal striae shallow and short. Abdomen with inner and outer marginal carinae of tergite IV – VI forming broad triangular ridge at lateral margins; tergite IV longest, about twice length of tergite V; discal carinae present.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC3FFE2FF47FA515C44778E.taxon	description	Description. Body flat and broad; length about 2.4. Head triangular, lacking frontal rostrum, antennal tubercles indistinct; lacking frontal fovea; vertex slightly convex medially, vertexal foveae well-defined, connected by U-shaped sulcus, with short median carina (carina indistinct in male due to presence of sexual feature); postocular margins angularly expanded laterally; posterior margin triangularly impressed medially; with eleven antennomeres, clubs weakly formed by apical three antennomeres, antennomeres XI robust and conical; ocular-mandibular carinae well-defined; eyes oval; maxillary palpi with palpomeres III small and triangular, IV gradually narrowed in basal half; small gular foveae close. Pronotum transversely triangular; lateral antebasal foveae punctiform, lacking median antebasal foveae; with distinct median and lateral sulci and discal carinae; with large lateral and antebasal spines; lateral margins largely incised at level of antebasal foveae; disc slightly convex; basolateral foveae absent; paranotal carinae distinct, with additional pair of lateral carinae; lateral procoxal foveae present. Elytra with two basal foveae, discal striae weakly indicated; sutural and marginal striae extending to elytral apex; lacking subhumeral foveae. Thorax with small median and lateral mesoventral foveae; lateral mesocoxal foveae present; with lateral metaventral foveae; posterior margin narrowly notched medially. Legs with tarsomeres thickened, tarsomeres II and III subequal in length. Abdomen with lateral margins of tergite IV – VI produced as thick triangular ridge edged by inner and outer marginal carinae, ridges extending entire tergal length; tergite IV with large lateral foveae; discal carinae short and close, reaching one-fifth tergal length; tergites V – VII lacking mediobasal impression and foveae. Sternite IV with distinct median and two pairs of lateral foveae, short lateral carinae present; sternites V – VII lacking foveae. Male with vertex and mesotibiae modified. Aedeagus with paramere fused to median lobe to from elongate ventral lobe; with membranous dorsal lobe. Comparative notes. Most close to Ceroderma in having a flat and broad body form, the triangular head lacking a frontal fovea and with medially concaved posterior margin, the presence of pronotal lateral and antebasal spines and the lack of a median antebasal fovea on the pronotum. Cerochusa can be separated by the elytra with two basal foveae and shallow discal striae, the presence of short discal carinae on tergite IV, that tergite being twice length of tergite V, and the presence of mediobasal and basolateral foveae on sternite IV. While Ceroderma has three basal elytral foveae, the elytra lacks discal striae, tergite IV lacks discal carinae and is slightly longer than tergite V, and sternite IV lacks any foveae. Hingstoniella shares with Cerochusa the presence of two basal elytral foveae, and tergite IV being twice as long as tergite V, but the roundly narrowed postocular margins, the smooth pronotal lateral margins and the lack of discal carinae on tergite IV quickly separate Hingstoniella from Cerochusa.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC3FFE2FF47FA515C44778E.taxon	etymology	Etymology. The generic name is a combination of ‘ cer ’ (horn), and ‘ - chusa ’ (with), referring to the strongly spinose pronotal lateral margins. The species name is formed by ‘ cil ’ (hair) and ‘ ceps ’ (head), with respect to the male sexual character on the head.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC1FFE1FF47FF185D4074CB.taxon	materials_examined	Type material (SNUC). Holotype: CHINA: 3, labeled ‘ China: Hainan Prov. / Ledong County / Jianfengling N. R. / 19. v. 2011, alt. 1,000 m / Wen-Xuan BI leg. ’; Paratype: CHINA: Ƥ, same label data as holotype, except ‘ Jianfengling N. R. / core area No. V, 978 m / 3. xii. 2009 / 18.73236 ° N / 108.87023 ° E / Mei-Ying Lin leg. ’.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC1FFE1FF47FF185D4074CB.taxon	diagnosis	Diagnosis. Head triangular; vertex convex medially, U-shaped sulcus connecting well-defined foveae; postocular margins expanded laterally to form thick triangular tubercles; triangular pronotum broadly and deeply incised at level of lateral antebasal foveae; elytra barely constricted at base, with two large basal foveae; tergite IV about twice length of V.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC1FFE1FF47FF185D4074CB.taxon	description	Description. Male. BL 2.39. Body (Fig. 2) reddish brown, maxillary palpi and tarsi lighter; head, pronotum and elytra covered with dense setae, microtuberculate where setae arise. Head (Fig. 12) much wider than long, HL 0.36, HW 0.64; vertex with tuft of setae anterior to transverse arched median carina, vertexal foveae connected by U-shaped sulcus, sulcus broken at anterior margin; eyes well-developed, each composed of about 65 facets; antennomeres II – VIII (Fig. 13) of similar width, VIII – X distinctly transverse; clubs formed by enlarged antennomeres IX – XI. Pronotum much wider than long, PL 0.50, PW 0.83; with distinct longitudinal median and lateral sulci; with discal carinae at middle; lateral and antebasal spines large. Elytra wider than long, EL 0.76, EW 0.98; barely constricted at basal half. Metaventrite concave medially. Legs with thick femora; mesotibiae (Fig. 14) with tiny membranous apical projections; metatrochanters lack projection at ventral margin. Abdomen wider than long, AL 0.77, AW 0.89; tergites IV – VI with thick triangular lateral ridges; tergite VI (Fig. 15) with short discal carinae close, reaching one-fifth tergal length; tergite VII with small lateral protuberances. Sternite IV twice length of V at midline, with inner pair of basolateral foveae close to mediobasal foveae; sternites V – VII lacking foveae. Aedeagus (Figs. 16 – 18) well-sclerotized, length 0.33; basal bulb with large foramen, elongate median lobe sharpened apically. Female. Similar to male in general. Measurements: BL 2.39, HL 0.36, HW 0.63, PL 0.49, PW 0.82, EL 0.74, EW 0.97, AL 0.80, AW 0.92. Vertex lacking transverse carina and tufted setae. Each eye composed of about 60 facets. Mesotibiae lacking apical tubercle.	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
03F9B062FFC1FFE1FF47FF185D4074CB.taxon	distribution	Distribution. South China: Hainan Island. Host ant. Vollenhovia Mayr sp. (det. A. Radchenko).	en	Yin, Zi-Wei, Nomura, Shûhei, Li, Li-Zhen (2012): Redescription of Ceroderma asperata Raffray, and description of Cerochusa cilioceps gen. et sp. n. from Hainan, South China (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 3355: 62-68, DOI: 10.5281/zenodo.281580
