taxonID	type	description	language	source
03F9EF199F55FFC7FDE9FD625E4B2FE4.taxon	type_taxon	(type genus: Thaumastodus Champion, 1924 [junior synonym of Pseudeucinetus Heller, 1921]). Thaumastodinae – Spilman 1959: 111; 1972: 113 [key to genera]. — Skelley 2005: 119.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F55FFC7FDE9FD625E4B2FE4.taxon	diagnosis	Diagnosis Body elongate to oblong, slightly shining, closely covered with short suberect setae. Eyes (Fig. 2) large, situated on dorso-lateral parts of head, separated narrowly on vertex. Antennae short to relatively short, 7 - or 11 - segmented, compactly articulated in terminal 6 antennomeres. Basal margin of elytra not crenulate. Tarsal formula 4 - 4 - 4 or 4 - 5 - 5. Hind legs long, capable of jumping; hind tibiae bearing spines. Sexual dimorphism distinct in fore tarsi (genus Pseudeucinetus) or in sternite VII.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F53FFC4FDDFFE215E4B2CCE.taxon	type_taxon	(type species: Acontosceles hydroporoides Champion, 1924).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F53FFC4FDDFFE215E4B2CCE.taxon	diagnosis	Diagnosis Body elongate. Gena wide; antennal groove absent. Antennae slender, relatively long, 11 - segmented; antennal insertions situated near eyes. Legs long and slender; tarsal formula 4 - 5 - 5.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F53FFC4FDDFFE215E4B2CCE.taxon	discussion	Remarks This genus was subdivided into two species groups by Pütz (2008) mainly based on the male genital characters. However, the points differentiating the two groups are obvious and quantitative features (see also Yoshitomi & Putra 2011), and so the species-group concept needs to be revised. Biological notes The habitat of this genus (Fig. 4 A) is freshwater environments, i. e., waterfalls (type locality of A. chujoi Yoshitomi & Satô, 2005), rivers and streams. The adults are found on wet rocks or actively flying. The larval stages are unknown.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F51FFCAFDB4FEEB5A162CAC.taxon	description	urn: lsid: zoobank. org: act: 2582 F 411 - 8790 - 4937 - B 6 A 0 - 248899 FD 9208 Figs 5 A, 6	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F51FFCAFDB4FEEB5A162CAC.taxon	etymology	Etymology This species is named after its type locality.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F51FFCAFDB4FEEB5A162CAC.taxon	materials_examined	Material examined Holotype MALAYSIA • ♂; “ SARAWAK: Nanga Pelagus 7 - 14. VIII. ‘ 58 ”; “ T. C. Maa Collector BISHOP ”; BPBM. Paratypes MALAYSIA • 1 ♂, 2 ♀♀; same collection data as for holotype; BPBM • 1 ♀; same collection data as for preceding; EUMJ • 2 ♀♀; “ BORNEO: SARAWAK Merirai Valley nr Kapit, 30 - 300 m, 1 - 6. VII. 58 ”; “ T. C. Maa Collector BISHOP Mus. ”; BPBM • 1 ♀; “ BORNEO: SARAWAK Bau, Lake Area 29 - 30. VIII. 1958 ”; “ T. C. Maa Collector BISHOP ”; BPBM.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F51FFCAFDB4FEEB5A162CAC.taxon	description	Description Male BODY. Oblong, subparallel-sided, slightly convex dorsally, weakly shiny, closely covered with suberect silver setae. Coloration of body blackish brown, but mouthparts and legs yellowish brown; elytra with obscure silver spots consisting of erect setae. HEAD. Moderate in size; frons and clypeus closely covered with erect, long silver setae; clypeus with straight anterior margin. Eyes large, prominent dorso-laterally. ANTENNAE. Reaching about proximal half of pronotum. PRONOTUM. Widest at basal third, densely bearing short yellowish setae and minute scale-like setae; PW / PL 2.25 – 2.50 (2.34). Scutellum small, subtriangular. ELYTRA. Oblong, subparallel-sided near base to apical fifth, densely covered with short yellowish setae and minute scale-like setae, projecting ventrally in apical part; EL / EW 1.59 – 1.80 (1.72); EL / PL 4.17 – 4.63 (4.39); EW / PW 1.02 – 1.16 (1.10); TL / EW 1.97 – 2.22 (2.11). ABDOMEN. Sternite VII (Fig. 6 A) broad, gently arcuate along caudal margin, shallowly concave in median part, bearing about 20 long setae and three pairs of long and stout setae. Sternite VIII (Fig. 6 B) membranous, bearing minute spines in apical parts. Sternite IX (Fig. 6 C) moderately sclerotized, large, with small apical plates. AEDEAGUS (Fig. 6 D – E). Long, well sclerotized, strongly curved ventrally; basal piece oblong, expanded basally, evenly curved ventrally; lateral lobes long and slender, serrate on dorso-lateral margins, curved ventrally; median lobe long and slender, about 0.6 times as long as lateral lobe, gently curved ventrally; ML / BL 0.36; ML / LL 0.68. Female Sexual dimorphism distinct in the following characteristics: 1) frons and clypeus covered with short yellowish setae; 2) apical part of elytra projecting ventro-apically; 3) sternite VII (Fig. 6 F) rather pointed on caudal margin, bearing about 40 long setae. Urosternite (Fig. 6 G) well sclerotized, with long and slender apodeme; lateral projections stout, bifid in apical parts. Ovipositor (Fig. 6 H) well sclerotized; coxite sparsely punctuate, pointed at apices; approximate ratio of coxite and baculus (n = 1) of 1.0: 1.7. Measurements Unsexed (n = 4): TL 2.17 – 2.25 (2.21) mm; PW 0.90 – 1.00 (0.96) mm; PL 0.40 – 0.42 (0.41) mm; EL 1.75 – 1.85 (1.80) mm; EW 1.00 – 1.10 (1.05) mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F51FFCAFDB4FEEB5A162CAC.taxon	distribution	Distribution Malaysia (Sarawak).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F51FFCAFDB4FEEB5A162CAC.taxon	discussion	Remarks This species belongs to the hydroporoides species group judging from characteristics of the male genitalia.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5CFFC8FDF2FEEB59312D35.taxon	description	Fig. 5 D	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5CFFC8FDF2FEEB59312D35.taxon	materials_examined	Material examined LAOS • 12 ex.; “ [LA 8] Nam Xan, near Ban Nahoung, Xieng Khouang Prov., Laos, 19 ° 09.134 ″ N, 103 ° 43.037 ″ E, 1453 m, 25. V. 2013 H. Yoshitomi leg. ”; EUMJ • 23 ex.; “ [LA 23] 2 km W of Nam Chat, Laos, 19 ° 290137 ″ N, 102 ° 424900 ″ E, ca 814 m, 22. IV. 2018, H. Yoshitomi leg. ”; EUMJ. VIETNAM • 5 ex.; “ Tam Dao (950 - 1,050 m) Vinh Phu Prov., Vietnam, 19. V. 1995 M. Sato leg. ”; EUMJ • 11 ex.; “ [VN 6] Ban Khoang, Lao Cai Prov., Vietnam, N 22 24.423 E 103 47.309, ca 1284 m, 22. VI. 2012, H. Yoshitomi leg. ”; EUMJ.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5CFFC8FDF2FEEB59312D35.taxon	distribution	Distribution Laos, Vietnam (new record).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5CFFC8FD80FC645F6E28E0.taxon	description	Fig. 5 B	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5CFFC8FD80FC645F6E28E0.taxon	materials_examined	Material examined JAPAN • 3 ex.; “ Sate-gawa Kunigami, Okinawa 18 - VI- 2006 K. Takahashi leg. ”; EUMJ • 3 ex.; “ Hijikawa, Kunigami, Okinawa, Ryukyus 15 - VI- 2003 K. Takahashi leg. ”; EUMJ. TAIWAN • 1 ex.; “ Taiwan Nantou Chunyang VI / 11 -- VII / 9 / 2002 C. S. Lin & W. T. Yang Malaise trap (KCN) ”; “ NMNS ENT 5240 - 73 ”; NMNS • 1 ex.; “ Taiwan Nantou Chunyang V / 7 -- VI / 1 / 2002 C. S. Lin & W. T. Yang Malaise trap (KCN) ”; “ NMNS ENT 5237 - 3502 ”; NMNS • 1 ex.; “ Taiwan Nantou Chunyang V / 11 - VII / 3 / 2004 C. S. Lin & W. T. Yang Malaise trap (KCN) ”; “ NMNS ENT 5249 - 6740 ”; NMNS • 1 ex.; “ Taiwan Pingtung Manchou Lite No: 169 Hsiang Rd. 3.8 km IV / 19 -- V / 1 / 2009 M. L. Jeng & T. J. Chen Malaise trap ”; NMNS.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5CFFC8FD80FC645F6E28E0.taxon	distribution	Distribution Japan: Ryukyu Islands (Okinawa-jima, Ishigaki-jima, Iriomote-jima); Taiwan.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5CFFC9FDDFF9325A372A4E.taxon	description	Figs 5 E, 7	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5CFFC9FDDFF9325A372A4E.taxon	materials_examined	Material examined NEPAL • 1 ♂, 5 ex.; “ NEPAL, Prov. Bheri Distr. Surkhet, 20 km S Surkhet, Bheri-River 600 m NN, 03. VI. 1995 leg. M. Hartmann ”; NME • 1 ♂, 1 ♀; same collection data as for preceding; EUMJ.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5CFFC9FDDFF9325A372A4E.taxon	description	Redescription For a detailed description of the external features, see the original description (Pütz 2008). PW / PL 2.63; EL / EW 1.58; EL / PL 4.70; EW / PW 1.13; TL / EW 1.92. Male Sternite VII (Fig. 7 A) gently arcuate on caudal margin, bearing about 30 long setae in median part. Sternite VIII (Fig. 7 B) membranous. Sternite IX (Fig. 7 C) slightly sclerotized, with long apical plates bifid in apical parts. Aedeagus (Fig. 7 D – F) relatively short, well sclerotized, slightly curved ventrally; basal piece oblong, subparallel-sided; lateral lobes short and broad, arcuate on lateral margins, closely punctuate, with small projections in lateral parts of apices; median lobe slender, about 0.8 times as long as lateral lobe, slightly curved ventrally; ML / BL 0.40; ML / LL 0.80. Female Sexual dimorphism slight. Sternite VII (Fig. 7 G) arcuate on caudal margin, bearing about 40 long setae. Urosternite (Fig. 7 H) well sclerotized, with long and slender apodeme; lateral projections short, projecting antero-laterally. Ovipositor (Fig. 7 I) well sclerotized; coxite closely punctuate, pointed at apices; approximate ratio of coxite and baculus (n = 1) 1.0: 1.8. Measurements Male (n = 1): TL 2.28 mm; PW 1.05 mm; PL 0.40 mm; EL 1.88 mm; EW 1.19 mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5CFFC9FDDFF9325A372A4E.taxon	distribution	Distribution Nepal.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5DFFCFFD80FB9B592F2F27.taxon	description	Figs 5 F, 8	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5DFFCFFD80FB9B592F2F27.taxon	materials_examined	Material examined LAOS • 1 ♂; “ LAOS: Vientiane Prov. Ban Van Eue 15. XII. 1966 ”; “ Native Collector BISHOP ”; BPBM.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5DFFCFFD80FB9B592F2F27.taxon	description	Redescription Male For a detailed description of the external features, see the original description (Pütz 2008). PW / PL 2.83; EL / EW 1.48; EL / PL 4.93; EW / PW 1.18; TL / EW 1.78. Sternite VII (Fig. 8 A) arcuate on caudal margin, bearing about 20 long setae in median part. Sternite VIII (Fig. 8 B) membranous. Sternite IX (Fig. 8 C) slightly sclerotized, with long apical plates bifid in apical parts. Aedeagus (Fig. 8 D – F) short, well sclerotized, slightly curved ventrally; basal piece oblong, subparallel-sided; lateral lobes short and broad, straight on lateral margins, closely punctuate, minutely projecting ventro-laterally at apices; median lobe slender, about 0.8 times as long as lateral lobe, slightly curved ventrally; ML / BL 0.34; ML / LL 0.82. Female Unknown. Measurements Male (n = 1): TL 1.78 mm; PW 0.85 mm; PL 0.30 mm; EL 1.48 mm; EW 1.00 mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5DFFCFFD80FB9B592F2F27.taxon	distribution	Distribution Thailand, Laos (new record).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5BFFCCFDA4FE9B5E4B2AC9.taxon	type_taxon	(type species: Pseudeucinetus zygops Heller, 1921).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5BFFCCFDA4FE9B5E4B2AC9.taxon	diagnosis	Diagnosis Body oblong. Gena wide; shallow antennal groove present. Frons (Fig. 2 E – F) narrow, distinctly projecting anteriorly. Antennae stout and short, 11 - segmented; antennal insertions situated near mandibles. Legs relatively short and stout; tarsal formula 4 - 4 - 4; male fore tarsi enlarged (Fig. 3 E).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F5BFFCCFDA4FE9B5E4B2AC9.taxon	discussion	Remarks This genus is easily distinguishable from other genera of the subfamily Thaumastodinae by the unique shape of the head: frons distinctly projecting anteriorly and narrow between the eyes. Biological notes The habitat of this genus (Fig. 4 A) is freshwater environments, i. e., rivers and streams. The adults are found in the splash zone of wet rocks, and frequently actively flying. In Java, Indonesia, Pseudeucinetus javanicus and Acontosceles javanicus were collected in the same place (Yoshitomi & Putra 2011). The adults were also collected by light trap. The larval stages are unknown.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F58FFD3FDA9FB185FE32CA3.taxon	description	urn: lsid: zoobank. org: act: 978 D 5 D 6 D- 8 D 48 - 425 D-AB 1 B- 0536 A 2 E 10753 Figs 9 A, 10	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F58FFD3FDA9FB185FE32CA3.taxon	etymology	Etymology This species is named after its type locality.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F58FFD3FDA9FB185FE32CA3.taxon	materials_examined	Material examined Holotype PAPUA NEW GUINEA • ♂; “ NEW GUINEA NETH. Waris, S. of Hollandia, 450 – 500 m VIII-I- 2 - 1959 ”; “ T. C. Maa Collector BISHOP ”; “ At Light ”; BPBM.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F58FFD3FDA9FB185FE32CA3.taxon	description	Description Male BODY. Oval, slightly convex dorsally, shiny, closely covered with suberect silver setae. Coloration of body black; antennae and legs blackish brown. HEAD. Moderate in size; clypeus with straight anterior margin. Eyes large; distance between eyes about 3 eye facets. ANTENNAE. Short, reaching about anterior third of pronotum. PRONOTUM. Widest at base; lateral margins straight; posterior margin gently bisinuate; PW / PL 2.33. Scutellum small, square. ELYTRA. Oblong, widest at basal third; EL / EW 1.43; EL / PL 4.00; EW / PW 1.20; TL / EW 1.79. Fore tarsomere I largely expanded, longer than remaining fore tarsomeres combined. ABDOMEN. Sternite VII (Fig. 10 A) with three pairs of anterior extra setae (a) and two pairs of lateral extra setae (b). Sternite IX (Fig. 10 B) moderately sclerotized, with long and pointed apical plates. AEDEAGUS (Fig. 10 C – D). Long, well sclerotized; basal piece oblong, subparallel-sided; lateral lobes long, about 1.4 times as long as basal piece, straight on lateral margins, with a pair of obtuse subapical inner projections on apical fifth; median lobe long, about 0.9 times as long as lateral lobes, nipple-like at apex; ML / BL 1.29; ML / LL 0.89. Female Unknown. Measurements Male (n = 1): TL 2.15 mm; PW 1.00 mm; PL 0.43 mm; EL 1.72 mm; EW 1.20 mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F58FFD3FDA9FB185FE32CA3.taxon	distribution	Distribution Papua New Guinea.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F58FFD3FDA9FB185FE32CA3.taxon	discussion	Remarks This species is related to Pseudeucinetus novabritannica from the Bismarck Archipelago and P. solomonicus from the Solomon Islands, and differs from them in the following characteristics: 1) subapical inner projection of lateral lobe obvious (large and distinct in P. solomonicus); 2) subbasal inner projection of lateral lobe absent (present in P. novabritannica); and 3) basal piece long, about 0.7 times as long as lateral lobe (about 0.5 in P. novabritannica, about 0.8 in P. solomonicus).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F47FFD3FE05FCEE58522B7C.taxon	materials_examined	Material examined INDONESIA • 1 ♂; “ INDONESIA: Lombok I: Aik Bukak 2. VII. 1978 ”; “ J. T. Medler Coll. BISHOP Museum Acc. # 1979. 483 ”; BPBM.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F47FFD3FE05FCEE58522B7C.taxon	distribution	Distribution Indonesia (Java, Lombok).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F47FFD3FE05FCEE58522B7C.taxon	discussion	Remarks This is the first record of this species from Lombok Island.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F47FFD0FDA1FAAF59AE2F0F.taxon	description	Fig. 9 E	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F47FFD0FDA1FAAF59AE2F0F.taxon	materials_examined	Material examined INDONESIA • 1 ♂; “ INDONESIA Southwest Sulawesi Aele 6 – 8. II. 2003 (LT) S. Hori leg. ”; EUMJ. MALAYSIA • 1 ♂; “ MALAYA Selangor Connaught Bridge 14. III. 1958 ”; “ T. C. Maa Collector BISHOP 108 ”; BPBM. PHILIPPINES • 2 ♂♂, 2 ♀♀; “ [PALAWAN] Olanguan (0 m) 76 km N. of Puerto Pron-cesa 31. VIII. 1985 M. Sakai leg. ”; “ attracted to mercury lamp ”; EUMJ • 3 ♂♂; same collection data as for preceding but “ 26. VIII. 1985 ”; EUMJ • 1 ♀; “ [PALAWAN] Trident Mine, 500 m foot of Victoria Peak nr. Narra 4. IX. 1985 M. Sakai ”; EUMJ • 1 ♂; same collection data as for preceding but “ 5. IX. 1985 ”; EUMJ • 1 ♂; “ [PALAWAN] Olanguan (0 – 50 m) between Puerto Princesa & Roxas 1. IX. 1985 M. Sakai ”; EUMJ.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F47FFD0FDA1FAAF59AE2F0F.taxon	distribution	Distribution India, Indonesia (Sulawesi), Malaysian Peninsula, Bismarck Archipelago (New Ireland), the Philippines (Balabac, Palawan, Mindanao, Negros).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F44FFD0FD45FC0858632B26.taxon	materials_examined	Material examined INDONESIA • 1 ♀; “ IRIAN JAYA East of Serui Yapen Is. 28. VII. 1996 N. Ohbayashi ”; EUMJ. BRAZIL • 1 ♀; “ BRAZIL: Cabo Frio 1 m 20 – 21. XI. 1970 ”; “ T. C. Maa Collector BISHOP ”; BPBM.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F44FFD0FD45FC0858632B26.taxon	discussion	Remarks The single female specimen labelled as having been collected in Brazil is questionable, because this genus is otherwise known only from the Oriental Region.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F44FFD0FDDBFE5B593A2DCC.taxon	description	Fig. 9 B	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F44FFD0FDDBFE5B593A2DCC.taxon	materials_examined	Material examined MALAYSIA • 1 ♂; “ Miri Sarawak 11 - XII- 1968 M. Satô leg. ”; EUMJ • 1 ♂; “ MALAYSIA Mamut Ranau Sabah 27. Ii. 1995 [L. T.] YR. SAWADA leg. ”; EUMJ • 1 ♂; “ Malaysia SABAH Kg. Sukau, Sandakan 15 - 16. XII. 2009 H. Takizawa ”; ITBC • 1 ♀; “ Malaysia SABAH Kg. Tikolod, Salt trail Tambunan 11. XI. 2009 H. Takizawa ”; ITBC • 1 ♀; “ Malaysia SABAH Kg. Mansiang, Mengg-atal, Kota Kinabalu 21. XI. 2009 H. Takizawa ”; ITBC.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F44FFD0FDDBFE5B593A2DCC.taxon	distribution	Distribution Malaysia (Sarawak, Sabah).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F44FFD1FD89FA9F590B2B09.taxon	type_taxon	(type species: Mexico litoralis Spilman, 1972, by original designation).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F44FFD1FD89FA9F590B2B09.taxon	type_taxon	(type species: Babalimnichus taiwanus Satô, 1994: 174, by original designation). Syn. nov. Mexico – Skelley 2005: 119. Babalimnichus – Yoshitomi & Satô 2001: 472 [larval description].	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F44FFD1FD89FA9F590B2B09.taxon	diagnosis	Diagnosis Body oblong. Gena relatively narrow; shallow antennal groove present. Antennae stout and short, 11 - segmented; antennal insertions situated near mandibles. Legs relatively short and stout; tarsal formula 4 - 4 - 4.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F44FFD1FD89FA9F590B2B09.taxon	discussion	Remarks As already mentioned by Skelley (2005), Babalimnichus Satô, 1994 cannot be separated from Mexico, and it should be treated as a junior synonym of the latter. The male genital structures are very peculiar, i. e., with very short lateral lobes with apical setae, longer median lobe and the presence of ventral plates (Fig. 1 A). Biological notes The habitat of this genus (Fig. 4 B – C) is rocky seashores. The adults (Fig. 4 E) are found in the splash zone of wet rocks, actively flying when disturbed. The larval stage was described by Yoshitomi & Satô (2001).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F45FFD4FDBBFA585E2E2F27.taxon	description	urn: lsid: zoobank. org: act: 8 B 1989 C 3 - 5 D 1 B- 4 E 9 C-A 87 F- 0 ED 19524 F 6 D 4 Figs 4 C, 11 C, 12	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F45FFD4FDBBFA585E2E2F27.taxon	etymology	Etymology This species is named after its type locality.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F45FFD4FDBBFA585E2E2F27.taxon	materials_examined	Material examined Holotype JAPAN • ♂; “ [OG 2] Meijima, Hahajima-guntô, Ogasawara Isls., Japan, 26 ° 567418 ″ N, 142 ° 227674 ″ E, 23. IX. 2012, H. Yoshitomi leg. ”; EUMJ. Paratypes JAPAN • 4 ♂♂, 2 ex.; same collection data as for holotype; EUMJ • 1 ♂, 1 ♀; same collection data as for preceding; NSMT • 1 ♂, 1 ♀; same collection data as for preceding; NMW • 1 ♀; “ [OG 5] Minamizaki, Hahajima, Ogasawara Isls., Japan, 26 ° 37.491 ″ N, 142 ° 10.600 ″ E, 25. IX. 2012, H. Yoshitomi leg. ”; EUMJ • 4 ♂♂, 2 ex.; “ [OG 4] Hirashima, Hahajima-guntô, Ogasawara Isls., Japan, 26 ° 35.169 ″ N, 142 ° 09.177 ″ E, 24. IX. 2012, H. Yoshitomi leg. ”; EUMJ.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F45FFD4FDBBFA585E2E2F27.taxon	description	Description Male BODY. Oval, slightly convex dorsally, shiny, closely covered with suberect golden setae. Coloration of body black; elytra with indistinct and irregular silver spots consisting of erect setae. HEAD. Closely and finely punctate, slightly convex between eyes; distance between eyes as long as an eye diameter in dorsal view. ANTENNAE. Short, reaching about anterior third of pronotum. PRONOTUM. Closely punctate, gently arcuate on lateral margins; PW / PL 1.92 – 2.30 (2.09). Scutellum equilateral triangular, punctate as in pronotum. ELYTRA. Oval, widest at base, gently tapered posteriorly; caudal third of lateral margins slightly and indistinctly serrate; EL / EW 1.40 – 1.54 (1.48); EL / PL 3.25 – 4.00 (3.58); EW / PW 1.12 – 1.20 (1.16); TL / EW 1.80 – 1.96 (1.89). ABDOMEN. Sternite VII (Fig. 12 A) bearing about 40 long setae on mesal part, pointed at postero-lateral corners, slightly projecting on median part of caudal margin. Sternite VIII (Fig. 12 B) slightly sclerotized, small, Y-shaped. Sternite IX (Fig. 12 C) moderately sclerotized, relatively wide, strongly curved in basal part, with long and pointed apical plates. AEDEAGUS (Fig. 12 D – G). Long and slender, well sclerotized; basal piece oval; lateral lobes short and slender, with three apical setae; median lobe long and slender, slightly expanded in apical fifth, pointed at apex, finely punctuate; ventral plates slender, reaching about apical fifth of median lobe, mostly fused to median lobe; ML / BL 2.94; ML / LL 2.42. Female Sexual dimorphism slight. Sternite VII (Fig. 12 H) bearing about 30 long setae on mesal part, pointed but minutely concaved at postero-lateral corners, slightly projecting on median part of caudal margin. Urosternite (Fig. 12 I) well sclerotized, T-shaped, with long and slender apodeme. Ovipositor (Fig. 12 J) well sclerotized; coxite sparsely punctuate; apex of baculus bifid; approximate ratio of coxite and baculus (n = 1) 1.0: 6.1. Measurements Unsexed (n = 20): TL 2.10 – 2.65 (2.39) mm; PW 0.98 – 1.20 (1.09) mm; PL 0.45 – 0.60 (0.52) mm; EL 1.65 – 2.05 (1.86) mm; EW 1.10 – 1.40 (1.26) mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F45FFD4FDBBFA585E2E2F27.taxon	distribution	Distribution Japan: Ogasawara Islands (Hahajima, Meijima, Hirashima).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F45FFD4FDBBFA585E2E2F27.taxon	discussion	Remarks This species appears to be closely related to M. baliensis sp. nov. based on the shape of the aedeagus, but differs from it in the following characteristics: sternite IX broad (slender in M. baliensis sp. nov.), median lobe pointed at apex (somewhat obtuse in M. baliensis sp. nov.) and apical setae on lateral lobes long (short in M. baliensis sp. nov.). Biological notes The type series was collected from the splash zone of wet rocks on a rocky seashore (Fig. 4 C).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F41FFDBFD66FEEB5E5B2D14.taxon	description	urn: lsid: zoobank. org: act: 98 D 64 DE 9 - 6 A 33 - 4 EBF-A 7 F 9 - 26 F 9 A 7 BA 409 E Figs 4 B, 11 D, 13	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F41FFDBFD66FEEB5E5B2D14.taxon	etymology	Etymology This species is named after its type locality.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F41FFDBFD66FEEB5E5B2D14.taxon	materials_examined	Material examined Holotype INDONESIA • ♂; “ Bali Is. Indonesia Nusa Dua, Kuta South 16 - II- 2013 Kiyoshi ANDO leg. ”; EUMJ. Paratypes INDONESIA • 1 ♂, 3 ♀♀, 2 ex.; same collection data as for holotype; EUMJ • 3 ex.; “ Nusa Dua, North Island 8 ° 47 ′ 52.72 ″ S, 115 ° 14 ′ 9.31 ″ E, Bali, INDONESIA 7 – 8. VIII. 2010 H. Yoshitomi leg. ”; EUMJ • 1 ex.; same collection data as for preceding; NMW • 1 ex.; same collection data as for preceding; BPBM • 4 ♂♂, 4 ♀♀, 13 ex.; “ Nusa Dua, South Island 8 ° 48 ′ 6.81 ″ S, 115 ° 14 ′ 16.07 ″ E, Bali, INDONESIA 8. VIII. 2010 H. Yoshitomi leg. ”; 1 ♀ on slide no HY 1141; EUMJ • 3 ex.; same collection data as for preceding; NMW • 3 ex.; same collection data as for preceding; BPBM.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F41FFDBFD66FEEB5E5B2D14.taxon	description	Description Male BODY. Oval, slightly convex dorsally, shiny, closely covered with suberect golden setae. Coloration of body black; elytra with small, irregular silver spots consisting of erect setae. HEAD. Finely punctate, projecting anteriorly between eyes; distance between eyes about 0.8 times as long as eye diameter in dorsal view. ANTENNAE. Very short, reaching about anterior quarter of pronotum. PRONOTUM. Closely punctate, gently arcuate on lateral margins; PW / PL 1.75 – 2.55 (2.06). Scutellum square. ELYTRA. Oval, widest at base, gently tapered posteriorly; caudal third of lateral margins serrate; EL / EW 1.32 – 1.43 (1.38); EL / PL 2.53 – 4.00 (3.25); EW / PW 1.09 – 1.20 (1.14); TL / EW 1.67 – 1.88 (1.81). ABDOMEN. Sternite VII (Fig. 13 A) with two pairs (inner and outer) of long and stout setae and two extra pairs of such setae, pointed at postero-lateral corners, slightly projecting on median part of caudal margin. Sternite VIII (Fig. 13 B) slightly sclerotized, small, Y-shaped. Sternite IX (Fig. 13 C) moderately sclerotized, slender, curved in basal part; apical plates long, truncate at apices. AEDEAGUS (Fig. 13 D – G). Long and slender; basal piece oval, subparallel-sided; lateral lobes short and robust, with 2 – 3 apical setae; median lobe long and slender, subparallel-sided near base to apex, finely punctuate, pointed at apex; ventral plates long and slender, reaching about apical tenth of median lobe, mostly fused to median lobe; ML / BL 2.26; ML / LL 2.72. Female Sexual dimorphism slight. Sternite VII (Fig. 13 H) similar to that of male, but one pair, not two, of extra inner setae. Urosternite (Fig. 13 I) well sclerotized, Y-shaped, with long and slender apodeme. Ovipositor (Fig. 13 J) well sclerotized; coxite sparsely punctuate, rather pointed at apices; apex of baculus bifid; approximate ratio of coxite and baculus (n = 1) 1.0: 6.2. Measurements Unsexed (n = 20): TL 1.88 – 2.40 (2.19) mm; PW 0.95 – 1.13 (1.06) mm; PL 0.40 – 0.60 (0.52) mm; EL 1.48 – 1.80 (1.67) mm; EW 1.05 – 1.30 (1.21) mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F41FFDBFD66FEEB5E5B2D14.taxon	distribution	Distribution Indonesia (Bali Island).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F41FFDBFD66FEEB5E5B2D14.taxon	discussion	Remarks This species is similar to M. borneensis sp. nov. in the shape of the aedeagus, but differs from it in the following characteristics: median lobe obtuse at apex (pointed in M. borneensis sp. nov.); and apical setae on lateral lobes relatively short (long in M. borneensis sp. nov.). Biological notes The type series was collected from the surface of wet rocks on a seashore (Fig. 4 B) together with three species of Laius Guérin-Méneville, 1830 (Melyridae: L. baliensis Yoshitomi, 2014, L. satoi Yoshitomi, 2008 and L. pankowi Wittmer, 1999) and Parathroscinus sp. (Limnichidae) (Yoshitomi 2008, 2014 b).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4FFFD9FD9FFC4658892A69.taxon	description	urn: lsid: zoobank. org: act: DB 69 DE 47 - 18 E 6 - 4 FA 0 - 8 D 94 - A 9 A 95470 D 78 E Figs 11 B, 14	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4FFFD9FD9FFC4658892A69.taxon	etymology	Etymology This species is named after its type locality.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4FFFD9FD9FFC4658892A69.taxon	materials_examined	Material examined Holotype PAPUA NEW GUINEA • ♂; “ NEW GUINEA (NE) Dreikikir, Sepik Distr., 350 m 25. VI. 1961 ”; “ J. L. & M. Gressitt Collectors ”; BPBM. Paratypes PAPUA NEW GUINEA • 1 ♂, 1 ♀; same collection data as for holotype; BPBM • 1 ♀; same collection data as for holotype but “ 23. VI. 1961 ”; BPBM • 1 ♀; same collection data as for holotype but “ 24. VI. 1961 ”; BPBM • 1 ♀; “ NEW GUINEA: NETH. VOG ELKOP: Bomberi 700 – 900 m, VI- 9 - ‘ 59 ”; “ J. L. Gressitt Collector ”; BPBM • 1 ♂; “ NE New Guinea Morabe Distr. 10 km W Bulolo 780 m, 5 – 25. VIII, 1967 ”; “ Malaise tr. over stream ”; “ R. Straatman Collector BISHOP MUS. ”; BPBM • 1 ♂; “ NE NEW GUINEA Akivitana Riv. 1550 m, 10. I. 65 ”; “ J. & M. Sedlacek Collectors BISHOP MUSEUM ”; BPBM • 1 ♂; “ NEW GUINEA: NETH. Waris, S. of Hollandia, 450 – 500 m VIII- 1 - 2 - 1959 ”; “ At Light ”; “ T. C. Maa Collector BISHOP ”; BPBM.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4FFFD9FD9FFC4658892A69.taxon	description	Description Male BODY. Oval, relatively flat dorsally, shiny, closely covered with suberect golden setae. Coloration of body black; antennae and legs dark brown; elytra with indistinct and irregular silver spots consisting of erect setae. HEAD. Finely punctate, shallowly depressed on dorsum between eyes; distance between eyes as long as eye diameter in dorsal view. ANTENNAE. Slender and short, reaching about anterior third of pronotum. PRONOTUM. Closely and rugosely punctate, with lateral margins almost straight; PW / PL 2.51 – 2.71 (2.61). Scutellum small, square. ELYTRA. Oval, widest at base, gently tapered posteriorly; caudal quarter of lateral margins slightly and minutely serrate; EL / EW 1.35 – 1.40 (1.38); EL / PL 3.77 – 4.00 (3.88); EW / PW 1.05 – 1.11 (1.08); TL / EW 1.71 – 1.75 (1.73). ABDOMEN. Sternite VII (Fig. 14 A) with two (inner and outer) pairs of long and slender setae and two extra setae, pointed in postero-lateral corners, gently projecting triangularly on mesal part of caudal margin. Sternite VIII (Fig. 14 B) small, slightly sclerotized, Y-shaped. Sternite IX (Fig. 14 C) wide, gently curved in basal part; apical plates long and slender, pointed at apices. AEDEAGUS (Fig. 14 D – F). Long, almost symmetrical; basal piece oval; lateral lobes short, with 4 short apical setae; median lobe slender, sparsely punctuate in apical part, slightly expanded in apical quarters, pointed and somewhat prolonged at apex; ventral plates slender, expanded apically, separated to median lobe; ML / BL 2.35; ML / LL 2.61. Female Sexual dimorphism slight. PW / PL 2.38 – 2.45 (2.42); EL / EW 1.36 – 1.42 (1.40); EL / PL 3.70 – 4.11 (3.85); EW / PW 1.07 – 1.18 (1.14); TL / EW 1.73 – 1.79 (1.76). Sternite VII (Fig. 14 G) similar to that of male, but postero-lateral corners shallowly concave at apices. Urosternite (Fig. 14 H) well sclerotized, Y-shaped, with long and slender apodeme. Ovipositor (Fig. 14 I) well sclerotized; coxite sparsely punctuate, rather pointed at apices; apex of baculus bifid; approximate ratio of coxite and baculus (n = 1) 1.0: 5.6. Measurements Male (n = 2): TL 1.75 – 2.05 (1.90) mm; PW 0.95 – 1.08 (1.02) mm; PL 0.35 – 0.43 (0.39) mm; EL 1.40 – 1.62 (1.51) mm; EW 1.00 – 1.20 (1.10) mm. Female (n = 3): TL 1.88 – 2.30 (2.03) mm; PW 0.95 – 1.10 (1.01) mm; PL 0.40 – 0.45 (0.42) mm; EL 1.48 – 1.85 (1.61) mm; EW 1.05 – 1.30 (1.15) mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4FFFD9FD9FFC4658892A69.taxon	distribution	Distribution Papua New Guinea.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4FFFD9FD9FFC4658892A69.taxon	discussion	Remarks This species is distinct in the peculiar form of the male genitalia.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4DFFDFFD9AFBB958CB2A94.taxon	description	urn: lsid: zoobank. org: act: 9 A 734 C 2 D-BF 7 A- 4 F 05 - 8628 - 36655720 A 091 Figs 11 E, 15	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4DFFDFFD9AFBB958CB2A94.taxon	etymology	Etymology This species is named after its type locality.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4DFFDFFD9AFBB958CB2A94.taxon	materials_examined	Material examined Holotype PALAU • ♂; “ Carp Is. Palau 13 – 15. IX. 2002 M. Sato leg. ”; EUMJ. Paratypes PALAU • 49 ex.; same collection data as for holotype; EUMJ • 4 ex.; same collection data as for preceding; NMW • 4 ex.; same collection data as for preceding; BPBM • 1 ♀, 2 ex.; “ Carp Is. Palau 20 - VII- 2003 K. Takahashi leg. ”; EUMJ • 1 ♂, 4 ex.; “ Palau Angaur Is. 16 - VIII- 2003, M. Sato leg. ”; EUMJ • 1 ♂, 6 ex.; “ Palau Peleliu Is. 13 – 17. VIII. 2003 M. Sato leg. ”; 1 ♂ on slide no. HY 1038; EUMJ.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4DFFDFFD9AFBB958CB2A94.taxon	description	Description Male BODY. Oval, slightly convex dorsally, shiny, closely covered with suberect golden setae. Coloration of body black; elytral silver spots indistinct. HEAD. Finely punctate, slightly convex on dorsum between eyes; distance between eyes about 0.8 times as long as eye diameter in dorsal view. ANTENNAE. Very short, reaching about anterior quarter of pronotum. PRONOTUM. Closely punctate, gently arcuate on lateral margins; PW / PL 1.96 – 2.80 (2.22). Scutellum small, square. ELYTRA. Oval, widest at base, gently tapered posteriorly; caudal third of lateral margins slightly and indistinctly serrate; EL / EW 0.42 – 1.43 (1.30); EL / PL 1.13 – 4.13 (3.14); EW / PW 1.05 – 1.16 (1.09); TL / EW 0.79 – 1.87 (1.72). ABDOMEN. Sternite VII (Fig. 15 A) with two pairs of long and stout extra setae, bearing about 50 long setae, obtuse at postero-lateral corners, pentagonally projecting in median part of caudal margin. Sternite VIII (Fig. 15 B) slightly sclerotized, small, Y-shaped. Sternite IX (Fig. 15 C) well sclerotized, curved in basal part; apical plates long, pointed at apices. AEDEAGUS (Fig. 15 D – G). Long and slender, almost symmetrical; basal piece oval, gently tapering basally; lateral lobes relatively long and slender, with 2 – 3 very short apical setae, sparsely punctuate; median lobe long, subparallel-sided near base to apical fifth, sparsely punctuate, pointed at apex; ventral plates long and slender, reaching about apical fifth of median lobe, mostly fused to median lobe; ML / BL 2.06; ML / LL 1.92. Female Sexual dimorphism slight. Sternite VII (Fig. 15 H) similar to that of male. Urosternite (Fig. 15 I) well sclerotized, with long and slender apodeme, with slender lateral projections. Ovipositor (Fig. 15 J) well sclerotized; coxite punctuate, obtuse at apices; apex of baculus pointed; approximate ratio of coxite and baculus (n = 1) 1.0: 6.8. Measurements Unsexed (n = 20): TL 0.85 – 2.23 (1.93) mm; PW 0.82 – 1.12 (1.03) mm; PL 0.35 – 0.55 (0.47) mm; EL 0.45 – 1.68 (1.46) mm; EW 0.95 – 1.20 (1.12) mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4DFFDFFD9AFBB958CB2A94.taxon	distribution	Distribution Palau (Carp, Peleliu Islands).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4DFFDFFD9AFBB958CB2A94.taxon	discussion	Remarks This species is similar to M. borneensis sp. nov. in the shape of the aedeagus, but differs from it in the following characteristics: ventral plates separated from median lobe (attached in M. borneensis sp. nov.) and apical setae on lateral lobes short (long in M. borneensis sp. nov.).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4BFFDDFD95FAC5583D282A.taxon	description	urn: lsid: zoobank. org: act: C 119 FDE 3 - 9 A 59 - 4 A 07 - A 77 A-AF 4 C 067 C 5 D 1 B Figs 11 A, 16	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4BFFDDFD95FAC5583D282A.taxon	etymology	Etymology This species is named after its type locality.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4BFFDDFD95FAC5583D282A.taxon	materials_examined	Material examined Holotype MALAYSIA • ♂; “ BRITISH N BORNEO Labuan Island Nov. 28 – 29, 1958 ”; “ Intertidal Zone ”; “ L. W. Quate Collector ”; BPBM. Paratypes MALAYSIA • 1 ♀; same collection data as for holotype; BPBM • 1 ♂, 1 ex.; “ BRITISH N BORNEO Labuan Island Nov. 28 – 29, 1959 ”; “ Intertidal Zone ”; “ Maa Collector ”; BPBM.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4BFFDDFD95FAC5583D282A.taxon	description	Description Male BODY. Oval, slightly convex dorsally, shiny, closely covered with suberect golden setae. Coloration of body black; elytra with small, irregular silver spots consisting of erect setae. HEAD. Finely punctate, slightly convex on dorsum between eyes; distance between eyes about 0.8 times as long as eye diameter in dorsal view. ANTENNAE. Short, reaching about anterior third of pronotum. PRONOTUM. Closely punctate; lateral margins gently arcuate; PW / PL 2.04 – 2.30 (2.20). Scutellum small, square. ELYTRA. Oval, widest at base, gently tapered posteriorly; caudal quarter of lateral margins slightly and minutely serrate; EL / EW 1.33 – 1.48 (1.38); EL / PL 3.11 – 3.70 (3.35); EW / PW 1.09 – 1.14 (1.11); TL / EW 1.73 – 1.88 (1.79). ABDOMEN. Sternite VII (Fig. 16 A) with two pairs of long and stout extra setae, bearing about 20 long setae, bifid in postero-lateral corners, gently projecting triangularly in median part of caudal margin. Sternite VIII (Fig. 16 B) small, slightly sclerotized, Y-shaped. Sternite IX (Fig. 16 C) well sclerotized, lightly curved in basal part; apical plates long and stout, pointed at apices. AEDEAGUS (Fig. 16 D – F). Long and slender, almost symmetrical; basal piece oval; lateral lobes short, with 4 apical setae; median lobe long and slender, subparallel-sided near base to near apex, finely punctuate, with nipple-like apex; ventral plates long and slender, reaching about apical tenth of median lobe, mostly fused to median lobe; ML / BL 2.36; ML / LL 2.57. Female Sexual dimorphism slight. Sternite VII (Fig. 16 G) similar to that of male, but lacking extra setae. Urosternite (Fig. 16 H) well sclerotized, T-shaped, with long and slender apodeme. Ovipositor (Fig. 16 I) well sclerotized; coxite sparsely punctuate, obtuse at apices; apex of baculus bifid; approximate ratio of coxite and baculus (n = 1) 1.0: 5.5. Measurements Unsexed (n = 3): TL 1.70 – 1.88 (1.81) mm; PW 0.90 – 0.92 (0.91) mm; PL 0.40 – 0.45 (0.42) mm; EL 1.30 – 1.48 (1.39) mm; EW 0.98 – 1.05 (1.01) mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4BFFDDFD95FAC5583D282A.taxon	distribution	Distribution Malaysia (Labuan Island, off Borneo).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F4BFFDDFD95FAC5583D282A.taxon	discussion	Remarks This species is similar to M. baliensis sp. nov. in the shape of the aedeagus, but differs from it in the following characteristics: median lobe pointed at apex (obtuse in M. baliensis sp. nov.) and lateral lobes bearing 4 apical setae (3 in M. baliensis sp. nov.).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F49FFE3FDC0F9775E592B9F.taxon	description	Figs 11 G, 17	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F49FFE3FDC0F9775E592B9F.taxon	materials_examined	Material examined JAPAN • 17 ex.; “ Hikawa-hama Yonaguni-jima OKINAWA 18. VI. 2001 Koji TOYODA leg. ”; EUMJ • 4 ex.; “ Dannuhama Yonaguni-jima 26 - VIII- 1994 M. Sato leg. ”; EUMJ. TAIWAN • 1 ♂, 1 ♀, 5 ex.; “ Lung Keng Pintung Taiwan 10 - VIII- 2000 M. Sato leg. ”; EUMJ • 1 ex.; “ Mao tantou Pintung, Taiwan 10 - VIII- 2000 M. Sato leg. ”; EUMJ • 1 ex.; “ Lutao Taiwan 26 - 28. III. 1998 M. Sato leg. ”; EUMJ • 5 ex.; “ Lanhsu Taiwan 5 – 10. VIII. 1998 M. Sato leg. ”; EUMJ • 3 ex., in ethanol; “ Fushan Village Donghe Township, Taitung Country, Taiwan 1 - V- 2013 Tatsuya Niisato leg. ”; EUMJ.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F49FFE3FDC0F9775E592B9F.taxon	description	Redescription Male For a detailed description of the external features, see the original description (Satô 1994). PW / PL 1.67 – 2.44 (2.00); EL / EW 1.18 – 1.42 (1.32); EL / PL 2.33 – 3.40 (2.87); EW / PW 1.00 – 1.15 (1.09); TL / EW 1.59 – 1.92 (1.78). Sternite VII (Fig. 17 A) with two pairs of long, stout extra setae, bearing about 20 long setae, bifid in postero-lateral corners, gently projecting triangularly in median part of caudal margin. Sternite VIII (Fig. 17 B) small, slightly sclerotized, Y-shaped. Sternite IX (Fig. 17 C) slender, lightly curved in basal part; apical plates long and slender, pointed at apices. Aedeagus (Fig. 17 D – F) relatively short, asymmetrical; basal piece oval; lateral lobes short, with 2 – 4 apical setae; median lobe a little wide, sparsely punctuate in apical part, slightly expanded in apical quarter, pointed at apex; ventral plates relatively slender, separated from median lobe, left one with thumb-like projection on inner margin of apex; ML / BL 2.42; ML / LL 2.42. Female Sexual dimorphism slight. Sternite VII (Fig. 17 G) similar to that of male. Urosternite (Fig. 17 H) well sclerotized, with long and slender apodeme, T-shaped. Ovipositor (Fig. 17 I) well sclerotized; coxite sparsely punctuate, rather pointed at apices; apices of baculus bifid; approximate ratio of coxite and baculus (n = 1) 1.0: 5.3. Measurements Unsexed (n = 20): TL 1.68 – 2.17 (1.95) mm; PW 0.90 – 1.10 (1.01) mm; PL 0.40 – 0.60 (0.51) mm; EL 1.28 – 1.62 (1.44) mm; EW 1.00 – 1.15 (1.10) mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F49FFE3FDC0F9775E592B9F.taxon	distribution	Distribution Taiwan (including Lutao and Lanhsu Islands), Japan (Yonaguni-jima).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F49FFE3FDC0F9775E592B9F.taxon	discussion	Remarks This is the first record of this species from Lutao and Lanhsu Islands, off Taiwan, and Yonaguni-jima.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F77FFE6FDC7F9CB596F2DCB.taxon	description	Figs 11 F, 18 – 19	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F77FFE6FDC7F9CB596F2DCB.taxon	materials_examined	Material examined JAPAN • 7 ex.; “ Japan: Honshû Nanaura-kaigan Tachibana, Sado-shi Niigata, 5. vi. 2016 37.986591, 138.234143 Kamezawa H. leg. ”; EUMJ • 1 ♂; “ [IZ 9] Ishijirogawa, Shikinejima, Izu Isls., Japan, 34 ° 19.409 ″ N, 139 ° 13.308 ″ E, ca 16 m, 7. II. 2013, H. Yoshitomi leg. ”; EUMJ • 1 ♂, 1 ♀, 14 ex.; “ Kariya-kaigan Tokoname-shi Aichi Pref. 17. VIII. 1995, 17. VIII. 1995 H. Yoshitomi leg. ”; 1 ♂ and 1 ♀ on slides HY 989, 992 – 994; EUMJ • 1 ex.; “ Zoubigasaki, Murozumi Hikari-shi Yamaguchi Pref., Jpn 28. VII. 2005 Y. Fujitani leg. ”; EUMJ • 5 ex.; “ Bashayamamura Amami-oshima 3 - V- 1999 M. Sato leg. ”; EUMJ • 28 ex.; “ Kawata Okinawa 5 - V- 1999 M. Sato leg. ”; EUMJ • 9 ex.; “ Kim-kaigan Okinawa 28 - III- 1997 M. Satô leg. ”; EUMJ • 1 ex.; “ Onnason Okinawa-jima 26 - 28. IV. 2004 M. Sato leg. ”; EUMJ • 1 ex.; “ Kume-jima Ryukyu 30 - IV- 2004 M. Sato leg. ”; EUMJ • 2 ex.; “ Tomori Ryukyus Miyako-jima 7 - V- 1998 M. Satô leg. ”; EUMJ • 13 ex.; “ Uganzaki Ishigaki 22 - IX- 2003 M. Sato leg. ”; EUMJ • 7 ex., in ethanol; “ Sakaura, Izumo, Shimane 31. VII. 2006 M. Hayashi ”; EUMJ • 2 pupae, with larval skins, in ethanol; “ Nahama-kaigan, Nagai, Yokosuka-shi Kanagawa Pref., Japan 6. July 2007 M. Asano leg. ”; EUMJ.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F77FFE6FDC7F9CB596F2DCB.taxon	description	Redescription Male For a detailed description of the external features, see the original description (Satô 1994). PW / PL 1.67 – 2.22 (1.99); EL / EW 1.27 – 1.48 (1.36); EL / PL 2.64 – 3.29 (2.96); EW / PW 1.00 – 1.18 (1.10); TL / EW 1.71 – 1.95 (1.82). Sternite VII (Fig. 18 A) bearing about 30 long setae, lacking extra setae; postero-lateral corners short and pointed; median part of caudal margin gently projecting triangularly. Sternite VIII (Fig. 18 B) small, slightly sclerotized, Y-shaped. Sternite IX (Fig. 18 C) well sclerotized, curved in basal part, shallowly concave at apex; apical plates rather stout, pointed at apices. Aedeagus (Fig. 18 D – F) relatively short, asymmetrical; basal piece oval; lateral lobes short, with 3 – 4 apical setae; median lobe wide, sparsely punctuate in apical part, expanded in apical quarter, gently pointed at apex; ventral plates wide, separated from median lobe, left one long and curved interiorly in apical part, right one short and straight; ML / BL 2.24; ML / LL 2.43. Female Sexual dimorphism slight. Sternite VII (Fig. 18 G) similar to that of male. Urosternite (Fig. 18 H) well sclerotized, T-shaped, with long and slender apodeme. Ovipositor (Fig. 18 I) well sclerotized; coxite sparsely punctuate, rather pointed at apices; apex of baculus bifid; approximate ratio of coxite and baculus (n = 1) 1.0: 5.9. Pupae Body (Fig. 19) about 1.5 mm, cream-colored, bearing long setae on head, pronotum, elytra and abdomen. Measurements Unsexed (n = 20): TL 1.82 – 2.32 (2.04) mm; PW 0.90 – 1.10 (1.02) mm; PL 0.45 – 0.60 (0.52) mm; EL 1.32 – 1.72 (1.53) mm; EW 1.00 – 1.30 (1.12) mm.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F77FFE6FDC7F9CB596F2DCB.taxon	distribution	Distribution Japan (Honshu, Sadogashima, Izu Islands (Shikine-jima, Kozu-shima), Shikoku, Kyushu, the Ryukyus (Yakushima, Amami-Ôshima, Tokuno-shima, Okinoerabu-jima, Yoron-tô, Okinawa-jima, Irabu-jima, Kume-jima, Miyako-jima, Ishigaki-jima )).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F77FFE6FDC7F9CB596F2DCB.taxon	discussion	Remarks I re-examined the specimens from Yonaguni-jima (Yoshitomi & Arai 2004), and it is clear that they do not represent this species but the preceding one. This is the first record of this species from Kume-jima. This species has the northernmost distribution (Sado Island) in the subfamily Thaumastodinae (Kamezawa 2017). Biological notes Overwintering takes place in the larval stage in Chiba Prefecture (Ono et al. 2012) or the adult stage on the Izu Islands (Yoshitomi 2014 a).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F72FFE6FE26FC3F5FD22BA8.taxon	distribution	Distribution Tonga Islands.	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F72FFE6FE26FC3F5FD22BA8.taxon	discussion	Remarks This species was described based on the female holotype. No additional specimens have been found. Phylogeny As a result of my analysis, one most parsimonious tree was obtained (L = 17, Ci = 88, Ri = 60), shown in Fig. 20. The synapomorphies of the subfamily Thaumastodinae are as follows: oblong body shape (1 - 1); eyes large, situated on dorso-lateral side of head (3 - 1); and hind legs with tibial spurs (9 - 1).	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F7CFFE8FD89FEEB5F102AA5.taxon	description	Mexico baliensis sp. nov. (Indonesia [Bali Island]) urn: lsid: zoobank. org: act: 98 D 64 DE 9 - 6 A 33 - 4 EBF-A 7 F 9 - 26 F 9 A 7 BA 409 E Mexico borneensis sp. nov. (Malaysia [Borneo]) urn: lsid: zoobank. org: act: C 119 FDE 3 - 9 A 59 - 4 A 07 - A 77 A-AF 4 C 067 C 5 D 1 B Mexico litoralis Spilman, 1972 (Mexico) urn: lsid: zoobank. org: act: 10667682 - 8 B 8 C- 4 E 02 - 9 AC 3 - 678 D 8 B 291954 Mexico masamii (Satô, 1994) (Japan [Honshu, Izu Islands, Shikoku, Kyushu, the Ryukyus]) urn: lsid: zoobank. org: act: 85474634 - 5 E 5 A- 4 F 83 - AD 88 - EDA 6242183 A 0 Mexico morrisoni Skelley, 2005 (Bahamas) urn: lsid: zoobank. org: act: 0 D 1 B 37 BE-BF 21 - 4679 - A 711 - 2 B 0 C 15 A 6 AC 96 Mexico ogasawaraensis sp. nov. (Japan [Ogasawara Islands]) urn: lsid: zoobank. org: act: 8 B 1989 C 3 - 5 D 1 B- 4 E 9 C-A 87 F- 0 ED 19524 F 6 D 4 Mexico palauensis sp. nov. (Palau) urn: lsid: zoobank. org: act: 9 A 734 C 2 D-BF 7 A- 4 F 05 - 8628 - 36655720 A 091 Mexico papuanus sp. nov. (Papua New Guinea) urn: lsid: zoobank. org: act: DB 69 DE 47 - 18 E 6 - 4 FA 0 - 8 D 94 - A 9 A 95470 D 78 E Mexico splendens (Hernando & Ribera, 2003) (Tonga Islands) urn: lsid: zoobank. org: act: 119 CC 677 - 2 FBF- 4117 - B 0 DA- 3931 B 3 A 6 CDF 1 Mexico taiwanus (Satô, 1994) (Taiwan, Japan [Yonaguni-jima]) urn: lsid: zoobank. org: act: CBAA 7928 - BE 17 - 4 A 2 B-AF 43 - 1 E 2 D 35 FBD 09 E	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
03F9EF199F7CFFE8FDA4FAE65F2B2998.taxon	description	Pseudeucinetus javanicus Yoshitomi & Putra, 2010 (Indonesia [Java, Lombok]) urn: lsid: zoobank. org: act: 9766 FA 9 E- 388 C- 455 E- 883 E- 7 FEF 7 DE 27 D 2 A Pseudeucinetus novabritannica Delève, 1973 (Bismarck Archipelago [New Britain]) urn: lsid: zoobank. org: act: D 4 F 058 A 4 - 8841 - 48 EF-B 1 EA- 03 BA 40 B 1 B 6 E 2 Pseudeucinetus papuanus sp. nov. (Papua New Guinea) urn: lsid: zoobank. org: act: 978 D 5 D 6 D- 8 D 48 - 425 D-AB 1 B- 0536 A 2 E 10753 Pseudeucinetus solomonicus Yoshitomi & Putra, 2010 (Solomon Islands [Santa Catalina, Santa Anna]) urn: lsid: zoobank. org: act: 113 B 8 D 47 - F 5 AA- 4 EC 7 - 8 F 07 - 9921 CB 8347 D 8 Pseudeucinetus spilmani Spangler, 1995 (Indonesia [Bacan]) urn: lsid: zoobank. org: act: 9 DAB 932 B- 0495 - 4 EE 9 - B 97 D-AAA 873871 A 2 E Pseudeucinetus uenoi Spangler, 1995 (Malaysia [Borneo]) urn: lsid: zoobank. org: act: 5 CC 8 AB 17 - 8 B 02 - 4170 - 97 C 2 - 7 A 18 F 85 D 751 D Pseudeucinetus zygops Heller, 1921 (India, Malaysian Peninsula, Bismarck Islands [New Ireland], the Philippines [Balabac, Palawan, Mindanao, Negros], Indonesia [Sulawesi]) urn: lsid: zoobank. org: act: 7903 C 913 - 85 A 7 - 4 AE 6 - B 4 E 4 - 619 A 86 BB 46 CB	en	Yoshitomi, Hiroyuki (2019): Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera. European Journal of Taxonomy 583: 1-45, DOI: 10.5852/ejt.2019.583
