identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038687B3FFCAFF9BFF7116843D392507.text	038687B3FFCAFF9BFF7116843D392507.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidiella maculosa Araújo & Bravo 2019	<div><p>Lepidiella maculosa, sp. nov.</p><p>(Figs. 1 A–F)</p><p>Type material. Holotype, male: BRAZIL, Bahia, [Elísio Medrado], Serra da Jibóia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.466667&amp;materialsCitation.latitude=-12.85" title="Search Plazi for locations around (long -39.466667/lat -12.85)">Sede</a> GAMBA [Environmentalist Group of Bahia] [12° 51' S; 39 28' W], (ligth trap), 10.v.2017, Silva-Neto, Mendes &amp; Moura cols. (MZFS), specimen dissected, mounted on micro-slide.</p><p>Etymology. The specific epithet, from Latin maculosus, spotted, refers to the marks on hypandrium.</p><p>Diagnosis. Head without cornicula; eyes separated by 2.0 facet diameters; scape without internal protuberance; interocular suture inverted V-shaped; hypandrium with microtrichia arranged in circular groups; aedeagus symmetrical, bifid, digitiform with a medial external lobe; two pairs of parameres, external pair fused medially forming a U-shaped plate; internal pair well sclerotized, lanceolate.</p><p>Description. Male holotype. Head without corniculum. Vertex dorsally expanded. Eyes separated by 2.0 facet diameters; eye bridge with four facet rows; interocular suture inverted V-shaped; frons with hair patch not divided at center, extending dorsally and reaching the interocular suture (Fig. 1A). Scape subspherical, approximately the same length as pedicel, without internal protuberances. Antenna incomplete; first three flagellomeres fusiform, each one with pair of filiform ascoids (Fig. 1B). Relative palpomere proportion: 1.0:1.1:1.2:2.0 (Fig. 1C). Wing (Fig. 1D). 2.5 times longer than wide; wing membrane brownish, with costal cell darker; lighter spots between veins R 5 /M 1 to CuA; Sc not reaching C; sc-r absent; radial fork distal to medial fork; R 4 ending at wing tip; CuA not reaching wing margin. Male terminalia: hypandrium plate-like, subtriangular with microtrichia arranged in circular groups (Figs. 1E, 1F); length of gonocoxite 0.3 times the length of gonostylus; gonostyli narrow, basally inflated, bare, except in the base with patch of alveoli; aedeagus symmetrical, bifid, distally digitiform; two pairs of parameres, external pair fused forming a U-shaped plate; internal pair ending at the same place of aedeagus, well sclerotized, lanciform, ending in pointed apex; ejaculatory apodeme dorsoventrally flattened (Fig. 1E); epandrium 2.5 times wider than long, with two foramina and two distal patches of alveoli (Fig. 1G); cercus digitiform, 1.2 the length of epandrium; left cercus with 12 tenacula (Fig. 1F) and right cercus with 11 tenacula; hypoproct and epiproct subtriangular with apical micropilosity (Fig. 1G).</p><p>Female: Unknown</p><p>Remarks. Males of five species of Lepidiella have a head without cornicula: the Brazilian species L. flabellata Bravo &amp; Santos, 2011, L. maculata sp. nov., L. olgae Bravo &amp; Araújo, 2013 and L. spinosa Bravo, 2005, and one Costa Rican species, L. hansoni (Quate, 1996) . The males of these five species can be recognized using the male identification key proposed below.</p></div>	https://treatment.plazi.org/id/038687B3FFCAFF9BFF7116843D392507	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Araújo, Maíra Xavier;Bravo, Freddy	Araújo, Maíra Xavier, Bravo, Freddy (2019): Two new species of Lepidiella Enderlein, 1937 (Diptera: Psychodidae) from the Neotropical Region with taxonomic comments about the species of the genus. Zootaxa 4551 (4): 487-493, DOI: 10.11646/zootaxa.4551.4.9
038687B3FFC8FF9AFF7113FF3D9E2758.text	038687B3FFC8FF9AFF7113FF3D9E2758.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidiella wagneri Araújo & Bravo 2019	<div><p>Lepidiella wagneri, sp. nov.</p><p>(Figs. 2 A–J)</p><p>Type material. Holotype, male: BRAZIL, Bahia, [Elísio Medrado], Serra da Jibóia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.466667&amp;materialsCitation.latitude=-12.85" title="Search Plazi for locations around (long -39.466667/lat -12.85)">Sede</a> GAMBA [Environmentalist Group of Bahia] [12° 51' S; 39 28' W], (ligth trap), 10.v.2017, Silva-Neto, Mendes e Moura cols. (MZFS), specimen dissected, mounted on micro-slide. 8 paratypes females: same location, data and collectors as holotype (MZFS), specimens dissected, mounted on micro-slide.</p><p>Etymology. The specific epithet is dedicated to Dr. Rüdiger Wagner for his important contribution to the study of the taxonomy of Psychodidae .</p><p>Diagnosis. Male. Head with pair of three branched cornicula and a glandular structure at base of it; eyes separated by 1.5 facet diameters; scape with internal protuberance; interocular suture inverted Y shaped reaching the middle of the vertex; hypandrium subtriangular with microtrichia spread along the sclerite; aedeagus symmetrical, bifid, digitiform; two pairs of parameres, external pair fused apically, inverted V-shaped; internal pair knife-like, ending in curved pointed apex. Female. Cornicula absent; scape without internal protuberance; terminalia with hypopygium trapezoidal; hypogynial valve setose with rounded external margin; cerci two times the length of subgenital plate.</p><p>Description. Male holotype. Head with large cornicula, three terminal branches and one oval gland in the base. Vertex dorsally expanded, 1.5 larger than wide. Eyes separated by 1.5 facet diameters; eye bridge with four facet rows; interocular suture inverted Y-shaped ending at the middle of the vertex (Figs. 2A, 2B). Antenna (Figs. 2A, 2D, 2E) with scape longer than the pedicel, inflated on the inner side. Flagellum with 14 flagellomeres; first flagellomere fusiform, 1.2 longer than second flagellomere; flagellomeres 2–13 fusiform with short internodes; flagellomere 14 with apiculus; pairs of filiform ascoids observed in flagellomeres 2 – 11 (Figs. 2A, 2D). Relative palpomere proportions: 1.0:1.2:1.6:2.1 (Fig. 2A). Wing 3.3 times longer than wide; wing membrane brownish, with costal cell darker; Sc not reaching C; sc-r absent; radial fork distal to medial fork; R 4 ending at wing tip; CuA not reaching wing margin (Fig. 2F). Male terminalia: hypandrium subtriangular, apical margin rounded with microtrichia spread along the sclerite; length of gonocoxite 0.6 times the length of gonostylus; gonostyli bare except in the base with patch of alveoli; 1/3 basal of gonostyli inflated; aedeagus symmetrical, bifid, digitiform; two pairs of parameres, external pair shorter than the internal, fused apically forming an U-inverted shaped plate; internal parameres, narrow, well sclerotized, curved apically, ending in pointed apex; ejaculatory apodeme dorsoventrally flattened (Fig. 2G); epandrium two times wider than long, with two foramina and two apical patches of alveoli; cercus digitiform 2.2 times the length of epandrium; left cercus with 10 tenacula; right cercus with 11 tenacula; hypoproct and epiproct subtriangular with apical micropilosity (Figs. 2H, 2I).</p><p>Female. Morphologically similar to male, except for the following characteristics: cornicula absent; scape with shorter internal protuberance than male (Fig. 1C); terminalia with hypogynium trapezoidal; hypogynial valves setose with rounded external margin; cerci two times the length of hypogynium (Fig. 1J).</p><p>Remarks. Males of eleven species of Lepidiella have cornicula and six of them a scape with an internal basal lobe: L. albipeda, L. cervi, L. monteverdica, L. pickeringi, L. robusta and L. wagneri sp. nov. This group of species is identified in the male identification key below.</p></div>	https://treatment.plazi.org/id/038687B3FFC8FF9AFF7113FF3D9E2758	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Araújo, Maíra Xavier;Bravo, Freddy	Araújo, Maíra Xavier, Bravo, Freddy (2019): Two new species of Lepidiella Enderlein, 1937 (Diptera: Psychodidae) from the Neotropical Region with taxonomic comments about the species of the genus. Zootaxa 4551 (4): 487-493, DOI: 10.11646/zootaxa.4551.4.9
038687B3FFCEFF9DFF7112C63C8D23A2.text	038687B3FFCEFF9DFF7112C63C8D23A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidiella Enderlein 1937	<div><p>Checklist of Lepidiella Enderlein, 1937</p><p>Lepidiella albipeda (Rapp, 1945) . Distribution: Panama (Barro Colorado Is.), Nicaragua (Matagalpa Department),</p><p>Mexico (Hidalgo, Veracruz) (Ibáñez-Bernal 2008 describes the female of the species from Mexico) Lepidiella amaliae Collantes &amp; Martínez-Ortega, 1997 . Distribution: Nicaragua Lepidiella cervi (Satchell, 1955) . Distribution: Caribbean (Sta. Lucia Is.) Lepidiella flabellata Bravo &amp; Santos, 2011 . Distribution: Brazil (Espirito Santo); female unknown Lepidiella hansoni (Quate, 1996) . Distribution: Costa Rica Lepidiella lanuginosa Enderlein, 1937 . Distribution: Bolívia (Yungas), Peru (Cuzco); redescribed by Quate (1963);</p><p>female unknown Lepidiella larryi Ibañez-Bernal, 2010 . Distribution: Colombia (Magdalena) (writing error by Ibáñez-Bernal) Lepidiella maculosa sp. nov. Distribution: Brazil (Bahia) Lepidiella matagalpensis (Collantes &amp; Martínez-Ortega, 1998) . Distribution: Nicaragua (Matagalpa Department) Lepidiella monteverdica (Quate, 1996) . Distribution: Costa Rica (Puentarenas) Lepidiella niveitarsis (Enderlein, 1937) . Distribution: Peru (Cuzco); redescribed by Quate (1963) Lepidiella olgae Bravo &amp; Araújo, 2013 . Distribution: Brazil (Paraíba) Lepidiella pickeringi (Quate, 1999) . Distribution: Panama (Barro Colorado Is.) Lepidiella robusta Bravo &amp; Santos, 2011 . Distribution: Brazil (Espirito Santo); female unknown Lepidiella spinosa Bravo, 2005 . Distribution: Brazil (São Paulo) Lepidiella wagneri sp. nov. Distribution: Brazil (Bahia) Lepidiella zumbadoi (Quate, 1996) . Distribution: Costa Rica (Cartago)</p></div>	https://treatment.plazi.org/id/038687B3FFCEFF9DFF7112C63C8D23A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Araújo, Maíra Xavier;Bravo, Freddy	Araújo, Maíra Xavier, Bravo, Freddy (2019): Two new species of Lepidiella Enderlein, 1937 (Diptera: Psychodidae) from the Neotropical Region with taxonomic comments about the species of the genus. Zootaxa 4551 (4): 487-493, DOI: 10.11646/zootaxa.4551.4.9
038687B3FFCEFF9CFF71158D3AB924A2.text	038687B3FFCEFF9CFF71158D3AB924A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidiella Enderlein 1937	<div><p>Key to males of Lepidiella Enderlein, 1937</p><p>1 Head without cornicula............................................................................... 2</p><p>- Head with multilobed cornicula......................................................................... 6</p><p>2 Internal paramere with bifurcated apex.................................. L. flabellata (Bravo &amp; Santos 2011, Fig. 6)</p><p>- Internal paramere not bifurcated, blade shaped............................................................. 3</p><p>3 Ejaculatory apodeme narrow in dorsal view........................................ L. spinosa (Bravo 2005, Fig. 5)</p><p>- Ejaculatory apodeme broad in dorsal view, wider than gonocoxite............................................... 4</p><p>4 Ejaculatory apodeme short, 0.7 times the length of gonocoxite.......................................... L. hansoni</p><p>- Ejaculatory apodeme long, 2 times the length of gonocoxite.................................................. 5</p><p>5 Margins of pedicel divergent distally larger than base; hypandrium subrectangular with microtrichia sparrow uniformly, not arranged in circular groups............................................. L. olgae (Bravo &amp; Araújo 2013, Fig. 2H)</p><p>- Margins of pedicel parallel; hypandrium wider in the middle portion with microtrichia arranged in circular groups............................................................................................ L. maculosa sp. nov.</p><p>6 Scape with a pronounced internal distal lobe............................................................... 7</p><p>- Scape cylindrical or with a short internal distal expansion................................................... 12</p><p>7 First flagellomere 3.0 X as long as the second....................................................... L. robusta - First flagellomere nearly the same length of second......................................................... 8</p><p>8 Scape 3.5 X as long as pedicel.............................................................. L. monteverdica</p><p>- Scape 2.0 – 2.5 times the length of pedicel................................................................ 9</p><p>9 Aedeagal apodeme narrow in dorsal view, narrower than the gonocoxite................ L. cervi (Satchell 1955, Fig. 5C)</p><p>- Aedeagal apodeme broad in dorsal view, with approximately the same width as the gonocoxite..................... 10</p><p>10 Eyes separated by 3.5 facet diameters............................................................. L. albipeda</p><p>- Eyes separated by 1.5 facet diameters................................................................... 11</p><p>11 Aedeagus digitiform; ejaculatory apodeme 3 times the length of gonocoxite; hypandrium triangular..... L. wagneri sp. nov.</p><p>- Aedeagus with ending in a dorsal spiraliform tip; ejaculatory apodeme 1.2 times the length of gonocoxite; hypandrium subrect- angular, narrow........................................................... L. pickeringi (Quate 1999, Fig. 2C)</p><p>12 Eyes separated by 1 facet diameters............................................................. L. niveitarsis</p><p>- Eyes separated by 2 or three facet diameters.............................................................. 13</p><p>13 Radial and medial forks at the same level; first flagellomere 2.0 X longer than second one................ L. lanuginosa</p><p>- Radial fork apical to medial fork; first flagellomere with the same length or 1.3 X longer than the second one.......... 14</p><p>14 Costal area convex, expanded........................... L. matagalpensis (Collantes &amp; Martínez-Ortega 1998, Fig. 4)</p><p>- Costal area narrow................................................................................... 15</p><p>15 Aedeagus and parameres asymmetrical........................................ L. zumbadoi (Quate 1996, Fig. 5D)</p><p>- Aedeagus and parameres symmetrical........................................ L. larryi (Ibañez-Bernal 2010, Fig. 6)</p></div>	https://treatment.plazi.org/id/038687B3FFCEFF9CFF71158D3AB924A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Araújo, Maíra Xavier;Bravo, Freddy	Araújo, Maíra Xavier, Bravo, Freddy (2019): Two new species of Lepidiella Enderlein, 1937 (Diptera: Psychodidae) from the Neotropical Region with taxonomic comments about the species of the genus. Zootaxa 4551 (4): 487-493, DOI: 10.11646/zootaxa.4551.4.9
