taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
038737309958FF87FF175239FB9BA387.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643244/files/figure.png	https://doi.org/10.5281/zenodo.15643244	Fig. 2. Rhyacoglanis annulatus, new species, ANSP 160625, holotype, 42.5 mm SL; río Orinoco, Amazonas, Venezuela.	Fig. 2. Rhyacoglanis annulatus, new species, ANSP 160625, holotype, 42.5 mm SL; río Orinoco, Amazonas, Venezuela.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
038737309958FF87FF175239FB9BA387.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643246/files/figure.png	https://doi.org/10.5281/zenodo.15643246	Fig. 3. Ossified portion of left pectoral-fin spine of a. Rhyacoglanis annulatus, holotype, ANSP 160625, 42.5 mm SL; b. R. epiblepsis, holotype, AMNH 266401, 53.3 mm SL; c. R. paranensis, holotype, MZUEL 14119, 61.9 mm SL; d. R. pulcher, syntype, BMNH 1880.12.8.105-107, 68.5 mm SL; e. R. seminiger, holotype, LIRP 12466. 74.2 mm SL. Scale bar = 2.0 mm.	Fig. 3. Ossified portion of left pectoral-fin spine of a. Rhyacoglanis annulatus, holotype, ANSP 160625, 42.5 mm SL; b. R. epiblepsis, holotype, AMNH 266401, 53.3 mm SL; c. R. paranensis, holotype, MZUEL 14119, 61.9 mm SL; d. R. pulcher, syntype, BMNH 1880.12.8.105-107, 68.5 mm SL; e. R. seminiger, holotype, LIRP 12466. 74.2 mm SL. Scale bar = 2.0 mm.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
038737309958FF87FF175239FB9BA387.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643250/files/figure.png	https://doi.org/10.5281/zenodo.15643250	Fig. 4. Map of central portions of South America showing distribution of Rhyacoglanis annulatus (squares; white square (1) = type locality); R. epiblepsis (white triangle (2) = type locality); R. paranensis (stars; white star (3) = type locality); R. pulcher (white circle (4) = type locality); R. cf. pulcher (diamond); and R. seminiger (pentagon; white pentagon (5) = type locality). Some symbols represent more than one locality and/or lot of specimens.	Fig. 4. Map of central portions of South America showing distribution of Rhyacoglanis annulatus (squares; white square (1) = type locality); R. epiblepsis (white triangle (2) = type locality); R. paranensis (stars; white star (3) = type locality); R. pulcher (white circle (4) = type locality); R. cf. pulcher (diamond); and R. seminiger (pentagon; white pentagon (5) = type locality). Some symbols represent more than one locality and/or lot of specimens.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
03873730995BFF8AFC4E55C6FCE6A387.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643254/files/figure.png	https://doi.org/10.5281/zenodo.15643254	Fig. 5. Rhyacoglanis epiblepsis, new species, AMNH 266401, holotype, 53.3 mm SL; río Itenez, Beni, Bolivia.	Fig. 5. Rhyacoglanis epiblepsis, new species, AMNH 266401, holotype, 53.3 mm SL; río Itenez, Beni, Bolivia.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
03873730995BFF8AFC4E55C6FCE6A387.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643258/files/figure.png	https://doi.org/10.5281/zenodo.15643258	Fig. 6. Rhyacoglanis epiblepsis, paratype, AMNH 40127, 52.0 mm SL; río Itenez, Beni, Bolivia; showing alternative coloration pattern.	Fig. 6. Rhyacoglanis epiblepsis, paratype, AMNH 40127, 52.0 mm SL; río Itenez, Beni, Bolivia; showing alternative coloration pattern.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
03873730995BFF8AFC4E55C6FCE6A387.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643246/files/figure.png	https://doi.org/10.5281/zenodo.15643246	Fig. 3. Ossified portion of left pectoral-fin spine of a. Rhyacoglanis annulatus, holotype, ANSP 160625, 42.5 mm SL; b. R. epiblepsis, holotype, AMNH 266401, 53.3 mm SL; c. R. paranensis, holotype, MZUEL 14119, 61.9 mm SL; d. R. pulcher, syntype, BMNH 1880.12.8.105-107, 68.5 mm SL; e. R. seminiger, holotype, LIRP 12466. 74.2 mm SL. Scale bar = 2.0 mm.	Fig. 3. Ossified portion of left pectoral-fin spine of a. Rhyacoglanis annulatus, holotype, ANSP 160625, 42.5 mm SL; b. R. epiblepsis, holotype, AMNH 266401, 53.3 mm SL; c. R. paranensis, holotype, MZUEL 14119, 61.9 mm SL; d. R. pulcher, syntype, BMNH 1880.12.8.105-107, 68.5 mm SL; e. R. seminiger, holotype, LIRP 12466. 74.2 mm SL. Scale bar = 2.0 mm.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
03873730995BFF8AFC4E55C6FCE6A387.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643250/files/figure.png	https://doi.org/10.5281/zenodo.15643250	Fig. 4. Map of central portions of South America showing distribution of Rhyacoglanis annulatus (squares; white square (1) = type locality); R. epiblepsis (white triangle (2) = type locality); R. paranensis (stars; white star (3) = type locality); R. pulcher (white circle (4) = type locality); R. cf. pulcher (diamond); and R. seminiger (pentagon; white pentagon (5) = type locality). Some symbols represent more than one locality and/or lot of specimens.	Fig. 4. Map of central portions of South America showing distribution of Rhyacoglanis annulatus (squares; white square (1) = type locality); R. epiblepsis (white triangle (2) = type locality); R. paranensis (stars; white star (3) = type locality); R. pulcher (white circle (4) = type locality); R. cf. pulcher (diamond); and R. seminiger (pentagon; white pentagon (5) = type locality). Some symbols represent more than one locality and/or lot of specimens.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
038737309956FF8DFC4F55C6FBF3A448.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643260/files/figure.png	https://doi.org/10.5281/zenodo.15643260	Fig. 7. Rhyacoglanis paranensis, new species, MZUEL 14119, holotype, 61.9 mm SL, rio Piracicaba, São Paulo, Brazil.	Fig. 7. Rhyacoglanis paranensis, new species, MZUEL 14119, holotype, 61.9 mm SL, rio Piracicaba, São Paulo, Brazil.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
038737309956FF8DFC4F55C6FBF3A448.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643264/files/figure.png	https://doi.org/10.5281/zenodo.15643264	Fig. 8. Live individual of Rhyacoglanis paranensis from Brazil, São Paulo, rio Paranapanema (not retained). Photo by Rony Suzuki.	Fig. 8. Live individual of Rhyacoglanis paranensis from Brazil, São Paulo, rio Paranapanema (not retained). Photo by Rony Suzuki.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
038737309956FF8DFC4F55C6FBF3A448.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643246/files/figure.png	https://doi.org/10.5281/zenodo.15643246	Fig. 3. Ossified portion of left pectoral-fin spine of a. Rhyacoglanis annulatus, holotype, ANSP 160625, 42.5 mm SL; b. R. epiblepsis, holotype, AMNH 266401, 53.3 mm SL; c. R. paranensis, holotype, MZUEL 14119, 61.9 mm SL; d. R. pulcher, syntype, BMNH 1880.12.8.105-107, 68.5 mm SL; e. R. seminiger, holotype, LIRP 12466. 74.2 mm SL. Scale bar = 2.0 mm.	Fig. 3. Ossified portion of left pectoral-fin spine of a. Rhyacoglanis annulatus, holotype, ANSP 160625, 42.5 mm SL; b. R. epiblepsis, holotype, AMNH 266401, 53.3 mm SL; c. R. paranensis, holotype, MZUEL 14119, 61.9 mm SL; d. R. pulcher, syntype, BMNH 1880.12.8.105-107, 68.5 mm SL; e. R. seminiger, holotype, LIRP 12466. 74.2 mm SL. Scale bar = 2.0 mm.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
038737309956FF8DFC4F55C6FBF3A448.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643250/files/figure.png	https://doi.org/10.5281/zenodo.15643250	Fig. 4. Map of central portions of South America showing distribution of Rhyacoglanis annulatus (squares; white square (1) = type locality); R. epiblepsis (white triangle (2) = type locality); R. paranensis (stars; white star (3) = type locality); R. pulcher (white circle (4) = type locality); R. cf. pulcher (diamond); and R. seminiger (pentagon; white pentagon (5) = type locality). Some symbols represent more than one locality and/or lot of specimens.	Fig. 4. Map of central portions of South America showing distribution of Rhyacoglanis annulatus (squares; white square (1) = type locality); R. epiblepsis (white triangle (2) = type locality); R. paranensis (stars; white star (3) = type locality); R. pulcher (white circle (4) = type locality); R. cf. pulcher (diamond); and R. seminiger (pentagon; white pentagon (5) = type locality). Some symbols represent more than one locality and/or lot of specimens.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
038737309951FF90FC815299FC3FA5C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643266/files/figure.png	https://doi.org/10.5281/zenodo.15643266	Fig. 9. Rhyacoglanis pulcher, BMNH 1880.12.8.105-107, syntype, 68.5 mm SL; Canelos, Ecuador.	Fig. 9. Rhyacoglanis pulcher, BMNH 1880.12.8.105-107, syntype, 68.5 mm SL; Canelos, Ecuador.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
038737309951FF90FC815299FC3FA5C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643270/files/figure.png	https://doi.org/10.5281/zenodo.15643270	Fig. 10. Original illustration of syntype of Rhyacoglanis pulcher showing entire specimen in lateral view and anterior portion in dorsal view (modified from Boulenger, 1887).	Fig. 10. Original illustration of syntype of Rhyacoglanis pulcher showing entire specimen in lateral view and anterior portion in dorsal view (modified from Boulenger, 1887).	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
038737309951FF90FC815299FC3FA5C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643246/files/figure.png	https://doi.org/10.5281/zenodo.15643246	Fig. 3. Ossified portion of left pectoral-fin spine of a. Rhyacoglanis annulatus, holotype, ANSP 160625, 42.5 mm SL; b. R. epiblepsis, holotype, AMNH 266401, 53.3 mm SL; c. R. paranensis, holotype, MZUEL 14119, 61.9 mm SL; d. R. pulcher, syntype, BMNH 1880.12.8.105-107, 68.5 mm SL; e. R. seminiger, holotype, LIRP 12466. 74.2 mm SL. Scale bar = 2.0 mm.	Fig. 3. Ossified portion of left pectoral-fin spine of a. Rhyacoglanis annulatus, holotype, ANSP 160625, 42.5 mm SL; b. R. epiblepsis, holotype, AMNH 266401, 53.3 mm SL; c. R. paranensis, holotype, MZUEL 14119, 61.9 mm SL; d. R. pulcher, syntype, BMNH 1880.12.8.105-107, 68.5 mm SL; e. R. seminiger, holotype, LIRP 12466. 74.2 mm SL. Scale bar = 2.0 mm.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
038737309951FF90FC815299FC3FA5C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643250/files/figure.png	https://doi.org/10.5281/zenodo.15643250	Fig. 4. Map of central portions of South America showing distribution of Rhyacoglanis annulatus (squares; white square (1) = type locality); R. epiblepsis (white triangle (2) = type locality); R. paranensis (stars; white star (3) = type locality); R. pulcher (white circle (4) = type locality); R. cf. pulcher (diamond); and R. seminiger (pentagon; white pentagon (5) = type locality). Some symbols represent more than one locality and/or lot of specimens.	Fig. 4. Map of central portions of South America showing distribution of Rhyacoglanis annulatus (squares; white square (1) = type locality); R. epiblepsis (white triangle (2) = type locality); R. paranensis (stars; white star (3) = type locality); R. pulcher (white circle (4) = type locality); R. cf. pulcher (diamond); and R. seminiger (pentagon; white pentagon (5) = type locality). Some symbols represent more than one locality and/or lot of specimens.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
03873730994CFF95FC495207FB53A306.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643272/files/figure.png	https://doi.org/10.5281/zenodo.15643272	Fig. 11. Rhyacoglanis seminiger, new species, LIRP 12466, holotype, 74.2 mm SL; rio Juruena, Mato Grosso, Brazil.	Fig. 11. Rhyacoglanis seminiger, new species, LIRP 12466, holotype, 74.2 mm SL; rio Juruena, Mato Grosso, Brazil.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
03873730994CFF95FC495207FB53A306.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643278/files/figure.png	https://doi.org/10.5281/zenodo.15643278	Fig. 12. Color variation in Rhyacoglanis seminiger, MZUSP 82085, paratypes; a. 70.4 mm SL; b. 48.6 mm SL; c. 44.4 mm SL; rio Sangue, tributary of rio Juruena, Mato Grosso, Brazil.	Fig. 12. Color variation in Rhyacoglanis seminiger, MZUSP 82085, paratypes; a. 70.4 mm SL; b. 48.6 mm SL; c. 44.4 mm SL; rio Sangue, tributary of rio Juruena, Mato Grosso, Brazil.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
03873730994CFF95FC495207FB53A306.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643246/files/figure.png	https://doi.org/10.5281/zenodo.15643246	Fig. 3. Ossified portion of left pectoral-fin spine of a. Rhyacoglanis annulatus, holotype, ANSP 160625, 42.5 mm SL; b. R. epiblepsis, holotype, AMNH 266401, 53.3 mm SL; c. R. paranensis, holotype, MZUEL 14119, 61.9 mm SL; d. R. pulcher, syntype, BMNH 1880.12.8.105-107, 68.5 mm SL; e. R. seminiger, holotype, LIRP 12466. 74.2 mm SL. Scale bar = 2.0 mm.	Fig. 3. Ossified portion of left pectoral-fin spine of a. Rhyacoglanis annulatus, holotype, ANSP 160625, 42.5 mm SL; b. R. epiblepsis, holotype, AMNH 266401, 53.3 mm SL; c. R. paranensis, holotype, MZUEL 14119, 61.9 mm SL; d. R. pulcher, syntype, BMNH 1880.12.8.105-107, 68.5 mm SL; e. R. seminiger, holotype, LIRP 12466. 74.2 mm SL. Scale bar = 2.0 mm.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
03873730994CFF95FC495207FB53A306.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643250/files/figure.png	https://doi.org/10.5281/zenodo.15643250	Fig. 4. Map of central portions of South America showing distribution of Rhyacoglanis annulatus (squares; white square (1) = type locality); R. epiblepsis (white triangle (2) = type locality); R. paranensis (stars; white star (3) = type locality); R. pulcher (white circle (4) = type locality); R. cf. pulcher (diamond); and R. seminiger (pentagon; white pentagon (5) = type locality). Some symbols represent more than one locality and/or lot of specimens.	Fig. 4. Map of central portions of South America showing distribution of Rhyacoglanis annulatus (squares; white square (1) = type locality); R. epiblepsis (white triangle (2) = type locality); R. paranensis (stars; white star (3) = type locality); R. pulcher (white circle (4) = type locality); R. cf. pulcher (diamond); and R. seminiger (pentagon; white pentagon (5) = type locality). Some symbols represent more than one locality and/or lot of specimens.	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
03873730994CFF95FC495207FB53A306.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643280/files/figure.png	https://doi.org/10.5281/zenodo.15643280	Fig. 13. Scatterplot of the second and third principal components calculated from combined samples of Rhyacoglanis annulatus (purple circle: n = 2), R. epiblepsis (green x: n = 25), R. paranensis (pink squares; n = 22), R. pulcher (red +; n = 3) and R. seminiger (open blue squares; n = 9).	Fig. 13. Scatterplot of the second and third principal components calculated from combined samples of Rhyacoglanis annulatus (purple circle: n = 2), R. epiblepsis (green x: n = 25), R. paranensis (pink squares; n = 22), R. pulcher (red +; n = 3) and R. seminiger (open blue squares; n = 9).	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
03873730994CFF95FC495207FB53A306.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15643282/files/figure.png	https://doi.org/10.5281/zenodo.15643282	Fig. 14. a. Unculiferous tubercle of Rhyacoglanis seminiger (u = unculus); b. detail of the same tubercle near the apex showing unculus with several rounded projections (rp).	Fig. 14. a. Unculiferous tubercle of Rhyacoglanis seminiger (u = unculus); b. detail of the same tubercle near the apex showing unculus with several rounded projections (rp).	2017-06-12	Shibatta, Oscar Akio;Vari, Richard P.		Zenodo	biologists	Shibatta, Oscar Akio;Vari, Richard P.			
