taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
038747324C7CC65D1EB72CBFFA5F22B2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C7CC65A1EB72934FCB3240A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264604/files/figure.png	https://doi.org/10.5281/zenodo.264604	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C7CC65A1EB72934FCB3240A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264599/files/figure.png	https://doi.org/10.5281/zenodo.264599	FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades.	FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C7CC65A1EB72934FCB3240A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264602/files/figure.png	https://doi.org/10.5281/zenodo.264602	FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm.	FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C7CC65A1EB72934FCB3240A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C7BC6591EB728FCFDFA2509.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264599/files/figure.png	https://doi.org/10.5281/zenodo.264599	FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades.	FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C7BC6591EB728FCFDFA2509.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C78C6471EB729E2FC722384.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264600/files/figure.png	https://doi.org/10.5281/zenodo.264600	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C66C6431EB72E14FCDB240A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264601/files/figure.png	https://doi.org/10.5281/zenodo.264601	FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale).	FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C66C6431EB72E14FCDB240A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264602/files/figure.png	https://doi.org/10.5281/zenodo.264602	FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm.	FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C66C6431EB72E14FCDB240A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264608/files/figure.png	https://doi.org/10.5281/zenodo.264608	FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL).	FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C66C6431EB72E14FCDB240A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264609/files/figure.png	https://doi.org/10.5281/zenodo.264609	FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai.	FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C66C6431EB72E14FCDB240A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C62C6401EB728FCFB6F2681.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C62C6401EB728FCFB6F2681.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264605/files/figure.png	https://doi.org/10.5281/zenodo.264605	FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm.	FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C62C6401EB728FCFB6F2681.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264606/files/figure.png	https://doi.org/10.5281/zenodo.264606	FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm.	FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C62C6401EB728FCFB6F2681.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264603/files/figure.png	https://doi.org/10.5281/zenodo.264603	FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9.	FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C62C6401EB728FCFB6F2681.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264604/files/figure.png	https://doi.org/10.5281/zenodo.264604	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C6DC6481EB72DBAFBA8218A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264605/files/figure.png	https://doi.org/10.5281/zenodo.264605	FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm.	FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C6DC6481EB72DBAFBA8218A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264604/files/figure.png	https://doi.org/10.5281/zenodo.264604	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C6DC6481EB72DBAFBA8218A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C69C6751EB72C61FB15228C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264606/files/figure.png	https://doi.org/10.5281/zenodo.264606	FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm.	FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C69C6751EB72C61FB15228C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264605/files/figure.png	https://doi.org/10.5281/zenodo.264605	FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm.	FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C69C6751EB72C61FB15228C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C54C6711EB72966FF2221A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C54C6711EB72966FF2221A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264607/files/figure.png	https://doi.org/10.5281/zenodo.264607	FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm.	FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C54C6711EB72966FF2221A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264604/files/figure.png	https://doi.org/10.5281/zenodo.264604	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C54C6711EB72966FF2221A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264608/files/figure.png	https://doi.org/10.5281/zenodo.264608	FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL).	FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C54C6711EB72966FF2221A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264603/files/figure.png	https://doi.org/10.5281/zenodo.264603	FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9.	FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C54C6711EB72966FF2221A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C50C67F1EB72C0FFB4522CC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264607/files/figure.png	https://doi.org/10.5281/zenodo.264607	FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm.	FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C50C67F1EB72C0FFB4522CC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264608/files/figure.png	https://doi.org/10.5281/zenodo.264608	FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL).	FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C50C67F1EB72C0FFB4522CC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264604/files/figure.png	https://doi.org/10.5281/zenodo.264604	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C50C67F1EB72C0FFB4522CC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C50C67F1EB72C0FFB4522CC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5EC67C1EB7292BFD3E27E9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264606/files/figure.png	https://doi.org/10.5281/zenodo.264606	FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm.	FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5EC67C1EB7292BFD3E27E9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264605/files/figure.png	https://doi.org/10.5281/zenodo.264605	FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm.	FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5EC67C1EB7292BFD3E27E9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264604/files/figure.png	https://doi.org/10.5281/zenodo.264604	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5EC67C1EB7292BFD3E27E9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5EC67C1EB7292BFD3E27E9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5AC67A1EB729EDFCB32348.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5AC67A1EB729EDFCB32348.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264609/files/figure.png	https://doi.org/10.5281/zenodo.264609	FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai.	FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5AC67A1EB729EDFCB32348.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264600/files/figure.png	https://doi.org/10.5281/zenodo.264600	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5BC6651EB72FAAFA9326FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264610/files/figure.png	https://doi.org/10.5281/zenodo.264610	FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN).	FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5BC6651EB72FAAFA9326FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264612/files/figure.png	https://doi.org/10.5281/zenodo.264612	FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm.	FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5BC6651EB72FAAFA9326FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264604/files/figure.png	https://doi.org/10.5281/zenodo.264604	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5BC6651EB72FAAFA9326FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264611/files/figure.png	https://doi.org/10.5281/zenodo.264611	FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm.	FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C5BC6651EB72FAAFA9326FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C45C6611EB72DBAFED12521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264610/files/figure.png	https://doi.org/10.5281/zenodo.264610	FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN).	FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C45C6611EB72DBAFED12521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264609/files/figure.png	https://doi.org/10.5281/zenodo.264609	FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai.	FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C45C6611EB72DBAFED12521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264613/files/figure.png	https://doi.org/10.5281/zenodo.264613	FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014).	FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C45C6611EB72DBAFED12521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264604/files/figure.png	https://doi.org/10.5281/zenodo.264604	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C45C6611EB72DBAFED12521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264612/files/figure.png	https://doi.org/10.5281/zenodo.264612	FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm.	FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C45C6611EB72DBAFED12521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264599/files/figure.png	https://doi.org/10.5281/zenodo.264599	FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades.	FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C45C6611EB72DBAFED12521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264601/files/figure.png	https://doi.org/10.5281/zenodo.264601	FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale).	FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C45C6611EB72DBAFED12521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C40C66E1EB72985FAE523DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264613/files/figure.png	https://doi.org/10.5281/zenodo.264613	FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014).	FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C40C66E1EB72985FAE523DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264612/files/figure.png	https://doi.org/10.5281/zenodo.264612	FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm.	FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C40C66E1EB72985FAE523DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264600/files/figure.png	https://doi.org/10.5281/zenodo.264600	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C40C66E1EB72985FAE523DC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C4FC66E1EB72FD1FAB4267D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C4FC6691EB72A7CFAA72543.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264614/files/figure.png	https://doi.org/10.5281/zenodo.264614	FIGURE 17. Adults in DV (left), VV (right). A: HT ♀ of Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov. (NHMUK 010289143; Anjanaharibe Sud). B: PT ♀ of Ht. tornado (Marojejy; KAP-MAD 1403). C: ♂ of Ht. tornado (‘ Mahassabe’) [from colour transparency]. D: ♀ of Ht. tornado (‘ Mahassabe’; BMNH (E) # 674881). Scale bar: 30 mm.	FIGURE 17. Adults in DV (left), VV (right). A: HT ♀ of Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov. (NHMUK 010289143; Anjanaharibe Sud). B: PT ♀ of Ht. tornado (Marojejy; KAP-MAD 1403). C: ♂ of Ht. tornado (‘ Mahassabe’) [from colour transparency]. D: ♀ of Ht. tornado (‘ Mahassabe’; BMNH (E) # 674881). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C4FC6691EB72A7CFAA72543.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C4FC6691EB72A7CFAA72543.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C4FC6691EB72A7CFAA72543.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264616/files/figure.png	https://doi.org/10.5281/zenodo.264616	FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm.	FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C4FC6691EB72A7CFAA72543.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264615/files/figure.png	https://doi.org/10.5281/zenodo.264615	FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm.	FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C4FC6691EB72A7CFAA72543.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264604/files/figure.png	https://doi.org/10.5281/zenodo.264604	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C48C6691EB729A7FABD24EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C48C6161EB728DCFCB92691.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264616/files/figure.png	https://doi.org/10.5281/zenodo.264616	FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm.	FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C48C6161EB728DCFCB92691.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264615/files/figure.png	https://doi.org/10.5281/zenodo.264615	FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm.	FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C48C6161EB728DCFCB92691.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C34C6111EB72C8FFDC9260E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264617/files/figure.png	https://doi.org/10.5281/zenodo.264617	FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees).	FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C34C6111EB72C8FFDC9260E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264618/files/figure.png	https://doi.org/10.5281/zenodo.264618	FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm.	FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C34C6111EB72C8FFDC9260E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264611/files/figure.png	https://doi.org/10.5281/zenodo.264611	FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm.	FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C34C6111EB72C8FFDC9260E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264615/files/figure.png	https://doi.org/10.5281/zenodo.264615	FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm.	FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C34C6111EB72C8FFDC9260E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C31C61E1EB72DBAFDD727ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264618/files/figure.png	https://doi.org/10.5281/zenodo.264618	FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm.	FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C31C61E1EB72DBAFDD727ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264615/files/figure.png	https://doi.org/10.5281/zenodo.264615	FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm.	FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C31C61E1EB72DBAFDD727ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264600/files/figure.png	https://doi.org/10.5281/zenodo.264600	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C31C61E1EB72DBAFDD727ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C3FC61A1EB72BCFFAD227D5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264619/files/figure.png	https://doi.org/10.5281/zenodo.264619	FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014).	FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C3FC61A1EB72BCFFAD227D5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264620/files/figure.png	https://doi.org/10.5281/zenodo.264620	FIGURE 23. Adults in views stated. A: ♂ PT genitalia of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; 33 DL), LV, SV. Scale bar: 3 mm. B: ♂ genitalia and sternal androconial scaling in Ht. roussettae (“ sp. 26 ” modified from Lees, 1997; not to scale), LV. C: ♂ genitalia and sternal androconial scaling in Ht. andasibe (modified from Lees, 1997: where labelled “ antsianakana ”; not to scale), LV.	FIGURE 23. Adults in views stated. A: ♂ PT genitalia of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; 33 DL), LV, SV. Scale bar: 3 mm. B: ♂ genitalia and sternal androconial scaling in Ht. roussettae (“ sp. 26 ” modified from Lees, 1997; not to scale), LV. C: ♂ genitalia and sternal androconial scaling in Ht. andasibe (modified from Lees, 1997: where labelled “ antsianakana ”; not to scale), LV.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C3FC61A1EB72BCFFAD227D5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264602/files/figure.png	https://doi.org/10.5281/zenodo.264602	FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm.	FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C3FC61A1EB72BCFFAD227D5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264600/files/figure.png	https://doi.org/10.5281/zenodo.264600	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C3FC61A1EB72BCFFAD227D5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264604/files/figure.png	https://doi.org/10.5281/zenodo.264604	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C3FC61A1EB72BCFFAD227D5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C3BC61A1EB72BC9FD9D2680.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C38C6061EB72DBAFE8226A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264621/files/figure.png	https://doi.org/10.5281/zenodo.264621	FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm.	FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C38C6061EB72DBAFE8226A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264622/files/figure.png	https://doi.org/10.5281/zenodo.264622	FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm.	FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C38C6061EB72DBAFE8226A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264623/files/figure.png	https://doi.org/10.5281/zenodo.264623	FIGURE 26. Live ♀ adults in natural posture. A: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 5 / 11 / 2006). B: Ht. andasibe (Lees, 2003) (Andasibe; 24 / 10 / 2014). Photos: D. C. Lees.	FIGURE 26. Live ♀ adults in natural posture. A: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 5 / 11 / 2006). B: Ht. andasibe (Lees, 2003) (Andasibe; 24 / 10 / 2014). Photos: D. C. Lees.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C38C6061EB72DBAFE8226A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264619/files/figure.png	https://doi.org/10.5281/zenodo.264619	FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014).	FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C38C6061EB72DBAFE8226A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264600/files/figure.png	https://doi.org/10.5281/zenodo.264600	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C38C6061EB72DBAFE8226A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C24C6021EB72BF9FBA92005.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264598/files/figure.png	https://doi.org/10.5281/zenodo.264598	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C24C6021EB72BF9FBA92005.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264621/files/figure.png	https://doi.org/10.5281/zenodo.264621	FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm.	FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C24C6021EB72BF9FBA92005.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264619/files/figure.png	https://doi.org/10.5281/zenodo.264619	FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014).	FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C24C6021EB72BF9FBA92005.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264616/files/figure.png	https://doi.org/10.5281/zenodo.264616	FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm.	FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C24C6021EB72BF9FBA92005.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264622/files/figure.png	https://doi.org/10.5281/zenodo.264622	FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm.	FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C24C6021EB72BF9FBA92005.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C23C6001EB72C94FE4726AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264624/files/figure.png	https://doi.org/10.5281/zenodo.264624	FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm.	FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C23C6001EB72C94FE4726AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C2EC60C1EB72DBAFC8B2602.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264625/files/figure.png	https://doi.org/10.5281/zenodo.264625	FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm.	FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C2EC60C1EB72DBAFC8B2602.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264626/files/figure.png	https://doi.org/10.5281/zenodo.264626	FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm.	FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C2EC60C1EB72DBAFC8B2602.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264600/files/figure.png	https://doi.org/10.5281/zenodo.264600	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV).	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
038747324C2EC60C1EB72DBAFC8B2602.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/264627/files/figure.png	https://doi.org/10.5281/zenodo.264627	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km.	2016-12-31	C, Lees David		Zenodo	biologists	C, Lees David			
