taxonID	type	description	language	source
038587F3FFE488695487FBEFFB3EF979.taxon	diagnosis	Diagnosis description Body minute and oval, distinctly convex; dorsum pubescent (Figure 2). Head transverse; small; clypeal and frontal regions usually prominent, sometimes rostrate (Figure 1 A). Eyes small and convex, coarsely faceted. Antenna 10 - segmented, with a distinct two-segmented club (Figure 1 C). Terminal palpomere elongate-conical (Figure 1 D). Pronotum always with distinct line or ridge separating anterior corners from pronotal disc, sometimes extending along lateral edge. Elytra with lateral margin slightly flared or with distinct flat ridge. Elytral punctures at least along suture in apparent rows; epipleuron narrow, incomplete apically, without cavities. Prosternum strongly reduced and narrow, prosternal process reduced to a narrow carina (Figure 1 E). Mesoventrite small and transverse between middle coxae; about three times as wide as long. Metaventrite broad and elevated, always strongly and densely punctured. Abdomen with six visible ventrites. Abdominal postcoxal line incomplete or reaching lateral margin, divided into two lines, usually with associated pits and pores (Figure 1 F). Tarsi four-segmented (Figures 1 G, H). Male genitalia asymmetrical; parameres reduced or absent (Figure 3 B – E). Female genitalia: ovipositor triangular elongate, lightly sclerotized bearing short styli; spermatheca composed of two or more than two round areas (Figure 1 I). Distribution China, Oriental region, Africa, Australia, Central America, Madagascar.	en	Wang, Xingmin, Ren, Shunxiang (2012): A review of Chinese Scymnomorphus Weise (Coleoptera: Coccinellidae) with description of five new species. Journal of Natural History 46 (31 - 32): 1905-1920, DOI: 10.1080/00222933.2012.693958, URL: http://dx.doi.org/10.1080/00222933.2012.693958
038587F3FFEF886454A1FF70FB8DFF24.taxon	description	(Figures 1 A – I, 2 A, 3 A – E, 6) Diagnosis This species is similar to S. bicolor in male genitalia, but it can be distinguished from the latter as follows: body uniformly brown, without other external spots (Figure 2 A), penis guide in lateral view slender, flattened and quadrate, with apex truncated (Figure 3 D), penis guide in ventral view elongate (Figure 3 E). In S. bicolor, head and pronotum are dark brown and elytra black, penis guide in lateral view is elongate with apex pointed, and penis guide in ventral view is stout. Description TL: 1.38 – 1.46 mm, TW: 1.04 – 1.09 mm, TH: 0.60 – 0.62 mm, TL / TW: 1.32 – 1.33; PL / PW: 0.58 – 0.61; EL / EW: 1.10 – 1.11. Body uniformly brown, without other external spots (Figure 2 A). Head yellowish brown. Underside brown, except mesoventrite and metaventrite dark brown. Legs yellowish brown. Body minute, short oval, distinctly convex; dorsum pubescent. Head small, 0.45 times elytral width (HW / EW = 1: 2.21); punctures on frons fine, separated by 0.8 – 3.0 times their diameter, with short setae in punctures; eyes small and coarsely faceted, widest interocular distance 0.63 times width of head. Pronotum 0.74 times elytral width (PW / EW = 1: 1.35), pronotal punctures fine, slightly smaller than those on head, separated by 1.5 – 5.0 times their diameter. Punctures on elytra larger than those on head, separated by 0.5 – 1.0 times their diameter. Surface of proventrite and mesoventrite shagreened. Punctures on metaventrite larger in middle, separated by 0.2 – 0.5 times their diameter, with short setae in punctures; becoming smaller towards lateral parts. Abdominal postcoxal line incomplete or reaching lateral margin smoothly, divided into two lines, with associated pits and pores (Figure 3 A). Male genitalia. Penis simple and slender, curved and narrowing toward apex; penis capsule with a short and stout outer process and a small inner one (Figure 3 B, C); penis guide in lateral view long, flattened and quadrate, six times as long as wide, with apex truncated; parameres very short and indistinct, with rather long setae (Figure 3 D); penis guide in ventral view elongate, widest near base and strongly narrowed at base, apex pointed and strongly bent (Figure 3 E). Type material Holotype: 1 ♂, China, Yunnan: Dajianshan, Pingbian, 2100 m, 20 April 2008, Wang XM leg. Paratypes: China, Yunnan:, 1 ♂ 1 ♀, same data as holotype; 4 ♂♂, Dajianshan, Pingbian, 2100 m, 22 May 2009, Wang XM leg; 3 ♂♂, Gaoligongshan, Baoshan, 19 September 2006, Wang XM leg. Distribution China (Yunnan). Etymology The specific epithet refers to the flattened and quadrate penis guide in lateral view.	en	Wang, Xingmin, Ren, Shunxiang (2012): A review of Chinese Scymnomorphus Weise (Coleoptera: Coccinellidae) with description of five new species. Journal of Natural History 46 (31 - 32): 1905-1920, DOI: 10.1080/00222933.2012.693958, URL: http://dx.doi.org/10.1080/00222933.2012.693958
038587F3FFEE886454C5FEC9FE55FB4F.taxon	description	(Figures 2 B, 3 F – J, 6)	en	Wang, Xingmin, Ren, Shunxiang (2012): A review of Chinese Scymnomorphus Weise (Coleoptera: Coccinellidae) with description of five new species. Journal of Natural History 46 (31 - 32): 1905-1920, DOI: 10.1080/00222933.2012.693958, URL: http://dx.doi.org/10.1080/00222933.2012.693958
038587F3FFEE886454C5FEC9FE55FB4F.taxon	diagnosis	Diagnosis This species can be distinguished from other Scymnomorphus by the extremely small body (Figure 2 B), and the unique male genitalia (Figure 3 F – J). Description For detailed description see Kamiya (1960: 24). Specimens examined China. Sichuan. 2 ♂♂ 1 ♀, Wangjiang Park, Chengdu, 3 November 1983, Pang XF leg. Distribution China (Sichuan; Taiwan), Japan.	en	Wang, Xingmin, Ren, Shunxiang (2012): A review of Chinese Scymnomorphus Weise (Coleoptera: Coccinellidae) with description of five new species. Journal of Natural History 46 (31 - 32): 1905-1920, DOI: 10.1080/00222933.2012.693958, URL: http://dx.doi.org/10.1080/00222933.2012.693958
038587F3FFEE886454C2FB14FEEDF949.taxon	description	China. Taiwan. 1 ♀, Kuraru, Hengchun Park, Taiwan, 250 m, 2 April 1965, Yoshimoto CM Leg. Distribution China (Taiwan).	en	Wang, Xingmin, Ren, Shunxiang (2012): A review of Chinese Scymnomorphus Weise (Coleoptera: Coccinellidae) with description of five new species. Journal of Natural History 46 (31 - 32): 1905-1920, DOI: 10.1080/00222933.2012.693958, URL: http://dx.doi.org/10.1080/00222933.2012.693958
038587F3FFED8867549EFF70FC2AF959.taxon	description	(Figures 2 C, 4 A – E, 6) Diagnosis This species is similar to S. japonicus in general appearance, but it can be distinguished from the latter by the elytron with large punctures and the unique male genitalia (Figures 2 C, 4 A – E). Description TL: 1.04 mm, TW: 0.76 mm, TH: 0.47 mm, TL / TW: 1.38; PL / PW: 0.56; EL / EW: 1.09. Body yellowish brown, without other spots (Figure 2 C). Head yellowish brown. Underside yellowish brown, except mesoventrite and metaventrite reddish brown. Legs yellow. Body very small, short oval, distinctly convex; dorsum pubescent. Head small, 0.52 times elytral width (HW / EW = 1: 1.93); frons smooth, convex, with fine punctures, separated by 1.0 – 3.0 times their diameter, with short sparse hairs on the apical half; eyes small and coarsely faceted, widest interocular distance 0.60 times width of head. Pronotum 0.74 times elytral width (PW / EW = 1: 1.35), pronotal punctures fine, similar to those on head, separated by 1.0 – 4.0 times their diameter. Punctures on elytra large, separated by 0.2 – 0.5 times their diameter. Surface of proventrite and mesoventrite slightly shagreened. Punctures on metaventrite moderately large, separated by 0.2 – 0.5 times their diameter, with short and sparse setae in punctures. Abdominal postcoxal line incomplete or reaching lateral margin smoothly, divided into two lines, with associated indistinctly pits and pores (Figure 4 A). Male genitalia. Penis simple and slender, curved and narrowing toward apex; penis capsule indistinct (Figure 4 B, C); penis guide in lateral view short and stout, nearly quadrate, apex slightly bent; parameres short and indistinct, with rather long setae (Figure 4 D); penis guide in ventral view short, widest at base, narrowing apically, apex truncated; parameres in ventral view flattened and quadrate (Figure 4 E). Type material Holotype: 1 ♂, China, Tibet: Yarang Village, Motuo, 1030 m, 24 October 2007, Wang XM leg. Paratypes: 1 ♂, same data as holotype. Distribution China (Tibet). Etymology The specific epithet refers to the elytron with large punctures.	en	Wang, Xingmin, Ren, Shunxiang (2012): A review of Chinese Scymnomorphus Weise (Coleoptera: Coccinellidae) with description of five new species. Journal of Natural History 46 (31 - 32): 1905-1920, DOI: 10.1080/00222933.2012.693958, URL: http://dx.doi.org/10.1080/00222933.2012.693958
038587F3FFEC886154E3FF70FC63F9E6.taxon	description	(Figures 2 D, 4 F – J, 6) Diagnosis This species is similar to S. magnopunctatus sp. nov. in male genitalia, but can be distinguished from the latter as follows: body distinctly larger, brown, punctures on elytra fine and apex of penis guide in ventral view pointed (Figures 2 D, 4 J). In S. magnopunctatus, body is yellow brown, punctures on elytra are large and apex of penis guide in ventral view is truncated (Figures 2 C, 4 E). Description TL: 1.28 – 1.38 mm, TW: 0.8 – 0.96 mm, TH: 0.60 – 0.65 mm, TL / TW: 1.43 – 1.44; PL / PW: 0.52 – 0.54; EL / EW: 1.18 – 1.22. Body brown, except mesoventrite and metaventrite dark brown (Figure 2 D). Body minute, short oval, distinctly convex; dorsum pubescent. Head small, 0.43 times elytral width (HW / EW = 1: 2.31); frons convex, with fine punctures, separated by 0.8 – 2.0 times their diameter, with short and sparse hairs; eyes small and coarsely faceted, widest interocular distance 0.66 times width of head. Pronotum 0.76 times elytral width (PW / EW = 1: 1.32), pronotal punctures very fine, smaller than those on head, separated by 2.0 – 4.0 times their diameter. Punctures on elytra moderately large, separated by 0.3 – 1.0 times their diameter. Surface of proventrite and mesoventrite slightly shagreened. Punctures on metaventrite fine, separated by 0.3 – 1.5 times their diameter, with short and sparse setae in punctures. Abdominal incomplete or reaching lateral margin smoothly, divided into two lines, with many associated pits and pores (Figure 4 F). Male genitalia. Penis simple and slender, curved and narrowing toward apex; penis capsule indistinct (Figure 4 G, H); penis guide in lateral view short and stout, apex pointed; parameres short and indistinct, with four or five long setae (Figure 4 I); penis guide in ventral view short and cuspate, widest at base (Figure 4 J). Type material Holotype: 1 ♂, China, Tibet: Xiayadong Town, Yadong, 2800 m, 29 September 2009, Wang XM leg. Paratypes: China, Tibet:, 1 ♂ 1 ♀, same data as holotype; 1 ♂, Hanmi Village, Motuo, 2100 m, 13 October 2009, Wang XM leg. Distribution China (Tibet). Etymology The specific epithet refers to the cuspate penis guide in ventral view.	en	Wang, Xingmin, Ren, Shunxiang (2012): A review of Chinese Scymnomorphus Weise (Coleoptera: Coccinellidae) with description of five new species. Journal of Natural History 46 (31 - 32): 1905-1920, DOI: 10.1080/00222933.2012.693958, URL: http://dx.doi.org/10.1080/00222933.2012.693958
038587F3FFEB886254A7F98BFD21FD5B.taxon	description	(Figures 2 E, 5 A – E, 6) Diagnosis This species can be identified by its elongate oval body, fine punctures on elytron (Figure 2 E), incomplete abdominal postcoxal line, reaching lateral margin and bulging out in middle (Figure 5 A). The male genitalia are also diagnostic (Figure 5 B – E). Figure 6. Distribution map. (A) Scymnomorphus isolateralis sp. nov.; (Δ) Scymnomorphus japonicus (Kamiya); (◦) Scymnomorphus magnopunctatus sp. nov.; (•) Scymnomorphus cuspidatus sp. nov.; (♦) Scymnomorphus yadongensis sp. nov.; (Ɨ) Scymnomorphus xiaomengyangus sp. nov.; (*) Scymnomorphus bicolor (Kamiya). Description TL: 1.25 mm, TW: 0.83 mm, TH: 0.42 mm, TL / TW: 1.50; PL / PW: 0.60; EL / EW: 1.16. Body uniformly reddish brown, minute, elongate oval, distinctly convex; dorsum pubescent (Figure 2 E). Head small, 0.50 times elytral width (HW / EW = 1: 2.0); frons convex, with fine punctures, separated by 2.0 – 4.0 times their diameter, with short and sparse hairs, eyes small and coarsely faceted, widest interocular distance 0.69 times width of head. Pronotum 0.78 times elytral width (PW / EW = 1: 1.28), pronotal punctures fine, slightly smaller than those on head, separated by 1.0 – 3.0 times their diameter. Punctures on elytra large, separated by 0.3 – 1.5 times their diameter. Surface of proventrite and mesoventrite slightly shagreened, with several setae. Punctures on metaventrite moderately large, separated by 0.8 – 3.0 times their diameter, with short and dense setae in punctures. Abdominal postcoxal line incomplete, reaching lateral margin and bulging out in middle, divided into two lines, with associated large pits and pores (Figure 5 A). Male genitalia. Penis simple and slender, curved and narrowing toward apex; penis capsule with a short and stout outer process and a small inner one (Figure 5 B, C); penis guide in lateral view flattened and nearly quadrate, six times as long as wide, inner margin bulging out at apical third (Figure 5 D); parameres short and indistinct, with five or six long setae; penis guide in ventral view slender, narrowing apical, apex pointed (Figure 5 E). Type material Holotype: 1 ♂, China, Tibet: Xiayadong Town, Yadong, 2800 m, 29 September 2009, Wang XM leg. Distribution China (Tibet). Etymology The specific epithet refers to the type locality.	en	Wang, Xingmin, Ren, Shunxiang (2012): A review of Chinese Scymnomorphus Weise (Coleoptera: Coccinellidae) with description of five new species. Journal of Natural History 46 (31 - 32): 1905-1920, DOI: 10.1080/00222933.2012.693958, URL: http://dx.doi.org/10.1080/00222933.2012.693958
038587F3FFE8887D54C5FD18FD65FD2E.taxon	description	(Figures 2 F, 5 F – J, 6) Diagnosis This species is similar to S. bicolor in general appearance and male genitalia, but based on Kamiya’s detailed descriptions and illustrations, it can be distinguished from the latter by the elytra with a red brown area (Figure 2 F). In S. bicolor, elytron is uniformly black. Their male genitalia are also distinctly diagnostic (Figures 5 F – J). Description TL: 1.02 mm, TW: 0.78 mm, TH: 0.47 mm, TL / TW: 1.30; PL / PW: 0.52; EL / EW: 0.90. Head yellow, with eyes black. Pronotum yellowish brown. Ground colour of elytra yellowish brown, with sutural area reddish brown (Figure 2 F). Underside yellowish brown, except mesoventrite and metaventrite dark brown. Legs yellow. Body minute, short oval, distinctly convex; dorsum pubescent. Head small, 0.47 times elytral width (HW / EW = 1: 2.14); frons convex, with fine punctures, separated by 1.5 – 3.0 times their diameter, with short and sparse hairs; eyes small and coarsely faceted, widest interocular distance 0.66 times width of head. Pronotum 0.77 times elytral width (PW / EW = 1: 1.30), pronotal punctures fine, similar to those on head, separated by 2.0 – 4.0 times their diameter. Punctures on elytra large, separated by 0.5 – 1.5 times their diameter. Surface of proventrite and mesoventrite slightly shagreened, with several setae. Punctures on metaventrite moderately large, separated by 1.0 – 3.0 times their diameter, with short and dense setae in punctures. Abdominal postcoxal line incomplete, reaching lateral margin smoothly, divided into two lines, with associated small pits and pores (Figure 5 F). Male genitalia. Penis simple and slender, curved and narrowing toward apex; penis capsule indistinct (Figure 5 G, H); penis guide in lateral view slender and almost straight, almost subparallel at basal five-sixths, somewhat bulging at apical one-sixth, apex pointed and strongly bent; parameres short and indistinct, with several setae (Figure 5 I); penis guide in ventral view stout and straight, narrowed at base, tapering apical, apex pointed and slightly bent (Figure 5 J). Type material Holotype: 1 ♂, China, Yunnan: Xiaomengyang Town, Xishuangbanna, 29 September 2009, Wang XM leg; Paratype: China, Yunnan: 1 ♂, Jiluo Town, Jinghong, 6 May 2009, Ren SX leg. Distribution China (Yunnan). Etymology The specific epithet refers to the type locality.	en	Wang, Xingmin, Ren, Shunxiang (2012): A review of Chinese Scymnomorphus Weise (Coleoptera: Coccinellidae) with description of five new species. Journal of Natural History 46 (31 - 32): 1905-1920, DOI: 10.1080/00222933.2012.693958, URL: http://dx.doi.org/10.1080/00222933.2012.693958
038587F3FFF7887D5499FCF3FE71FB45.taxon	distribution	Distribution China (Taiwan), Japan.	en	Wang, Xingmin, Ren, Shunxiang (2012): A review of Chinese Scymnomorphus Weise (Coleoptera: Coccinellidae) with description of five new species. Journal of Natural History 46 (31 - 32): 1905-1920, DOI: 10.1080/00222933.2012.693958, URL: http://dx.doi.org/10.1080/00222933.2012.693958
