identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0385FA5C9604FFB0FF3988E4D909C2C1.text	0385FA5C9604FFB0FF3988E4D909C2C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus opinatus (Karsch 1881) Arw	<div><p>The Thyropygus opinatus subgroup (including the T. bifurcus subgroup of Pimvichai et al. 2009 b)</p><p>This is a subgroup of the T. allevatus group, characterized by an additional projection on the anterior coxal fold (amp). Included species:</p><p>T. bearti Pimvichai et al., 2009a</p><p>T. bifurcus (Demange, 1986)</p><p>T. bispinispatula Pimvichai et al., 2009a</p><p>T. bispinus Pimvichai et al., 2009a</p><p>T. brachyacanthus Pimvichai et al., 2009a</p><p>T. casjeekeli Pimvichai et al., 2009b</p><p>T. chelatus Pimvichai et al., 2009a</p><p>T. cristagalli Pimvichai et al., 2009a</p><p>T. demangei Pimvichai et al., 2009b</p><p>T. enghoffi (Demange, 1989)</p><p>T. erectus Pimvichai et al., 2009a</p><p>T. floweri (Demange, 1961)</p><p>T. implicatus (Demange, 1961)</p><p>T. inflexus (Demange, 1989)</p><p>T. loxia Pimvichai et al., 2009a</p><p>T. opinatus (Karsch, 1881)</p><p>T. quadricuspis Pimvichai et al., 2009b</p><p>T. richardhoffmani Pimvichai et al., 2009b</p><p>T. cimi sp. nov.</p><p>T. culter sp. nov.</p><p>T. forceps sp. nov.</p><p>T. mesocristatus sp. nov.</p><p>T. navychula sp. nov.</p><p>T. planispina sp. nov.</p><p>T. sutchariti sp. nov.</p><p>T. undulatus sp. nov.</p><p>T. ursus sp. nov.</p><p>In addition to the diagnostic character combination given by Pimvichai et al. (2009a, b), all species in the opinatus subgroup share common characters of head, antennae, mandibles, gnathochilarium, collum, epiproct, paraprocts, hypoproct and first pair of male legs. We therefore give a general description of males of the opinatus subgroup, so that subsequent species descriptions (which are mainly based on males) can be kept brief.</p><p>General description of males of the T. opinatus subgroup</p><p>Head smooth, 5–8 supralabral setae, eyes moderate in size, ocelli in 7–8 horizontal rows consisting of 11–14, 10–13, 8–11, 7–9, 5–8, 3–6, 1–4 and 1–3 ocelli, total: 46–68.</p><p>Gnathochilarium: mentum smooth, at most with a few small setae distally and a large, horseshoe-shaped ridge opening distally. Stipites densely covered with spine-like setae, except for an irregular oblique band from c. middle of lateral margin to border of lamella lingualis; long setae present on the distolateral part; males (not females) distally with a small sclerotised ‘island’ with 1–3 spine-like setae in middle of an ovoid, poorly sclerotized, hairless area. Lamellae linguales with three long, apical setae and a number of short, basal, spine-like setae.</p><p>Sterna smooth, stigmatal grooves distinct, long. Legs relatively long; tip of postfemora, tibiae and tarsi visible from above when legs are extended laterad; coxae of anterior and posterior pair markedly dissimilar, posterior coxae humped; coxae with 2–8 setae, prefemora with 3–6 setae, femora with 2–5 setae in a median row and 2 larger, subapical setae; tarsi typically with 6–8 irregularly placed ventral setae and 1–2 large dorsal, apical spurs. Postfemora and tibiae without setae.</p><p>Ventral pads on postfemora and tibiae on all legs, except first three pairs.</p><p>Gonopods: sternum (st) triangular. Anterior coxal fold (ac) basally slender, becoming broader towards tip, lateral margins diverging; distally with two processes: a lateral process (alp) and a usually smaller mesal process (amp); the shape of these processes is species-specific. Posterior coxal (pc) fold much lower than anterior coxal fold, basally with moderately high lateral paracoxites (px), distally variously modified but always with a smooth area over which the telopodite shaft can slide. Telopodite with a single or double, well-developed femoral spine (fe) and a long, slender, curved tibial spine (ti). Many species with a characteristic spatulate lobe (sl) originating under the base of the tibial spine; lobe sometimes distally rounded, spoon-like, sometimes ending in a large, stout spine. Other species instead with a small and slightly folded lateral lamella (ll). Apical palette (pa) simple, forming a broad gutter, sometimes with a longitudinal crest in the concavity; apically with a row of 7–13 brownish blepharochaetae (bp).</p><p>Key to species of the T. opinatus subgroup (partly extracted and adapted from Pimvichai et al. 2009a, b)</p><p>1. Apical part of telopodite with spatulate lobe (sl)............................................................................2</p><p>− Apical part of telopodite with lateral lamella (ll)...........................................................................20</p><p>2. Spatulate lobe (sl) distally drawn out into one (rarely two) sharp dark brown spine(s)............3</p><p>− Spatulate lobe (sl) distally expanded and/or rounded, spoon-like, without a spine....................9</p><p>3. Spatulate lobe (sl) terminating in two sharp brown spines, the outer spine slightly smaller and shorter than the inner one; lateral process of anterior coxal fold (alp) slender, slightly curving mesad; mesal process of anterior coxal fold (amp) almost as long as alp, flattened ............................ .............................................................................................. T. bispinispatula Pimvichai et al., 2009a</p><p>− Spatulate lobe (sl) terminating in a single sharp dark brown spine.............................................4</p><p>4. Telopodite without a lobe distal to femoral spine (fe); lateral process of anterior coxal fold (alp) long, slender, regularly curved, tip close to tip of opposite alp, the two together forming a circle; mesal process of anterior coxal fold (amp) straight, shorter than alp; fe directed distad, pointed ...................................................................................................... T. erectus Pimvichai et al., 2009a</p><p>− Telopodite distally to fe with a large, round lobe (lo), projecting distolaterally.....................5</p><p>5. Lateral process of anterior coxal fold (alp) very slender, regularly curved...........................6</p><p>− Lateral process of anterior coxal fold (alp) different, broader and/or with several apical denticles...8</p><p>6. Mesal margin of lateral process of anterior coxal fold (alp) with fine serrations (Fig. 6A); mesal process of anterior coxal fold (amp) almost as long as alp, broadly expanded, apically sharp, straight distad, mesal margin forming a strong longitudinal crest (lc) in posterior view (Fig. 6B) ........................................................................................................................... T. navychula sp. nov.</p><p>− Mesal margin of lateral process of anterior coxal fold (alp) without serrations, tip of lateral process close to tip of the opposite side, the two together forming a circle..............................7</p><p>7. Mesal process of posterior coxal fold (pmp) strongly developed along anterior-posterior axis ..... ................................................................................................................. T. floweri (Demange, 1961)</p><p>− Mesal process of posterior coxal fold (pmp) slender, directed distolaterad (Fig. 4C) .................... ................................................................................................................................ T. forceps sp. nov.</p><p>8. Lateral process of anterior coxal fold (alp) broad, apically gradually narrowed; mesal process of anterior coxal fold (amp) almost as long as alp, slender, straight, terminally slightly curved, pointed ....................................................................................................... T. opinatus (Karsch, 1881)</p><p>− Lateral process of anterior coxal fold (alp) apically bent abruptly mesad, tip with serrate margins; mesal process of anterior coxal fold (amp) much shorter than alp, directed meso-distad, simple, pointed; mesal process of posterior coxal fold (pmp) strongly developed along anteriorposterior axis ................................................................................... T. implicatus (Demange, 1961)</p><p>9. Telopodite with a single femoral spine..........................................................................................10</p><p>− Telopodite with a double femoral spine..........................................................................................17</p><p>10. Lateral process of anterior coxal fold (alp) apically abruptly truncate .......................................... ......................................................................................................... T. bearti Pimvichai et al., 2009a</p><p>− Lateral process of anterior coxal fold (alp) apically pointed.........................................................11</p><p>11. Mesal process of anterior coxal fold (amp) very small; telopodite distally to fe with a small round lobe (lo) projecting distolaterally ......................................... T. loxia Pimvichai et al., 2009a</p><p>− Mesal process of anterior coxal fold (amp) relatively long....................................................12</p><p>12. Mesal process of anterior coxal fold (amp) shorter than lateral process (alp)...........................13</p><p>− Mesal process of anterior coxal fold (amp) as long as lateral process (alp)..............................14</p><p>13. Mesal process of anterior coxal fold (amp) directed obliquely disto-mesad, slender, straight .......... ...................................................................................................... T. chelatus Pimvichai et al., 2009a</p><p>− Mesal process of anterior coxal fold (amp) directed distad, thicker, slightly sigmoid ....................... ........................................................................................ T. brachyacanthus Pimvichai et al., 2009a</p><p>14. Mesal process of anterior coxal fold (amp) directed obliquely disto-mesad, tip overlapping tip of opposite amp (Fig. 8A); lateral process of posterior coxal fold (plp) a massive, broad lobe, projecting laterad (Fig. 8B) ................................................................ T. sutchariti sp. nov.</p><p>− Mesal process of anterior coxal fold (amp) directed distad.........................................................15</p><p>15. Lateral process of anterior coxal fold (alp) apically without a crest; telopodite distally with a rounded lobe (lo); margins of spatulate lobe (sl) terminally meeting in a distinct angle ............... .................................................................................................... T. bispinus Pimvichai et al., 2009a</p><p>− Lateral process of anterior coxal fold (alp) apically with a crest.............................................16</p><p>16. Mesal process of anterior coxal fold (amp) apically irregularly tuberculate; telopodite distally without a rounded lobe (lo) ..................................................................... T. inflexus (Demange, 1989)</p><p>− Mesal process of anterior coxal fold (amp) slender, straight, its tip pointed, its mesal margin forming a strong longitudinal crest (lc) in posterior view (Fig. 5B) .......... T. mesocristatus sp. nov.</p><p>17. Anterior coxal fold (ac) with an additional spine-like process (aip) between lateral process of anterior coxal fold (alp) and mesal process of anterior coxal fold (amp); alp broad, mesal margin concave, tip with serrate margins, cockscomb-like; amp much shorter than alp, directed meso-distad, simple, pointed; both femoral spines (fe) slender, long ............ T. cristagalli Pimvichai et al., 2009a</p><p>− Anterior coxal fold (ac) without an additional spine-like process (aip) between alp and amp ...18</p><p>18. Lateral process of anterior coxal fold (alp) apically without a crest, flattened, slightly curved, its laterodistal margin coarsely dentate, terminating in a short, sharp, pointed spine (Fig. 3A); mesal process (amp) much shorter than alp, directed distad, tip curving mesad, pointed; both femoral spines (fe 1, fe 2) long, curving backward; tibial spine (ti) long, not curving in horizontal plane (Fig. 3C) ......... ................................................................................................................................... T. culter sp. nov.</p><p>− Lateral process of anterior coxal fold (alp) apically with a crest extending caudad...............19</p><p>19. Lateral process of anterior coxal fold (alp) flattened, curving mesad, laterodistal margin coarsely dentate, terminating in a short spine, tip curving against the tip of opposite side (Fig. 9A); mesal process of anterior coxal fold (amp) much shorter than alp, slender, curving mesad; both femoral spines (fe 1, fe 2) broad, long; tibial spine (ti) long, curving in horizontal plane, not ending in a sharp spine (Fig. 9D) .................................................................... T. undulatus sp. nov.</p><p>− Lateral process (alp) regularly curved, terminating in a sharp, slightly upward-pointing spine (Fig. 7A); mesal process (amp) slightly shorter than alp, flattend, straight, directed distad; tibial spine (ti) flattend, short, curving mesad (Fig. 7D) ........................................... T. planispina sp. nov.</p><p>20. Telopodite with a single femoral spine.......................................................................................21</p><p>− Telopodite with a double femoral spine.......................................................................................23</p><p>21. Lateral process of anterior coxal fold (alp) without an apical crest; mesal process of anterior coxal fold (amp) shorter than and as broad as alp, directed distad; femoral spine (fe) very long and slender .............................................................. T. casjeekeli Pimvichai et al., 2009b</p><p>− Lateral process of anterior coxal fold (alp) with a sharp crest on the posterior surface near the tip...22</p><p>22. Lateral process of anterior coxal fold (alp) flattened, slightly curved, inflexed; femoral spine (fe) very long, slender, with an additional lamella at base ... T. quadricuspis Pimvichai et al., 2009b</p><p>− Lateral process of anterior coxal fold (alp) regularly curved, basally broad, gradually tapering towards end and ending in sharp point (Fig. 2A); femoral spine (fe) very long, slender, without an additional lamella at base (Fig. 2D) ............................................................................ T. cimi sp. nov.</p><p>23. Lateral process of anterior coxal fold (alp) broader..................................................................24</p><p>− Lateral process of anterior coxal fold (alp) slender, regularly curved, sickle-shaped............25</p><p>24. Lateral process of anterior coxal fold (alp) terminating in a very short external spine and a very long internal one; mesal process of anterior coxal fold (amp) as long as alp; first femoral spine (fe 1) very short, pointed; second femoral spine (fe 2) very long, as long as tibial spine (ti); an additional lamella at both sides of base of fe 2 ........................ ... T. richardhoffmani Pimvichai et al., 2009b</p><p>− Lateral process of anterior coxal fold (alp) flattened, apically curved laterad as a short spine, lateral margin of alp slightly folded; mesal process of anterior coxal fold (amp) shorter than alp, slender, straight, directed distad, pointed (Fig. 10A); the first femoral spine (fe 1) very short, directed upward, situated above the second femoral spine (fe 2); fe 2 very long, slender, curved downward (Fig. 10C) .................................................................................. T. ursus sp. nov.</p><p>25. Mesal margin of lateral process of anterior coxal fold (alp) simple, without a caudad spine or crest; mesal process of anterior coxal fold (amp) much shorter than alp, curved, pointed ............................. ................................................................................................................ T. enghoffi (Demange, 1989)</p><p>− Mesal margin of lateral process of anterior coxal fold (alp) caudad with a small spine or crest......26</p><p>26. Mesal margin of lateral process of anterior coxal fold (alp) with a small caudad crest; mesal process of anterior coxal fold (amp) slightly shorter than alp, slightly sigmoid, pointed ............... ............................................................................................................ ... T. bifurcus (Demange, 1986)</p><p>− Mesal margin of lateral process of anterior coxal fold (alp) with a short curved caudad spine; mesal process of anterior coxal fold (amp) as long as alp, straight ... T. demangei Pimvichai et al., 2009b</p></div>	https://treatment.plazi.org/id/0385FA5C9604FFB0FF3988E4D909C2C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak;Backeljau, Thierry	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9600FFB2FD008E36DEA3C2DC.text	0385FA5C9600FFB2FD008E36DEA3C2DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus cimi Pimvichai & Enghoff & Panha & Backeljau 2016	<div><p>Thyropygus cimi sp. nov.</p><p>urn:lsid:zoobank.org:act: 0252646C-AD4E-4B8B-8A7C-AEEA80DD4A72</p><p>Fig. 2 A–E</p><p>Diagnosis</p><p>A species of the opinatus subgroup. Lateral process of anterior coxal fold (alp) regularly curved, basally broad, gradually tapering towards end and ending in sharp point, with a lateral crest near tip. Similar in this respect to T. bifurcus . Differs from this species by having the mesal process of anterior coxal fold (amp) straight, directed distad, by having the mesal process of posterior coxal fold (pmp) slender, directed anteriad, and by having only one femoral spine (fe).</p><p>Etymology</p><p>This species is named after the organization “ Centre International de Myriapodologie – CIM” (www. myriapodology.org) in recognition of its immense importance for inspiring and supporting research on myriapods.</p><p>Material examined</p><p>Holotype THAILAND: ♂, Nakhonsrithammarat Province, Lanska District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.82833&amp;materialsCitation.latitude=8.340278" title="Search Plazi for locations around (long 99.82833/lat 8.340278)">Namwang Srithammasokrach</a>, 8°20'25" N, 99°49'42" E, 16 May 2010, leg. S. Panha, P. Pimvichai and members of the Animal Systematics Research Unit (CUMZ-D00086).</p><p>Paratypes THAILAND: 2 ♂♂, 1 ♀, same data as holotype (CUMZ-D00096); 1 ♂, same data as holotype (ZMUC).</p><p>Description</p><p>Adult males with 65–68 podous rings, no apodous rings. Length 13–16 cm, width 7.6–8.3 mm. Adult female with 65 podous rings, no apodous rings. Length 13 cm, width 7.4 mm. Colour in life unknown;</p><p>preserved specimens with head, antennae, prozona, middorsal metazona and legs brown; metazona, epiproct, paraprocts and hypoproct dark brown.</p><p>GONOPODS (Fig. 2 A–E). Anterior coxal fold (ac; Fig. 2A): lateral process (alp) regularly curved, basally broad, gradually tapering towards end and ending in sharp point, with lateral crest near tip (Fig. 2C, arrow); mesal process (amp) slightly shorter than alp, straight, directed distad. Posterior coxal fold (pc; Fig. 2B) basally with moderately high lateral paracoxites (px), distally truncate, forming shelf for accommodation of telopodite, mesal process (pmp) slender, directed anteriad. Telopodite (Fig. 2 D–E) leaving coxite over shelf of posterior coxal fold; femoral spine (fe) very long, slender, curving downward, in situ resting between alp and amp; tibial spine (ti) long, slender, curving in horizontal plane, its tip in situ resting close to base of fe; apical part: lamellar lobe (ll) broadly expanded, bent down; palette (pa) simple, gutter-like, with longitudinal rounded crest (cr) at middle; distally with about ten brownish blepharochaetae (bp).</p><p>DNA barcode</p><p>The GenBank accession number of the barcode of the holotype is KU306519 (voucher code CUMZ-D00086).</p><p>Distribution (Fig. 12)</p><p>Known only from the type locality.</p><p>Remarks</p><p>Coexisting with the smaller T. forceps sp. nov.</p></div>	https://treatment.plazi.org/id/0385FA5C9600FFB2FD008E36DEA3C2DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak;Backeljau, Thierry	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9602FFACFDFF8937DE26C26C.text	0385FA5C9602FFACFDFF8937DE26C26C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus culter Pimvichai & Enghoff & Panha & Backeljau 2016	<div><p>Thyropygus culter sp. nov.</p><p>urn:lsid:zoobank.org:act: 0093C39D-F1FC-4CAD-9DA0-0492F45E7370</p><p>Fig. 3 A–D</p><p>Diagnosis</p><p>A species of the opinatus subgroup. Lateral process of anterior coxal fold (alp) flattened, slightly curved, its laterodistal margin coarsely dentate. Similar in this respect to T. cristagalli, T. implicatus and T. undulatus sp. nov. Differs from these species by having the mesal process of posterior coxal fold (pmp) very high, pointed-triangular, directed almost straightly distad, and by having the tibial spine (ti) recurved.</p><p>Etymology</p><p>The name is a Latin noun in apposition, meaning “knife”, and refers to the knifelike second femoral spine (fe 2).</p><p>Material examined</p><p>Holotype THAILAND: ♂, Krabi Province, Khlong Thom District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.204994&amp;materialsCitation.latitude=7.935" title="Search Plazi for locations around (long 99.204994/lat 7.935)">Rorn waterfall</a>, 7°56'06" N, 99°12'18" E, 15 Jan. 2009, leg. P. Pimvichai, C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D00091).</p><p>Paratypes THAILAND: 3 ♂♂, 2 ♀♀, 1 subadult ♀, same data as holotype (CUMZ-D00078); 1 ♂, 1 ♀, same data as holotype (ZMUC).</p><p>Description</p><p>Adult males with 58–62 podous rings, no apodous rings. Length 12–14 cm, width 7.1–8.0 mm. Adult females with 59–62 podous rings, no apodous rings. Length 11–13 cm, width 6.9–7.9 mm. Colour in life unknown; preserved specimens with head, antennae, prozona, middorsal metazona and legs brown; metazona, epiproct, paraprocts and hypoproct reddish brown.</p><p>GONOPODS (Fig. 3 A–D). Anterior coxal fold (ac; Fig. 3A): lateral process (alp) flattened, slightly curved, its laterodistal margin coarsely dentate, terminating in short, sharp, pointed spine; mesal process (amp) much shorter than alp, directed distad, tip curving mesad, pointed. Posterior coxal fold (pc; Fig. 3B) basally with lateral paracoxites (px) quite low, distally with two processes: mesal process (pmp) very high, pointed-triangular, directed almost straightly distad; lateral process (plp) much shorter, digitiform, directed distad and slightly laterad. Telopodite (Fig. 3 C–D) leaving coxite over shelf of posterior coxal fold; femoral spine (fe) duplicated, fe 1 projecting above top of telopodite curvature, curving backward, ending in slender barb, fe 2 basally broad, situated under fe 1, curving backward, ending in slender barb; tibial spine (ti) long, slender and recurved (not curving in horizontal plane); apical part: spatulate lobe (sl) slender, rounded; with slender, long spine (sls) at base of apical part, opposite origin of tibial spine; palette (pa) simple, distally with about twelve brownish blepharochaetae (bp).</p><p>DNA barcode</p><p>The GenBank accession number of the barcode of one of the paratypes is KC519535 (voucher code CUMZ-D00078).</p><p>Distribution (Fig. 12)</p><p>Known only from the type locality.</p></div>	https://treatment.plazi.org/id/0385FA5C9602FFACFDFF8937DE26C26C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak;Backeljau, Thierry	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961CFFAEFDF688A7D851C281.text	0385FA5C961CFFAEFDF688A7D851C281.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus forceps Pimvichai & Enghoff & Panha & Backeljau 2016	<div><p>Thyropygus forceps sp. nov.</p><p>urn:lsid:zoobank.org:act: 11BAAEC5-404D-4E15-982C-2F3E51DEA56B</p><p>Figs 4 A–E, 11B</p><p>Diagnosis</p><p>A species of the opinatus subgroup. Lateral process of anterior coxal fold (alp) long, slender, regularly curved, tip close to tip of opposite alp, the two together forming a circle. Similar in this respect to T. erectus and T. floweri . Differs from the former by having a telopodite lobe (lo). Particularly similar to T. floweri, differing from it by having the mesal process of posterior coxal fold (pmp) slender, directed distolaterad, laterally with a digitiform process (plp), and by not having pmp strongly developed along the anterior-posterior axis.</p><p>Etymology</p><p>The name is a Latin noun in apposition, referring to the forceps-like gonopod coxae.</p><p>Material examined</p><p>Holotype THAILAND: ♂, Nakhonsrithammarat Province, Lanska District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.82833&amp;materialsCitation.latitude=8.340278" title="Search Plazi for locations around (long 99.82833/lat 8.340278)">Namwang Srithammasokrach</a>, 8°20'25" N, 99°49'42" E, 16 May 2010, leg. S. Panha, P. Pimvichai and members of the Animal Systematics Research Unit (CUMZ-D00092).</p><p>Paratypes THAILAND: 2♂♂, 2subadult ♂♂, same data as holotype (CUMZ-D00073); 5♂♂, Nakhonsrithammarat Province, Ronpiboon District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.8675&amp;materialsCitation.latitude=8.245" title="Search Plazi for locations around (long 99.8675/lat 8.245)">Tham Pha Deang temple</a>, 8°14'42" N, 99°52'03" E, 14 May 2012, leg. C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D00093); 2 ♂♂, same data as preceding (ZMUC).</p><p>Description</p><p>Adult males with 54–62 podous rings, no apodous rings. Length 9–12 cm, width 4.6–5.8 mm. Overall colour of living animal (Fig. 11B) brown. Legs, antennae, middorsal metazona, epiproct, paraprocts and hypoproct brownish orange; preserved specimens with head, antennae, prozona, legs, middorsal metazona, epiproct, paraprocts and hypoproct brown; metazona dark brown.</p><p>GONOPODS (Fig. 4 A–E). Anterior coxal fold (ac; Fig. 4A): distinctly crenulated along convex part of lateral margin (smooth in specimens from Tham Pha Deang temple), lateral process (alp) long, slender, regularly curved, tip close to tip of opposite alp, the two together forming a circle; mesal process (amp) slightly shorter than alp, directed obliquely disto-mesad (in specimens from Tham Pha Deang temple lateral process (alp) long, slender, crossing over with opposite tip, the two together forming a circle; Fig. 4B); mesal process (amp) slightly shorter than alp, directed distad, slightly sigmoid. Posterior coxal fold (pc; Fig. 4C) basally with moderately high lateral paracoxites (px), distally with two processes: mesal process (pmp) slender, directed distolaterad; lateral process (plp) digitiform. Telopodite (Fig. 4 D–E) leaving coxite over shelf of posterior coxal fold; femoral spine (fe) massive, curving in horizontal plane and curving against ti; telopodite distally to fe with large, round lobe (lo) projecting distolaterally; tibial spine (ti) very long, slender, curving in horizontal plane, its tip resting against base of fe; apical part: spatulate lobe (sl) with sharp dark brown spine at tip; palette (pa) simple, distally with about nine brownish blepharochaetae (bp).</p><p>DNA barcode</p><p>The GenBank accession number of the barcode of one of the paratypes is KC519531 (voucher code CUMZ-D00073).</p><p>Distribution (Fig. 12)</p><p>Known only from Namwang Srithammasokrach and Tham Pha Deang temple in Nakhonsrithammarat Province.</p><p>Remarks</p><p>Coexisting with the larger T. cimi sp. nov. at Namwang Srithammasokrach.</p></div>	https://treatment.plazi.org/id/0385FA5C961CFFAEFDF688A7D851C281	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak;Backeljau, Thierry	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961EFFA9FDCF889BDE26C24F.text	0385FA5C961EFFA9FDCF889BDE26C24F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus mesocristatus Pimvichai & Enghoff & Panha & Backeljau 2016	<div><p>Thyropygus mesocristatus sp. nov.</p><p>urn:lsid:zoobank.org:act: 5D8F7830-2E8E-46E3-B2BE-830FC31F290D</p><p>Fig. 5 A–E</p><p>Diagnosis</p><p>A species of the opinatus subgroup. Lateral process of anterior coxal fold (alp) slender, curving mesad; mesal process of anterior coxal fold (amp) as long as alp, slender, straight, directed distad. Similar in these respects to T. demangei . Differs from this species by having mesal margin of amp forming a strong longitudinal crest (lc) in posterior view, by having only one femoral spine (fe) and by having the spatulate lobe (sl) broad and rounded.</p><p>Etymology</p><p>The name is a Latin adjective, referring to the longitudinal crest on the mesal process of the anterior coxal fold.</p><p>Material examined</p><p>Holotype THAILAND: ♂, Songkhla Province, Rattaphum District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.14&amp;materialsCitation.latitude=7.005" title="Search Plazi for locations around (long 100.14/lat 7.005)">Srikasorn</a>, 7°00'18" N, 100°08'24" E, 12 Jan. 2009, leg. P. Pimvichai, C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D00094).</p><p>Paratype THAILAND: ♂, same data as holotype (CUMZ-D00077).</p><p>Description</p><p>Adult males with 61–66 podous rings, no apodous rings. Length 16–17 cm, width 9.2–9.5 mm. Colour in life unknown; preserved specimens with head, antennae, prozona and legs brown; metazona, epiproct, paraprocts and hypoproct reddish brown.</p><p>GONOPODS (Fig. 5 A–E). Anterior coxal fold (ac; Fig. 5A): lateral process (alp) slender, curving mesad, with lateral serrate crest near tip (Fig. 5C, arrow); mesal process (amp) as long as alp, slender, straight, directed distad, tip sharp, pointed, in posterior view mesal margin of amp forming strong longitudinal crest (lc). Posterior coxal fold (pc; Fig. 5B) basally with moderately high lateral paracoxites (px), distally with two processes: mesal process (pmp) slender, directed anteriad; lateral process (plp) short, erect, digitiform; forming shelf for accommodation of telopodite. Telopodite (Fig. 5 D–E) leaving coxite between pmp and plp; femoral spine (fe) long, curved downward, with expanded lamella at base; tibial spine (ti) very long, curving in horizontal plane, tip close to basal part of fe; small spine (ss) at base of apical part, opposite origin of tibial spine; spatulate lobe (sl) broad, rounded; a small accessory lobe (al) at base of sl; palette (pa) simple, gutter-like, with longitudinal crest (cr) at middle, distally with about twelve brownish blepharochaetae (bp).</p><p>DNA barcode</p><p>The GenBank accession number of the barcode of the paratype is KC519534 (voucher code CUMZ-D00077).</p><p>Distribution (Fig. 12)</p><p>Known only from the type locality.</p></div>	https://treatment.plazi.org/id/0385FA5C961EFFA9FDCF889BDE26C24F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak;Backeljau, Thierry	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9618FFABFDDB8B3ADE26C6AF.text	0385FA5C9618FFABFDDB8B3ADE26C6AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus navychula Pimvichai & Enghoff & Panha & Backeljau 2016	<div><p>Thyropygus navychula sp. nov.</p><p>urn:lsid:zoobank.org:act: 01AC06C9-EEF0-4211-9837-28FE2C12C68E</p><p>Figs 6 A–D, 11A</p><p>Diagnosis</p><p>A species of the opinatus subgroup. Mesal margin of anterior coxal fold (alp) with fine serrations. Similar in this respect to T. opinatus . Differs from this species by having the lateral process (alp)</p><p>slender, regularly curved and by having the mesal process (amp) broadly expanded and forming a strong longitudinal crest (lc) in posterior view.</p><p>Etymology</p><p>The species is named after the Royal Thai Navy, in recognition of their kind assistance which enabled us to pursue the necessary fieldwork at the type locality, and also after Chulalongkorn University where pink is the symbolic colour of the university, indirectly referring to the pink legs of the species; the species name is treated as a noun in apposition.</p><p>Material examined</p><p>Holotype THAILAND: ♂, Phang-Nga Province, Khuraburi District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.86056&amp;materialsCitation.latitude=9.405555" title="Search Plazi for locations around (long 97.86056/lat 9.405555)">Surin Islands</a>, 9°24'20" N, 97°51'38" E, 7 Apr. 2012, leg. S. Panha, P. Pimvichai and members of the Animal Systematics Research Unit (CUMZ-D00095).</p><p>Paratypes THAILAND: 5 ♂♂, 1 ♀, same data as holotype (CUMZ-D00089); 2 ♂♂, same data as holotype (ZMUC).</p><p>Description</p><p>Adult males with 58–62 podous rings, no apodous rings. Length 11–12 cm, width 6.2–7.2 mm. Adult female with 60 podous rings, no apodous rings. Length 14 cm, width 8.0 mm. Overall colour of living animal (Fig. 11A) dark brown, shiny. Legs and antennae pink; preserved specimen with head, antennae, prozona, legs, epiproct, paraprocts and hypoproct brown; metazona dark brown.</p><p>GONOPODS (Fig. 6 A–D). Anterior coxal fold (ac; Fig. 6A): lateral process (alp) slender, regularly curved, mesal margin with fine serrations; mesal process (amp) almost as long as alp, broadly expanded, apically sharp, straight distad, in posterior view mesal margin of amp forming strong longitudinal crest (lc). Posterior coxal fold (pc; Fig. 6B) basally with moderately high lateral paracoxites (px), distally with two processes: mesal process (pmp) shorter than plp, curving behind plp; lateral process (plp) flattened, directed distad. Telopodite (Fig. 6 C–D) leaving coxite between pmp and plp; femoral spine (fe) very long, curving almost exclusively in horizontal plane; tibial spine (ti) very long, curving in horizontal plane close to basal part of fe; spatulate lobe (sl) terminating in sharp spine; palette (pa) simple, gutterlike, distally with about twelve brownish blepharochaetae (bp), laterally with few sharp spines.</p><p>DNA barcode</p><p>The GenBank accession number of the barcode of one of the paratype is KU306522 (voucher code CUMZ-D00089).</p><p>Distribution (Fig. 12)</p><p>Known only from the type locality.</p></div>	https://treatment.plazi.org/id/0385FA5C9618FFABFDDB8B3ADE26C6AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak;Backeljau, Thierry	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961BFFA5FDDA8CE2DE26C497.text	0385FA5C961BFFA5FDDA8CE2DE26C497.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus planispina Pimvichai & Enghoff & Panha & Backeljau 2016	<div><p>Thyropygus planispina sp. nov.</p><p>urn:lsid:zoobank.org:act: 2F5A25A4-44CC-4B93-AC17-9A9B6E6E9E1F</p><p>Fig. 7 A–E</p><p>Diagnosis</p><p>A species of the opinatus subgroup. Differs from all other species in the subgroup by having the lateral process of the anterior coxal fold (alp) regularly curved, terminating in a sharp spine pointing slightly distad, and by having the tibial spine (ti) short and flattened.</p><p>Etymology</p><p>The name is a Latin noun in apposition and refers to the flattened tibial spine (ti).</p><p>Material examined</p><p>Holotype THAILAND: ♂, Krabi Province, Muang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.924995&amp;materialsCitation.latitude=8.125" title="Search Plazi for locations around (long 98.924995/lat 8.125)">Tham Sua temple</a>, 8°07'30" N, 98°55'30" E, 15 Jan. 2012, leg. P. Pimvichai, C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D00088).</p><p>Paratypes THAILAND: 3 ♀♀, same data as holotype (CUMZ-D00097).</p><p>Description</p><p>Adult male with 64 podous rings, no apodous rings. Length 13 cm, width 7.6 mm. Adult females with 60–63 podous rings, no apodous rings. Length 13–14 cm, width 7.8–8.6 mm. Colour in life unknown; preserved specimens with head, antennae, legs, metazona, epiproct, paraprocts and hypoproct brown; prozona yellowish brown.</p><p>GONOPODS (Fig. 7 A–E). Anterior coxal fold (ac; Fig. 7A): lateral process (alp) regularly curved, terminating in sharp spine pointing slightly distad, with lateral short, sharp crest at middle of alp (Fig. 7C, arrow); mesal process (amp) slightly shorter than alp, flattened, straight, directed distad. Posterior coxal fold (pc; Fig. 7B) basally with moderately high lateral paracoxites (px), distally truncate, forming shelf for accommodation of telopodite, mesal process (pmp) triangular, directed distad, lateral process (plp) erect, digitiform. Telopodite (Fig. 7 D–E) leaving coxite over shelf of posterior coxal fold; femoral spine double, fe 1 very long, curved downward, situated above fe 2, in situ resting at middle of alp, fe 2 long, very slender, curved downward; tibial spine (ti) flattened, short, curving mesad; apical part: spatulate lobe (sl) small, rounded; with slender, long spine (sls) at base of apical part; palette (pa) simple, gutter-like, distally with about nine brownish blepharochaetae (bp).</p><p>DNA barcode</p><p>The GenBank accession number of the barcode of the holotype is KU306521 (voucher code CUMZ-D00088).</p><p>Distribution (Fig. 12)</p><p>Known only from the type locality.</p></div>	https://treatment.plazi.org/id/0385FA5C961BFFA5FDDA8CE2DE26C497	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak;Backeljau, Thierry	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9615FFA7FDE08E8ADE26C31C.text	0385FA5C9615FFA7FDE08E8ADE26C31C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus sutchariti Pimvichai & Enghoff & Panha & Backeljau 2016	<div><p>Thyropygus sutchariti sp. nov.</p><p>urn:lsid:zoobank.org:act: 6804DF2C-7787-46FE-9185-2E5C40AF5ED3</p><p>Fig. 8 A–D</p><p>Diagnosis</p><p>A species of the opinatus subgroup. Mesal process of anterior coxal fold (amp) directed slightly obliquely disto-mesad, tip overlapping tip of opposite amp. Similar in this respect to T. chelatus . Differs from this species by having amp as long as alp and by having the lateral process of the posterior coxal fold (plp) as a massive, broad lobe.</p><p>Etymology</p><p>The species is named in honour of Chirasak Sutcharit in recognition of his devotion to collecting millipedes.</p><p>Material examined</p><p>Holotype THAILAND: ♂, Phetchaburi Province, Kaeng Krachan District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.59889&amp;materialsCitation.latitude=12.901944" title="Search Plazi for locations around (long 99.59889/lat 12.901944)">Kaeng Krachan</a>, 12°54'07" N, 99°35'56" E, 14 May 2012, leg. C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D00090).</p><p>Paratypes THAILAND: 2 ♂♂, 2 ♀♀, 1 subadult ♀, same data as holotype (CUMZ-D00098); 1 ♂, same data as holotype (ZMUC).</p><p>Description</p><p>Adult males with 63–65 podous rings, no apodous rings. Length 10–11 cm, width 5.4–5.7 mm. Adult females with 61–63 podous rings, no apodous rings. Length 12–13 cm, width 6.7–6.9 mm. Colour in life unknown; preserved specimens with head, antennae, prozona, legs, epiproct, paraprocts and hypoproct brown; metazona reddish brown.</p><p>GONOPODS (Fig. 8 A–D). Anterior coxal fold (ac; Fig. 8A): lateral process (alp) slender, regularly curved; mesal process (amp) as long as alp, directed slightly obliquely disto-mesad, tip overlapping tip of opposite amp. Posterior coxal fold (pc; Fig. 8B) basally with moderately high lateral paracoxites (px), distally with two processes: mesal process (pmp) a rounded lobe; lateral process (plp) a massive, broad lobe, projecting laterad. Telopodite (Fig. 8 C–D) leaving coxite between pmp and plp; femoral spine (fe) long, curved downward; telopodite distally to fe with large, round lobe (lo) projecting distolaterally; tibial spine (ti) very long, curving in horizontal plane, tip close to basal part of lo; spatulate lobe (sl) broad, rounded; palette (pa) simple, gutter-like, distally with about ten brownish blepharochaetae (bp).</p><p>DNA barcode</p><p>The GenBank accession number of the barcode of the holotype is KU306524 (voucher code CUMZ-D00090).</p><p>Distribution (Fig. 12)</p><p>Known only from the type locality.</p></div>	https://treatment.plazi.org/id/0385FA5C9615FFA7FDE08E8ADE26C31C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak;Backeljau, Thierry	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9617FFA1FDE48977DE26C157.text	0385FA5C9617FFA1FDE48977DE26C157.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus undulatus Pimvichai & Enghoff & Panha & Backeljau 2016	<div><p>Thyropygus undulatus sp. nov.</p><p>urn:lsid:zoobank.org:act: 972296F7-CF7B-4AD4-A2AA-3DCE22596B3E</p><p>Fig. 9 A–E</p><p>Diagnosis</p><p>A species of the opinatus subgroup. Lateral process (alp) flattened, slightly curved, its laterodistal margin coarsely dentate. Similar in this respect to T. cristagalli, T. implicatus and T. culter sp. nov. Differs from these species by having the mesal process of the posterior coxal fold (pmp) with two rounded distal lobes, visible in anterior view between alp and amp, and by having the tibial spine (ti) not ending in a sharp point.</p><p>Etymology</p><p>The name is a Latin adjective, referring to the undulate/coarsely dentate laterodistal margin of the lateral process of the anterior coxal fold.</p><p>Material examined</p><p>Holotype</p><p>THAILAND: ♂, Krabi Province, Muang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.924995&amp;materialsCitation.latitude=8.275" title="Search Plazi for locations around (long 98.924995/lat 8.275)">Khao Phanom Bencha</a>, 8°16'30" N, 98°55'30" E, 16 Jan. 2009, leg. P. Pimvichai, C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D00087).</p><p>Paratypes THAILAND: 1 ♀, same data as holotype (CUMZ-D00099).</p><p>Description</p><p>Adult male with 63 podous rings, no apodous rings. Length 15 cm, width 8.7 mm. Adult female with 64 podous rings, no apodous rings. Length 14 cm, width 8.5 mm. Colour in life unknown; preserved specimens with head, legs, middorsal metazona, epiproct, paraprocts and hypoproct brown; antennae and metazona dark brown; prozona whitish-brown.</p><p>GONOPODS (Fig. 9 A–E). Anterior coxal fold (ac; Fig. 9A): lateral process (alp) flattened, curving mesad, its laterodistal margin coarsely dentate, terminating in a short spine, tip curving against tip of opposite side, lateral margins with serrate crest (Fig. 9C, arrow); mesal process (amp) much shorter than alp, slender, curving mesad. Posterior coxal fold (pc; Fig. 9B) basally with moderately high lateral paracoxites (px); mesal process (pmp) distally with two rounded lobes, visible in anterior view between alp and amp; lateral process (plp) erect, digitiform. Telopodite (Fig. 9 D–E) leaving coxite over shelf of posterior coxal fold; femoral spine (fe) duplicated, fe 1 relatively long, projecting above top of telopodite, curving backward, ending in slender barb, fe 2 basally broad, situated under fe 1, curving backward, ending in slender barb; tibial spine (ti) long, curving in horizontal plane, not ending in sharp spine; apical part: spatulate lobe (sl) rounded, with slender long spine (sls) at base of apical part, opposite origin of tibial spine; palette (pa) simple, distally with about eleven brownish blepharochaetae (bp).</p><p>DNA barcode</p><p>The GenBank accession number of the barcode of the holotype is KU306520 (voucher code CUMZ-D00087).</p><p>Distribution (Fig. 12)</p><p>Known only from the type locality.</p></div>	https://treatment.plazi.org/id/0385FA5C9617FFA1FDE48977DE26C157	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak;Backeljau, Thierry	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9611FFA0FDF88B4ADE26C5CA.text	0385FA5C9611FFA0FDF88B4ADE26C5CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus ursus Pimvichai & Enghoff & Panha & Backeljau 2016	<div><p>Thyropygus ursus sp. nov.</p><p>urn:lsid:zoobank.org:act: D3E091F2-C075-458B-A9B5-DA35B673309B</p><p>Fig. 10 A–D</p><p>Diagnosis</p><p>A species of the opinatus subgroup. Differs from all other species in the subgroup by having the lateral process of the anterior coxal fold (alp) flattened, broad, apically curved caudad and ending in a short spine, as well as by the first femoral spine (fe 1) being very short and erect.</p><p>Etymology</p><p>The name is a Latin noun in apposition, meaning “bear”, and refers to the (somewhat) bearhead-like profile of the lateral process of the anterior coxal fold (alp).</p><p>Material examined</p><p>Holotype THAILAND: ♂, Krabi Province, Koh Lanta District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.04195&amp;materialsCitation.latitude=7.6555557" title="Search Plazi for locations around (long 99.04195/lat 7.6555557)">Lanta Islands</a>, 7°39'20" N, 99°02'31" E, 2 Jan. 2009, leg. B. Däubl and E. Haring (NMHW-Inv.7855).</p><p>Paratypes THAILAND: 1 ♀, same data as holotype (NMHW).</p><p>Description</p><p>Adult male with 62 podous rings, no apodous rings. Length 11 cm, width 6.4 mm. Adult female with 61 podous rings, no apodous rings. Length 12 cm, width 7.4 mm. Colour in life unknown; preserved specimens with head, antennae, legs, metazona, epiproct, paraprocts and hypoproct brown; prozona yellowish brown.</p><p>GONOPODS (Fig. 10 A–D).Anterior coxal fold (ac; Fig. 10A): lateral process (alp) flattened, broad, apically curved caudad, tip with short spine, lateral margin slightly folded; mesal process (amp) shorter than alp, slender, straight, directed distad, pointed. Posterior coxal fold (pc; Fig. 10B) basally with moderately high paracoxites (px), distally truncate, forming shelf to accommodate telopodite, mesal process (pmp) very small, directed distolaterad. Telopodite (Fig. 10 C–D) leaving coxite over shelf of posterior coxal fold; femoral spine double, fe 1 very short, erect, situated above fe 2, fe 2 very long, slender, curved downward, in situ resting behind alp; tibial spine (ti) long, slender, curving in horizontal plane, its tip close to base of fe; apical part: lamellar lobe (ll) broad, bent down; palette (pa) simple, gutter-like, with small crest (cr) near tip; distally with about twelve brownish blepharochaetae (bp).</p><p>DNA barcode</p><p>The GenBank accession number of the barcode of the holotype is KU306523 (voucher code NMHW-Inv.7855).</p><p>Distribution (Fig. 12)</p><p>Known only from the type locality.</p></div>	https://treatment.plazi.org/id/0385FA5C9611FFA0FDF88B4ADE26C5CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak;Backeljau, Thierry	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
