taxonID	type	description	language	source
0385FA5C9604FFB0FF3988E4D909C2C1.taxon	description	General description of males of the T. opinatus subgroup Head smooth, 5 – 8 supralabral setae, eyes moderate in size, ocelli in 7 – 8 horizontal rows consisting of 11 – 14, 10 – 13, 8 – 11, 7 – 9, 5 – 8, 3 – 6, 1 – 4 and 1 – 3 ocelli, total: 46 – 68. Gnathochilarium: mentum smooth, at most with a few small setae distally and a large, horseshoe-shaped ridge opening distally. Stipites densely covered with spine-like setae, except for an irregular oblique band from c. middle of lateral margin to border of lamella lingualis; long setae present on the distolateral part; males (not females) distally with a small sclerotised ‘ island’ with 1 – 3 spine-like setae in middle of an ovoid, poorly sclerotized, hairless area. Lamellae linguales with three long, apical setae and a number of short, basal, spine-like setae. Sterna smooth, stigmatal grooves distinct, long. Legs relatively long; tip of postfemora, tibiae and tarsi visible from above when legs are extended laterad; coxae of anterior and posterior pair markedly dissimilar, posterior coxae humped; coxae with 2 – 8 setae, prefemora with 3 – 6 setae, femora with 2 – 5 setae in a median row and 2 larger, subapical setae; tarsi typically with 6 – 8 irregularly placed ventral setae and 1 – 2 large dorsal, apical spurs. Postfemora and tibiae without setae. Ventral pads on postfemora and tibiae on all legs, except first three pairs. Gonopods: sternum (st) triangular. Anterior coxal fold (ac) basally slender, becoming broader towards tip, lateral margins diverging; distally with two processes: a lateral process (alp) and a usually smaller mesal process (amp); the shape of these processes is species-specific. Posterior coxal (pc) fold much lower than anterior coxal fold, basally with moderately high lateral paracoxites (px), distally variously modified but always with a smooth area over which the telopodite shaft can slide. Telopodite with a single or double, well-developed femoral spine (fe) and a long, slender, curved tibial spine (ti). Many species with a characteristic spatulate lobe (sl) originating under the base of the tibial spine; lobe sometimes distally rounded, spoon-like, sometimes ending in a large, stout spine. Other species instead with a small and slightly folded lateral lamella (ll). Apical palette (pa) simple, forming a broad gutter, sometimes with a longitudinal crest in the concavity; apically with a row of 7 – 13 brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9600FFB2FD008E36DEA3C2DC.taxon	description	urn: lsid: zoobank. org: act: 0252646 C-AD 4 E- 4 B 8 B- 8 A 7 C-AEEA 80 DD 4 A 72 Fig. 2 A – E	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9600FFB2FD008E36DEA3C2DC.taxon	diagnosis	Diagnosis A species of the opinatus subgroup. Lateral process of anterior coxal fold (alp) regularly curved, basally broad, gradually tapering towards end and ending in sharp point, with a lateral crest near tip. Similar in this respect to T. bifurcus. Differs from this species by having the mesal process of anterior coxal fold (amp) straight, directed distad, by having the mesal process of posterior coxal fold (pmp) slender, directed anteriad, and by having only one femoral spine (fe).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9600FFB2FD008E36DEA3C2DC.taxon	etymology	Etymology This species is named after the organization “ Centre International de Myriapodologie – CIM ” (www. myriapodology. org) in recognition of its immense importance for inspiring and supporting research on myriapods.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9600FFB2FD008E36DEA3C2DC.taxon	materials_examined	Material examined Holotype THAILAND: ♂, Nakhonsrithammarat Province, Lanska District, Namwang Srithammasokrach, 8 ° 20 ' 25 " N, 99 ° 49 ' 42 " E, 16 May 2010, leg. S. Panha, P. Pimvichai and members of the Animal Systematics Research Unit (CUMZ-D 00086). Paratypes THAILAND: 2 ♂♂, 1 ♀, same data as holotype (CUMZ-D 00096); 1 ♂, same data as holotype (ZMUC).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9600FFB2FD008E36DEA3C2DC.taxon	description	Description Adult males with 65 – 68 podous rings, no apodous rings. Length 13 – 16 cm, width 7.6 – 8.3 mm. Adult female with 65 podous rings, no apodous rings. Length 13 cm, width 7.4 mm. Colour in life unknown; preserved specimens with head, antennae, prozona, middorsal metazona and legs brown; metazona, epiproct, paraprocts and hypoproct dark brown. GONOPODS (Fig. 2 A – E). Anterior coxal fold (ac; Fig. 2 A): lateral process (alp) regularly curved, basally broad, gradually tapering towards end and ending in sharp point, with lateral crest near tip (Fig. 2 C, arrow); mesal process (amp) slightly shorter than alp, straight, directed distad. Posterior coxal fold (pc; Fig. 2 B) basally with moderately high lateral paracoxites (px), distally truncate, forming shelf for accommodation of telopodite, mesal process (pmp) slender, directed anteriad. Telopodite (Fig. 2 D – E) leaving coxite over shelf of posterior coxal fold; femoral spine (fe) very long, slender, curving downward, in situ resting between alp and amp; tibial spine (ti) long, slender, curving in horizontal plane, its tip in situ resting close to base of fe; apical part: lamellar lobe (ll) broadly expanded, bent down; palette (pa) simple, gutter-like, with longitudinal rounded crest (cr) at middle; distally with about ten brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9600FFB2FD008E36DEA3C2DC.taxon	materials_examined	DNA barcode The GenBank accession number of the barcode of the holotype is KU 306519 (voucher code CUMZ-D 00086).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9600FFB2FD008E36DEA3C2DC.taxon	distribution	Distribution (Fig. 12) Known only from the type locality.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9600FFB2FD008E36DEA3C2DC.taxon	discussion	Remarks Coexisting with the smaller T. forceps sp. nov.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9602FFACFDFF8937DE26C26C.taxon	description	urn: lsid: zoobank. org: act: 0093 C 39 D-F 1 FC- 4 CAD- 9 DA 0 - 0492 F 45 E 7370 Fig. 3 A – D	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9602FFACFDFF8937DE26C26C.taxon	diagnosis	Diagnosis A species of the opinatus subgroup. Lateral process of anterior coxal fold (alp) flattened, slightly curved, its laterodistal margin coarsely dentate. Similar in this respect to T. cristagalli, T. implicatus and T. undulatus sp. nov. Differs from these species by having the mesal process of posterior coxal fold (pmp) very high, pointed-triangular, directed almost straightly distad, and by having the tibial spine (ti) recurved.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9602FFACFDFF8937DE26C26C.taxon	etymology	Etymology The name is a Latin noun in apposition, meaning “ knife ”, and refers to the knifelike second femoral spine (fe 2).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9602FFACFDFF8937DE26C26C.taxon	materials_examined	Material examined Holotype THAILAND: ♂, Krabi Province, Khlong Thom District, Rorn waterfall, 7 ° 56 ' 06 " N, 99 ° 12 ' 18 " E, 15 Jan. 2009, leg. P. Pimvichai, C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D 00091). Paratypes THAILAND: 3 ♂♂, 2 ♀♀, 1 subadult ♀, same data as holotype (CUMZ-D 00078); 1 ♂, 1 ♀, same data as holotype (ZMUC).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9602FFACFDFF8937DE26C26C.taxon	description	Description Adult males with 58 – 62 podous rings, no apodous rings. Length 12 – 14 cm, width 7.1 – 8.0 mm. Adult females with 59 – 62 podous rings, no apodous rings. Length 11 – 13 cm, width 6.9 – 7.9 mm. Colour in life unknown; preserved specimens with head, antennae, prozona, middorsal metazona and legs brown; metazona, epiproct, paraprocts and hypoproct reddish brown. GONOPODS (Fig. 3 A – D). Anterior coxal fold (ac; Fig. 3 A): lateral process (alp) flattened, slightly curved, its laterodistal margin coarsely dentate, terminating in short, sharp, pointed spine; mesal process (amp) much shorter than alp, directed distad, tip curving mesad, pointed. Posterior coxal fold (pc; Fig. 3 B) basally with lateral paracoxites (px) quite low, distally with two processes: mesal process (pmp) very high, pointed-triangular, directed almost straightly distad; lateral process (plp) much shorter, digitiform, directed distad and slightly laterad. Telopodite (Fig. 3 C – D) leaving coxite over shelf of posterior coxal fold; femoral spine (fe) duplicated, fe 1 projecting above top of telopodite curvature, curving backward, ending in slender barb, fe 2 basally broad, situated under fe 1, curving backward, ending in slender barb; tibial spine (ti) long, slender and recurved (not curving in horizontal plane); apical part: spatulate lobe (sl) slender, rounded; with slender, long spine (sls) at base of apical part, opposite origin of tibial spine; palette (pa) simple, distally with about twelve brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9602FFACFDFF8937DE26C26C.taxon	materials_examined	DNA barcode The GenBank accession number of the barcode of one of the paratypes is KC 519535 (voucher code CUMZ-D 00078).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9602FFACFDFF8937DE26C26C.taxon	distribution	Distribution (Fig. 12) Known only from the type locality.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961CFFAEFDF688A7D851C281.taxon	description	urn: lsid: zoobank. org: act: 11 BAAEC 5 - 404 D- 4 E 15 - 982 C- 2 F 3 E 51 DEA 56 B Figs 4 A – E, 11 B	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961CFFAEFDF688A7D851C281.taxon	diagnosis	Diagnosis A species of the opinatus subgroup. Lateral process of anterior coxal fold (alp) long, slender, regularly curved, tip close to tip of opposite alp, the two together forming a circle. Similar in this respect to T. erectus and T. floweri. Differs from the former by having a telopodite lobe (lo). Particularly similar to T. floweri, differing from it by having the mesal process of posterior coxal fold (pmp) slender, directed distolaterad, laterally with a digitiform process (plp), and by not having pmp strongly developed along the anterior-posterior axis.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961CFFAEFDF688A7D851C281.taxon	etymology	Etymology The name is a Latin noun in apposition, referring to the forceps-like gonopod coxae.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961CFFAEFDF688A7D851C281.taxon	materials_examined	Material examined Holotype THAILAND: ♂, Nakhonsrithammarat Province, Lanska District, Namwang Srithammasokrach, 8 ° 20 ' 25 " N, 99 ° 49 ' 42 " E, 16 May 2010, leg. S. Panha, P. Pimvichai and members of the Animal Systematics Research Unit (CUMZ-D 00092). Paratypes THAILAND: 2 ♂♂, 2 subadult ♂♂, same data as holotype (CUMZ-D 00073); 5 ♂♂, Nakhonsrithammarat Province, Ronpiboon District, Tham Pha Deang temple, 8 ° 14 ' 42 " N, 99 ° 52 ' 03 " E, 14 May 2012, leg. C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D 00093); 2 ♂♂, same data as preceding (ZMUC).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961CFFAEFDF688A7D851C281.taxon	description	Description Adult males with 54 – 62 podous rings, no apodous rings. Length 9 – 12 cm, width 4.6 – 5.8 mm. Overall colour of living animal (Fig. 11 B) brown. Legs, antennae, middorsal metazona, epiproct, paraprocts and hypoproct brownish orange; preserved specimens with head, antennae, prozona, legs, middorsal metazona, epiproct, paraprocts and hypoproct brown; metazona dark brown. GONOPODS (Fig. 4 A – E). Anterior coxal fold (ac; Fig. 4 A): distinctly crenulated along convex part of lateral margin (smooth in specimens from Tham Pha Deang temple), lateral process (alp) long, slender, regularly curved, tip close to tip of opposite alp, the two together forming a circle; mesal process (amp) slightly shorter than alp, directed obliquely disto-mesad (in specimens from Tham Pha Deang temple lateral process (alp) long, slender, crossing over with opposite tip, the two together forming a circle; Fig. 4 B); mesal process (amp) slightly shorter than alp, directed distad, slightly sigmoid. Posterior coxal fold (pc; Fig. 4 C) basally with moderately high lateral paracoxites (px), distally with two processes: mesal process (pmp) slender, directed distolaterad; lateral process (plp) digitiform. Telopodite (Fig. 4 D – E) leaving coxite over shelf of posterior coxal fold; femoral spine (fe) massive, curving in horizontal plane and curving against ti; telopodite distally to fe with large, round lobe (lo) projecting distolaterally; tibial spine (ti) very long, slender, curving in horizontal plane, its tip resting against base of fe; apical part: spatulate lobe (sl) with sharp dark brown spine at tip; palette (pa) simple, distally with about nine brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961CFFAEFDF688A7D851C281.taxon	materials_examined	DNA barcode The GenBank accession number of the barcode of one of the paratypes is KC 519531 (voucher code CUMZ-D 00073).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961CFFAEFDF688A7D851C281.taxon	distribution	Distribution (Fig. 12) Known only from Namwang Srithammasokrach and Tham Pha Deang temple in Nakhonsrithammarat Province.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961CFFAEFDF688A7D851C281.taxon	discussion	Remarks Coexisting with the larger T. cimi sp. nov. at Namwang Srithammasokrach.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961EFFA9FDCF889BDE26C24F.taxon	description	urn: lsid: zoobank. org: act: 5 D 8 F 7830 - 2 E 8 E- 46 E 3 - B 2 BE- 830 FC 31 F 290 D Fig. 5 A – E	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961EFFA9FDCF889BDE26C24F.taxon	diagnosis	Diagnosis A species of the opinatus subgroup. Lateral process of anterior coxal fold (alp) slender, curving mesad; mesal process of anterior coxal fold (amp) as long as alp, slender, straight, directed distad. Similar in these respects to T. demangei. Differs from this species by having mesal margin of amp forming a strong longitudinal crest (lc) in posterior view, by having only one femoral spine (fe) and by having the spatulate lobe (sl) broad and rounded.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961EFFA9FDCF889BDE26C24F.taxon	etymology	Etymology The name is a Latin adjective, referring to the longitudinal crest on the mesal process of the anterior coxal fold.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961EFFA9FDCF889BDE26C24F.taxon	materials_examined	Material examined Holotype THAILAND: ♂, Songkhla Province, Rattaphum District, Srikasorn, 7 ° 00 ' 18 " N, 100 ° 08 ' 24 " E, 12 Jan. 2009, leg. P. Pimvichai, C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D 00094). Paratype THAILAND: ♂, same data as holotype (CUMZ-D 00077).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961EFFA9FDCF889BDE26C24F.taxon	description	Description Adult males with 61 – 66 podous rings, no apodous rings. Length 16 – 17 cm, width 9.2 – 9.5 mm. Colour in life unknown; preserved specimens with head, antennae, prozona and legs brown; metazona, epiproct, paraprocts and hypoproct reddish brown. GONOPODS (Fig. 5 A – E). Anterior coxal fold (ac; Fig. 5 A): lateral process (alp) slender, curving mesad, with lateral serrate crest near tip (Fig. 5 C, arrow); mesal process (amp) as long as alp, slender, straight, directed distad, tip sharp, pointed, in posterior view mesal margin of amp forming strong longitudinal crest (lc). Posterior coxal fold (pc; Fig. 5 B) basally with moderately high lateral paracoxites (px), distally with two processes: mesal process (pmp) slender, directed anteriad; lateral process (plp) short, erect, digitiform; forming shelf for accommodation of telopodite. Telopodite (Fig. 5 D – E) leaving coxite between pmp and plp; femoral spine (fe) long, curved downward, with expanded lamella at base; tibial spine (ti) very long, curving in horizontal plane, tip close to basal part of fe; small spine (ss) at base of apical part, opposite origin of tibial spine; spatulate lobe (sl) broad, rounded; a small accessory lobe (al) at base of sl; palette (pa) simple, gutter-like, with longitudinal crest (cr) at middle, distally with about twelve brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961EFFA9FDCF889BDE26C24F.taxon	materials_examined	DNA barcode The GenBank accession number of the barcode of the paratype is KC 519534 (voucher code CUMZ-D 00077).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961EFFA9FDCF889BDE26C24F.taxon	distribution	Distribution (Fig. 12) Known only from the type locality.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9618FFABFDDB8B3ADE26C6AF.taxon	description	urn: lsid: zoobank. org: act: 01 AC 06 C 9 - EEF 0 - 4211 - 9837 - 28 FE 2 C 12 C 68 E Figs 6 A – D, 11 A	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9618FFABFDDB8B3ADE26C6AF.taxon	diagnosis	Diagnosis A species of the opinatus subgroup. Mesal margin of anterior coxal fold (alp) with fine serrations. Similar in this respect to T. opinatus. Differs from this species by having the lateral process (alp) slender, regularly curved and by having the mesal process (amp) broadly expanded and forming a strong longitudinal crest (lc) in posterior view.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9618FFABFDDB8B3ADE26C6AF.taxon	etymology	Etymology The species is named after the Royal Thai Navy, in recognition of their kind assistance which enabled us to pursue the necessary fieldwork at the type locality, and also after Chulalongkorn University where pink is the symbolic colour of the university, indirectly referring to the pink legs of the species; the species name is treated as a noun in apposition.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9618FFABFDDB8B3ADE26C6AF.taxon	materials_examined	Material examined Holotype THAILAND: ♂, Phang-Nga Province, Khuraburi District, Surin Islands, 9 ° 24 ' 20 " N, 97 ° 51 ' 38 " E, 7 Apr. 2012, leg. S. Panha, P. Pimvichai and members of the Animal Systematics Research Unit (CUMZ-D 00095). Paratypes THAILAND: 5 ♂♂, 1 ♀, same data as holotype (CUMZ-D 00089); 2 ♂♂, same data as holotype (ZMUC).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9618FFABFDDB8B3ADE26C6AF.taxon	description	Description Adult males with 58 – 62 podous rings, no apodous rings. Length 11 – 12 cm, width 6.2 – 7.2 mm. Adult female with 60 podous rings, no apodous rings. Length 14 cm, width 8.0 mm. Overall colour of living animal (Fig. 11 A) dark brown, shiny. Legs and antennae pink; preserved specimen with head, antennae, prozona, legs, epiproct, paraprocts and hypoproct brown; metazona dark brown. GONOPODS (Fig. 6 A – D). Anterior coxal fold (ac; Fig. 6 A): lateral process (alp) slender, regularly curved, mesal margin with fine serrations; mesal process (amp) almost as long as alp, broadly expanded, apically sharp, straight distad, in posterior view mesal margin of amp forming strong longitudinal crest (lc). Posterior coxal fold (pc; Fig. 6 B) basally with moderately high lateral paracoxites (px), distally with two processes: mesal process (pmp) shorter than plp, curving behind plp; lateral process (plp) flattened, directed distad. Telopodite (Fig. 6 C – D) leaving coxite between pmp and plp; femoral spine (fe) very long, curving almost exclusively in horizontal plane; tibial spine (ti) very long, curving in horizontal plane close to basal part of fe; spatulate lobe (sl) terminating in sharp spine; palette (pa) simple, gutterlike, distally with about twelve brownish blepharochaetae (bp), laterally with few sharp spines.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9618FFABFDDB8B3ADE26C6AF.taxon	materials_examined	DNA barcode The GenBank accession number of the barcode of one of the paratype is KU 306522 (voucher code CUMZ-D 00089).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9618FFABFDDB8B3ADE26C6AF.taxon	distribution	Distribution (Fig. 12) Known only from the type locality.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961BFFA5FDDA8CE2DE26C497.taxon	description	urn: lsid: zoobank. org: act: 2 F 5 A 25 A 4 - 44 CC- 4 B 93 - AC 17 - 9 A 9 B 6 E 6 E 9 E 1 F Fig. 7 A – E	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961BFFA5FDDA8CE2DE26C497.taxon	diagnosis	Diagnosis A species of the opinatus subgroup. Differs from all other species in the subgroup by having the lateral process of the anterior coxal fold (alp) regularly curved, terminating in a sharp spine pointing slightly distad, and by having the tibial spine (ti) short and flattened.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961BFFA5FDDA8CE2DE26C497.taxon	etymology	Etymology The name is a Latin noun in apposition and refers to the flattened tibial spine (ti).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961BFFA5FDDA8CE2DE26C497.taxon	materials_examined	Material examined Holotype THAILAND: ♂, Krabi Province, Muang District, Tham Sua temple, 8 ° 07 ' 30 " N, 98 ° 55 ' 30 " E, 15 Jan. 2012, leg. P. Pimvichai, C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D 00088). Paratypes THAILAND: 3 ♀♀, same data as holotype (CUMZ-D 00097).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961BFFA5FDDA8CE2DE26C497.taxon	description	Description Adult male with 64 podous rings, no apodous rings. Length 13 cm, width 7.6 mm. Adult females with 60 – 63 podous rings, no apodous rings. Length 13 – 14 cm, width 7.8 – 8.6 mm. Colour in life unknown; preserved specimens with head, antennae, legs, metazona, epiproct, paraprocts and hypoproct brown; prozona yellowish brown. GONOPODS (Fig. 7 A – E). Anterior coxal fold (ac; Fig. 7 A): lateral process (alp) regularly curved, terminating in sharp spine pointing slightly distad, with lateral short, sharp crest at middle of alp (Fig. 7 C, arrow); mesal process (amp) slightly shorter than alp, flattened, straight, directed distad. Posterior coxal fold (pc; Fig. 7 B) basally with moderately high lateral paracoxites (px), distally truncate, forming shelf for accommodation of telopodite, mesal process (pmp) triangular, directed distad, lateral process (plp) erect, digitiform. Telopodite (Fig. 7 D – E) leaving coxite over shelf of posterior coxal fold; femoral spine double, fe 1 very long, curved downward, situated above fe 2, in situ resting at middle of alp, fe 2 long, very slender, curved downward; tibial spine (ti) flattened, short, curving mesad; apical part: spatulate lobe (sl) small, rounded; with slender, long spine (sls) at base of apical part; palette (pa) simple, gutter-like, distally with about nine brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961BFFA5FDDA8CE2DE26C497.taxon	materials_examined	DNA barcode The GenBank accession number of the barcode of the holotype is KU 306521 (voucher code CUMZ-D 00088).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C961BFFA5FDDA8CE2DE26C497.taxon	distribution	Distribution (Fig. 12) Known only from the type locality.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9615FFA7FDE08E8ADE26C31C.taxon	description	urn: lsid: zoobank. org: act: 6804 DF 2 C- 7787 - 46 FE- 9185 - 2 E 5 C 40 AF 5 ED 3 Fig. 8 A – D	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9615FFA7FDE08E8ADE26C31C.taxon	diagnosis	Diagnosis A species of the opinatus subgroup. Mesal process of anterior coxal fold (amp) directed slightly obliquely disto-mesad, tip overlapping tip of opposite amp. Similar in this respect to T. chelatus. Differs from this species by having amp as long as alp and by having the lateral process of the posterior coxal fold (plp) as a massive, broad lobe.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9615FFA7FDE08E8ADE26C31C.taxon	etymology	Etymology The species is named in honour of Chirasak Sutcharit in recognition of his devotion to collecting millipedes.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9615FFA7FDE08E8ADE26C31C.taxon	materials_examined	Material examined Holotype THAILAND: ♂, Phetchaburi Province, Kaeng Krachan District, Kaeng Krachan, 12 ° 54 ' 07 " N, 99 ° 35 ' 56 " E, 14 May 2012, leg. C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D 00090). Paratypes THAILAND: 2 ♂♂, 2 ♀♀, 1 subadult ♀, same data as holotype (CUMZ-D 00098); 1 ♂, same data as holotype (ZMUC).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9615FFA7FDE08E8ADE26C31C.taxon	description	Description Adult males with 63 – 65 podous rings, no apodous rings. Length 10 – 11 cm, width 5.4 – 5.7 mm. Adult females with 61 – 63 podous rings, no apodous rings. Length 12 – 13 cm, width 6.7 – 6.9 mm. Colour in life unknown; preserved specimens with head, antennae, prozona, legs, epiproct, paraprocts and hypoproct brown; metazona reddish brown. GONOPODS (Fig. 8 A – D). Anterior coxal fold (ac; Fig. 8 A): lateral process (alp) slender, regularly curved; mesal process (amp) as long as alp, directed slightly obliquely disto-mesad, tip overlapping tip of opposite amp. Posterior coxal fold (pc; Fig. 8 B) basally with moderately high lateral paracoxites (px), distally with two processes: mesal process (pmp) a rounded lobe; lateral process (plp) a massive, broad lobe, projecting laterad. Telopodite (Fig. 8 C – D) leaving coxite between pmp and plp; femoral spine (fe) long, curved downward; telopodite distally to fe with large, round lobe (lo) projecting distolaterally; tibial spine (ti) very long, curving in horizontal plane, tip close to basal part of lo; spatulate lobe (sl) broad, rounded; palette (pa) simple, gutter-like, distally with about ten brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9615FFA7FDE08E8ADE26C31C.taxon	materials_examined	DNA barcode The GenBank accession number of the barcode of the holotype is KU 306524 (voucher code CUMZ-D 00090).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9615FFA7FDE08E8ADE26C31C.taxon	distribution	Distribution (Fig. 12) Known only from the type locality.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9617FFA1FDE48977DE26C157.taxon	description	urn: lsid: zoobank. org: act: 972296 F 7 - CF 7 B- 4 AD 4 - A 2 AA- 3 DCE 22596 B 3 E Fig. 9 A – E	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9617FFA1FDE48977DE26C157.taxon	diagnosis	Diagnosis A species of the opinatus subgroup. Lateral process (alp) flattened, slightly curved, its laterodistal margin coarsely dentate. Similar in this respect to T. cristagalli, T. implicatus and T. culter sp. nov. Differs from these species by having the mesal process of the posterior coxal fold (pmp) with two rounded distal lobes, visible in anterior view between alp and amp, and by having the tibial spine (ti) not ending in a sharp point.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9617FFA1FDE48977DE26C157.taxon	etymology	Etymology The name is a Latin adjective, referring to the undulate / coarsely dentate laterodistal margin of the lateral process of the anterior coxal fold.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9617FFA1FDE48977DE26C157.taxon	materials_examined	Material examined Holotype THAILAND: ♂, Krabi Province, Muang District, Khao Phanom Bencha, 8 ° 16 ' 30 " N, 98 ° 55 ' 30 " E, 16 Jan. 2009, leg. P. Pimvichai, C. Sutcharit and members of the Animal Systematics Research Unit (CUMZ-D 00087). Paratypes THAILAND: 1 ♀, same data as holotype (CUMZ-D 00099).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9617FFA1FDE48977DE26C157.taxon	description	Description Adult male with 63 podous rings, no apodous rings. Length 15 cm, width 8.7 mm. Adult female with 64 podous rings, no apodous rings. Length 14 cm, width 8.5 mm. Colour in life unknown; preserved specimens with head, legs, middorsal metazona, epiproct, paraprocts and hypoproct brown; antennae and metazona dark brown; prozona whitish-brown. GONOPODS (Fig. 9 A – E). Anterior coxal fold (ac; Fig. 9 A): lateral process (alp) flattened, curving mesad, its laterodistal margin coarsely dentate, terminating in a short spine, tip curving against tip of opposite side, lateral margins with serrate crest (Fig. 9 C, arrow); mesal process (amp) much shorter than alp, slender, curving mesad. Posterior coxal fold (pc; Fig. 9 B) basally with moderately high lateral paracoxites (px); mesal process (pmp) distally with two rounded lobes, visible in anterior view between alp and amp; lateral process (plp) erect, digitiform. Telopodite (Fig. 9 D – E) leaving coxite over shelf of posterior coxal fold; femoral spine (fe) duplicated, fe 1 relatively long, projecting above top of telopodite, curving backward, ending in slender barb, fe 2 basally broad, situated under fe 1, curving backward, ending in slender barb; tibial spine (ti) long, curving in horizontal plane, not ending in sharp spine; apical part: spatulate lobe (sl) rounded, with slender long spine (sls) at base of apical part, opposite origin of tibial spine; palette (pa) simple, distally with about eleven brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9617FFA1FDE48977DE26C157.taxon	materials_examined	DNA barcode The GenBank accession number of the barcode of the holotype is KU 306520 (voucher code CUMZ-D 00087).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9617FFA1FDE48977DE26C157.taxon	distribution	Distribution (Fig. 12) Known only from the type locality.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9611FFA0FDF88B4ADE26C5CA.taxon	description	urn: lsid: zoobank. org: act: D 3 E 091 F 2 - C 075 - 458 B-A 9 B 5 - DA 35 B 673309 B Fig. 10 A – D	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9611FFA0FDF88B4ADE26C5CA.taxon	diagnosis	Diagnosis A species of the opinatus subgroup. Differs from all other species in the subgroup by having the lateral process of the anterior coxal fold (alp) flattened, broad, apically curved caudad and ending in a short spine, as well as by the first femoral spine (fe 1) being very short and erect.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9611FFA0FDF88B4ADE26C5CA.taxon	etymology	Etymology The name is a Latin noun in apposition, meaning “ bear ”, and refers to the (somewhat) bearhead-like profile of the lateral process of the anterior coxal fold (alp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9611FFA0FDF88B4ADE26C5CA.taxon	materials_examined	Material examined Holotype THAILAND: ♂, Krabi Province, Koh Lanta District, Lanta Islands, 7 ° 39 ' 20 " N, 99 ° 02 ' 31 " E, 2 Jan. 2009, leg. B. Däubl and E. Haring (NMHW-Inv. 7855). Paratypes THAILAND: 1 ♀, same data as holotype (NMHW).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9611FFA0FDF88B4ADE26C5CA.taxon	description	Description Adult male with 62 podous rings, no apodous rings. Length 11 cm, width 6.4 mm. Adult female with 61 podous rings, no apodous rings. Length 12 cm, width 7.4 mm. Colour in life unknown; preserved specimens with head, antennae, legs, metazona, epiproct, paraprocts and hypoproct brown; prozona yellowish brown. GONOPODS (Fig. 10 A – D). Anterior coxal fold (ac; Fig. 10 A): lateral process (alp) flattened, broad, apically curved caudad, tip with short spine, lateral margin slightly folded; mesal process (amp) shorter than alp, slender, straight, directed distad, pointed. Posterior coxal fold (pc; Fig. 10 B) basally with moderately high paracoxites (px), distally truncate, forming shelf to accommodate telopodite, mesal process (pmp) very small, directed distolaterad. Telopodite (Fig. 10 C – D) leaving coxite over shelf of posterior coxal fold; femoral spine double, fe 1 very short, erect, situated above fe 2, fe 2 very long, slender, curved downward, in situ resting behind alp; tibial spine (ti) long, slender, curving in horizontal plane, its tip close to base of fe; apical part: lamellar lobe (ll) broad, bent down; palette (pa) simple, gutter-like, with small crest (cr) near tip; distally with about twelve brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9611FFA0FDF88B4ADE26C5CA.taxon	materials_examined	DNA barcode The GenBank accession number of the barcode of the holotype is KU 306523 (voucher code NMHW-Inv. 7855).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
0385FA5C9611FFA0FDF88B4ADE26C5CA.taxon	distribution	Distribution (Fig. 12) Known only from the type locality.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2016): A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae). European Journal of Taxonomy 199: 1-37, DOI: 10.5852/ejt.2016.199
