identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038287A1F312FFAC6A5DFC3AFBE9CAAF.text	038287A1F312FFAC6A5DFC3AFBE9CAAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichogaster (Diplothecodrilus) annae (Horst 1983)	<div><p>Dichogaster (Diplothecodrilus) annae (Horst, 1983)</p> <p>Benhamia annae Horst 1893, p. 32.</p> <p>Dichogaster (Diplothecodrilus) annae: Csuzdi 1995, p. 112; Csuzdi 1996, p. 357; Csuzdi 2000, p. 59 (for complete synonymy).</p> <p>Material examined</p> <p>HNHM AF/3541, 2 exemplars, BMNH 1997.1552 – 1559, 8 exemplars. D. R. Congo, Kivu Region, Butembo, in the stem of a banana tree, leg. Gasana N’Doli, 15 July 1973.</p> </div>	https://treatment.plazi.org/id/038287A1F312FFAC6A5DFC3AFBE9CAAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
038287A1F312FFAC6ABBFD64FD41CDE7.text	038287A1F312FFAC6ABBFD64FD41CDE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichogaster Beddard 1888	<div><p>Genus Dichogaster Beddard, 1888</p> <p>Subgenus Dichogaster (Diplothecodrilus) Csuzdi, 1996 Dichogaster Beddard 1888, p. 251, Michaelsen 1900, p. 334 (part), Omodeo 1955, p. 224 (part), Omodeo 1958, p. 61 (part). Dichogaster (Diplothecodrilus): Csuzdi 1996, p. 355.</p> </div>	https://treatment.plazi.org/id/038287A1F312FFAC6ABBFD64FD41CDE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
038287A1F312FFAA6A57FB62FE92CA7D.text	038287A1F312FFAA6A57FB62FE92CA7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichogaster (Diplothecodrilus) cagnii Cognetti 1906	<div><p>Dichogaster (Diplothecodrilus) cagnii Cognetti, 1906</p> <p>(Figure 1A–D)</p> <p>Dichogaster cagnii Cognetti 1906, p. 2; Cognetti 1909, p. 12.</p> <p>Dichogaster (Diplothecodrilus) cagnii: Csuzdi 1995, p. 112; Csuzdi 1996, p. 356.</p> <p>Material examined</p> <p>Dichogaster cagnii Holotype, MZUT OL56, Ruwenzori, Ibanda, Toro (1800 m), leg. Duca degli Abruzzi, July 1906. Dichogaster cagnii Paratype, MZUT OL54, Ruwenzori, Lago Kianga, leg. Duca degli Abruzzi, July 1906. Dichogaster cagnii Paratype, MZUT OL 55, Ruwenzori, Toro, leg. Duca degli Abruzzi, July 1906.</p> <p>New records. HNHM AF/3549, one exemplar, BMNH 1997.1605 – 1606 two exemplars. D. R. Congo, Kivu Region, between Kagundu and Gitovu, leg. Gasana N’Doli, 17 July 1973. HNHM AF/3569, one exemplar, BMNH 1997.1563 – 1564, two exemplars. D. R. Congo, Kivu, Butembo, in the stem of a banana tree, leg. Gasana N’Doli, 15 July 1973. HNHM AF/3556, two exemplars, BMNH 1997.1485 – 1488, four exemplars. D. R. Congo, Kivu, Mewa, leg. Gasana N’Doli, 17 July 1973. HNHM AF/3554, three exemplars, BMNH 1997.1500 – 1505, six exemplars. D. R. Congo, Kivu, Katana, in stems of banana trees, leg. Gasana N’Doli, 05 August 1973. HNHM AF/3562, five exemplars, BMNH 1997.1596 – 1604, nine exemplars. D. R. Congo, Kivu, Lwiro, leg. Gasana N’Doli, 10 May 1973. HNHM AF/3546, five exemplars, BMNH, 1997.1583– 1592, 10 exemplars. D. R. Congo, Kivu, river Mululu, leg. Gasana N’Doli, 15 July 1973.</p> <p>Diagnosis</p> <p>Length 30–63 mm, D. 2–2.5 mm, no. of segments 110–120. Pigmentation brownish. First dorsal pore in 5/6. Clitellum 13–20. ♀ 14 paired before setae a–a. Prostatic pores 17, 19, genital field oblong. Spermathecal pores 7/8, 8/9, spermathecae with elongated ampoule, and an equally long duct. The distal part of the ampoule bears a round and stalked diverticulum. Gizzard in 5/6, last pair of hearts in 12. Calciferous glands in 15–17, their size slightly increasing backwards. Excretory system meroic with five or six meronephridia per side. Penial setae are of two types, larger L. 1.1– 1.3 mm, D. 0.02 mm, tip sharply pointed, ornamentation strong spines. Smaller penial setae L. 0.60–0.65 mm, D. 0.01 mm, tip rounded, ornamentation lacking.</p> <p>Remarks</p> <p>This species was described by Cognetti (1906) with a unique type of penial setae, therefore previously I placed it in the congica species group (Csuzdi 1996). An examination of the type material (MZUT OL 54, OL 55) revealed that the species possesses two different types of penial setae and consequently it belongs in the bolaui species group.</p></div> 	https://treatment.plazi.org/id/038287A1F312FFAA6A57FB62FE92CA7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
038287A1F314FFAB6A39FBB7FBDCCAF0.text	038287A1F314FFAB6A39FBB7FBDCCAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichogaster (Diplothecodrilus) foetida Michaelsen 1932	<div><p>Dichogaster (Diplothecodrilus) foetida Michaelsen, 1932</p> <p>(Figure 2A–D)</p> <p>Dichogaster foetida Michaelsen 1932, p. 5.</p> <p>Dichogaster rubella Michaelsen 1935, p. 64; 1936b, p. 46; 1937a, p. 508; 1937b, p. 1, syn. nov.</p> <p>Dichogaster (Diplothecodrilus) rubella: Csuzdi 1995, p. 13; Csuzdi 1996, p. 357; Csuzdi 2000, p. 66.</p> <p>Material examined</p> <p>Dichogaster rubella, ZMUH V 12615, Kamatembe 2100 m. Dichogaster rubella, BMNH 1934.4.23.1 Uganda, Kigezi, Kayansa forest 7500 ft (22,500 m), leg. Pittman.</p> <p>New records. HNHM AF/3540, one exemplar, AF3445, two exemplars, BMNH 1997.1475 – 1476, two exemplars. D. R. Congo, Kivu Region, Ngungu, leg. Gazsana N’Doli 16 July 1973. BMNH 1997.1499, one exemplar. D. R. Congo, Kivu Region, Ngungu, in the swamp, leg. Gasana N’Doli 16 July 1973. BMNH 1997.1519, one exemplar. D. R. Congo, Kivu Region, Ngungu, in the swamp, leg. Gasana N’Doli 16 July 1973. BMNH 1997.1520 – 1523, four exemplars. D. R. Congo, Kivu Region, Ngungu, leg. Gasana N’Doli 16 July 1973. HNHM AF/3553, one exemplar, BMNH 1997.1545 – 1547, three exemplars. D. R. Congo, Kivu Region, Kajabwe, leg. Gasana N’Doli 30 May 1973.</p> <p>Diagnosis</p> <p>Length 50–80 mm, D. 3–4 mm, no. of segments 74–115. Pigmentation brownish. First dorsal pore in 5/6. Clitellum 13–20, 21. ♀ 14 unpaired ventral-median. Prostatic pores 17, 19, genital field oblong. Spermathecal pores 7/8, 8/9, ampoule of the spermatheca elongated with small distal part hardly detached from the long duct. The distal part of the ampoule bears a round and stalked diverticulum. Gizzard in 5/ 6, last pair of hearts in 12. Calciferous glands in 15–17 of equal size. Excretory system meroic with six or seven meronephridia per side. Penial seta L. 1.8–2 mm, D. 0.01 mm, tip hook-shaped, ornamentation a few very fine teeth.</p> <p>Remarks</p> <p>Dichogaster (Dt.) rubella was described by Michaelsen (1935). Its description agrees completely with that of D. (Dt.) foetida Michaelsen, 1932 except in the size of the penial setae, which were somewhat larger (1.8–2 mm in D. rubella and 1.6 mm in D. foetida). However, the type specimen of D. (Dt.) foetida was extremely macerated and from the four penial setal bundles only one seta could have been examined. As the mature setae are usually protruding from the prostatic pores, in macerated specimens they are frequently lost. In this case only the immature replacement setae could be found inside the body and naturally they are of shorter length.</p> <p>According to the original descriptions both species possess smooth penial setae, but Michaelsen (1935, p. 65) writes ‘‘ Da die sehr zarte Borste sich nicht rein aus der Borstenscheide herauspräparieren und frei untersuchen liess, so möchte ich sicher behaupten, dass eine Ornamentierung fehle.’’ (Since this fragile seta could not be prepared from the setal sac and completely cleared and freely examined, I’m not sure whether ornamentation is lacking.) Examining the penial setae of the present material at higher magnification a few very small teeth were observed at the middle of the seta (Figure 2C). This fine ornamentation can easily be overlooked by light microscopy.</p></div> 	https://treatment.plazi.org/id/038287A1F314FFAB6A39FBB7FBDCCAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
038287A1F315FFA96ABCFB42FBE1CA42.text	038287A1F315FFA96ABCFB42FBE1CA42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichogaster (Diplothecodrilus) gasanai Csuzdi 2008	<div><p>Dichogaster (Diplothecodrilus) gasanai sp. nov.</p> <p>(Figure 3A–D)</p> <p>Material examined</p> <p>Holotype. BMNH 1997.1561. D. R. Congo, Kivu Region, Butembo, in stems of banana trees, leg. Gasana N’Doli, 15 July 1973.</p> <p>Paratypes. HNHM AF/3564, one exemplar, BMNH 1997.1562, one exemplar. Locality same as that of the Holotype. HNHM AF/3561, ten exemplars, BMNH 1997.1618 – 1629, 12 exemplars D. R. Congo, Kivu Region, Bugabo, leg. Gasana N’Doli, 25 July 1973. HNHM AF/3550, four exemplars, BMNH 1997.1607 – 1616, ten exemplars D. R. Congo, Kivu Region, between Kagundu and Gitovu, leg. Gasana N’Doli, 17 July 1973. HNHM AF/3551, four exemplars, BMNH 1997.1574 – 1582, nine exemplars. D. R. Congo, Kivu Region, River Mululu, leg. Gasana N’Doli, 15. July 1973. HNHM AF/3557, two exemplars, BMNH 1997.1479 – 1484, six exemplars. D. R. Congo, Kivu Region, Mewa, leg. Gasana N’Doli, 17 July 1973.</p> <p>Derivatio nominis</p> <p>The specific epithet is referring to the collector of this interesting material:Gasana N’Doli.</p> <p>Diagnosis</p> <p>Length 32–56 mm, D. 1.8–2 mm, no. of segments 75–112. Pigmentation lacking. First dorsal pore in 5/6. Clitellum 12– K 21. ♀ 14 paired between setae a–a. Prostatic pores 17, 19. Spermathecal pores 7/8, 8/9, spermathecae with elongated ampoule, and a shorter duct, half as long as the ampoule. Gizzard in 5/6, last pair of hearts in 13. Calciferous glands in 15–17, with slightly backwards increasing size. Excretory system meroic with four meronephridia per side. Penial seta L. 1.3–1.4 mm, D. 0.006 mm, tip sharply pointed, ornamentation small teeth.</p> <p>External characters</p> <p>Holotype. Length 48 mm, D. 2 mm, number of segments 96.</p> <p>Paratypes. 32–56 mm in length, 1.8–2 mm in diameter, number of segments 75–112. Colour pale, pigmentation lacking. Prostomium schizolobous. First dorsal pore in 5/6. Setae all ventral, setal arrangement after the clitellum aa:ab:bc:cd: dd52.5:1.25:3.75:1:25. Clitellum extends over segments 12– K 21, with a butterfly-shaped male field. Prostatic pores paired on 17, 19 in the setal line b. Male pores externally not visible, they are on 18, in the seminal grooves. Female pores paired on 14 between setae a–a. Spermathecal pores paired in 7/8, 8/9 near to the setae b. Porophores absent (Figure 3A).</p> <p>Internal characters</p> <p>The first visible septum 6/7, thickened septa lacking. Oesophageal gizzards small, in segment 5 and 6. Three pairs of calciferous glands located in segment 15–17, their size slightly increasing backwards. Excretory system meroic, with four meronephridia on each side. Paired hearts are present in segments 10–13. Typhlosole small, begins in segment 20 and accompanied by a pair of auxiliary typhlosoles between segments 27 and 33. Testes in 10 and 11 enclosed in perioesophageal testis sacs. Seminal vesicles in 11, 12, and a pair of large ovaries in segment 13. Seminal duct convoluted, discharging in segment 18. Two pairs of prostatic glands are present in 17 and 19, slightly intruding on the neighbouring segments and each with a penial setal sac containing a mature and several immature penial setae. The mature penial seta is somewhat undulated, about 1.4–1.5 mm long and 0.006 mm wide with a slightly hooked tip. Ornamentation consists of scattered small teeth (Figure 3B,C). There are two pairs of spermathecae in segment 8 and 9. The ampoule is elongated dumb-bell-shaped and on its basis bears a small rounded diverticulum. The duct is narrow and half as long as the ampoule (Figure 3D).</p> <p>Remarks</p> <p>This species belongs to the affinis species group and, by the penial setae, resembles D. (Dt.) crateris Michaelsen, 1912. Nevertheless it differs from the latter (and from all other member of the affinis group) in the more backward location of the last pair of hearts, in the position of the clitellum and in the shape of the spermathecae.</p> </div>	https://treatment.plazi.org/id/038287A1F315FFA96ABCFB42FBE1CA42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
038287A1F317FFA76A83FBD3FD83CC90.text	038287A1F317FFA76A83FBD3FD83CC90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichogaster (Diplothecodrilus) paulclarki Csuzdi 2008	<div><p>Dichogaster (Diplothecodrilus) paulclarki sp. nov.</p> <p>(Figure 4A–D)</p> <p>Material examined</p> <p>Holotype. BMNH 1997.1628 D. R. Congo, Kivu Region, Bugabo, leg. Gasana N’Doli, 25 July 1973.</p> <p>Paratypes. HNHM AF/3544, one exemplar, BMNH 1997.1629, one exemplar. Locality same as that of the holotype. HNHM AF/3571, one exemplar, BMNH 1997.1593, one exemplar. D. R. Congo, Kivu Region, River Mululu, leg. Gasana N’Doli, 15 July 1973.</p> <p>Diagnosis</p> <p>Length 41–50 mm, D. 2–2.3 mm, no. of segments 95–115. Pigmentation lacking. First dorsal pore in 5/6. Clitellum 13–20. ♀ 14 paired before setae a–a. Prostatic pores 17, 19. Spermathecal pores 7/8, 8/9, spermathecae with a subdivided irregular ampoule, and an equally long duct. Gizzard in 5/6, last pair of hearts in 13. Calciferous glands in 15–17 of equal size. Excretory system meroic with four meronephridia per side.</p> <p>Penial seta L. 1.7–1.8 mm, D. 0.015 mm, tip sharply pointed, ornamentation small thorns.</p> <p>Derivatio nominis</p> <p>The specific epithet refers to Dr Paul Clark (The Natural History Museum, London) for always being helpful throughout my stay in London.</p> <p>External characters</p> <p>Holotype. Length 46 mm, diameter 2 mm, no. of segments 115.</p> <p>Paratypes. 41–50 mm in length, 2–2.3 mm in diameter, number of segments 95–115. Colour pale, pigmentation lacking. Prostomium schizolobous. First dorsal pore in 5/ 6. Setae all ventral, setal arrangement after the clitellum aa:ab:bc:cd:dd54:1.1:4:1:34. Clitellum extends over segments 13–20, male field oblong. Prostatic pores paired on 17, 19 in the setal line b. Male pores externally not visible, they are on 18, in the seminal groves. Female pores paired on 14 just before the setae aa. Spermathecal pores paired in 7/8, 8/9 near to the setae b. Porophores absent (Figure 4A).</p> <p>Internal characters</p> <p>The first visible septum 6/7, thickened septa lacking. Oesophageal gizzards small, in segment 5 and 6. Three pairs of calciferous glands in segment 15–17 of equal size. Excretory system meroic, with four meronephridia on each side. Paired hearts are present in segments 10–13. Typhlosole very small, begins in segment 21, auxiliary typhlosole lacking. Testes in 10 and 11 enclosed in perioesophageal testis sacs. Seminal vesicles in 11, 12, and a pair of large ovaries in segment 13. Seminal duct convoluted, discharging in segment 18. Two pairs of prostatic glands are present in 17 and 19, slightly intruding into the neighbouring segments and each with a penial setal sac containing a mature and several immature penial setae. The mature penial seta is undulated at the ectal third, about 1.7–1.8 mm long and 0.015 mm wide with a sharply pointed tip. Ornamentation consists of scattered thorns (Figure 4B,C). There are two pairs of spermathecae in segment 8 and 9. The ampoule is subdivided. The ental part small and turns back to the pear-shaped ectal part which bears a small rounded diverticulum. The duct is narrow and half as long as the ampoule (Figure 4D).</p> <p>Remarks</p> <p>This species belongs to the affinis species group and, by the penial setae, resembles D. (Dt.) kigogoana Stephenson, 1933. Nevertheless it differs from the latter in the position of the last pair of hearts, in the shape of the spermathecae and in the absence of a thickened septa.</p> </div>	https://treatment.plazi.org/id/038287A1F317FFA76A83FBD3FD83CC90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
038287A1F319FFA46A49FCADFC88C867.text	038287A1F319FFA46A49FCADFC88C867.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichogaster (Diplothecodrilus) silvestris (Michaelsen 1896)	<div><p>Dichogaster (Diplothecodrilus) silvestris (Michaelsen, 1896)</p> <p>(Figure 5A–D)</p> <p>Benhamia silvestris Michaelsen 1896, p. 24.</p> <p>Dichogaster silvestris: Michaelsen 1900, p. 352, Michaelsen 1905, p. 310. Dichogaster seriatospina Michaelsen 1912, p. 30 syn. nov.</p> <p>Dichogaster anguana Michaelsen 1915, p. 209 syn. nov.</p> <p>Dichogaster (Diplothecodrilus) seriatospina: Csuzdi 1995, p. 114, Csuzdi 1996, p. 357. Dichogaster (Diplothecodrilus) anguana: Csuzdi 1995, p. 112, Csuzdi 1996, p. 357. Dichogaster (Diplothecodrilus) silvestris: Csuzdi 1995, p. 114, Csuzdi 1996, p. 357, Csuzdi 2000, p. 67.</p> <p>Material examined</p> <p>Dichogaster silvestris Holotype, ZMUH V295, D.-O. Afrika, Ru-nsosro, leg. Stuhlmann.</p> <p>New records. HNHM 3537, four exemplars, BMNH 1997.1468 – 1474, seven exemplars, BMNH 1997.1524 – 1525, two exemplars. D. R. Congo, Kivu Region, Ngungu, leg. Gasana N’Doli, 16 July 1973, HNHM 3547, one exemplar, BMNH 1997.1509 – 1511, three exemplars. D. R. Congo, Kivu Region, Katana, in stems of banana trees, leg. Gasana N’Doli, 5 August 1973. HNHM AF/3552, two exemplars, BMNH 1997.1648 – 1650, three exemplars. D. R. Congo, Kivu Region, Idjiri island, leg. Gasana N’Doli, 2 July 1973. HNHM AF/3559, one exemplar, BMNH 1997.1677 – 1678, two exemplars. D. R. Congo, Kivu Region, Lwiro, leg. Gasana N’Doli, 19 January 1973. HNHM AF/3563, three exemplars, BMNH 1997.1493 – 1498, six exemplars. D. R. Congo, Kivu Region, Ngungu, in the swamp, leg. Gasana N’Doli, 16 July 1973. HNHM AF/3567, 10 exemplars, BMNH 1997.1652 – 1661, ten exemplars. D. R. Congo, Kivu Region, Lwiro, leg. Gasana N’Doli, 16 February 1973.</p> <p>Diagnosis</p> <p>Length 35–60 mm, D. 3–4 mm, no. of segments 85–112. Pigmentation brownish. First dorsal pore in 5/6. Clitellum 13–20, K 21, 21. ♀ 14 paired between setae a–a. Prostatic pores 17, 19, genital field oblong with rounded edges. Spermathecal pores 7/8, 8/9, ampoule of the spermatheca bulb-shaped with small distal part bearing a round and stalked diverticulum. The duct is slightly convoluted and much longer than the ampoule. Gizzard in 5/6, last pair of hearts in 12. Calciferous glands in 15–17 of equal size. Excretory system meroic with six meronephridia per side. Penial seta L. 4–5 mm, D. 0.02 mm, ectal third undulated, tip hook-shaped, ornamentation fine teeth.</p> <p>Remarks</p> <p>This species was described on a pre-adult specimen with not fully developed clitellum on segment 13–19. Dichogaster (Dt.) seriatospina Michaelsen, 1912 differs from D. (Dt.) silvestris only in the position of the clitellum therefore it must be regarded as a synonym of D. (Dt.) silvestris. Dichogaster anguana Michaelsen, 1915 was also described on a juvenile specimen; however, its spermathecae and the structure of the penial setae are identical to those of D. (Dt.) silvestris. It differs only in the length of the penial setae, which is because of the juvenile stage of the type specimen, therefore D. (Dt.) anguana is also a synonym of D. (Dt.) silvestris.</p> </div>	https://treatment.plazi.org/id/038287A1F319FFA46A49FCADFC88C867	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
038287A1F31AFFA56A21F9B9FD3FCB95.text	038287A1F31AFFA56A21F9B9FD3FCB95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichogaster (Diplothecodrilus) toroensis Cognetti 1907	<div><p>Dichogaster (Diplothecodrilus) toroensis Cognetti 1907</p> <p>(Figure 6A–D) Dichogaster toroensis Cognetti 1907, p. 3, Cognetti 1909, p. 23, Csuzdi and Zicsi 1994, p. 52. Dichogaster (Diplothecodrilus) toroensis: Csuzdi 1995, p. 114, Csuzdi 1996, p. 357.</p> <p>Material examined</p> <p>Dichogaster toroensis Syntypes. OL 93, Fort Portal, Toro, 1532 m.</p> <p>New records. HNHM AF/3573, one exemplar, BMNH 1997.1630, one exemplar. D. R. Congo, Kivu Region, Bugabo, leg. Gasana N’Doli, 25 July 1973.</p> <p>Diagnosis</p> <p>Length 20–26 mm, D. 1–1.5 mm, no. of segments 63–110. Pigmentation grey. First dorsal pore in 5/6. Clitellum 13–20. ♀ 14 paired between setae a–a. Prostatic pores 17, 19, genital field oblong. Spermathecal pores 7/8, 8/9, ampoule of the spermatheca irregular pear-shaped with smaller distal part bearing a roundish diverticulum. The duct is slightly convoluted and somewhat longer than the ampoule. Gizzard in 5/6, last pair of hearts in 12. Calciferous glands in 15–17 of equal size. Excretory system meroic with four meronephridia per side. Penial seta L. 0.8 mm, D. 0.008 mm, tip rounded, ornamentation serrated teeth.</p></div> 	https://treatment.plazi.org/id/038287A1F31AFFA56A21F9B9FD3FCB95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
038287A1F31BFFA56AC6F9D9FB90C8C4.text	038287A1F31BFFA56AC6F9D9FB90C8C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pygmaeodrilus Michaelsen 1890	<div><p>Genus Pygmaeodrilus Michaelsen, 1890</p> <p>Pygmaeodrilus Michaelsen 1890, p. 11, Michaelsen 1900, p. 385, Michaelsen 1912, p. 41, Michaelsen 1936a, p. 222, Jamieson 1957, p. 449, Zicsi 1996, p. 34.</p> </div>	https://treatment.plazi.org/id/038287A1F31BFFA56AC6F9D9FB90C8C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
038287A1F31CFFA26A6BFC56FC0CC86C.text	038287A1F31CFFA26A6BFC56FC0CC86C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pygmaeodrilus kiwuensis Michaelsen 1910	<div><p>Pygmaeodrilus kiwuensis Michaelsen, 1910</p> <p>(Figure 7A–B)</p> <p>Pygmaeodrilus kiwuensis Michaelsen 1912, p. 46. Jamieson 1957, p. 466.</p> <p>Material examined</p> <p>HNHM AF/3565, five exemplar, BMNH 1997.1535 – 1544, ten exemplars. D. R. Congo, Kivu Region, Kajabwe, in the swamp, leg. Gasana N’Doli, 30 May 1973.</p> <p>Diagnosis</p> <p>Length 25–33 mm, D. 1–1.5 mm, no. of segments 70–93. Pigmentation lacking, dorsal pore none. Clitellum 13–18, ♀ 14 paired just before the setae a–a. Prostatic pores 17, genital field pentangular with two protuberances corresponding to prostate pores. Spermathecal pores 8/9 just lateral to b. Ampoule of the spermatheca roundish duct narrow. The distalmost part of the duct is encircled by several small diverticuli. Gizzard lacking, one pair of club-shaped calciferous glands in 9. Excretory system holoic, Prostates long slender, penial setae lacking.</p></div> 	https://treatment.plazi.org/id/038287A1F31CFFA26A6BFC56FC0CC86C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
038287A1F31CFFA36A69F9B9FBACCB9B.text	038287A1F31CFFA36A69F9B9FBACCB9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pygmaeodrilus modestus Michaelsen 1910	<div><p>Pygmaeodrilus modestus Michaelsen, 1910</p> <p>(Figure 8A–B)</p> <p>Pygmaeodrilus bipunctatus var. modesta Michaelsen 1910, p. 41.</p> <p>Pygmaeodrilus modestus: Michaelsen 1936a, p. 225, Jamieson 1957, p. 51.</p> <p>Material examined</p> <p>HNHM AF/3543, one exemplar, BMNH 1997.1512 – 1513, two exemplars. D. R. Congo, Kivu Region, Katana, in stems of banana trees, leg. Gasana N’Doli, 5 August 1973. HNHM AF/3548, two exemplars, BMNH 1997.1489 – 1492, four exemplars. D. R. Congo, Kivu Region, Mewa, leg. Gasana N’Doli, 17 July 1973. HNHM AF/3568, four exemplars, BMNH 1997.1567 – 1573, seven exemplars. D. R. Congo, Kivu Region, river Mululu, leg. Gasana N’Doli, 15 July 1973. HNHM AF/ 3542, four exemplars, BMNH 1997.1631 – 1637, seven exemplars. D. R. Congo, Kivu Region, Tshibati, in the swamp, Gasana N’Doli, 5 May 1973.</p> <p>Diagnosis</p> <p>Length 40–60 mm, D. 1–1.5 mm, no. of segments 75–105. Pigmentation lacking, dorsal pore none. Clitellum 13–18, K 19. ♀ 14 paired just before the setae a–b. Prostatic pores 17, genital field ovate with two close papillae corresponding to prostate pores. Spermathecal pores 8/9 just medial to a. Ampoule of the spermatheca pear-shaped duct narrow. The distalmost part of the duct bears two pairs of digitiform diverticuli. Gizzard lacking, one pair of club-shaped calciferous glands in 9. Excretory system holoic, Prostates long slender, penial setae lacking.</p></div> 	https://treatment.plazi.org/id/038287A1F31CFFA36A69F9B9FBACCB9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Csuzdi, Csaba	Csuzdi, Csaba (2008): Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae). Journal of Natural History 42 (33 - 34): 2227-2241, DOI: 10.1080/00222930802277616, URL: http://dx.doi.org/10.1080/00222930802277616
