taxonID	type	description	language	source
038387C8A85D040CFF22F98B0DA9F81B.taxon	materials_examined	Type species. Catops strigosus Kraatz, 1852, fixed by subsequent designation by Jeannel (1922: 41). Synonymy. Namadeus Jeannel, 1936: 203 (valid genus; species included: acicularis, anatolicus, creticus, cribratostriatus, graecus, leonhardi, macedo, pellitus; Type Species: Catops acicularis Kraatz, fixed by original designation); Giachino & Vailati, 1993: 62 (synonym with Anemadus).	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85E040FFF22FF780C79FE2B.taxon	diagnosis	Diagnosis. Small size compared to other congeners; protarsi weakly expanded and expansion of mesotarsi imperceptible; median lobe of aedeagus bifid, apical indentation wide or reduced to a simple incision longitudinally short (Perreau, 2002).	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85E040FFF22FF780C79FE2B.taxon	distribution	Distribution. China (Distribution map as shown in Fig. 1).	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85F0408FF22FE0B0DC5FE9E.taxon	materials_examined	Type material. Holotype: CHINA, Sichuan: ♂, Baoxing, Ruobigou, 1630 m, Cyclobalanopsis forest, from fallen wood, 10. VIII. 2003, Jie Wu leg. (IZ-CAS). Paratypes: 1 ♀, same data as holotype (IZ-CAS); 1 ♀, Baoxing, Ganyanggou, 2000 m, Cyclobalanopsis forest, from fallen wood, 9. VIII. 2003, Jie Wu leg. (IZ-CAS).	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85F0408FF22FE0B0DC5FE9E.taxon	description	Description. Male. EBL: 3.00 mm. Length of different body parts: HL: AL: PL: ELL = 0.56: 1.14: 0.68: 1.61 mm; width: HW: EW: PW: ELW = 0.70: 0.12: 1.16: 1.42 mm. Proportion of antennomeres from base to tip in µm (length × width): 111 × 62, 147 × 50, 119 × 43, 74 × 42, 79 × 47, 64 × 51, 94 × 71, 41 × 68, 85 × 81, 83 × 89, 155 × 88. Habitus (Fig. 2 A) elongated oval, regularly convex and sublustrous. Moderately pigmented: mostly chestnut brown; mouthparts, basal four or five antennomeres and apical half of ultimate antennomere, areas near hind corners of pronotum, elytral apices, and tarsi more or less yellowish. Dorsum continually covered with fine, recumbent, sallow pubescence. Insertions of pubescence on elytra aligned along transverse striolations. Head retractile, HW / HL = 1.24. Surface covered with strong round punctures, separated about 2.0 – 4.0 times of their diameter; interspaces smooth, without visable microsculpture. Clypeofrontal suture distinct. Clypeus trapezoidal, anterior margin almost straight. Compound eyes well developed, EW / HW = 0.12. Antennae (Fig. 3 A) long and slender, AL / HW = 1.63; length of 2 th / 3 th = 1.24; 8 th asymmetrical, wider than long; 10 th slightly wider than long; 11 th elongated pear-like. Pronotum (Fig. 3 D) transverse, widest at about basal 2 / 5, PW / PL = 1.69. Sides fairly curved, strongly narrowing forward but moderately and almost rectilinearly narrowing backward from widest; hind corners obtuse. Posterior edge slightly bisinuate. Surface covered with large and rough punctures, separated about 1.0 – 2.0 times of their diameter; interspaces smooth, without visable microsculpture. Postero-lateral impressions absent. Elytra oval, widest at about basal 3 / 8, ELL / EW = 1.13. Sides regularly curved, gradually narrowing from widest to apices; apices rounded (Fig. 3 E). Sculpture of type « a » (sensu Giachino & Vailati, 1993): transverse striolations well-marked and irregular, longitudinal striae invisible. When immersed in glycerine, large primitive punctures visible within interior of cuticule, arranging into 8 irregular longitudinal rows (imperceptible in dry specimens) (Fig. 2 D). Sutural striae present but inconspicuous. Interspaces of striolations with microtrichiae, aligning into ca. 5 transverse lines. Metathoracic wings fully developed. Prolegs slender, with basal three protarsomeres weakly expanded (Fig. 3 B): TW / BTW = 1.49. Protibiae with outer margin almost straight, while inner margin slightly expanded around middle. Profemora wider than protibiae. Mesotibiae distinctly curved, expansion of basal two mesotarsomeres imperceptible. Metatibiae almost straight. Genital segment (Fig. 3 G) as typical for the genus, tergite IX with a pair of setae long and strong, as well as several fine setae apically. Aedeagus with median lobe moderately wide, weakly and almost rectilinearly narrowed towards a bifid apex in dorsal view, apical indentation wide, deep and subtriangular, and both branches rounded at apex (Fig. 3 H). Paramere constricted subapically, apex dentiform, pointing outwards and bearing 5 setae, their arrangement as shown in Figs. 3 K & L. In lateral view, median lobe almost straight, parameres wide and rounded at apex (Fig. 3 J). Endophallus of holotype everted, with a pair of spine rows on dorsal wall and a pair of large teeth pointing upwards at end (Figs. 3 H – J). Female. Similar to male in general appearance (Figs. 2 B & C), but can be distinguished from following characteristics: pro- and mesotarsi simply linear (Fig. 3 C); ventrite VIII with spiculum ventrale short and subtriangularly narrowed anteriorly (Fig. 3 F).	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85F0408FF22FE0B0DC5FE9E.taxon	distribution	Distribution. China (Sichuan).	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85F0408FF22FE0B0DC5FE9E.taxon	etymology	Etymology. The specific epithet is dedicated to Dr. Michel Perreau (Université Paris 7, Paris, France), a famous taxonomist on Leiodidae, for his excellent work on Chinese fauna and generous help to our study. Remarks. This new species well resembles A. ruzickai Perreau and A. wolongianus Perreau in general appearance and their type localities also with geographical proximity, but it is easily to distinguish it from the latter two by the combination of the following characteristics: antennomere length of 2 th / 3 th = 1.24; aedeagus with median lobe moderately wide, weakly and almost rectilinearly narrowed towards apex when viewed dorsally, apical indentation wide, deep and subtriangular, and both branches rounded at apex; paramere constricted subapically, apex dentiform, pointing outwards when viewed dorsally, and wide and rounded at apex when viewed laterally; each paramere with 5 setae on inner margin of apex.	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85A040BFF22FF780D7FF862.taxon	materials_examined	Type material. Holotype: CHINA, Sichuan: ♂, Baoxing, Pujigou, 2400 m, forest of Manchurian walnut, from fallen wood, 11. VIII. 2003, Jie Wu leg. (IZ-CAS). Paratypes: 1 ♀, same data as holotype (IZ-CAS).	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85A040BFF22FF780D7FF862.taxon	description	Description. Male. EBL: 3.32 mm. Length of different body parts: HL: AL: PL: ELL = 0.53: 1.21: 0.69: 1.89 mm; width: HW: EW: PW: ELW = 0.68: 0.07: 1.13: 1.33 mm. Proportion of antennomeres from base to tip in µm (length × width): 130 × 64, 152 × 51, 132 × 47, 82 × 40, 83 × 45, 60 × 53, 94 × 66, 44 × 64, 81 × 76, 84 × 78, 166 × 80. Habitus (Fig. 2 E) elongated oval, regularly convex and sublustrous. Moderately pigmented: mostly chestnut brown; mouthparts, basal two antennomeres and apical half of ultimate antennomere, area near hind corners of pronotum, elytral apices, and tarsi more or less yellowish. Dorsum continually covered with fine, recumbent, sallow pubescence. Insertions of pubescence on elytra aligned along transverse striolations. Head retractile, HW / HL = 1.28. Surface covered with strong round punctures, separated about 1.5 – 3.0 times of their diameter. Clypeofrontal suture distinct. Clypeus transverse, anterior margin subrounded. Compound eyes well developed, EW / HW = 0.10. Antennae (Fig. 4 A) very long and slender, AL / HW = 1.78; all antennomeres longer than wide except 8 th; length of 2 th / 3 th = 1.15; 11 th rather long, pear-like. Pronotum (Fig. 4 D) transverse, widest behind middle, PW / PL = 1.64. Sides fairly curved, strongly narrowing forward and moderately narrowing backward from widest; hind corners bluntly rounded and slightly protruded. Posterior edge slightly bisinuate. Surface covered with very large and rough punctures, separated about 0.5 – 2.0 times of their diameter, some of them even confluent; interspaces smooth, without visable microsculpture. Posterolateral impressions absent. Elytra oval and very elongated, widest at about basal 1 / 3, ELL / EW = 1.42. Sides regularly curved, gradually narrowing from widest to apices; apices much narrowly rounded (Fig. 4 E). Sculpture of type « a » (sensu Giachino & Vailati, 1993): transverse striolations well-marked and irregular, longitudinal striae invisible. When immersed in glycerine, large primitive punctures visible within interior of cuticule, arranging into 8 irregular longitudinal rows (imperceptible in dry specimens) (Fig. 2 G). Sutural striae present but inconspicuous. Interspaces of striolations with microtrichiae, aligning into ca. 5 transverse lines. Metathoracic wings fully developed. Prolegs slender, with basal three protarsomeres weakly expanded (Fig. 4 B): TW / BTW = 1.34. Protibiae with outer margin almost straight, while inner margin slightly expanded around middle. Profemora wider than protibiae. Mesotibiae distinctly curved, expansion of basal two mesotarsomeres imperceptible. Metatibiae straight. Genital segment (Fig. 4 G) as typical for the genus, tergite IX with 4 setae long and strong, as well as several fine setae apically. Aedeagus with median lobe moderately wide, weakly widened to subapex and then narrowed towards a bifid apex in dorsal view (Fig. 4 H), apical indentation deep but not very wide, and both branches obliquely truncated and subacute at apex. Paramere constricted subapically, apex expanded, subrounded and bearing 5 setae, their arrangement as shown in Figs. 4 K & L. In lateral view, median lobe gently bent ventrad at apical part, parameres very wide and turned ventrad at apex (Fig. 4 J). Endophallus of holotype everted, with a pair of spine rows on dorsal wall, abundant long spines at apical part, and a pair of large teeth pointing downwards at end (Figs. 4 H – J). Female. Similar to male in general appearance (Fig. 2 F), but can be distinguished from following characteristics: pro- and mesotarsi simply linear (Fig. 4 C); ventrite VIII with spiculum ventrale short and subrounded anteriorly (Fig. 4 F).	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85A040BFF22FF780D7FF862.taxon	distribution	Distribution. China (Sichuan).	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85A040BFF22FF780D7FF862.taxon	etymology	Etymology. The specific epithet is from the Chinese name (Pinyin) of the type locality “ Sichuan ”, a province in China.	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
038387C8A85A040BFF22FF780D7FF862.taxon	discussion	Remarks. This new species also closely allied to A. ruzickai Perreau, A. wolongianus Perreau, as well as A. perreaui sp. nov., but it is easily to distinguish it from the latter three by the combination of the following characteristics: antennomere length of 2 th / 3 th = 1.15; aedeagus with median lobe moderately wide, weakly widened to subapex and then narrowed towards apex when viewed dorsally, apical indentation deep but not very wide, and both branches obliquely truncated and subacute at apex; paramere constricted subapically, apex expanded, subrounded when viewed dorsally, and very wide and turned ventrad at apex when viewed laterally; each paramere with 5 setae at apex, 3 inner and 2 outer.	en	Wang, Cheng-Bin, Zhou, Hong-Zhang (2016): Two new species of the Anemadus taiwanus species-group (Coleoptera: Leiodidae: Cholevinae: Anemadini) from China. Zootaxa 4072 (2): 282-290, DOI: 10.11646/zootaxa.4072.2.9
