taxonID	type	description	language	source
038387F69D66184FFF22F89E1FA29EE7.taxon	materials_examined	Type material. Holotype: CHINA (Gansu Province): ♂, two labels: " CHINA: S-Gansu [CH 12 - 13] / Mts. 36 km SE Longnan, / 33 º 13 ' 03 '' N, 105 º 14 ' 55 '' E, 2080 m, / N-slope with mixed pine and birch / forest, litter and mushrooms sifted, / 4. VIII. 2012, leg. M. Schülke " [white, printed]; " CEPHENNODES (s. str.) / intricatus m. / det. P. Jałoszyński, ' 15 / HOLOTYPUS " [red, printed] (cMS).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D66184FFF22F89E1FA29EE7.taxon	diagnosis	Diagnosis. Body uniformly dark brown; elytra in male modified, with subapical circumsutural flattening covered with dense setae and apical impressions, which are suboval and asetose except for several long setae along posterior margins, area lateral to each impression bears thick setal brush directed mesally; aedeagus with very long and slender apical projection, which in lateral view is weakly curved dorsally at widely obtuse angle.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D66184FFF22F89E1FA29EE7.taxon	description	Description. Body of male (Fig. 1) strongly convex, oval with feebly marked constriction between pronotum and elytra, uniformly dark brown with slightly lighter appendages, covered with light brown vestiture; BL 1.50 mm. Head broadest at moderately large, strongly convex and coarsely faceted eyes, HL 0.20 mm, HW 0.35 mm; vertex and frons confluent, weakly convex; supraantennal tubercles weakly raised. Punctures on vertex and frons distinct and dense but small, separated by spaces subequal to diameters of punctures; setae sparse, short, suberect. Antennae long and slender, gradually thickened distally, AnL 0.85 mm, antennomeres I – VIII elongated, VII distinctly longer than VIII, antennomeres IX and X about as long as broad, XI about as long as IX – X together, 2.3 × as long as broad, with blunt apex. Pronotum semioval, strongly convex at middle and distinctly flattened near hind corners, broadest near middle; PL 0.50 mm, PW 0.65 mm. Anterior margin weakly arcuate; lateral margins strongly rounded in anterior half and very weakly in posterior third, posterior pronotal corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae narrow and sharply demarcated from disc in posterior third; lateral antebasal pits shallow but distinct, each equally distant from posterior and lateral pronotal margin. Punctures fine and inconspicuous, much denser near each anterior pronotal corner, but even there shallow and small, not coarse. Setae on pronotal disc moderately long and sparse, suberect. Elytra about as convex as pronotum, subrectangular, broadest distinctly in front of middle; EL 0.80 mm, EW 0.73 mm, EI 1.10; subhumeral lines non-carinate, each developed as sharp border between more convex humeral and less convex adsutural region, as long as only 0.28 EL and distinctly divergent caudad; basal fovea on each elytron located closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices (Figs 2 – 3) broadly rounded, each with subapical adsutural area flattened and surrounded by dense setae (forming single circumsutural flattening), apical part of each elytron sharply demarcated and impressed, impressed area oval in shape and asetose, with additional C-shaped ridge on its surface, area lateral to impression bears long setal brush directed mesally. Punctures on elytra slightly more distinct than those on pronotum but superficial and inconspicuous; setae shorter and denser than those on pronotum, nearly recumbent, with several longer suberect setae standing out on each elytron. Hind wings not studied. Metaventrite without lateral impressions, convex. Legs moderately long and slender; all tibiae nearly straight. Aedeagus (Figs 11 – 12) elongate, AeL 0.38 mm; capsular part o median lobe in ventral view symmetrical with asymmetrical assemblage of apical projections, of which dorsal component is strikingly long but in lateral view only slightly curved dorsally; parameres slender, each with two robust apical setae. Female. Unknown.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D66184FFF22F89E1FA29EE7.taxon	distribution	Distribution. Central China: Gansu Province.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D66184FFF22F89E1FA29EE7.taxon	etymology	Etymology. The specific epithet intricatus refers to the complex modification of elytral apices.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D66184FFF22F89E1FA29EE7.taxon	discussion	Remarks. Within the excavatus species group most species have the dorsal apical projection of the aedeagus strongly curved dorsally and in lateral view visible as an apical hook. Only a few species have the dorsal projection weakly curved and not forming a hook. Besides C. intricatus, also C. hamatus, C. spinosus and C. uenoi have such a feebly curved projection, but each of them clearly differs from C. intricatus in external characters, including elytral modifications, and the shape and details of the aedeagus. Among other characters, Cephennodes intricatus is uniformly dark brown, while C. hamatus and C. spinosus have the head and pronotum nearly black and the elytra reddish-brown and C. uenoi has the head and pronotum very dark reddish-brown and elytra light reddish-brown. Moreover, males of C. hamatus and C. uenoi have a shallow posterolateral or lateral impression on each elytron covered with modified setae (lacking in C. intricatus) and males of C. spinosus have the last visible abdominal sternite with a short posteromedian mucro (lacking in C. intricatus). The most externally similar species, C. cryptointricatus s. n. described below, differs from C. intricatus in slightly shorter pronotum (the pronotal width is the same), antennomeres IX and X distinctly elongate (and not about as long as broad) and the aedeagus with strongly curved dorsal apical hook-like projection.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D631849FF22F8DC1D839D27.taxon	materials_examined	Type material. Holotype: CHINA (Gansu Province): ♂, two labels: " CHINA: S-Gansu [CH 12 - 03] / W-Qinling Shan, 43 km N Chengxian, / 34 º 08 ' 24 '' N, 105 º 46 ' 43 '' E, 1750 m, / moist valley with creek and ponds, / meadow with Artemisia, 28. VII. / 2012, leg. M. Schülke " [white, printed]; " CEPHENNODES (s. str.) / cryptointricatus m. / det. P. Jałoszyński, ' 15 / HOLOTYPUS " [red, printed] (cMS). Paratype: 1 ♂, same data as holotype except " [CH 12 - 06] 2012 ", " 47 km N Chengxian, 34 º 10 ' 20 '' N, 105 º 42 ' 19 '' E, 1890 m, creek valley with second [ary]. deciduous forest, litter sifted, 29. VII " (cPJ).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D631849FF22F8DC1D839D27.taxon	diagnosis	Diagnosis. Externally nearly identical with C. intricatus but pronotum distinctly shorter (0.45 mm vs. 0.50 mm, width the same in both species), antennomeres IX and X distinctly elongate (and not as long as broad) and aedeagus with strongly curved, hook-like dorsal apical projection.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D631849FF22F8DC1D839D27.taxon	description	Description. Body of male almost identical as that of C. intricatus, inlcuding elytral modifications; only differences are given here. BL 1.39 – 1.45 mm; HL 0.19 – 0.20 mm, HW 0.34 – 0.35 mm, AnL 0.83 mm; PL 0.45 mm, PW 0.65 mm; EL 0.75 – 0.80 mm, EW 0.74 – 0.75 mm, EI 1.02 – 1.07; subhumeral line on each elytron as long as 0.33 EL. Antennomeres IX and X distinctly elongate. Aedeagus (Figs 13 – 14) elongate, AeL 0.33 mm; capsular part of median lobe in ventral view symmetrical with asymmetrical assemblage of apical projections, of which dorsal component is short and strongly curved dorsally forming robust apical hook; parameres slender, each with two robust apical setae. Female. Unknown.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D631849FF22F8DC1D839D27.taxon	distribution	Distribution. Central China: Gansu Province.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D631849FF22F8DC1D839D27.taxon	etymology	Etymology. The specific epithet cryptointricatus refers to the similarity of this species to C. intricatus.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D631849FF22F8DC1D839D27.taxon	discussion	Remarks. Cephennodes cryptointricatus is strikingly similar to C. intricatus; see remarks for the latter species.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D621849FF22FCAD1C9C9AA8.taxon	materials_examined	Material studied. 2 ♂♂, China, S Shaanxi, Qinling Shan, 42 km SW of Meixian, 34 º 01 ' 32 '' N, 107 º 24 ' 13 '' E, 1875 m, northern slope, secondary deciduous forest, near creek, litter and grass sifted, 26. VII. 2012, leg. M. Schülke (cMS, cPJ).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D621849FF22FCAD1C9C9AA8.taxon	discussion	Remarks. Cephennodes subcaudatus was previously known from only a single specimen (holotype male) collected also in Qinling Shan.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D62184AFF22F8CD18089846.taxon	materials_examined	Type material. Holotype: CHINA (Shaanxi Province): ♂, two labels: " CHINA: S-Shaanxi [CH 12 - 32] / Micang Shan, 42 km S Hanzhong, / 32 º 40 ' 43 '' N, 106 º 48 ' 33 '' E, 1090 m, / stream valley, shady S-slope, sec. / mixed forest, litter, grass & herbs nr. / path sifted, 17. VIII. 2012, M. Schülke " [white, printed]; " CEPHENNODES (s. str.) / monofoveatus m. / det. P. Jałoszyński, ' 15 / HOLOTYPUS " [red, printed] (cMS). Paratype: 1 ♂, same data as holotype except " [CH 12 - 29] ", " 30 km S Hangzhong, 32 º 45 ' 56 '' N, 106 º 53 ' 57 '' E, 1070 m, stream valley, litter and soil sifted, 15. VIII. 2012 " (cPJ).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D62184AFF22F8CD18089846.taxon	diagnosis	Diagnosis. This is the only member of the taurus group with a small round, shallow and indistinctly demarcated impression on the frons in males.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D62184AFF22F8CD18089846.taxon	description	Description. Body of male (Fig. 4) strongly convex, oval with feebly marked constriction between pronotum and elytra, dark brown with slightly lighter legs and palps, covered with yellowish vestiture; BL 1.63 – 1.68 mm. Head (Fig. 5) broadest at moderately large, strongly convex and coarsely faceted eyes, HL 0.23 mm, HW 0.40 – 0.43 mm; vertex and frons confluent and flattened, frons with large but shallow and diffused median impression about as long as broad and larger than eye. Punctures on vertex and frons very fine but dense; setae sparse and moderately long, suberect to erect, also median impression with similar setae. Rod-like modified setae much longer than eye. Antennae moderately long, five terminal antennomeres distinctly and gradually broadening, AnL 0.85 mm, antennomere I shorter than broad, II distinctly but weakly elongate, III indistinctly transverse, IV – VI each about as long as broad, VII distinctly elongate, VIII – X each slightly transverse, XI about as long as IX – X together, 2.2 × as long as broad, pointed at apex. Pronotum in strictly dorsal view subtrapezoidal, strongly convex at middle and distinctly flattened near hind corners, broadest near middle; PL 0.53 mm, PW 0.73 – 0.75 mm. Anterior margin nearly straight; lateral margins strongly rounded in anterior half and weakly in posterior fourth, only slightly convergent toward nearly rightangled hind pronotal corners; posterior margin distinctly bisinuate; lateral marginal carinae indistinctly thickened and near posterior pronotal corners indistinctly demarcated from disc; lateral antebasal pits shallow but distinct, each located much closer to posterior than to lateral pronotal margin. Punctures on median part of pronotal disc small but distinct, moderately deep and with sharp margins, separated by spaces equal to or slightly wider than diameters of punctures; punctures toward anterior and posterior pronotal margins reducing in size and depth, those near lateral margins becoming denser but not larger or deeper, area adjacent to each anterior corner covered with fine and dense but not coarse punctures. Setae on pronotal disc long and sparse, suberect. Elytra as convex as pronotum, oval, broadest near anterior third; EL 0.88 – 0.93 mm, EW 0.80 mm, EI 1.09 – 1.16; subhumeral lines carinate, sharply marked, as long as 0.54 EL and distinctly divergent caudad; basal fovea on each elytron located closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, separately rounded. Punctures on elytra much less distinct than those on pronotal disc, superficial, small but dense; setae similar to those on pronotum. Hind wings well developed, about twice as long as elytra. Metaventrite without lateral impressions, convex. Legs moderately long and slender; all tibiae nearly straight. Aedeagus (Figs 15 – 16) simonis form, AeL 0.35 mm; median lobe in ventral view drop-shaped with narrow subtriangular and strongly protruding apex; apical projections short, with short apical hook weakly and evenly curved dorsally; dorsal wall with only very short and sparse, barely discernible subapical setae; parameres slender, each with one apical and two subapical setae. Female. Unknown.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D62184AFF22F8CD18089846.taxon	distribution	Distribution. Central China: Shaanxi Province.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D62184AFF22F8CD18089846.taxon	etymology	Etymology. The specific epithet monofoveatus refers to the single frontal pit.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D62184AFF22F8CD18089846.taxon	discussion	Remarks. Within the taurus species group, only three species have single (and not paired or absent) impression on the frons: C. taurus, C. qiongdaoanus and C. clypeicornis. However, each of these species has a broad, transverse impression across frons, and not circular shallow pit at middle of frons, as C. monofoveatus.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D601844FF22FE161C369CC2.taxon	materials_examined	Type material. Holotype: CHINA (Sichuan Province): ♂, four labels: " CHINA: W-Sichuan / Ya'an Pref., Tianquan Co. / Jiajin Shan, valley above Labahe " [white, printed]; " N. R. Station, 57 kmW Ya'an / 30.06.63 N, 102.25.18 E, light forest / 1800 m, 12. VII. 1999, leg. A. Pütz " [white, printed]; " Sammlung / Andreas Pütz / Eisenhüttenstadt " [yellow, printed]; " CEPHENNODES (s. str.) / pinnapes m. / det. P. Jałoszyński, ' 15 / HOLOTYPUS " [red, printed] (cAP). Paratype: 1 ♂, same data as holotype (cPJ).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D601844FF22FE161C369CC2.taxon	diagnosis	Diagnosis. Abdomen of male strongly modified, with sternite V bearing median lamina strongly projecting ventrally; elytral subhumeral lines much shorter than half EL; aedeagus in ventral view with the distal margin of apical projection perpendicular to the long axis of aedeagus and slightly recurved.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D601844FF22FE161C369CC2.taxon	description	Description. Body of male (Fig. 6) slightly flattened, oval with feebly marked constriction between pronotum and elytra, moderately dark brown with slightly lighter legs and distinctly lighter palps, covered with light brown vestiture. BL 1.09 – 1.19 mm. Head broadest at moderately large but strongly convex and coarsely faceted eyes, HL 0.14 mm, HW 0.30 mm; vertex and frons confluent and weakly convex, supraantennal tubercles distinct. Punctures on frons and vertex very fine but dense, separated by spaces subequal to diameters of punctures; setae sparse, short, suberect. Antennae moderately long, four terminal antennomeres distinctly and gradually broadening, AnL 0.60 – 0.61 mm, antennomeres I – II elongate, III slightly transverse, IV – VI each about as long as broad, VII slightly elongate, VIII – X each slightly transverse, XI about as long as IX – X together, twice as long as broad, with rounded, blunt apex. Pronotum semioval, convex at middle and slightly flattened near hind corners, broadest between middle and posterior third but only slightly narrowing posteriorly; PL 0.33 – 0.35 mm, PW 0.48 – 0.50 mm. Anterior margin slightly arcuate; lateral margins strongly rounded in anterior half and nearly straight in posterior third; hind pronotal corners nearly right-angled; posterior margin shallowly bisinuate; lateral carinae indistinct, not delimited from disc and not thickened; lateral antebasal pits shallow but distinct, each nearly equally distant from posterior and lateral pronotal margin. Punctures on median part of pronotal disc as fine and dense as those on frons and vertex, punctures becoming denser, but not larger or deeper, toward lateral pronotal margins. Setae on pronotal disc dense and moderately long, suberect. Elytra slightly more convex than pronotum, oval, broadest between middle and anterior third; EL 0.63 – 0.70 mm, EW 0.53 – 0.59 mm, EI 1.19; each subhumeral line marked as sharp border between more convex humeral and less convex adsutural region, as long as only 0.32 EL, lines weakly divergent caudad; basal elytral fovea on each elytron located closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, rounded together. Punctures on elytra as fine and dense as those on pronotum; setae dense, moderately long, suberect. Hind wings well-developed, about twice as long as elytra. Metaventrite with large and deep postmesocoxal impressions. Abdomen (Figs 7 – 8) modified, sternite V with median lamina strongly protruding ventrally with lateral pair of large cavities at base of lamina. Legs moderately long and slender; pro- and mesotibiae distinctly recurved, metatibiae with large apical broadening. Aedeagus (Figs 17 – 18) simonis form, AeL 0.15 mm; median lobe in ventral view drop-shaped with apical portion subtriangular; apical projection short and broad, with distal margin perpendicular to the long axis of aedeagus and slightly recurved; parameres slender, each with one apical and one subapical seta. Female. Unknown.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D601844FF22FE161C369CC2.taxon	distribution	Distribution. Central-southern China: Sichuan Province.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D601844FF22FE161C369CC2.taxon	etymology	Etymology. The specific epithet pinnapes means " winged on the feet, having winged feet " and refers to the distal broadening and flattening of the metatibiae.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D601844FF22FE161C369CC2.taxon	discussion	Remarks. Within the inflatipes species group, the Vietnamese C. patellatus and C. pseudopatellatus have strikingly long elytral subhumeral lines (about as long as 0.7 EL), while in all Chinese species, including the newly described C. pinnapes, the subhumeral lines are as long as only about 0.3 EL. Males of C. inflatipes and C. kopeipes differ from C. pinnapes in an unmodified abdomen. The three species with modified abdomen, C. abdominalis, C. parabdominalis and C. pinnapes can be distinguished on the basis of body proportions and, first of all, the aedeagi (examined in strictly ventral view). The distal margin of the apical projection in C. abdominalis is evenly convex; in C. parabdominalis it is nearly straight; in C. pinnapes the margin is weakly recurved. The apex of apical projection in C. abdominalis is triangular and pointed, whereas apices in C. parabdominalis and C. pinnapes are rounded and blunt. The distal margin of the apical projection in C. abdominalis and C. parabdominalis is oblique in relation to the long axis of the aedeagus, forming with this axis about 45 º-angle, whereas in C. pinnapes the apical margin is perpendicular to the long axis of the aedeagus. Moreover, the apex of median lobe in ventral view in C. abdominalis and C. parabdominalis forms a projection parallel to the long axis of the aedeagus, whereas in C. pinnapes the apex of median lobe is protruding obliquely outward from the longitudinal axis.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6F1844FF22FC8E1FFE9A44.taxon	materials_examined	Material studied. 1 ♂, 3 ♀♀, China, Shaanxi, Qin Ling Shan, 110.06 E, 34.27 N, Hua Shan Mt. N valley, 1200 - 1400 m, 118 km E Xian, sifted, 18 - 20. VIII. 1995, leg. A. Pütz (cAP, cPJ); 1 ♂, China, Shaanxi, Qin Ling Shan, 108.49 E, 33.55 N, river valley 40 km S Xian, river bank, 1200 m, 31. VIII. 1995, leg. A. Pütz (cPJ).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6F1844FF22FC8E1FFE9A44.taxon	discussion	Remarks. Cephennodes parabdominalis was described on the basis of specimens collected also in Qin Ling Shan, probably from the same collecting site.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6F1845FF22F92E19CC9EE7.taxon	materials_examined	Material studied. 3 ♂♂, 5 ♀♀, China, Sichuan, Emei Shan, Wannian, 1050 m, 19 – 30. III. 1999, leg. W. Schawaller (SMNS, cPJ); 1 ♂, Sichuan, Ya'an Pref., Tianquan County, Jiajin Shan, valley above Labahe N. R. station, 57 km W of Ya'an, 30.06.63 N, 102.25.18 E, forest, 1800 m, 12. VII. 1999, leg. A. Pütz (cAP).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6F1845FF22F92E19CC9EE7.taxon	discussion	Remarks. Cephennodes pteroscapus was described on the basis of a single specimen (holotype male) collected also in Sichuan, but in its western part, in the Ganzi Tibet Autonomous Prefecture.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6E1846FF22FEA41FCC9AA7.taxon	materials_examined	Type material. Holotype: CHINA (Sichuan Province): ♂, three labels: " CHINA - S Sichuan, 3000 m / pass N of Jinyang, Rhododendron / primary coniferous forest / 15 - 19. VI. 2004, leg. R. Fabbri " [white, printed]; " MUSEO GENOVA / Acquisto V. 2005 / da R. Fabbri " [white, printed]; " CEPHENNODES (s. str.) / elegantissimus m. / det. P. Jałoszyński, ' 15 / HOLOTYPUS " [red, printed] (MSNG). Paratypes: 3 ♀♀, same data as for holotype (MSNG, cPJ).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6E1846FF22FEA41FCC9AA7.taxon	diagnosis	Diagnosis. Strongly elongate, slender and strongly convex species with head and pronotum darker than elytra; elytral subhumeral lines shorter than 0.2 EL; aedeagus with apical projections bent dorsally at nearly right angle and in strictly ventral view not visible; each paramere in addition to robust apical setae with short submedian seta inserted on short protuberance.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6E1846FF22FEA41FCC9AA7.taxon	description	Description. Body of male (Fig. 9) strongly convex, oval with shallow constriction between pronotum and elytra; head and pronotum dark reddish-brown, elytra light reddish-brown with slightly darkened suture, legs except tarsi and antennae slightly darker than elytra; vestiture of setae light brown. BL 1.83 mm. Head broadest at moderately large but strongly convex and coarsely faceted eyes, HL 0.25 mm, HW 0.38 mm; vertex and frons confluent and weakly convex; supraantennal tubercles feebly marked. Punctures on frons and vertex very fine but dense, separated by spaces subequal to diameters of punctures; setae sparse, short and suberect. Antennae long and slender and gradually thickening from antennomere VII, AnL 0.93 mm, antennomeres I – VIII each distinctly elongate, IX and X about as long as broad, XI about as long as IX – X together, 2.2 × as long as broad, with pointed apex. Pronotum semioval, strongly convex and only slightly flattened near hind corners, broadest near middle; PL 0.53 mm, PW 0.65 mm. Anterior margin arcuate; lateral margins strongly rounded in anterior half and weakly rounded in posterior third, weakly convergent toward nearly right-angled posterior pronotal corners; posterior margin shallowly bisinuate; lateral carinae narrow and sharply delimited from disc in posterior half of pronotum; lateral antebasal pits small but distinct, each located closer to lateral than to posterior pronotal margin. Punctures on pronotal disc fine and very shallow but sharply marked, at middle separated by spaces equal to 1.5 or slightly more diameters of punctures, also area near each anterior pronotal corner covered with very fine and inconspicuous punctures, so that entire pronotal disc is glossy. Setae short, sparse, suberect. Elytra about as convex as pronotum, oval, broadest near anterior third; EL 1.05 mm, EW 0.75 mm, EI 1.40; subhumeral lines carinate, moderately sharply marked, extremely short, equal to only 0.17 of EL and weakly divergent caudad; basal fovea on each elytron located closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, separately rounded. Punctures on elytra larger and denser than those on pronotum but superficial and diffused, and therefore indistinct; setae short, moderately dense, suberect. Hind wings absent. Metaventrite without lateral impressions, convex. Legs moderately long and slender; pro- and mesotibiae slightly curved inward. Aedeagus (Figs 19 – 20) stout and darkly sclerotized, AeL 0.50 mm; capsular part of median lobe in ventral view oval and elongate, with diaphragm located on subapical area of ventral wall and exposed; apical projections so strongly bent ventrally that in ventral view not visible; parameres short and strongly curved, asymmetrical, each with two robust apical setae and one short median seta inserted on short protuberance. Female. Externally similar to male, with less curved apical portion of protibia. BL 1.65 – 1.74 mm; HL 0.23 – 0.24 mm, HW 0.35 – 0.36 mm, AnL 0.83 mm; PL 0.50 – 0.53 mm, PW 0.64 – 0.68 mm; EL 0.93 – 0.98 mm, EW 0.73 – 0.74 mm, EI 1.28 – 1.32.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6E1846FF22FEA41FCC9AA7.taxon	distribution	Distribution. Central-southern China: Sichuan Province.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6E1846FF22FEA41FCC9AA7.taxon	etymology	Etymology. The epithet elegantissimus was chosen to underline the elegant body shape of this interesting species.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6E1846FF22FEA41FCC9AA7.taxon	discussion	Remarks. No other species of Cephennodes has such an elongate and strongly convex body with head and pronotum darker than elytra, very short subhumeral lines and the aedeagus with its apical projections bent ventrally at nearly right angle. An interesting character of C. elegantissimus is the prosternal intercoxal process, which is distinctly narrower than that of any other known species of Cephennodes. The mesoventral process is also somewhat narrower than in other species, and it seems that this is a consequence of a general body shape, which is very slender and elongate as for Cephennodes.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6D1847FF22FB231F9D9ADA.taxon	materials_examined	Type material. Holotype: CHINA (Gansu Province): ♂, two labels: " CHINA: S-Gansu [CH 12 - 17] / Min Shan, 38 km SSW Longnan, / 33 º 05 ' 24 '' N, 104 º 45 ' 13 '' E, 1500 m, / N-slope, macchia, litter and moss / sifted, 6. VIII. 2012, leg. M. Schülke " [white, printed]; " CEPHENNODES (s. str.) / palpalis m. / det. P. Jałoszyński, ' 15 / HOLOTYPUS " [red, printed] (cMS).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6D1847FF22FB231F9D9ADA.taxon	diagnosis	Diagnosis. Maxillary palps strikingly long (longer than head), palpomere III broadest near apex and palpomere IV shorter than broad; metaventrite with deep postmesocoxal impressions; elytra in lateral view slightly flattened around suture in subapical region and then rapidly declining; aedeagus with drop-shaped median lobe and slender, triangular apical portion, subtriangular apical projection forming weakly curved apical hook.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6D1847FF22FB231F9D9ADA.taxon	description	Description. Body of male (Fig. 10) moderately convex, oval with barely marked constriction between pronotum and elytra, light brown with slightly lighter appendages, covered with yellowish-brown vestiture. BL 1.05 mm. Head broadest at moderately large but strongly convex and coarsely faceted eyes, HL 0.14 mm, HW 0.28 mm; vertex and frons confluent, weakly convex; supraantennal tubercles indistinct. Punctures on frons and vertex fine and dense but superficial and inconspicuous; setae short, sparse and suberect. Antennae moderately long, strongly and gradually broadening from antennomere VIII, AnL 0.60 mm, antennomeres I – II elongate, III slightly transverse, IV – VI each about as long as broad, VII elongate, VIII about as long as broad, IX – X slightly transverse, XI slightly longer than IX – X together, about 1.3 × as long as broad, with pointed apex. Pronotum semioval, convex at middle and slightly flattened near hind corners, broadest slightly in front of middle; PL 0.35 mm, PW 0.45 mm. Anterior margin arcuate; lateral margins strongly rounded in anterior half and nearly straight in posterior third, indistinctly convergent toward slightly obtuse-angled hind pronotal corners; posterior margin shallowly bisinuate; lateral carinae very narrow and indistinctly demarcated from disc; lateral antebasal pits shallow but distinct, each located indistinctly closer to posterior than lateral pronotal margin. Punctures inconspicuous, very fine and superficial, even those near anterior pronotal corners are very fine; setae moderately sparse and long, suberect. Elytra about as convex as pronotum, oval, broadest near anterior third and strongly narrowing caudad; EL 0.56 mm, EW 0.48 mm, EI 1.18; subhumeral lines developed as fine grooves as long as 0.35 EL and distinctly divergent caudad; basal fovea on each elytron located slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices slightly modified, with circumsutural flattening in subapical region and then rapidly lowering caudad, rounded together. Punctures on elytra inconspicuous, superficial and fine; setae sparse, moderately long, suberect. Hind wings not studied. Metaventrite with deep and large postmesocoxal impressions. Legs moderately long and slender; pro- and mesotibiae recurved. Aedeagus (Figs 21 – 22) simonis form, AeL 0.24 mm; median lobe in ventral view drop-shaped, with narrowly subtriangular apical portion; apical projections short, subtriangular, dorsal projection weakly curved dorsally; parameres slender, each with three apical setae. Female. Unknown.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6D1847FF22FB231F9D9ADA.taxon	distribution	Distribution. Central China: Gansu Province.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6D1847FF22FB231F9D9ADA.taxon	etymology	Etymology. The name palpalis refers to the unusual structure of maxillary palps in this species.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6D1847FF22FB231F9D9ADA.taxon	discussion	Remarks. The structure of maxillary palpomeres III and IV in this species is unique among Cephennodes. Typically, the palpomere III is elongate and broadest near distal third, narrowing slightly toward apex, and palpomere IV is elongate or about as long as broad, subconical with truncate apex. In C. palpalis the palpomere III is stout, broadest at apex, and palpomere IV is very broad and short, resembling that known in the ' Cephennomicrus group' of genera. However, all other characters (studied by low-pressure SEM in uncoated specimen) do not differ from those in Cephennodes, and also the aedeagus represents clearly a relatively weakly transformed simonis form, with asymmetrical, drop-shaped median lobe surrounded by asymmetrical parameres and with subtriangular assemblage of apical projections in which the dorsal projection forms the apical hook. The structure of the maxillary palp is an important character in the taxonomy of Scydmaeninae and such a deviation from ' normal' form within any genus of Cephenniini has not been known. This structure in C. palpalis certainly deserves attention, but I prefer to place this unusual species in Cephennodes s. str. (on the basis of its genital structures) than to establish a new genus-group name.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6C1842FF22F92318419D0A.taxon	materials_examined	Type material. Holotype: CHINA (Sichuan Province): ♂, four labels: " CHINA: W-Sichuan / Ya'an Pref., Tianquan / Co., E Erlang Shan Pass " [white, printed]; " 2900 m, 20. VI. 1999 / 29.52.36 N, 102.17.82 E / leg. A. Pütz " [white, printed]; " Sammlung / Andreas Pütz / Eisenhüttenstadt " [yellow, printed]; " CEPHENNODES / (FUSIONODES) / corpulentus m. / det. P. Jałoszyński, ' 15 / HOLOTYPUS " [red, printed] (cAP).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6C1842FF22F92318419D0A.taxon	diagnosis	Diagnosis. Aedeagus in ventral view with nearly egg-shaped capsular part, distal columella weakly protruding; dorsal subapical projection in dorsal view with large oval ' head' and in lateral view strongly protruding dorsally and curved distally.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6C1842FF22F92318419D0A.taxon	description	Description. Body of male (Fig. 23) strongly convex, oval with distinct but shallow constriction between pronotum and elytra, uniformly light brown, covered with yellowish vestiture. BL 1.58 mm. Head broadest at moderately large but strongly convex and coarsely faceted eyes, HL 0.25 mm, HW 0.35 mm; vertex and frons confluent, weakly convex; supraantennal tubercles weakly raised. Punctures on frons and vertex distinct, moderately large but shallow and with moderately sharp margins, separated by spaces equal to or slightly wider than diameters of punctures; setae short, sparse, suberect. Antennae moderately long, gradually broadening distally from antennomere VII, AnL 0.80 mm, antennomeres I and II strongly elongate, III – V each indistinctly elongate, VI about as long as broad, VII distinctly elongate, VIII distinctly shorter than VII and about as long as broad, IX and X slightly transverse, XI about as long as IX – X together, twice as long as broad, with pointed apex. Pronotum semioval, strongly convex and distinctly flattened near hind corners, broadest between middle and anterior third; PL 0.50 mm, PW 0.66 mm. Anterior margin arcuate; lateral margins strongly rounded in anterior half and nearly straight in posterior third, distinctly convergent toward nearly right-angled posterior pronotal corners; posterior margin shallowly bisinuate; lateral carinae very narrow and not demarcated from disc; lateral antebasal pits large and deep, each located slightly closer to lateral than to posterior pronotal margin. Punctures on pronotal disc very fine and shallow but dense, those near each anterior pronotal corner appear as slightly coarse scale-like but very fine microsculpture; setae moderately dense and long, only slightly suberect. Elytra slightly more convex than pronotum, oval, broadest near anterior third and strongly narrowing caudad; EL 0.83 mm, EW 0.73 mm, EI 1.14; subhumeral lines carinate, moderately sharply marked, as long as 0.33 EL and distinctly divergent caudad; basal fovea on each elytron located slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, separately rounded. Punctures on elytra slightly more distinct than those on pronotal disc but still superficial and unremarkable; setae similar to those on pronotum. Hind wings not studied. Metaventrite with short setose postmesocoxal impressions. Legs moderately long and slender; all tibiae nearly straight. Aedeagus (Figs 26 – 28) stout, AeL 0.40 mm; median lobe in ventral view with egg-shaped capsular part, with lateral apical lobes weakly protruding laterally and distal columella weakly protruding distally; columella with obtuse-angled distal margin; subapical dorsal projection in dorsal view with broad oval head and broad subtriangular base, with constriction in between, in lateral view dorsal subapical projection protruding strongly dorsally and curved distally; dorsal subtriangular projections robust and long; parameres strongly curved mesally and touching at middle at base of distal columella, numerous parameral setae located in subapical region. Female. Unknown.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6C1842FF22F92318419D0A.taxon	distribution	Distribution. Central-southern China: Sichuan Province.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6C1842FF22F92318419D0A.taxon	etymology	Etymology. The name corpulentus (Latin " corpulent, fleshy, fat ") refers to the stout body of this species.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D6C1842FF22F92318419D0A.taxon	discussion	Remarks. External morphology of species included in the subgenus Fusionodes is usually unremarkable, whereas the aedeagi are most complicated among Cephenniini of the Northern Hemisphere and show an enormous array of often monstrous apical lobes and projections, which are strikingly different even between externally almost identical species. The aedeagus of C. corpulentus is most similar to that of C. serratus (which is also known from Sichuan); no other species of Fusionodes has a similar lateral apical lobes surrounding the central apical columella, subtriangular ' fins' or robust lobes projecting from dorsal wall distally to the orifice at the parameral bases and a tail-like dorsal subapical projection. The aedeagus of C. corpulentus is stouter than that of C. serratus, in ventral view the distal margin of columella is obtuse-angled (distinctly subtriangular, nearly right-angled in C. serratus), and in dorsal view the dorsal subapical projection has a broad oval head and is strongly constricted in its sub-basal region (broadest at base and narrowing toward subtriangular, rounded apex in C. serratus). Cephennodes corpulentus is also distinctly smaller than C. serratus, with BL 1.58 mm vs. 1.82 – 1.84 mm, respectively.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D691843FF22FB381FE59AF6.taxon	materials_examined	Type material. Holotype: CHINA (Shaanxi Province): ♂, two labels: " CHINA: Shaanxi, Qin Ling Shan / 108.47 E, 33.51 N, Mountain W Pass / at Autoroute km 70, 47 km S Xian / 2300 - 2500 m, sifted / 26. - 30.08.1995. leg. A. Pütz " [white, printed]; " CEPHENNODES / (FUSIONODES) / gutta m. / det. P. Jałoszyński, ' 15 / HOLOTYPUS " [red, printed] (cAP). Paratype: 1 ♂, same data as for holotype (cPJ).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D691843FF22FB381FE59AF6.taxon	diagnosis	Diagnosis. Aedeagus in ventral view with apical assemblage of projections with small proximal lobes and apical lamina subrectangular with concave lateral margins; dorsal subapical projection bifurcate; in lateral view apical lamina and dorsal subapical projection strongly protruding as slender and curved tail-like structures; each paramere with three setae.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D691843FF22FB381FE59AF6.taxon	description	Description. Body of male (Fig. 24) moderately convex, oval with indistinct constriction between pronotum and elytra, uniformly dark brown, covered with yellowish vestiture. BL 1.63 – 1.78 mm. Head broadest at moderately large but strongly convex and coarsely faceted eyes, HL 0.20 – 0.25 mm, HW 0.38 – 0.40 mm; vertex and frons confluent, weakly convex; supraantennal tubercles weakly raised. Punctures on frons and vertex distinct but small, shallow and with moderately sharp margins, separated by spaces subequal to diameters of punctures; setae short, sparse, suberect. Antennae moderately long, gradually broadening distally from antennomere VII, AnL 0.85 – 0.88 mm, antennomeres I – VII each distinctly elongate, VIII about as long as broad, IX and X slightly transverse, XI about as long as IX – X together, about twice as long as broad, with pointed apex. Pronotum semioval, strongly convex at middle and distinctly flattened near hind corners, broadest slightly in front of middle; PL 0.50 – 0.55 mm, PW 0.78 – 0.80 mm. Anterior margin broadly rounded; lateral margins strongly rounded in anterior half and indistinctly concave in posterior third, slightly convergent toward nearly right-angled posterior pronotal corners; posterior margin shallowly bisinuate; lateral carinae not demarcated from disc; lateral antebasal pits large and deep, each located slightly closer to posterior than to lateral pronotal margin. Punctures on pronotum distinct, small and shallow but dense, those in central region of disc separated by spaces 1 – 1.5 times as wide as diameters of punctures, those near each anterior pronotal corner distinctly denser, but not larger or deeper, appear as slightly coarse scale-like and fine microsculpture; setae moderately dense and long, only slightly suberect. Elytra as convex as pronotum, oval, broadest near anterior fourth and strongly narrowing caudad; EL 0.93 – 0.98 mm, EW 0.78 – 0.80 mm, EI 1.19 – 1.22; subhumeral lines indistinctly carinate, as long as 0.25 EL and only slightly divergent caudad; basal fovea on each elytron located slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, separately rounded. Punctures and setae on elytra similar to those on median part of pronotal disc. Hind wings not studied. Metaventrite with short setose postmesocoxal impressions. Legs moderately long and slender; all tibiae nearly straight. Aedeagus (Figs 29 – 31) relatively slender, AeL 0.55 mm; median lobe in ventral view with oval capsular part with broadly rounded apical margin; apical assemblage of projections complicated, in ventral view with small proximal lobes and long apical lamina with concave lateral margins; dorsal component with bifurcate dorsal subapical projection, both apical lamina and dorsal subapical projection in lateral view visible as slender and curved tail-like structures; each paramere with one apical and two subapical setae. Female. Unknown.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D691843FF22FB381FE59AF6.taxon	distribution	Distribution. Central-eastern China: Shaanxi Province.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D691843FF22FB381FE59AF6.taxon	etymology	Etymology. The name gutta (Latin " a drop of a fluid ", a noun in apposition) refers to the drop-like body shape of adults of this species.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D691843FF22FB381FE59AF6.taxon	discussion	Remarks. Within the graeseri species group, C. gutta is the only species with the aedeagus in lateral view showing two tail-like projections curved dorsally (the apical lamina and dorsal subapical projection), also the apical lamina in ventral view with both lateral margins concave is a unique character. This component in all other species of this group is variously twisted around the long axis of the aedeagus and then one of its sides is concave and the other one convex, with exception of C. medusa sp. n. described below, in which the apical lamina is not twisted and has both lateral margins convex.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D68185DFF22FADD1DBB9EC5.taxon	materials_examined	Type material. Holotype: CHINA (Sichuan Province): ♂, two labels: " CHINA: N-Sichuan [CH 12 - 26] / 70 km N Songpan, road S 301, above / Gan lake, 33 º 15 ' 26 '' N, 103 º 46 ' 03 '' E, / 2700 m, spruce forest with birch, / litter, mushrooms, moss, and dead / wood sifted, 12. VIII. 2012, M. Schülke " [white, printed]; " CEPHENNODES / (FUSIONODES) / medusa m. / det. P. Jałoszyński, ' 15 / HOLOTYPUS " [red, printed] (cMS).	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D68185DFF22FADD1DBB9EC5.taxon	diagnosis	Diagnosis. Externally unremarkable species; aedeagus in ventral view with apical assemblage of projections with large proximal lobes and apical lamina broadly rounded at apex and with convex lateral margins; dorsal subapical projection with rounded apex; each paramere with numerous setae.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D68185DFF22FADD1DBB9EC5.taxon	description	Description. Body of male (Fig. 25) moderately convex, oval with indistinct constriction between pronotum and elytra, uniformly dark brown, covered with light brown vestiture. BL 1.75 mm. Head broadest at moderately large but strongly convex and coarsely faceted eyes, HL 0.25 mm, HW 0.40 mm; vertex and frons confluent, weakly convex; supraantennal tubercles weakly raised. Punctures on frons and vertex distinct, moderately large and deep with sharp margins, separated by spaces subequal to diameters of punctures; setae short, sparse, suberect. Antennae moderately long, gradually broadening distally from antennomere VII, AnL 0.93 mm, antennomeres I – III distinctly elongate, IV – VI each about as long as broad, VII slightly elongate, VIII – X each slightly transverse, XI about as long as IX – X together, about twice as long as broad, with pointed apex. Pronotum subrectangular, strongly convex at middle and distinctly flattened near hind corners, broadest near middle but only indistinctly narrowing posteriorly; PL 0.53 mm, PW 0.80 mm. Anterior margin broadly and weakly rounded; lateral margins strongly rounded in anterior half and nearly straight in posterior third; posterior pronotal corners nearly right-angled; posterior margin shallowly bisinuate; lateral carina narrow and sharply demarcated from disc on its entire length; lateral antebasal pits developed as very large elongate and deep impressions each adjacent to lateral marginal carina and slightly wider separated from posterior pronotal margin, impressions are as long as more than one-third of PL. Punctures on pronotal disc very small and shallow but distinct and dense, those in median region separated by spaces subequal to diameters of punctures, those near each anterior pronotal corner denser but not larger or deeper, appear as slightly coarse scale-like but fine microsculpture; setae moderately dense and long, only slightly suberect. Elytra as convex as pronotum, oval, broadest near anterior fourth and strongly narrowing caudad; EL 0.98 mm, EW 0.85 mm, EI 1.15; subhumeral lines sharply but finely carinate, as long as 0.25 EL and weakly divergent caudad; basal fovea on each elytron located slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, separately rounded. Punctures on elytra slightly less distinct than those on pronotal disc, superficial; setae similar to those on pronotum but nearly recumbent. Hind wings not studied. Metaventrite with very short setose postmesocoxal impressions. Legs moderately long and slender; all tibiae nearly straight. Aedeagus (Figs 32 – 34) relatively stout, AeL 0.48 mm; median lobe in ventral view with oval capsular part with its apical margin forming abruptly delimited subtriangular and rounded apical lobe; apical assemblage of projections complicated, in ventral view with large proximal lobes and moderately long apical lamina with convex lateral and apical margins; dorsal component with broadly rounded dorsal subapical projection; each paramere with numerous robust apical and subapical setae. Female. Unknown.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D68185DFF22FADD1DBB9EC5.taxon	distribution	Distribution. Central China: Sichuan Province.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D68185DFF22FADD1DBB9EC5.taxon	etymology	Etymology. The name medusa (a noun in apposition) refers to the striking multiple waving and disordered parameral setae, just like the mythical Medusa's snake hairs.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
038387F69D68185DFF22FADD1DBB9EC5.taxon	discussion	Remarks. Within the graeseri species group, C. medusa is the only species with multisetose parameres; all allied species have 3 – 4 setae only. Moreover, the shape of apical lamina in ventral and dorsal views is also unique, no other similar species has the lamina symmetrical and with both lateral margins convex. See also remarks for C. gutta.	en	Jałoszyński, Paweł (2015): The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4033 (3): 393-410, DOI: 10.11646/zootaxa.4033.3.5
