taxonID	type	description	language	source
03812A52FFCE7949F3B1FEB27B63E13E.taxon	materials_examined	Type series. Holotype: female, slide 9029, Russia, Yaroslavl Province, Breitovo District, the Kamenka stream near the village of Sit’ – Pokrovskoe, depth 0.3 – 0.5 m; substrates: stones, sand and mosses, 8. 08. 2004, coll. P. V. Tuzovskij. The holotype is deposited in the collection of Institute for Biology of Inland Waters (Borok, Russia). Paratypes: one female, one male and six larvae from the same locality and collection date as holotype. Larvae reared from one female (holotype) in laboratory. The duration of the embryonic period at room temperature was 12 days. Additional material. 17 females and 12 males, Russia, Yaroslavl Province, Nekouz District, the Ild river near the village of Mar’ino, June – August 2004 – 2005, coll. P. V. Tuzovskij.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFCE7949F3B1FEB27B63E13E.taxon	diagnosis	Diagnosis. Adults: posterior edge of coxal plate IV obliquely – posteromediallydirected, tarsus of female leg I long and slender (ratio L tibia / tarsus 1.25 – 1.32), pedipalpal femur of male with large ventrodistal projection; larva: dorsal shield elongate (ratio length / width 1.64 – 1.70) with parallel lateral edges, posterior margin of basal segments of chelicera convex with small median incision, seta C 3 considerably shorter than C 4.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFCE7949F3B1FEB27B63E13E.taxon	description	Description. Female. Color yellowish – brown, mature specimens nearly black. The body flat, oval, integument soft with very fine strips. Setae Fch longest and heaviest of all dorsal setae (Fig. 1). Setae Oe, Hi and He shorter than Fch but considerably longer and thicker than other idiosomal setae. Setae Fp and Oi without glandularia, other dorsal setae associated with glandularia. First pair of lyriform organs (i 1) situated behind eye lenses; i 2, in interval between setae Ve and He; i 3, behind humeral row of setae (Hi, He); i 4, more or less corresponding to lumbar row of setae (Li, Le); i 5 in the posterior part of the dorsum or ventrum. In mature specimens coxal plates covering less than half of the ventral surface of body (Fig. 2). Posteromedial margin of coxal plates I + II usually rounded, but sometimes straight, with short median process and two moderately large apodemes; median suture line between coxal plates I distinct. The seta and glandularium Hv situated in posterolateral part of coxal plate II. Medial margin of coxal plates III + IV rounded, setae Pe situated near middle of anterior margin of coxal plate IV. Posterior edge of coxal plate IV obliquely-posteromedially directed. The setae Pi without glandularia and usually separated from setae Ci, but sometimes both setae located on a common sclerite. Other ventral idiosomal setae associated with glandularia. Excretory pore unsclerotized, placed between or slightly behind setae Ci. Gonopore and genital plates subequal in length, or gonopore slightly longer than plates. Each genital plate with 3 acetabula and 16 – 22 fine setae. Genital plates narrow elongate, ratio length / width 3.1 – 3.4; genital acetabula in arc line, posterior acetabulum larger than anterior and median acetabula (Figs 3 – 4), but occasionally genital plate with two acetabula (Fig. 5). Anterior genital sclerite considerably wider than posterior sclerite. Chelicera (Fig. 6) with large basal segment and short crescent-shaped stylet. Cheliceral stylet with small teeth on concave side. Pedipalp (Fig. 7) slender; trochanter short, with a single dorsodistal seta; femur with straight ventral margin with 5 unequal dorsal setae. Pedipalpal genu with straight ventral margin, with 4 thick and 4 – 6 thin setae. Bases of ventral setae divide tibia into three approximately equal sectors (1: 1: 1), ventrolateral sword-like seta rather large and placed near middle of segment in space between ventral setae; dorsolateral setae relatively not numerous and located mainly in distal half of segment. Pedipalpal tarsus slightly tapering distally, with one solenidion, 4 thick spines and 5 fine setae; all setae located in distal half of segment. Capitulum (Fig. 9) elongate, rostrum short, not reaching anterior edge of capitulum; dorsal hypostomal setae longer than ventral setae. All legs without swimming setae. Tibia of leg I (Fig. 8) slightly thickened distally; bases of setae S 1 and S 2 separated, narrowed proximally; seta S 2 shorter and wider than S 1, both setae more or less pointed. Tarsus I long and slender, ratio length tibia / tarsus 1.25 – 1.32; with convex dorsal and concave ventral margins; slightly thickened proximally. Posterior legs, in particular legs IV, very slender; distal ends of genu and tibia with rather long thick unequal setae (Fig. 10). Claws of leg I (Fig. 11) smaller than claws of legs II – IV (Fig. 12). All leg claws with two denticles, internal denticle slightly longer than external denticle; lamella with slightly concave ventral margin. Measurements (n = 10). Length of body 750 – 1250; length of setae Fch 140 – 160, length of setae Oe Hi and He 80 – 110, length of coxae I + II 250 – 300, width 340 – 365; length of medial portion of coxae I 115 – 125; length of coxae III + IV 245 – 270, width 220 – 245; lengths of genital plate 142 – 150, width 42 – 48; length / width of genital acetabula (ac. 1 – 3): 42 – 45 / 24 – 27, 40 – 42 / 30 – 36, 48 – 55 / 40 – 48; length of basal segment of chelicera 210 – 215, length of cheliceral stylet 78 – 85; length of pedipalpal segments (P 1 – 5): 35 – 42, 78 – 90, 108 – 125, 115 – 140, 30 – 36; length / width of sword seta on P 4 33 – 39 / 6 – 7; lengths of leg segments I – Leg. 1 – 6: 65 – 78, 105 – 138, 150 – 180, 225 – 265, 225 – 270, 180 – 205; II – Leg. 1 – 6: 60 – 72, 90 – 105, 125 – 150, 185 – 205, 190 – 205, 185 – 195; III – Leg. 1 – 6: 65 – 85, 95 – 110, 125 – 155, 195 – 235, 205 – 240, 190 – 220; IV – Leg. 1 – 6: 125 – 150, 125 – 155, 195 – 235, 270 – 315, 275 – 335, 230 – 275; tibia of leg I: length of seta S 1 130 – 140, width 12 – 13; length of seta S 2 102 – 115, width 17 – 19; distance between bases of setae S 1 – S 2 24 – 30. Male. Dorsal surface, coxae of legs similar to female, from which it differs in smaller size, as well as shape of genital field, pedipalp and terminal segments of leg I. Coxae of legs covering larger than one half of ventral surface. Suture line between coxae I clearly developed, central sub – dermal projection short; apodeme of coxal plates II rather long, reaching anteromedial margin of coxal plates III (Fig. 15). Posteromedial margin of coxae I convex. Setae Hv located in lateral portion of coxae II. Coxae IV trapezoidal, their medial margins rather wide and rounded; glandularia Pe situated near anterior margins of coxae IV. Posterior edge of coxal plate IV obliquely – posteromedially directed. Setae Pi without glandularia and usually separated from setae Ci, but sometimes both setae located on a common sclerite, excretory pore unsclerotized. Genital plates fused to each other by anterior and posterior ends and forming small anterior and rather deep posterior median incisions (Fig. 13 – 14). Posterior pair of genital acetabula large, oval or triangular and considerably larger than two anterior pairs of acetabula. Genital plate with 23 – 32 fine setae on each side, three of them longer than others; genital opening narrow. Genital plate usually wider than long (Fig. 13), but occasionally length and width nearly subequal (Fig. 14). Pedipalp (Fig. 16) stout; trochanter short, with a single short dorsodistal seta; femur with widely rounded ventrodistal protrusion (hump) and relatively large angular projection (protuberance) which reaches well beyond the protrusion, with five dorsal (two proximal and three distal) unequal setae. Pedipalpal genu with slightly concave ventral margin and convex dorsal margin, two proximal and two distal thick unequal setae and 4 – 5 fine setae. Pedipalpal tibia narrowed proximally with convex dorsal edge and very numerous dorsolateral setae, bases of two ventral setae divide tibia into three unequal sectors (1 – 1 – 2); sword – like seta rather large, considerably longer than distance between ventral setae. Genu of leg I (Fig. 17) thickened distally; tibia with nearly parallel ventral and dorsal margins; seta S 1 longer and thinner than S 2, both these setae more or less pointed, narrowed proximally; tarsus relatively short, slightly thickened proximally, with slightly convex dorsal margin and concave ventral one. Measurements (n = 10). Length of body 630 – 750; length of setae Fch 150 – 190, length of setae Oe Hi and He 95 – 130; length of coxae I + II 250 – 275, width 285 – 385; length of medial portion of coxae I 115 – 135; length of coxae III + IV 190 – 230, width 145 – 230; lengths of genital plate 100 – 125, width 125 – 140; length / width of genital acetabula (ac. 1 – 3): 36 – 42 / 28 – 30, 36 – 42 / 24 – 30, 48 – 65 / 35 – 42; length of basal segment of chelicera 150 – 160, length of cheliceral stylet 65 – 70; lengths of pedipalpal segments (P – 1 – 5): 30 – 36, 65 – 80, 70 – 85, 90 – 105, 29 – 31; length / width of sword seta on P 4 36 – 42 / 6 – 7; lengths of leg segments: I – Leg. 1 – 6: 60 – 66, 90 – 105, 120 – 140, 180 – 210, 185 – 215,135 – 155; II – Leg. 1 – 6: 55 – 72, 70 – 85, 100 – 120, 130 – 155, 140 – 160, 135 – 155; III – Leg. 1 – 6: 55 – 75, 75 – 90, 105 – 130, 160 – 185, 180 – 200, 150 – 170; IV – Leg. 1 – 6: 115 – 140, 105 – 125, 165 – 190, 160 – 250, 225 – 260, 190 – 205; tibia of leg I: length of seta S 1 100 – 120, width 9 – 11; length of seta S 2 90 – 96, width 12 – 14, distance between setae S 1 and S 2 18 – 30. Larva. Idiosoma flat and oval in shape, dorsal shield in unengorged larva covers nearly all dorsum (Fig. 18). Dorsal shield elongate, ratio length / width 1.64 – 1.70, with straight or slightly convex anterior margin, indented anterolaterally, with parallel lateral edges, and bears four pairs of setae in anterior portion: two pairs of simple setae (Fch, Vi) and two pairs of trichobothria (Fp, Oi). Simple setae thick and long, anterior setae (Fch) considerably shorter than posterior setae (Vi). Both pairs of trichobothria short, thin and subequal in length. Seven pairs of setae (Oe, Hi, He, Sci, Sce, Li, Si) are situated on soft wrinkled membrane. Setae Hi and Sci longest and heaviest of all dorsal setae situated on soft membrane. Setae Oe slightly shorter than setae Hi and Sci but longer than other dorsal setae situated on soft membrane. Setae Hi and He located close to each other, setae Sci and Sce divided by wide interspace. Dorsal shield with reticulated pattern consisting of elongated cells. Coxae I – III fused to each other on each side, suture line between them developed only in their lateral parts (Fig. 19). Coxal plates I – III straight posteromedially and fitting closely with anterior margin of excretory pore plate. Both setae (C 1, C 2) on coxae I short and approximately subequal in length. Lateral seta (C 3) considerably shorter than medial seta on coxae IV (C 4). Setae C 4 not extending to posterior margin of coxal plates I – III. All coxal plates with reticulated pattern consisting of elongated cells. Transverse muscle attachment scar on coxae III not developed. Urstigma with cap, very small, occupies lateral position on border between coxae I and II. Setae Ci very long, whip-like, well thickened and borne on rather large projections. Setae Le and Se fine and short. Excretory pore plate (Fig. 20) large, nearly twice as wide as long and bearing four pairs of setae (Ai, Ae, Pi, Pe). Both pairs of anal setae (Ai, Ae) reduced and represented by alveoli. Distance between setae Ae – Ae slightly longer than distance between setae Ai – Ai. Setae Pi slightly shorter than setae Pe, bases of both pairs of these setae placed on anterior margin of excretory pore plate. Excretory pore situated proximally to middle of excretory pore plate near setae Ae. Capitulum (Fig. 21) elongate with rather wide base and narrow rostrum, anterior and posterior pairs of setae approximately subequal. Basal part of capitulum twice as long as rostrum. Basal segments of chelicerae fused to each other medially, their posterior edges convex with small median incision (Fig. 22). Cheliceral stylet small, crescent – shaped with one subapical tooth (Fig. 23). Pedipalps relatively short (Fig. 24). Trochanter very short, fused with femur but suture line between these segments present. Pedipalpal femur with one short, thin dorsodistal seta; genu with very long, thick proximal seta and short, fine dorsodistal one; tibia with two long unequal setae and massive dorsodistal claw; tarsus small, with one rather long solenidion and short, thin simple unequal setae. Legs 5 – segmented and without swimming setae. Shape and arrangement of setae on legs segments as shown in Figs 25 – 27. Total number of leg setae, excluding eupathidia, as follows (specialized setae indicated in parenthesis): I – Leg. 1 – 5: 1, 7, 5 (s), 11 (2 s), 14 (s, ac); II – Leg. 1 – 5: 1, 7, 5 (s), 11 (2 s), 14 (s, ac); III – Leg. 1 – 5: 1, 6, 5 (s), 10 (s), 11 (ac). Trochanter of legs I with thick and relatively short seta, trochanter of legs II and III each with rather long thin seta. Solenidion on genu leg I and leg II considerably longer than solenidion on genu III. Solenidia on tibia of leg I and II located at distal end of segment. Proximal solenidion on tibia of leg I and II longer than distal solenidion. Solenidion on tarsus of leg I occupying dorsodistal position, solenidion on tarsus II situated proximally. Genu of leg III with short solenidion and tibia III with long solenidion. Acanthoid seta located distally on tarsus of all legs. Claws of legs III (Fig. 28) larger than claws of legs I and II (Fig. 29). Central claw and lateral claws each pair of legs nearly subequal in size. All claws with a few fine lateral teeth. Measurements (n = 6). Length of dorsal plate 230 – 245, width 145 – 150; length of setae Fch 29 – 38, length of setae Fp and Oi 9 – 13, length of setae Vi 60 – 70, length of setae Oe 50 – 60, length of setae Hi 64 – 70, length of setae He 25 – 32, length of setae Sci 65 – 74, length of setae Sce 22 – 29, length of setae Li 22 – 26, length of setae Si 22 – 29, length of setae Ci 145 – 157, length of setae Se 25 – 29, length of setae Le 22 – 26, length of setae Pi 15 – 19, length of setae Pe 16 – 23, length of setae C 1 16 – 23, length of setae C 2 16 – 21, length of setae C 3 38 – 48, length of setae C 4 64 – 75; length of medial edges of coxae I – III 155 – 170; length of capitular bay 41 – 45; diameter of urstigma 11 – 13; length of excretory pore plate 40 – 45, width 75 – 80; length of capitulum 70 – 76; length of basal segment of chelicera 64 – 75, length of cheliceral stylet 18 – 23; length of pedipalpal segments (P – 1 – 5): 4 – 5, 32 – 35, 16 – 19, 8 – 10, 4 – 5; length of basal projection for insertion Ci 12 – 16, diameter of basal projection for insertion Ci 9 – 11; distance between setae Ai – Ai 14 – 17, distance between setae Ae – Ae 19 – 23; length of legs segments: I – Leg. 1 – 5: 25 – 32, 32 – 35, 29 – 35, 38 – 42, 44 – 50; II – Leg. 1 – 5: 28 – 32, 32 – 35, 32 – 35, 41 – 45, 50 – 55; III – Leg. 1 – 5: 35 – 38, 41 – 45, 41 – 45, 54 – 58, 55 – 60. Differential diagnosis. The present species is closely related to Atractides nodipalpis Thor, 1899 and A. magadanensis Tuzovskij, 2010. The new species differs from A. nodipalpis in the following characters (character states of adults and larva A. nodipalpis are indicated in parenthesis from Gerecke 2003 and Wainstein 1980 respectively): both sexes — posterior edge of coxal plate IV obliquely posteromedially directed (transverse); male — pedipalpal femur with the ventrodistal protrusion consisting of a widely rounded medial hump and relatively large angular protuberance (blindly pointed medial hump and an equally convex short lateral protuberance), P- 2 ventral margin 1: 1: 2 (2: 1: 2), sword seta placed near proximodistal seta (near proximoventral seta); female — genital acetabula in arc line (in an obtuse angle); larva — dorsal shield relatively narrow with parallel lateral margins, ratio length / width 1.64 – 1.70 (rather wide with convex lateral margins, ratio length / width 1.38), posterior margin of basal segments of chelicera convex with small median incision (straight, without median incision), seta C 3 considerably shorter than C 4 (setae C 3 and C 4 equal in length). The new species differs from A. magadanesnsis by the male with posterior pair of the genital acetabula rectangular, P- 2 of female with convex ventral margin, leg claws with short blade, both claw denticles approximately subequal in length; larva - the capitular base nearly as long as wide, the excretory pore located in the centre of the excretory pore plate (Tuzovskij 2010 a). In contrast, the male of A. yaroslavlensis has the posterior pair of genital acetabulum oval or triangular, P- 2 of female with straight or slightly concave ventral margin, leg claws with relatively long blade, an internal denticle longer than external denticle; larva - the capitular base longer than wide, the excretory pore located in anterior half of the excretory pore plate.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFCE7949F3B1FEB27B63E13E.taxon	etymology	Etymology. The species epithet yaroslavlensis is derived from the name of the Province where it was collected (Yaroslavl). Habitat. Running waters.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFCE7949F3B1FEB27B63E13E.taxon	distribution	Distribution. Europe (Russia, Yaroslavl Province).	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC6794CF3B1FC7A7B63E155.taxon	materials_examined	Type series. Holotype: female, slide 8858, Russia, Yaroslavl Province, Nekouz District, Ild stream near village Mar’ino, 9.06. 2003, coll. P. V. Tuzovskij. The river bottom: stones, pebbles and sand, depth 0.3 – 0.5 m. The holotype is deposited in the collection of Institute for Biology of Inland Waters (Borok, Russia). Paratypes: two females is collected in Ild stream, same locality as holotype, 21.07. 2003 (coll. P. V. Tuzovskij).	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC6794CF3B1FC7A7B63E155.taxon	diagnosis	Diagnosis. Female: setae Fch heaviest and longer of all dorsal setae; setae Oe, Hi and He shorter and thinner than Fch; other idiosomal setae short hair – like; caudal margin of coxal plates I slightly convex, apodemes of coxal plates II in obtuse angle, pedipalpal femur with ventrodistal protruding; bases of ventral setae divide pedipalpal tibia into three approximately equal sectors (1: 1: 1); sword seta on P – IV shorter than distance between bases of ventral setae, base of sword seta placed near ventrodistal seta.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC6794CF3B1FC7A7B63E155.taxon	description	Description. Female. Color yellowish – brown. The body flat oval, integument soft with very fine strips. Number and position of idiosomal setae typical for representatives of the genus Atractides. Setae Fch heaviest and longer of all dorsal ones (Fig. 30). Setae Oe, Hi and He shorter and thinner than Fch; other idiosomal setae short hair – like. Setae Fp and Oi without glandularia, other dorsal setae associated with glandularia. First pair of lyriform organs (i 1) situated close to posterior eye lenses; i 2, in interval between setae Ve and He; i 3, behind humeral row of setae (Hi, He); i 4, more or less corresponding to lumbar row of setae (Li, Le); i 5 in the posterior part of the dorsum. Coxal plates covering less than half of the ventral surface of body. Posteromedial margin of coxal plates I + II wide slightly convex, with short wide median process and two short apodemes which forming obtuse angle (Fig. 31). First coxal plates fused to each other, but median suture line between them distinct. The seta and glandularium Hv situated in posterolateral part of coxal plate II. Medial margin of coxal plate IV oblique and nearly straight, directed posteromedially; seta Pe situated near middle of anterior margin of coxal plate IV; posterior margin of coxal plate IV slightly convex. Capitulum elongate, rostrum rather long, slightly reaching anterior edge of the capitulum. The setae Ci and Pi separated (Fig. 32). Setae Pi without glandularia, other ventral idiosomal setae associated with glandularia. Excretory pore unsclerotized. Genital plates (Fig. 33) elongate and narrow (ratio length / width 2.7 – 3.5) with 3 acetabula and 15 – 19 fine setae each; genital acetabula in arc line, posterior acetabulum larger than anterior and median acetabula. Gonopore longer than genital plates, anterior genital sclerite considerably wider than posterior sclerite. Chelicera with large basal segment and short stylet (Fig. 34). Cheliceral stylet with a few small teeth on dorsal side. Pedipalp (Fig. 35) slender; trochanter short, with a single short dorsodistal seta; femur with ventrodistal protruding, with 3 short dorsoproximal setae and two long unequal dorsodistal setae. Pedipalpal genu slightly longer than femur with straight ventral margin, with 4 thick unequal dorsal setae and 4 – 6 thin subequal setae. Pedipalpal tibia very slender, with slightly convex dorsal margin; bases of ventral setae divide tibia into three approximately equal sectors (1: 1: 1); with a few dorsolateral setae which located mainly in distal half of segment. Ventrolateral sword-like seta moderately in length and shorter than distance between bases of ventral setae, base of sword-like seta placed near ventrodistal seta. Pedipalpal tarsus slightly tapering distally, all spines and thin setae located in distal half of segment. Genu of leg I (Fig. 36) with parallel dorsal and ventral margins, with two short ventroproximal spine near middle of segment, two rather long ventrodistal spines and a few fine setae. Tibia of leg I very slightly thickened distally; bases of setae S 1 and S 2 separated; S 2 shorter and slightly wider than seta S 1, both setae more or less pointed. Tarsus I short, ratio length tibia / tarsus = 1.37 – 1.39; with weakly convex dorsal and concave ventral margins; slightly thickened proximally and distally, with minimum thickness near middle of the segment. Posterior legs, in particular legs IV, very slender; genu and tibia with rather numerous thin and thick setae, tarsus with single ventral spine near middle of segment (Fig. 37). Claws of leg I (Fig. 38) smaller than claws of legs II – IV (Fig. 39). All claws with two pointed denticles, an internal denticle shorter than external denticle, claw lamella with concave ventral margin. Measurements (n = 3). Length of body 785 – 1090; length of setae Fch 135 – 150, length of setae Oe, Hi and He 90 – 105; length of coxae I + II 245 – 270, width 365 – 475; length of medial portion of coxae I 100 – 120; length of coxae III + IV 255 – 290, width 215 – 240; lengths of genital plate 100 – 125, width 36 – 42; length / width of genital acetabula (ac. 1 – 3): 36 – 42 / 24 – 30, 36 – 42 / 24 – 30, 38 – 45 / 36 – 42; length of basal segment of chelicera 113 – 115, length of cheliceral stylet 41 – 43; lengths of pedipalpal segments (P – 1 – 5): 33 – 36, 90 – 96. 102 – 108, 125 – 132, 33 – 38; length / width of sword seta on P – 4: 24 – 30 / 6 – 7; lengths of leg segments: I – Leg. 1 – 6 — 65 – 72, 100 – 110, 135 – 145, 210 – 215, 215 – 220, 155 – 165; II – Leg. 1 – 6 — 65 – 70, 85 – 95, 100 – 125, 180 – 195, 190 – 205, 175 – 185; III – Leg. 1 – 6 — 65 – 72, 90 – 95, 135 – 140, 210 – 225, 225 – 245, 180 – 210; IV – Leg. 1 – 6 — 130 – 155, 135 – 150, 215 – 230, 240 – 290, 300 – 325, 240 – 260; tibia of leg I: length of seta S 1 95 – 102, width 8 – 9; length of seta S 2 72 – 78, width 10 – 12, distance between bases of setae S 1 and S 2 18 – 24. Differential diagnosis. The new species is closely related to Atractides clavipalpis (Lundblad, 1956) and A. samsonellus Tuzovskij, 2006. The new species differs from A. clavipes in the following characters (character states of female A. clavipalpis are indicated in parenthesis from Gerecke 2003): the caudal margin of coxal plates I slightly convex (strongly convex), the apodemes of coxal plates II in obtuse angle (in acute angle), the tibia of leg I without distally protruding (with distally protruding near insertion S 1); the pedipalp rather long, total length 385 – 410 µm (relatively short, total length 364 µm); the sword setae (S 1 and S 2) on tibia of leg I short, respectively 95 – 102 and 72 – 78 µm (relatively long, respectively 121 and 92 µm), the pedipalpal femur with ventrodistal protruding (without ventrodistal protruding). The gonopore and genital plates in female A. samsonellus subequal in length, the excretory pore plate is surrounded by narrow sclerotized ring, the setae S 1 and S 2 on tibia of leg I nearly subequal in shape, leg claws with short internal denticle with rounded tip and long pointed external denticle, P- 2 with convex ventrodistal protrusion (Tuzovskij 2006); in contrast, in female A. ildicus the gonopore longer than genital plates, the excretory pore plate unsclerotized, the setae S 1 and S 2 on tibia of leg I heteromorphic, leg claws with approximately subequal pointed denticles, P- 2 with flat ventrodistal protrusion.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC6794CF3B1FC7A7B63E155.taxon	etymology	Etymology. The species is named after the stream where it was collected (Ild). Habitat. Running waters.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC6794CF3B1FC7A7B63E155.taxon	distribution	Distribution. Europe (Russia, Yaroslavl Province).	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC3794FF3B1FC597B20E075.taxon	materials_examined	Material examined. Holotype female, Russia, Far East, Primory Territory, Dalnegorsk District, Ussuri National Park, the Komarovka River, 10. 07 I. 1984, coll. T. Vshivkova, slide 4951. The holotype is deposited in the collection of Institute for Biology of Inland Waters (Borok, Russia).	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC3794FF3B1FC597B20E075.taxon	diagnosis	Diagnosis. Setae Fch thick, stick – shaped; setae Oe, Hi and He slightly longer but considerably thinner than Fch; rostrum long well outstanding to anterior margin of capitulum; apodemes of anterior coxal plates in right angle, posteromedial margin of coxae I wide and convex; gonopore considerably longer than genital plates; pedipalpal femur short with convex ventral margin but without ventrodistal expansion; sword seta on P – 4 rather long pointed, inserted near ventrodistal seta; tibia of leg I slightly thickened in distal half, with short ventral spine near S 1, seta S 1 longer than S 2 and separated by rather greater interspace, both these setae pointed and approximately equal in width.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC3794FF3B1FC597B20E075.taxon	description	Description. Female. Color yellow in preservative liquid. Body flat oval, integument soft with very fine strips. Number and position of idiosomal setae typical for representatives of the genus Atractides. Setae Fch (Fig. 40) relatively short, thick; setae Oe, Hi and He (Fig. 41) slightly longer but considerably thinner than Fch; trichobothria Fp and Oi without glandularia (Fig. 42), other idiosomal dorsal setae associated with glandularia short and hair-like (Fig. 43). Coxae of legs (Fig. 44) covering about half of ventral surface. Median suture between coxae I clearly developed; central sub – dermal projection and apodemes of anterior coxae short, apodemes in right angle. Posteromedial margin of coxae I wide and convex. Setae Hv located in lateral portion of coxae II. Coxae III + IV trapezoidal, their medial margin rather wide and rounded. Glandularia and seta Pe situated near middle of`anterior margins of coxae IV. Setae Pi without glandularia, their bases fused or separated from setae Ci, excretory pore unsclerotized (Fig. 45). Genital plates (Fig. 46) elongate (ratio length / width = 3.0) with 3 acetabula and 13 thin setae each. Genital acetabula arranged in an arc, all acetabula approximately subequal in length, but posterior pair of acetabula slightly wider than anterior and median acetabula. Gonopore considerably longer than genital plates, anterior and posterior genital sclerites nearly equal in width. Capitulum (Fig. 44) wide, ratio length / width 1.15. Rostrum well developed, outstanding to anterior margin of capitulum; ventral hypostomal setae shorter than dorsal setae. Basal segment of chelicera large, cheliceral stylet well developed, crescent with a few small teeth on concave side (Fig. 47). Pedipalp (Fig. 48) slender; trochanter short with single dorsodistal seta; femur with slightly convex ventral margin, five dorsal unequal setae; genu considerably longer than femur with straight ventral margin, two short proximal and two long stout dorsodistal setae and several subequal thin setae. Pedipalpal tibia straight, slightly longer than genu, with a few dorsal setae; bases of ventral setae divide tibia into three unequal sectors (1 – 2 – 2), sword seta rather large, placed near ventrodistal seta. Legs without swimming setae. Genu of leg I (Fig. 49) not thickened distally with two ventroproximal and two ventrodistal short spine-like setae. Tibia of leg I slightly thickened in distal half, with short ventral spine near S 1 and a few thin dorsal and ventral setae; seta S 1 longer than S 2 and separated by rather greater interspace, both these setae pointed and approximately equal in width; curved seta moderately developed. Tarsus of leg I slender, thin slightly curved dorsoventrally, its anterior, median and terminal portions approximately subequal in thickness. Posterior legs, especially legs IV (Fig. 50) slender, genu and tibia with a few thin and thick short setae, tibia with fine setae only. Claws of leg I lesser than claws of legs II – IV. All leg claws with two denticles, external denticle slightly longer than internal denticle, lamella rather long with concave ventral margin (Fig. 51). Measurements, n = 1. Length of body 975; length of setae Fch 55 – 60, length of setae Oe, Hi and He 65 – 75; length of coxae I + II 290, width 295; length of medial portion of coxae I 100; length of coxae III + IV 230, width 170; lengths of genital plate 110, width 35; length / width of genital acetabula (ac. 1 – 3): 33 – 36 / 24, 30 – 33 / 27 – 30, 30 – 33 / 33 – 35; length of capitulum 125, length of basal segment of chelicera 155, length of cheliceral stylet 65; lengths of pedipalpal segments (P 1 – 5): 32, 66, 80, 90, 30; length / width of sword seta on P 4 25 / 6; lengths of leg segments: I – Leg. 1 – 6 — 55, 90, 120, 190, 185, 145; II – Leg. 1 – 6 — 55, 65, 100, 130, 130, 120; III – Leg. 1 – 6: 50, 60, 95, 155, 170, 140; IV – Leg. 1 – 6 — 95, 85, 150, 215, 230, 180; tibia of leg I: length of seta S 1 100, width 13; length of seta S 2 70, width 15, distance between setae S 1 and S 2 30. Differential diagnosis. The present species is similar to Atractides inflatipalpis (K. Viets, 1950), A. inflatipes (Lundblad, 1956) and A. fluviatilis (Szalay, 1929). The new species differs from Atractides inflatipalpis in the following characters (character states of female A. inflatipalpis are indicated in parenthesis from Gerecke 2003): integument striated (lineated), pedipalpal femur convex without ventrodistal expansion (straight with ventrodistal expansion), length of sword seta on P – 4 nearly equal distance between ventral setae (considerably shorter), tibia and tarsus of leg I relatively short, respectively 185 and 145 µm (rather long, respectively 234 and 194 µm), pedipalp relatively short, total length 296 µm (rather long, total length 363 µm). The gonopore and genital plates of female A. inflatipes approximately subequal in length, the anterior genital sclerite considerable wider than posterior sclerite, setae S 1 and S 2 on tibia of leg I heteromorphic with obtuse tips (Gerecke 2003); in contrast, in female A. komarovkaensis the gonopore longer than genital plates, the anterior and posterior genital sclerites approximately subequal in width, setae S 1 and S 2 on tibia of leg I pointed and nearly subequal in shape. The genital plates of female A. fluviatilis wide and longer than gonopore, acetabula in obtuse triangular; P- 2 with ventrodistal protruding, ventral margin P- 4 slightly protruding near proximoventral seta (Gerecke 2003); in contrast, in female A. komarovkaensis sp. n. the genital plates relatively narrow and shorter than gonopore, acetabula in an arc line; P- 2 without ventrodistal seta protruding, ventral margin P- 4 not protruding near proximoventral seta.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC3794FF3B1FC597B20E075.taxon	etymology	Etymology. The species is named after the river where it was collected (Komarovka). Habitat. Running waters.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC3794FF3B1FC597B20E075.taxon	distribution	Distribution. Asia (Russia, Far East, Primory Territory).	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC07952F3B1FDB97B20E37E.taxon	materials_examined	Material examined. Holotype female, Russia, Primorie Territory, Octyabrsky District, Razdolnaya River, 4. 06. 1990, coll. T. S. Vshivkova, slide 6308. The holotype is deposited in the collection of Institute for Biology of Inland Waters (Borok, Russia).	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC07952F3B1FDB97B20E37E.taxon	diagnosis	Diagnosis. Setae Fch thick, stick – shaped with parallel lateral margins; setae Oe, Hi and He slightly longer but considerably thinner than Fch; the tibia of leg I well thickened in distal half, with not parallel dorsal and ventral margins, both the sword setae (S 1 and S 2) wide in proximal half and not narrowed near their bases; tarsus of leg I moderately curved, ratio length tibia / length tarsus 1.32; genital plates longer than gonopore, all genital acetabula approximately subequal in length, anterior and posterior genital sclerites approximately equal in width.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC07952F3B1FDB97B20E37E.taxon	description	Description. Female. Color yellow in preservative liquid. Body flat oval, integument smooth with very fine strips. Number and position of idiosomal setae typical for representatives of the genus Atractides. Setae Fch (Fig. 52) thick, stick – shaped with more or less rounded tips; setae Oe, Hi and He (Fig. 53) slightly longer but considerably thinner than Fch; trichobothria Fp and Oi thin without glandularia (Fig. 54), other dorsal idiosomal setae hair – like and associated with glandularia short (Fig. 55). Coxal plates of legs I – IV covering about half of ventral surface. Median suture line between coxae I clearly developed; median sub – dermal process short, apodemes of anterior coxal plates rather long, slightly curved and forming right angle (Fig. 56). Posteromedial margin of coxal plates I wide and convex. Setae Hv located in lateral portion of coxal plates II. Coxal plate IV trapezoidal, its medial margins slightly concave. Glandularia Pe situated at anterior margins of coxae IV. Setae Pi without glandularia, their bases separated from setae Ci, excretory pore unsclerotized (Fig. 57). Genital plates (Fig. 58) elongate (ratio length / width 2.6 – 2.8), with 3 acetabula and 12 thin setae each; genital acetabula arranged in an arc, all acetabula approximately subequal in length, but posterior pair of acetabula wider than anterior and median acetabula. Genital plates longer than gonopore, anterior and posterior genital sclerites approximately equal in width. Basal segment of chelicera (Fig. 59) large with straight dorsal margin, cheliceral stylet crescent and relatively short, ratio length of basal segment of chelicera / length cheliceral stylet 2.4. Cheliceral stylet with a few small teeth on concave side. Pedipalps (Fig. 60) slender; trochanter short with single dorsodistal seta; femur with straight ventral margin and rounded ventrodistal edge, with five dorsal unequal setae; genu considerably longer than femur with straight ventral margin, with two short proximal and two long stout dorsodistal unequal setae and several hair-like setae. Pedipalpal tibia slightly longer than genu, with a few thin dorsal setae; bases of two ventral setae divide tibia into three sectors approximately as 2 – 3 – 2; sword seta pointed, shorter than distance between ventral setae and placed near ventrodistal seta. Legs without swimming setae. Genu of leg I (Fig. 61) slightly thickened distally with a few thin setae and one short ventral spine near middle of segment. Tibia of leg I thickened in distal half with maximum thickness near proximal sword seta; setae S 1 and S 2 separated by rather greater interspace, both these setae approximately equal in width but S 1 longer than S 2, S 2 pointed, S 1 with rounded tip; curved seta moderately developed. In addition, tibia I with short ventral spine near proximal sword seta and with a few thin setae. Tarsus of leg I (Fig. 62) moderately equally curved dorsoventrally, thickened basally and distally with minimum thickness near middle of segment. Posterior legs, especially legs IV (Fig. 63), slender; genu and tibia with a few thin setae and several spines, tarsus with a few thin setae and one ventral spine near middle of segment. Claws of legs I lesser than claws II – IV. Claws all legs with two denticles, external denticle longer than internal denticle; claw lamella well developed with slightly concave ventral margin (Fig. 64). Measurements, n = 1. Length of body 770; length of setae Fch 72, length of setae Oe, Hi and He 78 – 95; diameter or length of sclerite bearing setae Fch 24, diameter of sclerite bearing setae Le 21, diameter or length of sclerite bearing other idiosomal setae Fch 18; length of coxae I + II 210, width 370; length of coxae III + IV 230, width 180; lengths of genital plate 95 – 100, width 35; length / width of genital acetabula (ac. 1 – 3): 33 – 36 / 27 – 30, 36 – 42 / 27 – 30, 36 – 37 / 30 – 32; length of capitulum 125, length of basal segment of chelicera 155, length of cheliceral stylet 65; lengths of pedipalpal segments (P – 1 – 5): 30, 66 – 70, 90 – 95, 95 – 102, 30; length / width of sword seta on P – 4 24 / 9; lengths of leg segments: I – Leg. 1 – 6 — 48, 95, 145, 215, 205, 155; II – Leg. 1 – 6: 55, 78, 108, 150, 150, 145; III – Leg. 1 – 6: 60, 65, 115, 160, 175, 150; IV – Leg. 1 – 6 — 125, 110, 180, 235, 260, 200; tibia of leg I: length of seta S 1 100, width 15; length of seta S 2 78, width 16; distance between setae S 1 and S 2 35. Differential diagnosis. The present species is similar to Atractides montanus (Halbert, 1911), from which it is easily distinguishable by the structure of the tibia and tarsus of leg I, small genital acetabula and arranged of anterior coxal apodemes. The female of A. montanus characterized by follows features: the tibia of leg I with nearly parallel dorsal and ventral margins, both the sword setae (S 1, S 2) slightly narrowed near their base, interspace between setae S 1 and S 2 26 µm; tarsus of leg I strongly curved, ratio length tibia / length tarsus 1.43; posterior acetabula considerably larger than anterior and median acetabula, length of acetabula (ac. 1 – 3): 39, 43, 61 µm, anterior genital sclerite considerably wider than posterior sclerite (Gerecke 2003). In contrast, in the female A. primoryensis the tibia of leg I without parallel dorsal and ventral margins, both the sword setae (S 1, S 2) wide and not narrowed near their bases, interspace between setae S 1 and S 2 35 µm; the tarsus of leg I moderately curved, the ratio length tibia / length tarsus 1.32; all acetabula approximately subequal in length, length of acetabula (ac. 1 – 3): 33 – 36, 38 – 42, 33 – 37 µm, anterior and posterior genital sclerites equal in width.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC07952F3B1FDB97B20E37E.taxon	etymology	Etymology. The species epithet primoryensis is derived from the name of the Province where it was collected (Primory). Habitat. Running waters.	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
03812A52FFC07952F3B1FDB97B20E37E.taxon	distribution	Distribution. Asia (Russia, Far East, Primory Territory).	en	Tuzovskij, Petr V. (2011): Four new water mite species of the genus Atractides Koch (Acari: Hydrachnidia: Hygrobatidae) from Russia. Zootaxa 2738: 41-59, DOI: 10.5281/zenodo.206972
