taxonID	type	description	language	source
A0A24C0AE674173E2B812FB31B0B9625.taxon	description	Figures 2 D, 4, 5, 6 D, 8 D, 9 D, 13	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A0A24C0AE674173E2B812FB31B0B9625.taxon	diagnosis	Diagnosis. Palpus broad, flat, with a few moderate setae on outer lateral surface (Figure 9 D). Paramere apically truncated in lateral view (Figure 13 K). Aedeagus apically finely serrated along ventrolateral margin (Figure 13 I). Nucleotide status in COI sequence = C and T at sites 391 and 589, respectively (Table 3).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A0A24C0AE674173E2B812FB31B0B9625.taxon	description	Description (characters in common with D. puberula sp. n. not repeated). Adult ♂. Head (Figures 2 D, 4 - 6 D, 8 D, 9 D): Arista with eight dorsal and three ventral branches. Supracervical setae 24 - 27 per side; postocular setae 14 - 17 per side. Cibarium with ca. nine medial and ca. 13 posterior sensilla per side. Prementum with five (one proximal, two lateral, and two distal) pairs of setae. Thorax (Figure 2 D): Postpronotal lobe with two prominent and three short setae. Anterior dorsocentral setae slightly convergent; posterior dorso-central setae nearly parallel. Thoracic pleura with four indistinct stripes. Legs (Figures 2 D): First tarsomeres of all legs slightly shorter than total length of four succeeding tarsomeres. Male terminalia (Figure 13): Cercus with 29 - 32 setae, including ca. ten ventral, small ones. Surstylus with ca. ten prensisetae and ca. eleven ventral spines. Hypandrium not pubescent. Paramere apically rounded in ventral view, with 1 - 2 sensilla. Aedeagus apically not darkened. Measurements (in mm): BL = 2.50 in holotype, ThL = 1.08, WL = 2.10, WW = n / a. Indices. FW / HW = 0.59, ch / o = 0.09, prorb = 0.63, rcorb = 0.24, vb = n / a, orbito = 0.50, dcl = 0.67, sctl = 1.02, sterno = 0.70, dcp = 0.50, sctlp = 0.91, C = 1.71, 4 c = 1.31, 4 v = 1.65, 5 x = 1.38, ac = 2.83, M = 0.69, C 3 F = 0.49.	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A0A24C0AE674173E2B812FB31B0B9625.taxon	distribution	Distribution. Indonesia (West Java).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A0A24C0AE674173E2B812FB31B0B9625.taxon	etymology	Etymology. Referring to the white palpus in the new species.	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
953F02FC13AA6B8D3601FDEE406E0DAD.taxon	description	Figures 3 C, 4 G, 5 G, 7 C, 8 G, 9 G, 16	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
953F02FC13AA6B8D3601FDEE406E0DAD.taxon	diagnosis	Diagnosis. Palpus short, club-shaped, medio- to baso-laterally dark grayish brown, with one prominent seta apically and several long setae ventrally (Figure 9 G). Cercus without caudoventral process (Figure 16 E). Paramere apically somewhat quadrate in lateral view (Figure 16 J), ventroapically with three sensilla (Figure 16 I, J). Aedeagus distally dilated laterally, somewhat lunate in lateral view (Figure 16 F-H).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
953F02FC13AA6B8D3601FDEE406E0DAD.taxon	description	Description (characters in common with D. orthophallata sp. n. not repeated). Adult ♂. Head (Figures 3 C, 4 G, 5 G, 7 C, 8 G, 9 G): Longest axis of eye nearly rectangular to body axis. Frontal vitta grayish white. Fronto-orbital plate slightly grayish; anterior reclinate orbital seta situated between proclinate and posterior reclinate orbital setae. Occiput and postgena dark brown, marginally milky white. Arista with six dorsal and three ventral branches. Supracervical setae 16 - 19 per side; postocular setae 17 - 19 per side. Cibarium slightly thickened on anterior margin; medial sensilla ca. nine per side and posterior sensilla ca. nine per side; first and second medial sensilla weaker than and anteriorly apart from others. Clypeus not thickened at median portion, laterally dark brown. Thorax (Figure 3 C): Postpronotal lobe pale brownish white in upper half, dark brown in lower half; setae broken. Dorsocentral and scutellar setae broken. Notopleuron pale brownish white. Thoracic pleura nearly entirely dark brown, without stripes. Wing (Figure 3 C) slightly wrinkled at basal portion of R 4 + 5; longitudinal veins pale brown except for basal section of M 1 (brown) and CuA 1 (brown). Legs (Figure 3 C) pale grayish yellow to pale yellow. Foreleg first tarsomere shorter than total length of four succeeding tarsomeres. Mid- and hind-leg first tarsomeres slightly shorter than total length of four succeeding tarsomeres, without subproximal spine. Abdomen (Figure 3 C): Tergites pale brown, each with dark brown caudal band narrower than ca. 1 / 2 of tergite but laterally extended anteriorly. Male terminalia (Figure 16): Epandrium with ca. two and ca. 16 long setae per side on caudodorsal and ventral portions, respectively. Cercus with 30 - 31 setae. Surstylus with ca. eight prensisetae and ca. ten ventral spines; basal sclerotized ridge indistinct. Tenth sternite flat, not wrinkled. Hypandrium somewhat oval in anterior portion. Aedeagal apodeme apically not flattened, slightly shorter than aedeagus. Measurements (in mm): BL = 2.33 in holotype, ThL = 1.04, WL = 2.35, WW = 1.16. Indices. FW / HW = 0.60, ch / o = 0.08, prorb = n / a, rcorb = n / a, vb = n / a, orbito = 0.78, dcl = n / a, sctl = n / a, sterno = 0.62, dcp = 0.65, sctlp = 0.94, C = 2.09, 4 c = 1.21, 4 v = 1.95, 5 x = 1.24, ac = 2.83, M = 0.64, C 3 F = 0.53.	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
953F02FC13AA6B8D3601FDEE406E0DAD.taxon	distribution	Distribution. China (Yunnan).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
953F02FC13AA6B8D3601FDEE406E0DAD.taxon	etymology	Etymology. Referring to the morphological difference from the other species in the subgenus Dudaica.	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
28AB4757E0A1616C2D254AB43FC8D5C4.taxon	diagnosis	Diagnosis. Head, scutum, and scutellum mostly milky white, contrasting with mostly dark brown thoracic pleura (Figures 2 - 4). Scutellum large, more or less rounded apically in dorsal view (Figures 2, 3). Wing fuscous, somewhat wavy (Figures 2, 3).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
28AB4757E0A1616C2D254AB43FC8D5C4.taxon	description	Common characters. Head (Figures 2 - 9): Eye red, with dense interfacetal setulae; longest axis of eye slightly oblique (nearly rectangular in Drosophila dissimilis sp. n.) against body axis. Ocellar triangle convex; ocellar setae inserted outside triangle made by ocelli. Anterior reclinate orbital seta situated slightly before proclinate orbital seta or just lateral to it (between proclinate and posterior reclinate orbital setae in Drosophila dissimilis sp. n.) (Figure 4). Pedicel dorsolaterally dark brown (Figure 4); first flagellomere pubescent; arista with 6 - 8 dorsal and 2 - 4 ventral branches in addition to terminal bifurcation; terminal bifurcation moderate. Facial carina high, broad. Gena anteriorly dark brown. Occiput ventrally dark brown. Postgena medially dark brown. Postocellar setae present (Figure 5). Supracervical setae tapered, thin, apically curved and slightly blunt (Figure 5). Cibarium not thickened on anterior margin in lateral view (slightly thickened in Drosophila dissimilis sp. n.); anterior portion slightly dilated; anterolateral corners slightly protruded; dorsal wall pear-shaped, with posterior portion oval; anterior sensilla four, arranged in square; medial sensilla apically sharp, arranged in anteriorly slightly convergent rows; posterior sensilla apically blunt, arranged in anteriorly divergent rows (Figures 6, 7). Clypeus thick at median portion (except for Drosophila dissimilis sp. n.) (Figures 6, 7). Palpus distinctly large, long and broad (Figures 4, 9) (except for Drosophila dissimilis sp. n.), pubescent, setigerous, distally flat (except for Drosophila dissimilis sp. n.) (Figures 9). Prementum dark brown, swollen at the distal end in lateral view (Figure 8). Labellum with five pseudotracheae per side. Thorax (Figures 2, 3): Postpronotal lobe with 1 - 3 prominent (lowermost the longest) and 0 - 4 short setae. Scutum narrowly dark brown along anteromost margin. Thoracic pleura mostly covered with broad, dark brown, more or less blurry, longitudinal stripes (except for Drosophila dissimilis sp. n.). Basal scutellar setae divergent; apical scutellar setae cruciate. Acrostichal setulae in six, somewhat regular rows. Mid katepisternal seta much shorter than anterior and posterior ones, but distinct from setulae in row below it; caudoventral corner of katepisternum with one long, prominent seta. Wing (Figures 2, 3) pale grayish yellow to grayish yellow, elliptic, rounded distally (not so elliptic in Drosophila dissimilis sp. n.), wrinkled especially at basal portion of R 4 + 5; longitudinal veins brown except for R 2 + 3 (pale brown), basal section of M 1 (dark brown), and CuA 1 (dark brown) (except for Drosophila dissimilis sp. n.); crossveins shaded at r-m and dm-cu; C 1 setae two, subequal; R 2 + 3 distally slightly curved to costa; R 4 + 5 basally diverged from M 1, distally nearly parallel with M 1; M 1 more or less sinuate; A 1 well developed, as stout as other veins. Halter entirely grayish yellow to grayish brown. Legs (Figures 2, 3, 10 A, 11 A) pale grayish yellow to grayish brown. Preapical dorsal setae present on tibiae of all legs; apical setae present on fore- and mid-leg tibiae. Foreleg first tarsomere with one subproximal and one apical, short, black spines. Abdomen (Figures 2, 3): Tergite 1 nearly entirely dark brown, 2 to 6 pale yellow, each with anterior and caudal dark brown bands; anterior bands medially sometimes interrupted; caudal bands medially and laterally extended anteriad (except for Drosophila dissimilis sp. n.). Female tergite 7 nearly entirely pale yellow. Sternites somewhat quadrate, grayish brown to dark brown. Male terminalia (Figures 10 B-K, 11 B-L, 12 A-K, 13, 14 A-K, 15 A-H, 16): Epandrium pale brown, pubescent except for anterior sub-dorsal to - ventral margin, ventrally narrowed, ventroapically rounded, with setae on caudodorsal and ventral portions; lobe-like apodeme present anterosubdorsally. Cercus dark brown, nearly entirely pubescent, separated from epandrium, caudoventrally with distinct process (except for Drosophila senilis and Drosophila dissimilis sp. n.). Surstylus more or less quadrangular; dorsoproximal portion broadly fused to epandrium, with sclerotized ridge connecting epandrium and surstylus (except for Drosophila dissimilis sp. n.); outer surface not pubescent, anterosubmedially concaved; caudal margin with a slightly sinuate row of peg-like, apically more or less roundish prensisetae decreasing in size ventrally; ventral portion with apically pointed spines on either inner or outer surface: spines on inner, subventral surface longer, somewhat curved upwards. Tenth sternite pale brown, moderately sclerotized, anteromedially wrinkled (flat in Drosophila dissimilis sp. n.). Hypandrium narrowly triangular (except for Drosophila orthophallata sp. n. and Drosophila dissimilis sp. n.), anteriorly with narrow, well developed apodeme, not pubescent (except for Drosophila malayana, Drosophila gracilipalpis sp. n., and Drosophila puberula sp. n.), caudolaterally with a pair of somewhat expanded lobes; a pair of paramedian setae present on the portion fused to paramere. Paramere elongated, apically rounded in ventral view (except for Drosophila puberula sp. n.), ventrosubapically with 1 - 2 sensilla (three in Drosophila dissimilis sp. n.), basally fused to aedeagus, ventrally fused to hypandrium. Aedeagus rod-like, fused to aedeagal apodeme, pale brown to brown, apically darker, slightly curved dorsad (straight in Drosophila orthophallata sp. n.); aedeagal guide and basal processes absent; apodeme anteriorly expanded in lateral view, shorter than aedeagus. Female terminalia (Figures 10 L-N, 11 M-O, 12 L-N, 14 L-N, 15 I, J): Tergite 8 pale brown, pubescent dorsally to caudolaterally. Epiproct dark brown, entirely pubescent. Hypoproct dark brown, laterally pubescent. Sternite 7 grayish brown, caudally darker, caudomedially deeply notched, nearly entirely pubescent; setae on caudal portion rather long. Oviscapt valve with stout lateral and marginal ovisensilla increasing in size posteriad; apical ovisensillum stout and the largest, bent outwards. Spermathecal capsule pale brown, spherical.	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A015F54B391A0F8622BC954EC26B1746.taxon	description	Figures 2 B, 4, 5, 6 B, 8 B, 9 B, 11	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A015F54B391A0F8622BC954EC26B1746.taxon	diagnosis	Diagnosis. Palpus long, sinuate, with several stout setae only on basal portion (Figure 9 B). Foreleg tibia with one apical seta (Figure 11 A). Cercus with distinct caudoventral process (Figure 11 F). Hypandrium pubescent, but very indistinctly, on caudolateral lobes (Figure 11 J).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A015F54B391A0F8622BC954EC26B1746.taxon	description	Description (characters in common with D. malayana not repeated). Adult ♂ and ♀. Head (Figures 2 B, 4 - 6 B, 8 B, 9 B): Arista with 7 - 8 dorsal and 3 - 4 ventral branches. Supracervical setae 26 - 27 per side; postocular setae 12 - 14 per side. Cibarium with ca. ten medial and ca. nine posterior sensilla per side. Prementum with six (one proximal, three lateral, and two distal) pairs of setae. Thorax (Figure 2 B): Postpronotal lobe with two prominent and rarely 1 - 2 short setae. Stripes on thoracic pleura mostly confluent with each other. Abdomen (Figure 2 B): Anterior bands on tergites 5 and / or 6 (and 7 in female) medially broadly interrupted. Male terminalia (Figure 11 B-L): Epandrium with ca. two and 10 - 12 long setae per side on caudodorsal and ventral portions, respectively. Cercus with 22 - 24 setae. Surstylus with 8 - 9 prensisetae and ca. seven ventral spines. Paramere ventrosubapically with one sensillum (longer in Indonesian specimen than in Chinese ones). Aedeagal apodeme ca. 1 / 2 length of aedeagus. Female terminalia (Figure 11 M-O): Tergite 8 with 4 - 5 small setae on ventral portion but no on caudodorsal portion. Oviscapt valve with three lateral and 12 - 13 marginal ovisensilla. Spermathecal capsule apically slightly narrowed, slightly broader than long. Measurements (in mm): BL = 2.20 in holotype (range in 8 ♂ / 2 ♀ paratypes: 1.63 - 2.50 / 2.03 - 2.63), ThL = 1.00 (0.88 - 1.06 / 0.98 - 1.06), WL = 1.84 (1.60 - 2.15 / 2.03 - 2.10), WW = 0.92 (0.83 - 1.04 / 1.02 - 1.06). Indices. FW / HW = 0.59 (range in 8 ♂ and 2 ♀, or less if noted, paratypes: 0.58 - 0.63), ch / o = 0.12 (0.07 - 0.13), prorb = 0.65 (6 ♂, 2 ♀: 0.46 - 0.63), rcorb = 0.30 (7 ♂, 2 ♀: 0.20 - 0.37), vb = 0.45 (7 ♂, 2 ♀: 0.22 - 0.60), orbito = 0.63 (0.50 - 0.64), dcl = 0.72 (0.63 - 0.79), sctl = 1.26 (7 ♂, 1 ♀: 0.94 - 1.12), sterno = 0.56 (0.44 - 0.71), dcp = 0.38 (0.41 - 0.57), sctlp = 0.95 (0.91 - 1.00), C = 1.70 (1.56 - 2.00), 4 c = 1.45 (1.32 - 1.58), 4 v = 2.05 (1.83 - 2.26), 5 x = 2.38 (1.80 - 2.43), ac = 2.90 (2.87 - 3.29), M = 0.90 (0.75 - 0.95), C 3 F = 0.56 (0.52 - 0.59).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A015F54B391A0F8622BC954EC26B1746.taxon	distribution	Distribution. Indonesia (West Java), China (Yunnan).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A015F54B391A0F8622BC954EC26B1746.taxon	etymology	Etymology. Referring to the long and thin palpus.	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
6F482B466F2A6F3F77E25C24717F863D.taxon	description	Figures 2 A, 4, 5, 6 A, 8 A, 9 A, 10	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
6F482B466F2A6F3F77E25C24717F863D.taxon	materials_examined	Specimens examined. MALAYSIA: 2 ♂ (holotype and paratype of Zygothrica malayana Takada, 1976), near Kuala Lumpur, Peninsular Malaysia, 3. vii. 1972, H Takada (SEHU); 2 ♂, Poring, Sabah, 16. iii. 1999, MJ Toda (KPSP, SEHU); 1 ♂ (# 03904), same except for 13. iii. 2008 (KIZ); 1 ♂ (# 03903), same except for 20. iii. 2008 (SEHU). INDONESIA: 1 ♂, 1 ♀, Gunung Poteng, West Kalimantan, 4. xii. 1996, MJ Toda (MZB, SEHU).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
6F482B466F2A6F3F77E25C24717F863D.taxon	diagnosis	Diagnosis. Palpus long, sinuate, with several stout setae only on basal portion (Figure 9 A). Foreleg tibia with two apical setae (Figure 10 A). Cercus with distinct caudoventral process (Figure 10 F). Hypandrium pubescent on caudolateral lobes (Figure 10 J).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
6F482B466F2A6F3F77E25C24717F863D.taxon	description	Supplementary and revised description. Adult ♂ and ♀. Head (Figures 2 A, 4 - 6 A, 8 A, 9 A): Arista with 7 - 8 dorsal and four ventral branches. Supracervical setae 25 - 30 per side; postocular setae 15 - 16 per side. Cibarium with ca. eight medial and ca. ten posterior sensilla per side. Prementum with five (one proximal, two lateral, and two distal) pairs of setae. Thorax (Figure 2 A): Postpronotal lobe milky white in upper half, dark brown in lower half, with 2 - 3 prominent but no short setae. Right and left dorsocentral setae nearly parallel. Notopleuron milky white. Thoracic pleura with three, sometimes rather indistinct stripes. Legs (Figures 2 A, 10 A): Foreleg first tarsomere shorter than total length of four succeeding tarsomeres. Mid- and hind-leg first tarsomeres shorter than or as long as total length of four succeeding tarsomeres. Abdomen (Figure 2 A): Anterior bands on tergite 6 medially broadly interrupted. Male terminalia (Figure 10 B-K): Epandrium with ca. three and ca. 13 long setae per side on caudodorsal and ventral portions, respectively. Cercus with 23 - 24 setae. Surstylus with ca. nine prensisetae and 7 - 8 ventral spines. Paramere apically rounded in lateral view, with 1 - 2 sensilla. Aedeagus dorsoapically with a small, acute claw; apodeme slightly longer than 1 / 2 length of aedeagus. Female terminalia (Figure 10 L-N): Tergite 8 with two small setae on ventral portion but no on caudodorsal portion. Oviscapt valve yellowish brown, with three lateral, 13 marginal ovisensilla, and four (three dorsal, one ventral) subterminal, inner, trichoid ovisensilla. Spermathecal capsule slightly longer than broad; introvert ca. 1 / 5 height of capsule. Measurements (in mm): BL (straight distance from anterior edge of pedicel to tip of abdomen) = 2.13 - 2.39 / 2.27 (range in 2 ♂ / 1 ♀ specimens), ThL (distance from anterior notal margin to apex of scutellum) = 0.98 - 1.08 / 1.01, WL (distance from humeral cross vein to wing apex) = 1.78 - 1.92 / 1.73, WW (maximum wing width) = 0.89 - 0.96 / 0.86. Indices. FW / HW (frontal width / head width) = 0.56 - 0.60 (range in 2 ♂ and 1 ♀ specimens), ch / o (maximum width of gena / maximum diameter of eye) = 0.08 - 0.12, prorb (proclinate orbital seta / posterior reclinate orbital seta in length) = 0.61 - 0.62, rcorb (anterior reclinate orbital seta / posterior reclinate orbital seta in length) = 0.29 - 0.32, vb (subvibrissa / vibrissa in length) = 0.41 - 0.59, orbito (distance between proclinate and posterior reclinate orbital setae / distance between inner vertical and posterior reclinate orbital setae) = 0.50 - 0.67, dcl (anterior dorsocentral seta / posterior dorsocentral seta in length) = 0.64 - 0.71, sctl (basal scutellar seta / apical scutellar seta in length) = 0.95 - 0.98, sterno (anterior katepisternal seta / posterior katepisternal seta in length) = 0.59 - 0.74, dcp (distance between ipsilateral dorsocentral setae / distance between anterior dorsocentral setae) = 0.39 - 0.44, sctlp (distance between ipsilateral scutellar setae / distance between apical scutellar setae) = 0.94 - 1.09, C (2 nd costal section between subcostal break and R 2 + 3 / 3 rd costal section between R 2 + 3 and R 4 + 5) = 1.75 - 1.81, 4 c (3 rd costal section between R 2 + 3 and R 4 + 5 / M 1 between r-m and dm-cu) = 1.25 - 1.36, 4 v (M 1 between dm-cu and wing margin / M 1 between r-m and dm-cu) = 1.75 - 1.95, 5 x (CuA 1 between dm-cu and wing margin / dm-cu between M 1 and CuA 1) = 2.12 - 2.25, ac (3 rd costal section between R 2 + 3 and R 4 + 5 / distance between distal ends of R 4 + 5 and M 1) = 2.56 - 3.00, M (CuA 1 between dm-cu and wing margin / M 1 between r-m and dm-cu) = 0.75 - 0.86, C 3 F (length of heavy setation in 3 rd costal section / length of 3 rd costal section) = 0.50 - 0.62.	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
6F482B466F2A6F3F77E25C24717F863D.taxon	distribution	Distribution. Malaysia (Peninsular Malaysia, Sabah *), Indonesia * (West Kalimantan). * New records.	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
EC2AB67D24A721AB4D3CBB2329D31BFD.taxon	description	Figures 3 B, 4 F, 5 F, 7 B, 8 F, 9 F, 15	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
EC2AB67D24A721AB4D3CBB2329D31BFD.taxon	diagnosis	Diagnosis. Palpus broad, flat, with a few moderate setae on outer lateral surface (Figure 9 F). Paramere apically somewhat truncated in lateral view (Figure 15 G). Aedeagus straight, subapically swollen in lateral view, apically without small, acute claw (Figure 15 G).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
EC2AB67D24A721AB4D3CBB2329D31BFD.taxon	description	Description (characters in common with D. puberula sp. n. not repeated). Adult ♂ and ♀. Head Figures 3 B, 4 F, 5 F, 7 B, 8 F, 9 F): Arista with 6 - 7 dorsal and 2 - 3 ventral branches. Supracervical setae 25 - 26 per side; postocular setae 16 - 18 per side. Cibarium with ca. nine medial and ca. ten posterior sensilla per side. Prementum with five (one proximal, two lateral, and two distal) pairs of setae. Thorax (Figure 3 B): Postpronotal lobe with 1 - 2 prominent and 1 - 3 short setae. Thoracic pleura with four, slightly indistinct stripes. Legs (Figure 3 B): Foreleg first tarsomere shorter than or as long as total length of four succeeding tarsomeres. Mid- and hind-leg first tarsomeres as long as total length of four succeeding tarsomeres. Male terminalia (Figure 15 A-H): Epandrium with ca. three and ca. 13 long setae per side on caudodorsal and ventral portions, respectively. Cercus with 26 - 28 setae. Surstylus with ca. eight prensisetae and 7 - 8 ventral spines. Tenth sternite damaged. Hypandrium somewhat triangular, anteriorly round, not pubescent. Aedeagus apically not darkened; apodeme ca. 2 / 5 length of aedeagus. Female terminalia (Figure 15 I, J): Tergite 8 with 3 - 4 and ca. three setae on ventral and caudodorsal portions, respectively. Oviscapt valve with three lateral and 17 - 19 marginal ovisensilla. Data of spermatheca unavailable. Measurements (in mm): BL = 2.34 in holotype (range in 2 ♀ paratypes: 2.20 - 2.60), ThL = 1.18 (1.18 - 1.40), WL = 2.00 (2.35 - 2.63), WW = 1.06 (1.28 - 1.37). Indices. FW / HW = 0.56 (range in 2 ♀, or less if noted, paratypes: 0.56 - 0.63), ch / o = 0.09 (0.12), prorb = 0.68 (1 ♀: 0.67), rcorb = 0.34 (1 ♀: 0.31), vb = n / a (1 ♀: 0.50), orbito = 0.65 (0.40 - 0.55), dcl = 0.65 (0.71 - 0.75), sctl = 1.00 (1 ♀: 1.04), sterno = 0.58 (0.63 - 0.71), dcp = 0.44 (0.43), sctlp = 0.72 (0.73 - 0.85), C = 1.72 (1.76 - 1.78), 4 c = 1.25 (1.27 - 1.28), 4 v = 1.61 (1.59 - 1.67), 5 x = 1.33 (1.33 - 1.44), ac = 3.56 (2.93 - 3.04), M = 0.63 (0.67 - 0.72), C 3 F = 0.56 (0.54 - 0.58).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
EC2AB67D24A721AB4D3CBB2329D31BFD.taxon	distribution	Distribution. Malaysia (Sabah).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
EC2AB67D24A721AB4D3CBB2329D31BFD.taxon	etymology	Etymology. Referring to the straight aedeagus in the new species.	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
5F614E24C984F1327DC32C0B58D21BC4.taxon	description	Figures 2 C, 4, 5, 6 C, 8 C, 9 C, 12	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
5F614E24C984F1327DC32C0B58D21BC4.taxon	diagnosis	Diagnosis. Palpus slightly shorter than arista, with 3 - 4 prominent, stout setae on outer lateral surface (Figure 9 C). Hypandrium pubescent on portion fused to paramere (Figure 12 I, J). Paramere apically sharp (Figure 12 I-K).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
5F614E24C984F1327DC32C0B58D21BC4.taxon	description	Description (characters in common with D. gracilipalpis sp. n. not repeated). Adult ♂ and ♀. Head (Figures 2 C, 4 - 6 C, 8 C, 9 C): Arista with 6 - 8 dorsal and 2 - 3 ventral branches. Supracervical setae 20 - 21 per side; postocular setae 15 - 18 per side. Cibarium with ca. nine medial and ca. eight posterior sensilla per side. Prementum with 5 - 6 (one proximal, 2 - 3 lateral, and two distal) pairs of setae. Thorax (Figure 2 C): Postpronotal lobe entirely dark brown, with 1 - 3 prominent and 2 - 4 short setae. Right and left dorsocentral setae slightly convergent. Notopleuron dark brown. Thoracic pleura with four stripes. Legs (Figure 2 C): First tarsomeres of all legs shorter than total length of four succeeding tarsomeres; mid-leg first tarsomere with one subproximal short, black spine. Male terminalia (Figure 12 A-K): Epandrium with ca. three and 10 - 13 long setae per side on caudodorsal and ventral portions, respectively. Cercus with 33 - 35 setae. Surstylus with ca. nine prensisetae and ca. eight ventral spines. Hypandrium not pubescent on caudolateral lobes. Paramere apically with one sensillum. Aedeagus apically darkened; apodeme laterally flat in muscle-attaching portion. Female terminalia (Figure 12 L-N): Tergite 8 with ca. four small setae on ventral portion but no on caudodorsal portion. Oviscapt valve with three lateral and 13 - 14 marginal ovisensilla. Spermathecal capsule apically round, not narrowed. Measurements (in mm): BL = 2.30 in holotype (range in 5 ♂ / 1 ♀ paratypes: 1.92 - 2.43 / 2.69), ThL = 0.88 (0.98 - 1.14 / 1.30), WL = 1.86 (1.82 - 2.10 / 2.35), WW = 0.98 (0.88 - 1.10 / 1.20). Indices. FW / HW = 0.57 (range in 5 ♂ and 1 ♀, or less if noted, paratypes: 0.55 - 0.58), ch / o = 0.11 (0.07 - 0.18), prorb = 0.61 (0.54 - 0.58), rcorb = 0.30 (5 ♂: 0.27 - 0.38), vb = 0.34 (0.33 - 0.41), orbito = 0.63 (0.50 - 0.75), dcl = 0.65 (4 ♂, 1 ♀: 0.55 - 0.74), sctl = 1.05 (4 ♂, 1 ♀: 1.00 - 1.10), sterno = 0.67 (5 ♂: 0.42 - 0.67), dcp = 0.52 (0.42 - 0.55), sctlp = 1.05 (0.78 - 1.00), C = 1.67 (1.50 - 1.79), 4 c = 1.43 (1.36 - 1.58), 4 v = 2.00 (1.75 - 2.16), 5 x = 1.70 (1.79 - 2.11), ac = 3.00 (2.83 - 3.58), M = 0.81 (0.76 - 0.89), C 3 F = 0.55 (0.53 - 0.59).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
5F614E24C984F1327DC32C0B58D21BC4.taxon	distribution	Distribution. China (Yunnan).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
5F614E24C984F1327DC32C0B58D21BC4.taxon	etymology	Etymology. Referring to the pubescence of hypandrium on the portion fused to paramere in the new species.	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A90219302E04F582A327385D39003DA5.taxon	description	Figures 3 A, 4 E, 5 E, 7 A, 8 E, 9 E, 14	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A90219302E04F582A327385D39003DA5.taxon	diagnosis	Diagnosis. Palpus slender, not so flat, somewhat rod-shaped in lateral view, with a few moderate setae on outer lateral surface (Figure 9 E). Paramere apically rounded in lateral view (Figure 14 K). Nucleotide status in COI sequence = C and C at sites 92 and 226, respectively (Table 3).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A90219302E04F582A327385D39003DA5.taxon	description	Description (characters in common with D. puberula sp. n. not repeated). Adult ♂ and ♀. Head (Figures 3 A, 4 E, 5 E, 7 A, 8 E, 9 E): Arista with 6 - 8 dorsal and three ventral branches. Supracervical setae 23 - 24 per side; postocular setae 15 - 17 per side. Cibarium with ca. ten medial and ca. nine posterior sensilla per side. Prementum with five (one proximal, two lateral, and two distal) pairs of setae. Thorax (Figure 3 A): Postpronotal lobe with 1 - 2 prominent and 1 - 3 short setae. Legs (Figure 3 A): Foreleg first tarsomere shorter than total length of four succeeding tarsomeres. Mid- and hind-leg first tarsomeres slightly shorter than or nearly as long as total length of four succeeding tarsomeres. Male terminalia (Figure 14 A-K): Epandrium caudodorsally with 2 - 3 and ca. 16 long setae per side on caudodorsal and ventral portions, respectively. Cercus with 31 - 33 setae. Surstylus with ca. nine prensisetae and ca. ten ventral spines. Hypandrium not pubescent. Paramere apically rounded in ventral view, with one sensillum. Aedeagus apically not darkened, subapically sometimes scarcely serrated along ventrolateral margin. Female terminalia (Figure 14 L-N): Tergite 8 with 4 - 5 and ca. four setae on ventral and caudodorsal portions, respectively. Oviscapt valve with 3 - 4 lateral and ca. 14 marginal ovisensilla. Spermathecal introvert ca. 1 / 6 height of capsule. Measurements (in mm): BL = 2.72 in holotype (range in 5 ♂ / 4 ♀ paratypes: 1.99 - 2.36 / 2.08 - 3.00), ThL = 1.22 (0.90 - 0.98 / 0.82 - 1.19), WL = 2.30 (1.74 - 1.82 / 1.62 - 2.33), WW = 1.26 (0.88 - 0.96 / 0.82 - 1.24). Indices. FW / HW = 0.58 (range in 5 ♂ and 4 ♀, or less if noted, paratypes: 0.55 - 0.58), ch / o = 0.09 (0.08 - 0.13), prorb = 0.68 (5 ♂, 3 ♀: 0.50 - 0.80), rcorb = 0.36 (0.27 - 0.45), vb = 0.40 (0.32 - 0.55), orbito = 0.70 (0.50 - 0.67), dcl = 0.67 (3 ♂, 4 ♀: 0.58 - 0.68), sctl = 0.96 (0.89 - 1.06), sterno = 0.47 (0.54 - 0.68), dcp = 0.49 (0.42 - 0.52), sctlp = 0.83 (0.69 - 1.00), C = 1.66 (1.48 - 1.87), 4 c = 1.46 (1.20 - 1.61), 4 v = 1.88 (1.62 - 2.00), 5 x = 1.36 (1.45 - 1.90), ac = 2.92 (2.90 - 3.44), M = 0.73 (0.68 - 0.83), C 3 F = 0.61 (0.51 - 0.62).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A90219302E04F582A327385D39003DA5.taxon	distribution	Distribution. China (Hainan).	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
A90219302E04F582A327385D39003DA5.taxon	etymology	Etymology. Pertaining to the type locality, Hainan (formerly known as " Qiongzhou ").	en	Katoh, Takehiro K., Zhang, Guang, Toda, Masanori J., Suwito, Awit, Gao, Jian-Jun (2018): A revision of the subgenus Dudaica Strand of the genus Drosophila Fallen, with descriptions of six new species (Diptera, Drosophilidae). ZooKeys 781: 19-50, DOI: http://dx.doi.org/10.3897/zookeys.781.27354, URL: http://dx.doi.org/10.3897/zookeys.781.27354
