identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038E862E4471FF92FF0D4A4BFED909A8.text	038E862E4471FF92FF0D4A4BFED909A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Armadillidium Brandt 1831	<div><p>Genus Armadillidium Brandt, 1831</p><p>Type species: Armadillidium vulgare (Latreille, 1804) by subsequent designation (= Armadillidium commutatum Brandt, 1831)</p></div>	https://treatment.plazi.org/id/038E862E4471FF92FF0D4A4BFED909A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cifuentes, Julio;Robla, Jairo;Garcia, Lluc	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4471FF9AFF0D4B0AFBF10F8F.text	038E862E4471FF9AFF0D4B0AFBF10F8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Armadillidium boalense Cifuentes & Robla & Garcia 2024	<div><p>Armadillidium boalense sp. nov.</p><p>Figs 2–6</p><p>Diagnosis. An Armadillidium species with conglobation ability of eusphaeric type. Head of duplocarinate type, with scutellar and post-scutellar ridges. Dorsum smooth. Cuticular structures characterized by circular cavities and sharp scale-setae. Pleotelson triangular with rounded tip. Male pereopod 7 strongly differentiated on ischium, merus and carpus. First male pleopod with endopod straight, three times as long as the exopod, not narrowed, slightly bent apically; pleopod 1–2 exopod notched on outer margin of tracheal field, both in males and females.</p><p>Etymology</p><p>The name of the species comes from Boal (municipality of Asturias, Spain) (Fig. 1A) which is the area where the specimens were collected.</p><p>Material examined</p><p>Holotype: Spain • ♂; Asturias, Boal, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.8051753&amp;materialsCitation.latitude=43.427784" title="Search Plazi for locations around (long -6.8051753/lat 43.427784)">As Cabanas</a> (Os Chaos); 43º25′40.02′′ N, 6º48′18.63′′ W; 470 m a.s.l.; 20 Mar. 2023; G. Sánchez Jardón leg.; hand collected under stones of a granitic wall embedded; MNCN 20.04 /20717.</p><p>Paratypes: Spain • 5♂; same collection data as for holotype; MNCN 20718-22 • 4♀; same collection data as for holotype; MNCN 20723-26 • 2♂; same collection data as for holotype; MBCN 26863 and MBCN 26864 • 1♂ dissected and mounted into preparations; same collection data as for holotype; CLLG 1187 • 2♀; same collection data as for holotype; MBCN 26865 and MBCN 26866 • 1♀ partially dissected; same collection data as for holotype; CLLG 1187 B . • 2♂; same collection data as for holotype; CJC708 and CJC709 • 1♀; same collection data as for holotype; CJC710 .</p><p>Description</p><p>Body. Measurements. Maximum length observed 6.5 x 2.8 mm (male). Colour. Dark purplish brown (Figs. 2A–B, 3A–C), with yellow muscle spots disposed as follows: pereon -tergites with two paramedian large spots, lateral smaller spots more or less fused; pereon-epimera with small spots except on sevenths; pleon-tergites with some small paramedian spots and large ones on epimera (Figs. 2A–B, 3C). Antenna, pereopods and pleopods slightly to quite pigmented. Eusphaeric-type conglobation (Fig. 2B). Pereon and pleon epimera strongly arched, directed downwards vertically (Fig. 3A). Dorsum smooth, without granulations (Fig. 3A–C); integument with circular cavities; scale-setae tricorn type with elongated and sharp tip (Fig. 4A). Cephalon. Duplocarinate-type, with scutellar ridge more marked than post-scutellar ridge (Figs 3B, 4B–C). Frontal shield with concave sides, slightly raised above vertex, separated from it, forming subscutellar pit (Figs 3B, 4B–C). Eyes with 18–20 ommatidia (Figs. 3A–B, 4B–C). Pereon. Posterior margin of pereon-tergites 1–2 forming clear angles with corresponding epimera; tergite 1 with acute posterior corner; tergites 3–4 with slightly sinuous posterior margin; tergites 5–6 with straight posterior margin; tergite 7 posterior margin slightly angled at sides (Figs. 2A, 3A). Pleon. Pleon-tergites with epimera directed backwards; epimera of pleonites 3–4 with acute posterior corners; epimera 5 with more rounded posterior corners (Figs. 3C, 4D) Pleotelson. Triangular, about 1.4 times as wide as long, with rounded tip (Figs. 3C, 4D).</p><p>First Antenna. Three-jointed, with medial article about half as long as other two; distal article bearing 8–10 aesthetascs and small triangular tip (Fig. 4E). Second antenna. Reaching posterior margin of first pereon-tergite when extended backwards. Flagellum 2-jointed with proximal article three times shorter than distal (Fig. 4F). Distal article with two groups of tiny aesthetascs.</p><p>Mouthparts. Mandibles with dichotomized molar penicils; left mandible (Fig. 4G) with 2 penicils on hairy lobe and 4 free penicils; right mandible (Fig. 4H) with 1+1 penicils. Maxillula: outer branch (Fig. 4I) with 4 strong teeth, 5 cleft smaller teeth and one simple tooth; inner branch (Fig. 4J) with 2 long penicils and short posterior corner. Maxilla (Fig. 5A) with outer lobe twice as wide as inner one; 3+1 setae between lobes. Maxilliped (Fig. 5B): first palp article with 2 setae; second palp article with three strong setae and several smaller ones; distal article with short apical setae; endite sub-quadrangular with parallel sides, strong seta on caudal face, two teeth on anterior edge and protruding triangular anterolateral corner.</p><p>Pereopods. All pereopods sexually dimorphic, with different number of ventral setae in males than in females; dactyli with inner claw shorter than outer one. Pereopod 1 (Fig. 5D) with longitudinal antennal-grooming brush. Male pereopod 7 (Fig. 5E–F) specially modified.</p><p>Pleopods. Pleopod exopodites 1 and 2 with polyspiracular lungs and notched outer margin of tracheal field, in both males and females (Figs. 3F–H; 6A–D). Exopodites 3–5 subtriangular in males (Fig. 6E–G), subquadrangular in females.</p><p>Uropods. Endopods inserted proximally on medial margin of protopods; cylindrical with outer margin straight, inner margin slightly widened medially. Exopods plate-like, trapezoidal, slightly wider than long, with rounded posterior corners (Fig 5C).</p><p>Male</p><p>Pereopods 1 (Figs 3D, 5D) to 6 with ventral brush of setae on carpus; ventral setae on merus, more abundant on 5th and 6th pereopods. Pereopod 7 (Figs. 3E, 5E–F), ischium with enlarged distal half, ventral margin strongly concave with many piliform setae extended to frontal and caudal faces, 4–5 strong setae on dorsal edge; merus with protruded basal heel and several strong setae on ventral margin; carpus with ventral brush of strong setae and keeled dorsal edge. Pleopod 1 (Figs. 3F–G, 6A): exopod triangular with short posterior lobe, distally rounded and notched outer margin of tracheal field; endopod three times as long as exopod, straight, not progressively narrowed and weakly curved distally, with some apical spines. Pleopod 2 (Figs. 3H, 6B): exopod triangular, with long and straight posterior lobe and notched outer margin of tracheal field; endopod thin and long, surpassing exopod length.</p><p>Remarks</p><p>Due to its duplocarinate-type cephalic structure and eusphaeric conglobation ability, A. boalense sp. nov. should be included in the ‘ pictum ’ group, according to the definition of Vandel (1962). In the Iberian Peninsula three species belonging to this group have been recorded: A. pictum Brandt, 1833, A. album Dollfus, 1887, and A. galiciense Schmölzer, 1955 . Furthermore, in other regions of Western Europe close to the northern Iberian Peninsula (e.g., Ireland or France) an extra species has also been found: Armadillidium pulchellum (Zenker) . A. boalense sp. nov. is externally very similar to A. pictum . Both species show a very similar coloration, which can lead to confusion. However, the new species differs from A. pictum mainly in having the endopodite of the male pleopod 1 almost completely straight, instead of strongly bent from the middle; in the exopod of the pleopod 2, unbent and more distally narrowed instead of with the distal half bent outwards; and in having a greater differentiation of the male pereopod 7, with the ischium with a much larger setose area and a much more pronounced proximal heel than in A. pictum . The cephalic structure of both species is very similar, but in A. boalense the scutellar (frontal) ridge is more marked than the post-scutellar ridge, while it is the other way around in A. pictum . The new species differs from A. album in the exopodites of the male pleopods 1–2; in having triangular scale-setae instead of erect bristles; in the morphology of the male pereopod 7, and in the shape of telson, which is trapezoidal in A. album, instead of triangular. The coloration also differs because A. album is a whitish species with few spots. From A. galiciense differs in the shape of the male pleopods 1–2 and in the strong differentiation of the male pereopod 7. In A. galiciense, the exopod of the male pleopod 7 lacks a posterior lobe and the endopod is even straighter. Moreover, the male pereopod 7 of A. galiciense is not modified. Finally, it differs from A. pulchellum mainly by the endopod of the male pleopod 1; by the exopod of the male pleopod 2, and by the modifications of the male pereopod 7, with very different shape and spinnulation in both species.</p><p>Collection site and ecology</p><p>The population of Armadillidium boalense sp. nov. was located in Os Chaos (Boal) in the Principality of Asturias (northern Spain) (Fig. 1A). The collecting area has an Atlantic macrobioclimate with mesotemperate climate (Rivas-Martínez et al. 2017). The area of Boal mean annual temperature is 11.2 ºC and 1250 mm mean annual precipitation (data available at https://es.climate-data.org). This species can be categorized as an epigeous species. Specimens were found in a wall of granite stones which also composed the geological substrate of the study area (Fig. 1B–C). Woodlice were usually found among the stones of the wall, although some specimens were found directly on the ground. The wall is closer to a mature forest composed mainly by pedunculate oak ( Quercus robur L.) (Fig. 1B), which has in its vicinity a replantation of southern blue gum ( Eucalyptus globulus Labill.). Under the wall stones there were an important organic matter accumulation mainly composed of peduncle oak leaf litter and mosses. The species is locally abundant and share habitat with other arthropods such as the woodlouse Oniscus asellus L. or centipedes of the genus Lithobius L. The area presents a slight anthropization due to the traffic of people and, sometimes, vehicles, and due to the proximity to several urban areas and villages.</p></div>	https://treatment.plazi.org/id/038E862E4471FF9AFF0D4B0AFBF10F8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cifuentes, Julio;Robla, Jairo;Garcia, Lluc	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4479FF99FF0D4D2FFB980BC3.text	038E862E4479FF99FF0D4D2FFB980BC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Armadillidium galiciense , Schmolzer 1955	<div><p>Armadillidium galiciense Schmölzer, 1955</p><p>Fig. 7</p><p>Armadillidium galiciense, Schmölzer, 1955, p. 189–192, fig. 9</p><p>Material examined</p><p>Spain • ♂ Holotype, male; high-definition photographs of the slide containing male pleopods 1–2 and pereopods 1 and 7 (see comments and taxonomic remarks).</p><p>Additional material</p><p>Spain • 1♀; Galicia, Pontevedra, Cerdedo; 31 Apr. 1952; Franz leg.; Schmölzer det.; MNCN 20.04/9634.</p><p>Taxonomical remarks</p><p>Vandel (1962) considered A. galiciense a junior synonym of A. pictum, a criterion followed by other authors (Cruz 1993; Cifuentes 2019). However, A. galiciense is a valid and distinct species. A. galiciense differs from A. pictum mainly in the endopod of the male pleopod 1 (Fig. 7A), which is straight, progressively tapered, and only slightly bent and pointed distally; and in the exopod, which is longitudinally elongated, without posterior lobe and twice as wide as long. It also differs in the male pleopod 2 (Fig. 7B) (not illustrated in the original description), by the exopod with a triangular posterior lobe, not bent nor narrowed as in A. pictum . According to the original description, the coloration of the body is also different. For all these reasons, it does not seem justified that A. galiciense is considered a junior subjective synonym of A. pictum and we can confirm the validity of this species. From A. boalense sp. nov. differs mainly in the exopods of the male pleopods 1–2 and in having the male pereopods 1 and 7 (Fig. 7C–D) without evident modifications, instead of strongly differentiated.</p><p>Distribution remarks</p><p>Schmölzer (1955) described A. galiciense as an endemic species of northwestern Spain, based on specimens from Galicia (Fig. 1A, 1D). The same author reported this species in other localities of the Cantabric region: Asturias and Cantabria (Schmölzer 1971) (Fig. 1A, 1D). This species has never been reported again since 1971.</p><p>Comments on holotype</p><p>According to the original description of A. galiciense, it is a clearly different species from A. pictum and from A. boalense sp. nov. The holotype slide (Fig. 7A–E) contained the male pleopods 1 and 2 (the left exopod 1 is missing), the genital papilla and the pereopods 1 and 7. It is hand-labelled (Fig. 7E) as following: « Armadillidium (A.) galiciensis (sic) Schmlzr (sic)» inked-written and « Holotypus » pencil-written. Additional information inked-written included «Sp 169/1F Leg. Franz» and «K. SCHMÖLZER INNSBRUCK». It is important to comment that in its original description, Schmölzer (1955) did not expressly designated an holotype but indicated «Cerdedo» (Pontevedra province, Galicia, Spain) as its type locality, corresponding to the locality «(Sp 179, loc. typ.), leg. F». Otherwise, the slide annotated as « holotypus » corresponded to the locality of Pyrnedo (sic), (i.e.,., Piornedo, province of Lugo, Galicia, Spain), recorded in the original description as «(loc. Sp 169), leg. F». So, a complete review of the type series of A. galiciense should be addressed in a subsequent work.</p></div>	https://treatment.plazi.org/id/038E862E4479FF99FF0D4D2FFB980BC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cifuentes, Julio;Robla, Jairo;Garcia, Lluc	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447AFF99FF0D4903FEB608AB.text	038E862E447AFF99FF0D4903FEB608AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Armadillidium album Dollfus 1887	<div><p>Armadillidium album Dollfus, 1887</p><p>Material examined</p><p>Spain • 5 specs; Asturias, Gozón, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.9140253&amp;materialsCitation.latitude=43.609764" title="Search Plazi for locations around (long -5.9140253/lat 43.609764)">Xagó beach</a>; 43º36′35.15′′ N, 5º54′50.49′′ W; 1 m a.s.l.; 10 Oct. 2021; J. Robla leg.; hand collected under logs and trunks in the sand, 10–30 m far from the coastline; CLLG. Population first photographed by César Fernández and Rafael González in Mar. 2013 .</p><p>Remarks and ecology</p><p>Halophilous species well distributed along the Mediterranean and Atlantic coasts of Europe and northern Africa. A. album has barely been recorded in the Iberian Peninsula and was only known from Almería (Cruz 1993), A Coruña (Galicia) and Valencia (Garcia 2005) and Mallorca (Balearic Islands) (Garcia et al. 2003) in Spain and several districts of Portugal (Dias et al. 2005; Gonçalves et al. 2009). Here we provided the first records of A. album for Asturias and expanded the distribution in the north of the Iberian Peninsula (Fig. 1A, 1D). Many specimens were located under the trunks and logs at the beginning of the dunar system and from 10 to 30 meters far from the coastline and along the entire beach. Specimens were always found attached to the inner side of the trunks and logs, but also directly on the sand or close to the only plant species present in this area, the marram grass Ammophila arenaria (L.) Link.</p></div>	https://treatment.plazi.org/id/038E862E447AFF99FF0D4903FEB608AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cifuentes, Julio;Robla, Jairo;Garcia, Lluc	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447BFF98FF0D4CA6FF280C74.text	038E862E447BFF98FF0D4CA6FF280C74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Armadillidium assimile Budde-Lund 1885	<div><p>Armadillidium assimile Budde-Lund, 1885</p><p>Material examined</p><p>Spain • 1♂; Asturias, Ribadeva, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.9219193&amp;materialsCitation.latitude=43.18702" title="Search Plazi for locations around (long -5.9219193/lat 43.18702)">Pimiango</a>; 6 Aug. 1984; J. Cifuentes leg; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.9219193&amp;materialsCitation.latitude=43.18702" title="Search Plazi for locations around (long -5.9219193/lat 43.18702)">in an Erica L. shrubland close to the coast</a>; JC202. • 6 specs; Asturias, Quirós, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.9219193&amp;materialsCitation.latitude=43.18702" title="Search Plazi for locations around (long -5.9219193/lat 43.18702)">Altu de la Cobertoria (Gamoniteiru)</a>; 43º11′13.26′′ N, 5º55′18.91′′ W; 1764 m a.s.l.; 17 Sep. 2021; J. Robla leg.; hand collected under embedded big stones; CLLG .</p><p>Remarks and ecology</p><p>This species is present in western Europe. It has been occasionally recorded in northern Spain and several Mediterranean coastal and inland provinces. This species had already been recorded in Asturias (Schmölzer 1955, 1971; Cruz 1990; Vázquez-Felechosa &amp; Anadón, 2001) (Fig. 1A). Here we provided two new localities, one representing one of the highest records of this species in the Iberian Peninsula (ca. 1800 m). Although it is said that this species tends to occur in humid and flooded areas like marshes, river areas or meadows (Cruz 1993; Vandel 1962) or even anthropic areas like gardens (Vandel 1962), specimens from Gamoniteiru were found in alpine meadows without shrub or tree vegetation under large stones strongly attached to the substrate and far from water sources.</p></div>	https://treatment.plazi.org/id/038E862E447BFF98FF0D4CA6FF280C74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cifuentes, Julio;Robla, Jairo;Garcia, Lluc	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447BFF98FF0D4F46FF320817.text	038E862E447BFF98FF0D4F46FF320817.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Armadillidium nasatum Budde-Lund 1885	<div><p>Armadillidium nasatum Budde-Lund, 1885</p><p>Material examined</p><p>Spain • 3♂ and 3♀; Asturias, Oviedo; Barros leg.; MNCN 20.04/11854 . • 6 specs; Asturias, Ribera de Arriba, Palomar; 43º18′29.66′′ N, 5º54′39.93′′ W; 130 m a.s.l.; 28 Aug. 2021; J. Robla leg.; hand collected under stones and roof tiles; CLLG. • 17 specs; Asturias, Lena, Pola de Lena (Mofusu forest); 43º8′59.05′′ N, 5º51′19.84′′ W; 325 m a.s.l.; 1 Oct. 2021; J. Robla leg.; hand collected under logs, trunks, rocks, and leaf litter in a mixed forest of pedunculate oak Quercus robur with common hazel Corylus avellana L., ash Fraxinus excelsior L. and common alder Alnus glutinosa (L.) Gaertn; CLLG. • 1 spec; Asturias, Grado, Somines; 43º22′0.29′′ N, 6º0′53.83′′ W; 204 m a.s.l.; 27 Sep. 2021; J. Robla leg.; hand collected under stones near a private house; CLLG . • 1 spec; Asturias, Oviedo, Pista Finlandesa; 43º23′8.73′′ N, 5º50′12.22′′ W; 303 m a.s.l.; 21 Aug. 2021; J. Robla leg.; hand collected under blue gum Eucalyptus globulus leaf litter; CLLG . • 3 spec; Asturias, Gozón, Verdicio beach; 43º22′0.29′′ N, 6º0′53.83′′ W; 7 m a.s.l.; 27 Aug. 2020; J. Robla leg.; hand collected under stones close to the dunar system; CLLG.</p><p>Remarks and ecology</p><p>Armadillidium nasatum is a species distributed from western to eastern areas of the Eurasian region. In Spain, this species is limited to northern Iberian Peninsula, from Galicia to Gerona (a summary of most of these records is included in Dollfus 1892; Fidalgo &amp; Herrera 1980; Cruz 1993, Schmölzer 1955, 1971; Vivar et al. 1994; Vázquez-Felechosa &amp; Anadón 2001, Cifuentes 2019) (Fig. 1A). It had already been recorded in Asturias (Schmölzer 1955, 1971; Vandel 1962; Vázquez Felechosa &amp; Anadón 2001). We provide here several new localities. This species was usually found in several habitats: different native forest masses, eucalyptus plantations, meadows, rural areas, gardens, rocky walls, and coastal habitats. A. nasatum was always located under logs, stones or other structures that create humid and shady microhabitats and our ecological observations agreed with those provided by Vandel (1962).</p></div>	https://treatment.plazi.org/id/038E862E447BFF98FF0D4F46FF320817	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cifuentes, Julio;Robla, Jairo;Garcia, Lluc	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447BFF9FFF0D4BA6FE5B0D43.text	038E862E447BFF9FFF0D4BA6FE5B0D43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Armadillidium pictum Brandt 1833	<div><p>Armadillidium pictum Brandt, 1833</p><p>Remarks and ecology</p><p>Species well distributed across Europe in different ecotypes. Rare and scarcely recorded species in the Iberian Peninsula, limited to northern areas, especially in the Pyrenean region: Huesca (Schmölzer 1971, as A. garumnicum); Navarra (Schmölzer 1971, as A. garumnicum; Fidalgo &amp; Herrera 1980; Vivar et al. 1984; Cruz 1993, as A. pictum); Andorra and Lleida (Cruz 1993, as A. pictum). The only two records in Asturias (Cruz 1991a; Vázquez-Felechosa &amp; Anadón 2001) need to be confirmed since they are based on female specimens that could have been confused with A. boalense sp. nov., which presents a similar habitus. The unique record in Galicia, also a female specimen (Cifuentes 2019), comes from the type locality of A. galiciense and could correspond to this species, now revalidated. During our samplings in Asturias, we have not found Armadillidium pictum . We hypothesised that could be more restricted to the Pyrenean region, being replaced by A. boalense and A. galiciense in northern Iberian Peninsula. Little is known about its ecology, although it is considered a silvicolous species well adapted to other natural and seminatural habitats.</p></div>	https://treatment.plazi.org/id/038E862E447BFF9FFF0D4BA6FE5B0D43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cifuentes, Julio;Robla, Jairo;Garcia, Lluc	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447CFF9FFF0D4E5BFBCE0887.text	038E862E447CFF9FFF0D4E5BFBCE0887.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Armadillidium vulgare (Latreille 1804)	<div><p>Armadillidium vulgare (Latreille, 1804)</p><p>Material examined</p><p>Spain • 84♂ and 127♀; Asturias, Oviedo; Barros leg.; MNCN 20.04/7892 . • 1♂ and 6♀; Asturias, Cangas del Narcea, Muniellos; 20 Aug. 1966; E. Ortiz de Vega leg.; MNCN 20.04/11551 . • 3♂ and 18♀; Asturias, Cangas del Narcea, Muniellos; 28 Aug. 1966; E. Ortiz de Vega leg.; MNCN 20.04/11553. • 2♂ and 5♀; Asturias, Llanes, Purón; 9 Aug. 1979; E. Ortiz de Vega leg.; MNCN 20.04/11537 . • 2♀; Asturias, Llanes, Purón; 19 Aug. 1979; E. Ortiz de Vega leg.; MNCN 20.04/11549. • 2♂, 5♀ and 2 inmature; Asturias, Llanes, Purón; 22 May. 1982; E. Ortiz de Vega leg.; MNCN 20.04/7389 . • 1♂ and 1♀; Asturias, Ribadeva, Pimiango; 6 Aug. 1984; J. Cifuentes leg.; JC145. • 6 specs; Asturias, Ribera de Arriba, Palomar; 43º18′29.66′′ N, 5º54′39.93′′ W; 130 m a.s.l.; 28 Aug. 2021; J. Robla leg.; hand collected under stones and roof tiles; CLLG . • 14 spec; Asturias, Grado, Somines; 43º22′0.29′′ N, 6º0′53.83′′ W; 204 m a.s.l.; 27 Sep. 2021; J. Robla leg.; hand collected under stones near a private house and in a Eucalyptus plantation; CLLG . • 4 spec; Asturias, Oviedo, Pista Finlandesa; 43º23′8.73′′ N, 5º50′12.22′′ W; 303 m a.s.l.; 21 Aug. 2021; J. Robla leg.; hand collected under blue gum Eucalyptus globulus leaf litter and in a mixed forest mainly compose of Quercus robur; CLLG. • 3 spec; Asturias, Gozón, Verdicio beach; 43º22′0.29′′ N, 6º0′53.83′′ W; 7 m a.s.l.; 27 Aug. 2020; J. Robla leg.; hand collected under stones close to the dunar system; CLLG . • 20 spec; Asturias, Mieres, Yana’l Monte (Turón); 43º12′24.36′′ N, 5º45′11.14′′ W; 303 m a.s.l.; 28 Aug. 2021; J. Robla leg.; hand collected under logs, trunks and leaf litter in a mixed forest composed by chestnut Castanea sativa Mill. and pedunculate oak; CLLG . • 5 spec; Asturias, Pola de Siero, El Rebollar Street; 43º23′39.90′′ N, 5º39′17.14′′ W; 239 m a.s.l.; 3 Sep. 2020; J. Robla leg.; hand collected in a meadow at night; • 5 spec; Asturias, Vegadeo, Piantón; 43º27′39.32′′ N, 7º1′43.95′′ W; 55 m a.s.l.; 13 Oct. 2021; D. Cabanillas leg.; hand collected in a private garden; CLLG . • 5 spec; Asturias, Selorio, Ría de Villaviciosa; 43º31′10.89′′ N, 5º23′2.72′′ W; 7 m a.s.l.; 24 Sep. 2021; J. Robla leg.; hand collected under a log in a marsh; CLLG . • 2 spec; Asturias, Coaña, Playa de Foxos; 43º33’19.7’’ N, 6º43’36.3’’ W; 0 m a.s.l.; 16 May. 2024; G. Sánchez-Jardón leg; hand collected under logs and stones in the sand of a beach; CJR. • 1 spec; Asturias, El Franco, Valdepares; 43º33’42.7’’ N, 6º52’3.6’’ W; 43 m a.s.l.; 19 May. 2024; G. Sánchez-Jardón leg; hand collected in a catch basin in the private garden of a house.</p><p>Remarks and ecology</p><p>A well-distributed species in Europe, the westernmost parts of Asia and some areas of north Africa. Furthermore, the most abundant and well distributed Armadillidium species throughout the Ibero-Balearic area. Species previously recorded in Asturias (Schmölzer 1955; 1971; Vázquez-Felechosa &amp; Anadón 2001; Vázquez-Felechosa et al. 2004) (Fig. 1A). We provide new localities that point out its broad ecological tolerance: forests (native and artificial plantations), meadows, marshes, riparian zones, coastal areas, and urban and rural areas. During the fieldwork, it has been observed that the dark coloration is the most commonly registered, but reddish or slightly pigmented specimens were sometimes detected. The most common species in urban areas. A. vulgare usually form large aggregations of individuals and are active during daytime, even in areas without great humidity.</p></div>	https://treatment.plazi.org/id/038E862E447CFF9FFF0D4E5BFBCE0887	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cifuentes, Julio;Robla, Jairo;Garcia, Lluc	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447DFF9DFF0D4CA6FAAA0F8F.text	038E862E447DFF9DFF0D4CA6FAAA0F8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Armadillidium Brandt 1831	<div><p>Key to the Iberian-Balearic species of the genus Armadillidium</p><p>Identification key of the Armadillidium species recorded in the Iberian Peninsula and the Balearic Islands. Mainly based on Schmölzer (1954; 1965); Vandel (1960; 1962); Cruz (1990; 1993); Cruz &amp; Dalens (1990); Garcia (2003; 2020); Garcia &amp; Cabanillas (2021); Cifuentes &amp; Escarabajal (2022a, 2022b) and the present study. A. scabrum Dollfus, 1892, (only known from Seville) is excluded due to insufficient information in its original description (Dollfus, 1892). The distribution is indicated only for the endemic species.</p><p>1. Head of duplocarinate type, with scutellar and post-scutellar ridges.............................................. 2.</p><p>- Head of ‘ Armadillidium -type’ without scutellar ridge (linea frontalis)........................................... 13.</p><p>2. Conglobation ability of eusphaeric type (living animals can roll into a perfect ball). Integument smooth or weakly granulated................................................................................................... 3.</p><p>- Conglobation ability of pseudo/mesosphaeric type (living animals can roll into an ovoid or imperfect ball, respectively). Integument smooth, granulated or with spiniform tubercles.................................................... 6.</p><p>3. Telson triangular. Living animals with a vivid coloration, usually with yellowish muscle spots on a dark background. Integument smooth or slightly granulated............................................................................ 4.</p><p>- Telson trapezoidal. Very little or no pigmented body. Integument smooth, covered with bristle-like scale-setae. Male pereopod 7 with ventrally concave ischium and a protruded distal knob on the basis. Strictly littoral species... A. album Dollfus, 1887 .</p><p>4. Male pleopod 1 endopod with distal third bent 90º outward. Exopod with notched tracheal field..... A. pictum Brandt, 1833 .</p><p>- Male pleopod 1 endopod long and straight, only slightly bent apically............................................ 5.</p><p>5. Male pleopod 1 exopod without posterior lobe, twice as wide as long and with notched tracheal field. Galicia and Cantabric region ...................................................................... A. galiciense Schmölzer, 1955 .</p><p>- Male pleopod 1 exopod with triangular posterior lobe, almost as long as wide, and with notched tracheal field. Asturias ......................................................................................... A. boalense sp. nov.</p><p>6. Integument smooth.................................................................................... 7.</p><p>- Integument with large or spiniform tubercles................................................................ 9.</p><p>7. Telson trapezoidal, with a wide posterior margin and straight lateral edges. Menorca (Balearic Islands).................................................................................................. A. strinatii Vandel, 1961 .</p><p>- Telson triangular...................................................................................... 8.</p><p>8. Lateral edge of first the pereon-tergite, angulate. Male pereopod 7 without distinct sexual modification.Mallorca and surrounding islets (Balearic Islands)................................................................ A. cruzi Garcia, 2003 .</p><p>- Lateral edge of the first pereon-tergite regularly curved. Male pereopods 6-7 with a marked distal knob on ischium; male pereopod 7 with ventrally concave ischium. Levantine Iberian region......................... A. espanyoli Cruz, 1992 .</p><p>9. Telson trapezoidal.................................................................................... 10.</p><p>- Telson triangular..................................................................................... 12.</p><p>10. Telson with narrow posterior edge, rounded posterior angles and concave lateral edges; back covered with large conical tubercles; scale-setae with forked setae. Male pleopod 1 exopodite with short posterior lobe and notched tracheal field. Endemic from Murcia region ............................................... A. ibericum Cifuentes &amp; Escarabajal Bernabé, 2022 .</p><p>- Telson with broad posterior edge, obtuse posterior angles and straight lateral edges; scale-setae tricorn-type, without forked setae.............................................................................................. 11.</p><p>11. Cephalon with angled postscutellar ridge. Pleonites with a transverse row of strong granulations. Male pleopod 1 exopodite twice as wide as long, with barely prominent posterior lobe. Menorca (Balearic Islands)..... A. serrai Cruz &amp; Dalens, 1990 .</p><p>- Cephalon with regularly arched post-scutellar ridge. Pleonites without apparent granulations. Exopodite of the male pleopod 1 with distinct posterior lobe. Pre-Pyrenean Iberian region................................... A. nahumi Garcia, 2020 .</p><p>12. Cephalon with strongly angulate post-scutellar ridge; scutellar ridge, complete. Back covered with hypertrophic spiniform tubercles. Telson broad and short, with broadly rounded apex. Mallorca (Balearic Islands).......... A. pretusi Cruz, 1992 .</p><p>- Cephalon with regularly arched post-scutellar ridge; scutellar ridge incomplete. Body covered with spiny tubercles. Telson narrow, with straight lateral sides................................................ A. serratum Budde-Lund, 1885 .</p><p>13. Telson trapezoidal.................................................................................... 14.</p><p>- Telson triangular..................................................................................... 15.</p><p>14. Conglobation ability of mesosphaeric type. Male pleopod 1 exopod with sinuous inner margin; endopod straight. Male pereopod 7 with a rounded distal bulge on ischium.............................................. A. sordidum Dollfus, 1887 .</p><p>- Conglobation ability of eusphaeric type. Male pleopod 1 exopod with convex inner margin; endopod distally curved. Male pereopod 7 without distal bulge on ischium........................................... A. vulgare (Latreille, 1804) .</p><p>15. Conglobation ability of mesosphaeric type. Back granulated. The posterior edge of the first pereon-tergite forms angles at the limit with the epimera............................................................ A. granulatum Brand, 1833.</p><p>- Back smooth or with very weak granulations.............................................................. 16.</p><p>16. Conglobation ability of pseudosphaeric type. Frontal shield almost twice as long as wide, very prominent above the vertex. The posterior margin of the first pereon-tergite forms angles at the limit with the epimera........ A. nasatum Budde-Lund, 1885 .</p><p>- Conglobation ability of mesosphaeric type. Frontal shield slightly prominent and directed backwards above the vertex. Posterior margin of first pereon-tergite regularly sinuous............................................................. 17.</p><p>17. Male pleopod 1 exopod with very short posterior lobe................................ A. assimile Budde-Lund, 1885 .</p><p>- Male pleopod 1 exopod with well-developed posterior lobe.............................. A. arcangelii Strouhal, 1929 .</p><p>* Note that Armadillidium mateui Vandel is not currently included in the genus Armadillidium . This species has been transferred to Alloschizidium Verhoeff (see Cifuentes 2022). Thus, it was not included in the identification key.</p></div>	https://treatment.plazi.org/id/038E862E447DFF9DFF0D4CA6FAAA0F8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cifuentes, Julio;Robla, Jairo;Garcia, Lluc	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
