taxonID	type	description	language	source
038E862E4471FF92FF0D4A4BFED909A8.taxon	type_taxon	Type species: Armadillidium vulgare (Latreille, 1804) by subsequent designation (= Armadillidium commutatum Brandt, 1831)	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4471FF9AFF0D4B0AFBF10F8F.taxon	description	Figs 2 – 6	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4471FF9AFF0D4B0AFBF10F8F.taxon	diagnosis	Diagnosis. An Armadillidium species with conglobation ability of eusphaeric type. Head of duplocarinate type, with scutellar and post-scutellar ridges. Dorsum smooth. Cuticular structures characterized by circular cavities and sharp scale-setae. Pleotelson triangular with rounded tip. Male pereopod 7 strongly differentiated on ischium, merus and carpus. First male pleopod with endopod straight, three times as long as the exopod, not narrowed, slightly bent apically; pleopod 1 – 2 exopod notched on outer margin of tracheal field, both in males and females.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4471FF9AFF0D4B0AFBF10F8F.taxon	etymology	Etymology The name of the species comes from Boal (municipality of Asturias, Spain) (Fig. 1 A) which is the area where the specimens were collected.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4471FF9AFF0D4B0AFBF10F8F.taxon	materials_examined	Material examined Holotype: Spain • ♂; Asturias, Boal, As Cabanas (Os Chaos); 43 º 25 ′ 40.02 ′′ N, 6 º 48 ′ 18.63 ′′ W; 470 m a. s. l.; 20 Mar. 2023; G. Sánchez Jardón leg.; hand collected under stones of a granitic wall embedded; MNCN 20.04 / 20717. Paratypes: Spain • 5 ♂; same collection data as for holotype; MNCN 20718 - 22 • 4 ♀; same collection data as for holotype; MNCN 20723 - 26 • 2 ♂; same collection data as for holotype; MBCN 26863 and MBCN 26864 • 1 ♂ dissected and mounted into preparations; same collection data as for holotype; CLLG 1187 • 2 ♀; same collection data as for holotype; MBCN 26865 and MBCN 26866 • 1 ♀ partially dissected; same collection data as for holotype; CLLG 1187 B. • 2 ♂; same collection data as for holotype; CJC 708 and CJC 709 • 1 ♀; same collection data as for holotype; CJC 710.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4471FF9AFF0D4B0AFBF10F8F.taxon	description	Description Body. Measurements. Maximum length observed 6.5 x 2.8 mm (male). Colour. Dark purplish brown (Figs. 2 A – B, 3 A – C), with yellow muscle spots disposed as follows: pereon - tergites with two paramedian large spots, lateral smaller spots more or less fused; pereon-epimera with small spots except on sevenths; pleon-tergites with some small paramedian spots and large ones on epimera (Figs. 2 A – B, 3 C). Antenna, pereopods and pleopods slightly to quite pigmented. Eusphaeric-type conglobation (Fig. 2 B). Pereon and pleon epimera strongly arched, directed downwards vertically (Fig. 3 A). Dorsum smooth, without granulations (Fig. 3 A – C); integument with circular cavities; scale-setae tricorn type with elongated and sharp tip (Fig. 4 A). Cephalon. Duplocarinate-type, with scutellar ridge more marked than post-scutellar ridge (Figs 3 B, 4 B – C). Frontal shield with concave sides, slightly raised above vertex, separated from it, forming subscutellar pit (Figs 3 B, 4 B – C). Eyes with 18 – 20 ommatidia (Figs. 3 A – B, 4 B – C). Pereon. Posterior margin of pereon-tergites 1 – 2 forming clear angles with corresponding epimera; tergite 1 with acute posterior corner; tergites 3 – 4 with slightly sinuous posterior margin; tergites 5 – 6 with straight posterior margin; tergite 7 posterior margin slightly angled at sides (Figs. 2 A, 3 A). Pleon. Pleon-tergites with epimera directed backwards; epimera of pleonites 3 – 4 with acute posterior corners; epimera 5 with more rounded posterior corners (Figs. 3 C, 4 D) Pleotelson. Triangular, about 1.4 times as wide as long, with rounded tip (Figs. 3 C, 4 D). First Antenna. Three-jointed, with medial article about half as long as other two; distal article bearing 8 – 10 aesthetascs and small triangular tip (Fig. 4 E). Second antenna. Reaching posterior margin of first pereon-tergite when extended backwards. Flagellum 2 - jointed with proximal article three times shorter than distal (Fig. 4 F). Distal article with two groups of tiny aesthetascs. Mouthparts. Mandibles with dichotomized molar penicils; left mandible (Fig. 4 G) with 2 penicils on hairy lobe and 4 free penicils; right mandible (Fig. 4 H) with 1 + 1 penicils. Maxillula: outer branch (Fig. 4 I) with 4 strong teeth, 5 cleft smaller teeth and one simple tooth; inner branch (Fig. 4 J) with 2 long penicils and short posterior corner. Maxilla (Fig. 5 A) with outer lobe twice as wide as inner one; 3 + 1 setae between lobes. Maxilliped (Fig. 5 B): first palp article with 2 setae; second palp article with three strong setae and several smaller ones; distal article with short apical setae; endite sub-quadrangular with parallel sides, strong seta on caudal face, two teeth on anterior edge and protruding triangular anterolateral corner. Pereopods. All pereopods sexually dimorphic, with different number of ventral setae in males than in females; dactyli with inner claw shorter than outer one. Pereopod 1 (Fig. 5 D) with longitudinal antennal-grooming brush. Male pereopod 7 (Fig. 5 E – F) specially modified. Pleopods. Pleopod exopodites 1 and 2 with polyspiracular lungs and notched outer margin of tracheal field, in both males and females (Figs. 3 F – H; 6 A – D). Exopodites 3 – 5 subtriangular in males (Fig. 6 E – G), subquadrangular in females. Uropods. Endopods inserted proximally on medial margin of protopods; cylindrical with outer margin straight, inner margin slightly widened medially. Exopods plate-like, trapezoidal, slightly wider than long, with rounded posterior corners (Fig 5 C). Male Pereopods 1 (Figs 3 D, 5 D) to 6 with ventral brush of setae on carpus; ventral setae on merus, more abundant on 5 th and 6 th pereopods. Pereopod 7 (Figs. 3 E, 5 E – F), ischium with enlarged distal half, ventral margin strongly concave with many piliform setae extended to frontal and caudal faces, 4 – 5 strong setae on dorsal edge; merus with protruded basal heel and several strong setae on ventral margin; carpus with ventral brush of strong setae and keeled dorsal edge. Pleopod 1 (Figs. 3 F – G, 6 A): exopod triangular with short posterior lobe, distally rounded and notched outer margin of tracheal field; endopod three times as long as exopod, straight, not progressively narrowed and weakly curved distally, with some apical spines. Pleopod 2 (Figs. 3 H, 6 B): exopod triangular, with long and straight posterior lobe and notched outer margin of tracheal field; endopod thin and long, surpassing exopod length.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4471FF9AFF0D4B0AFBF10F8F.taxon	discussion	Remarks Due to its duplocarinate-type cephalic structure and eusphaeric conglobation ability, A. boalense sp. nov. should be included in the ‘ pictum ’ group, according to the definition of Vandel (1962). In the Iberian Peninsula three species belonging to this group have been recorded: A. pictum Brandt, 1833, A. album Dollfus, 1887, and A. galiciense Schmölzer, 1955. Furthermore, in other regions of Western Europe close to the northern Iberian Peninsula (e. g., Ireland or France) an extra species has also been found: Armadillidium pulchellum (Zenker). A. boalense sp. nov. is externally very similar to A. pictum. Both species show a very similar coloration, which can lead to confusion. However, the new species differs from A. pictum mainly in having the endopodite of the male pleopod 1 almost completely straight, instead of strongly bent from the middle; in the exopod of the pleopod 2, unbent and more distally narrowed instead of with the distal half bent outwards; and in having a greater differentiation of the male pereopod 7, with the ischium with a much larger setose area and a much more pronounced proximal heel than in A. pictum. The cephalic structure of both species is very similar, but in A. boalense the scutellar (frontal) ridge is more marked than the post-scutellar ridge, while it is the other way around in A. pictum. The new species differs from A. album in the exopodites of the male pleopods 1 – 2; in having triangular scale-setae instead of erect bristles; in the morphology of the male pereopod 7, and in the shape of telson, which is trapezoidal in A. album, instead of triangular. The coloration also differs because A. album is a whitish species with few spots. From A. galiciense differs in the shape of the male pleopods 1 – 2 and in the strong differentiation of the male pereopod 7. In A. galiciense, the exopod of the male pleopod 7 lacks a posterior lobe and the endopod is even straighter. Moreover, the male pereopod 7 of A. galiciense is not modified. Finally, it differs from A. pulchellum mainly by the endopod of the male pleopod 1; by the exopod of the male pleopod 2, and by the modifications of the male pereopod 7, with very different shape and spinnulation in both species.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4471FF9AFF0D4B0AFBF10F8F.taxon	biology_ecology	Collection site and ecology The population of Armadillidium boalense sp. nov. was located in Os Chaos (Boal) in the Principality of Asturias (northern Spain) (Fig. 1 A). The collecting area has an Atlantic macrobioclimate with mesotemperate climate (Rivas-Martínez et al. 2017). The area of Boal mean annual temperature is 11.2 ºC and 1250 mm mean annual precipitation (data available at https: // es. climate-data. org). This species can be categorized as an epigeous species. Specimens were found in a wall of granite stones which also composed the geological substrate of the study area (Fig. 1 B – C). Woodlice were usually found among the stones of the wall, although some specimens were found directly on the ground. The wall is closer to a mature forest composed mainly by pedunculate oak (Quercus robur L.) (Fig. 1 B), which has in its vicinity a replantation of southern blue gum (Eucalyptus globulus Labill.). Under the wall stones there were an important organic matter accumulation mainly composed of peduncle oak leaf litter and mosses. The species is locally abundant and share habitat with other arthropods such as the woodlouse Oniscus asellus L. or centipedes of the genus Lithobius L. The area presents a slight anthropization due to the traffic of people and, sometimes, vehicles, and due to the proximity to several urban areas and villages.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4479FF99FF0D4D2FFB980BC3.taxon	description	Fig. 7	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4479FF99FF0D4D2FFB980BC3.taxon	materials_examined	Material examined Spain • ♂ Holotype, male; high-definition photographs of the slide containing male pleopods 1 – 2 and pereopods 1 and 7 (see comments and taxonomic remarks). Additional material Spain • 1 ♀; Galicia, Pontevedra, Cerdedo; 31 Apr. 1952; Franz leg.; Schmölzer det.; MNCN 20.04 / 9634.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4479FF99FF0D4D2FFB980BC3.taxon	diagnosis	Taxonomical remarks Vandel (1962) considered A. galiciense a junior synonym of A. pictum, a criterion followed by other authors (Cruz 1993; Cifuentes 2019). However, A. galiciense is a valid and distinct species. A. galiciense differs from A. pictum mainly in the endopod of the male pleopod 1 (Fig. 7 A), which is straight, progressively tapered, and only slightly bent and pointed distally; and in the exopod, which is longitudinally elongated, without posterior lobe and twice as wide as long. It also differs in the male pleopod 2 (Fig. 7 B) (not illustrated in the original description), by the exopod with a triangular posterior lobe, not bent nor narrowed as in A. pictum. According to the original description, the coloration of the body is also different. For all these reasons, it does not seem justified that A. galiciense is considered a junior subjective synonym of A. pictum and we can confirm the validity of this species. From A. boalense sp. nov. differs mainly in the exopods of the male pleopods 1 – 2 and in having the male pereopods 1 and 7 (Fig. 7 C – D) without evident modifications, instead of strongly differentiated.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4479FF99FF0D4D2FFB980BC3.taxon	distribution	Distribution remarks Schmölzer (1955) described A. galiciense as an endemic species of northwestern Spain, based on specimens from Galicia (Fig. 1 A, 1 D). The same author reported this species in other localities of the Cantabric region: Asturias and Cantabria (Schmölzer 1971) (Fig. 1 A, 1 D). This species has never been reported again since 1971.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E4479FF99FF0D4D2FFB980BC3.taxon	discussion	Comments on holotype According to the original description of A. galiciense, it is a clearly different species from A. pictum and from A. boalense sp. nov. The holotype slide (Fig. 7 A – E) contained the male pleopods 1 and 2 (the left exopod 1 is missing), the genital papilla and the pereopods 1 and 7. It is hand-labelled (Fig. 7 E) as following: « Armadillidium (A.) galiciensis (sic) Schmlzr (sic) » inked-written and « Holotypus » pencil-written. Additional information inked-written included « Sp 169 / 1 F Leg. Franz » and « K. SCHMÖLZER INNSBRUCK ». It is important to comment that in its original description, Schmölzer (1955) did not expressly designated an holotype but indicated « Cerdedo » (Pontevedra province, Galicia, Spain) as its type locality, corresponding to the locality « (Sp 179, loc. typ.), leg. F ». Otherwise, the slide annotated as « holotypus » corresponded to the locality of Pyrnedo (sic), (i. e.,., Piornedo, province of Lugo, Galicia, Spain), recorded in the original description as « (loc. Sp 169), leg. F ». So, a complete review of the type series of A. galiciense should be addressed in a subsequent work.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447AFF99FF0D4903FEB608AB.taxon	materials_examined	Material examined Spain • 5 specs; Asturias, Gozón, Xagó beach; 43 º 36 ′ 35.15 ′′ N, 5 º 54 ′ 50.49 ′′ W; 1 m a. s. l.; 10 Oct. 2021; J. Robla leg.; hand collected under logs and trunks in the sand, 10 – 30 m far from the coastline; CLLG. Population first photographed by César Fernández and Rafael González in Mar. 2013.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447AFF99FF0D4903FEB608AB.taxon	discussion	Remarks and ecology Halophilous species well distributed along the Mediterranean and Atlantic coasts of Europe and northern Africa. A. album has barely been recorded in the Iberian Peninsula and was only known from Almería (Cruz 1993), A Coruña (Galicia) and Valencia (Garcia 2005) and Mallorca (Balearic Islands) (Garcia et al. 2003) in Spain and several districts of Portugal (Dias et al. 2005; Gonçalves et al. 2009). Here we provided the first records of A. album for Asturias and expanded the distribution in the north of the Iberian Peninsula (Fig. 1 A, 1 D). Many specimens were located under the trunks and logs at the beginning of the dunar system and from 10 to 30 meters far from the coastline and along the entire beach. Specimens were always found attached to the inner side of the trunks and logs, but also directly on the sand or close to the only plant species present in this area, the marram grass Ammophila arenaria (L.) Link.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447BFF98FF0D4CA6FF280C74.taxon	materials_examined	Material examined Spain • 1 ♂; Asturias, Ribadeva, Pimiango; 6 Aug. 1984; J. Cifuentes leg; in an Erica L. shrubland close to the coast; JC 202. • 6 specs; Asturias, Quirós, Altu de la Cobertoria (Gamoniteiru); 43 º 11 ′ 13.26 ′′ N, 5 º 55 ′ 18.91 ′′ W; 1764 m a. s. l.; 17 Sep. 2021; J. Robla leg.; hand collected under embedded big stones; CLLG.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447BFF98FF0D4CA6FF280C74.taxon	discussion	Remarks and ecology This species is present in western Europe. It has been occasionally recorded in northern Spain and several Mediterranean coastal and inland provinces. This species had already been recorded in Asturias (Schmölzer 1955, 1971; Cruz 1990; Vázquez-Felechosa & Anadón, 2001) (Fig. 1 A). Here we provided two new localities, one representing one of the highest records of this species in the Iberian Peninsula (ca. 1800 m). Although it is said that this species tends to occur in humid and flooded areas like marshes, river areas or meadows (Cruz 1993; Vandel 1962) or even anthropic areas like gardens (Vandel 1962), specimens from Gamoniteiru were found in alpine meadows without shrub or tree vegetation under large stones strongly attached to the substrate and far from water sources.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447BFF98FF0D4F46FF320817.taxon	materials_examined	Material examined Spain • 3 ♂ and 3 ♀; Asturias, Oviedo; Barros leg.; MNCN 20.04 / 11854. • 6 specs; Asturias, Ribera de Arriba, Palomar; 43 º 18 ′ 29.66 ′′ N, 5 º 54 ′ 39.93 ′′ W; 130 m a. s. l.; 28 Aug. 2021; J. Robla leg.; hand collected under stones and roof tiles; CLLG. • 17 specs; Asturias, Lena, Pola de Lena (Mofusu forest); 43 º 8 ′ 59.05 ′′ N, 5 º 51 ′ 19.84 ′′ W; 325 m a. s. l.; 1 Oct. 2021; J. Robla leg.; hand collected under logs, trunks, rocks, and leaf litter in a mixed forest of pedunculate oak Quercus robur with common hazel Corylus avellana L., ash Fraxinus excelsior L. and common alder Alnus glutinosa (L.) Gaertn; CLLG. • 1 spec; Asturias, Grado, Somines; 43 º 22 ′ 0.29 ′′ N, 6 º 0 ′ 53.83 ′′ W; 204 m a. s. l.; 27 Sep. 2021; J. Robla leg.; hand collected under stones near a private house; CLLG. • 1 spec; Asturias, Oviedo, Pista Finlandesa; 43 º 23 ′ 8.73 ′′ N, 5 º 50 ′ 12.22 ′′ W; 303 m a. s. l.; 21 Aug. 2021; J. Robla leg.; hand collected under blue gum Eucalyptus globulus leaf litter; CLLG. • 3 spec; Asturias, Gozón, Verdicio beach; 43 º 22 ′ 0.29 ′′ N, 6 º 0 ′ 53.83 ′′ W; 7 m a. s. l.; 27 Aug. 2020; J. Robla leg.; hand collected under stones close to the dunar system; CLLG.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447BFF98FF0D4F46FF320817.taxon	discussion	Remarks and ecology Armadillidium nasatum is a species distributed from western to eastern areas of the Eurasian region. In Spain, this species is limited to northern Iberian Peninsula, from Galicia to Gerona (a summary of most of these records is included in Dollfus 1892; Fidalgo & Herrera 1980; Cruz 1993, Schmölzer 1955, 1971; Vivar et al. 1994; Vázquez-Felechosa & Anadón 2001, Cifuentes 2019) (Fig. 1 A). It had already been recorded in Asturias (Schmölzer 1955, 1971; Vandel 1962; Vázquez Felechosa & Anadón 2001). We provide here several new localities. This species was usually found in several habitats: different native forest masses, eucalyptus plantations, meadows, rural areas, gardens, rocky walls, and coastal habitats. A. nasatum was always located under logs, stones or other structures that create humid and shady microhabitats and our ecological observations agreed with those provided by Vandel (1962).	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447BFF9FFF0D4BA6FE5B0D43.taxon	discussion	Remarks and ecology Species well distributed across Europe in different ecotypes. Rare and scarcely recorded species in the Iberian Peninsula, limited to northern areas, especially in the Pyrenean region: Huesca (Schmölzer 1971, as A. garumnicum); Navarra (Schmölzer 1971, as A. garumnicum; Fidalgo & Herrera 1980; Vivar et al. 1984; Cruz 1993, as A. pictum); Andorra and Lleida (Cruz 1993, as A. pictum). The only two records in Asturias (Cruz 1991 a; Vázquez-Felechosa & Anadón 2001) need to be confirmed since they are based on female specimens that could have been confused with A. boalense sp. nov., which presents a similar habitus. The unique record in Galicia, also a female specimen (Cifuentes 2019), comes from the type locality of A. galiciense and could correspond to this species, now revalidated. During our samplings in Asturias, we have not found Armadillidium pictum. We hypothesised that could be more restricted to the Pyrenean region, being replaced by A. boalense and A. galiciense in northern Iberian Peninsula. Little is known about its ecology, although it is considered a silvicolous species well adapted to other natural and seminatural habitats.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447CFF9FFF0D4E5BFBCE0887.taxon	materials_examined	Material examined Spain • 84 ♂ and 127 ♀; Asturias, Oviedo; Barros leg.; MNCN 20.04 / 7892. • 1 ♂ and 6 ♀; Asturias, Cangas del Narcea, Muniellos; 20 Aug. 1966; E. Ortiz de Vega leg.; MNCN 20.04 / 11551. • 3 ♂ and 18 ♀; Asturias, Cangas del Narcea, Muniellos; 28 Aug. 1966; E. Ortiz de Vega leg.; MNCN 20.04 / 11553. • 2 ♂ and 5 ♀; Asturias, Llanes, Purón; 9 Aug. 1979; E. Ortiz de Vega leg.; MNCN 20.04 / 11537. • 2 ♀; Asturias, Llanes, Purón; 19 Aug. 1979; E. Ortiz de Vega leg.; MNCN 20.04 / 11549. • 2 ♂, 5 ♀ and 2 inmature; Asturias, Llanes, Purón; 22 May. 1982; E. Ortiz de Vega leg.; MNCN 20.04 / 7389. • 1 ♂ and 1 ♀; Asturias, Ribadeva, Pimiango; 6 Aug. 1984; J. Cifuentes leg.; JC 145. • 6 specs; Asturias, Ribera de Arriba, Palomar; 43 º 18 ′ 29.66 ′′ N, 5 º 54 ′ 39.93 ′′ W; 130 m a. s. l.; 28 Aug. 2021; J. Robla leg.; hand collected under stones and roof tiles; CLLG. • 14 spec; Asturias, Grado, Somines; 43 º 22 ′ 0.29 ′′ N, 6 º 0 ′ 53.83 ′′ W; 204 m a. s. l.; 27 Sep. 2021; J. Robla leg.; hand collected under stones near a private house and in a Eucalyptus plantation; CLLG. • 4 spec; Asturias, Oviedo, Pista Finlandesa; 43 º 23 ′ 8.73 ′′ N, 5 º 50 ′ 12.22 ′′ W; 303 m a. s. l.; 21 Aug. 2021; J. Robla leg.; hand collected under blue gum Eucalyptus globulus leaf litter and in a mixed forest mainly compose of Quercus robur; CLLG. • 3 spec; Asturias, Gozón, Verdicio beach; 43 º 22 ′ 0.29 ′′ N, 6 º 0 ′ 53.83 ′′ W; 7 m a. s. l.; 27 Aug. 2020; J. Robla leg.; hand collected under stones close to the dunar system; CLLG. • 20 spec; Asturias, Mieres, Yana’l Monte (Turón); 43 º 12 ′ 24.36 ′′ N, 5 º 45 ′ 11.14 ′′ W; 303 m a. s. l.; 28 Aug. 2021; J. Robla leg.; hand collected under logs, trunks and leaf litter in a mixed forest composed by chestnut Castanea sativa Mill. and pedunculate oak; CLLG. • 5 spec; Asturias, Pola de Siero, El Rebollar Street; 43 º 23 ′ 39.90 ′′ N, 5 º 39 ′ 17.14 ′′ W; 239 m a. s. l.; 3 Sep. 2020; J. Robla leg.; hand collected in a meadow at night; • 5 spec; Asturias, Vegadeo, Piantón; 43 º 27 ′ 39.32 ′′ N, 7 º 1 ′ 43.95 ′′ W; 55 m a. s. l.; 13 Oct. 2021; D. Cabanillas leg.; hand collected in a private garden; CLLG. • 5 spec; Asturias, Selorio, Ría de Villaviciosa; 43 º 31 ′ 10.89 ′′ N, 5 º 23 ′ 2.72 ′′ W; 7 m a. s. l.; 24 Sep. 2021; J. Robla leg.; hand collected under a log in a marsh; CLLG. • 2 spec; Asturias, Coaña, Playa de Foxos; 43 º 33 ’ 19.7 ’’ N, 6 º 43 ’ 36.3 ’’ W; 0 m a. s. l.; 16 May. 2024; G. Sánchez-Jardón leg; hand collected under logs and stones in the sand of a beach; CJR. • 1 spec; Asturias, El Franco, Valdepares; 43 º 33 ’ 42.7 ’’ N, 6 º 52 ’ 3.6 ’’ W; 43 m a. s. l.; 19 May. 2024; G. Sánchez-Jardón leg; hand collected in a catch basin in the private garden of a house.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447CFF9FFF0D4E5BFBCE0887.taxon	discussion	Remarks and ecology A well-distributed species in Europe, the westernmost parts of Asia and some areas of north Africa. Furthermore, the most abundant and well distributed Armadillidium species throughout the Ibero-Balearic area. Species previously recorded in Asturias (Schmölzer 1955; 1971; Vázquez-Felechosa & Anadón 2001; Vázquez-Felechosa et al. 2004) (Fig. 1 A). We provide new localities that point out its broad ecological tolerance: forests (native and artificial plantations), meadows, marshes, riparian zones, coastal areas, and urban and rural areas. During the fieldwork, it has been observed that the dark coloration is the most commonly registered, but reddish or slightly pigmented specimens were sometimes detected. The most common species in urban areas. A. vulgare usually form large aggregations of individuals and are active during daytime, even in areas without great humidity.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447DFF9DFF0D4CA6FAAA0F8F.taxon	distribution	Identification key of the Armadillidium species recorded in the Iberian Peninsula and the Balearic Islands. Mainly based on Schmölzer (1954; 1965); Vandel (1960; 1962); Cruz (1990; 1993); Cruz & Dalens (1990); Garcia (2003; 2020); Garcia & Cabanillas (2021); Cifuentes & Escarabajal (2022 a, 2022 b) and the present study. A. scabrum Dollfus, 1892, (only known from Seville) is excluded due to insufficient information in its original description (Dollfus, 1892). The distribution is indicated only for the endemic species.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
038E862E447DFF9DFF0D4CA6FAAA0F8F.taxon	discussion	* Note that Armadillidium mateui Vandel is not currently included in the genus Armadillidium. This species has been transferred to Alloschizidium Verhoeff (see Cifuentes 2022). Thus, it was not included in the identification key.	en	Cifuentes, Julio, Robla, Jairo, Garcia, Lluc (2024): Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae). Zootaxa 5497 (1): 83-99, DOI: 10.11646/zootaxa.5497.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5497.1.4
