taxonID	type	description	language	source
038EF23FFFEBFFFEFF22F596FBE8F817.taxon	type_taxon	Type species: Harnischia (Cladopelma) tethys Townes, 1945 by objective synonymy.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEBFFFEFF22F596FBE8F817.taxon	diagnosis	Diagnosis (As in Saether, 1977; Cranston et al., 1989; Zorina, 2006). Acrostichals present or absent; anal point arising from bluntly triangular projection of anal tergite; inferior volsella small, without setae, sometimes fully or partly fused with superior volsella; stout gonostylus widest distally in few species.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE8FFFDFF22F3E4FBADF86A.taxon	description	urn: lsid: zoobank. org: act: 4 ED 2 CD 46 - 2 EAB- 4 F 2 E- 9 DF 2 - 50 F 79 AB 607 CC	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE8FFFDFF22F3E4FBADF86A.taxon	materials_examined	Material examined. Holotype male, labelled ‘ Holotype Beckidia inflata sp. n., India, West Bengal, Purba Barddhaman [23.2446 ° N, 87.8496 ° E], 31. iii. 2022, Coll. B. Mukherjee’. Paratype 1 male, labelled ‘ Paratype Beckidia inflata sp. n., India, West Bengal, Purba Barddhaman [23.2446 ° N, 87.8496 ° E], 26. viii. 2022, Coll. B. Mukherjee’. Paratypes 2 males, labelled ‘ Paratype Beckidia inflata sp. n., India, west Bengal, Purba Barddhaman [23.2393 ° N, 87.8512 ° E], 02. ix. 2020, Coll. N. Hazra’. Paratype 1 male, labelled ‘ Paratype Beckidia inflata sp. n., India, West Bengal, Purba Barddhaman [23.1840 ° N, 88.0302 ° E], 20. ii. 2023, Coll. Sk. Mohim Ali’. Diagnostic characters. The new species can be separated from other species of Beckidia by the following combination of characters: acrostichals absent, anal point slightly widened apically, apically broadened superior volsella having 13 – 16 apical setae and covered with microtrichia, and gonostylus apically widened.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE8FFFDFF22F3E4FBADF86A.taxon	etymology	Etymology. The name ‘ inflata ’, originated from a Latin word ‘ inflatus ’, refers to the apically expanded superior volsella. Male (n = 5). Total length 2.75 – 3.07, 2.9 mm. Wing length 1.53 – 1.55, 1.54 mm. Costal length 1.47 – 1.49, 1.48 mm. Antennal length 0.82 – 0.85, 0.835 mm. Colouration. Thorax yellow with brown marking; leg and abdomen yellowish brown. Head. Head width 375 – 470, 422.5 µm. Temporal setae 7 – 8 (IV 2, OV 3, Po 2 – 3). Clypeal setae 10 – 12. Frontal tubercles absent. Eyes bare with dorsomedial extension of 100 – 110, 105 µm. AR 1.58 – 1.63, 1.61; ultimate flagellomere 520 – 530, 525 µm long. Palpomere lengths (I – V) (µm): 34.5: 27.6 – 28, 28.8: 106 – 110; 108: 140 – 146, 143: 200. CA 0.46 – 0.55, 0.50. Thorax. Scutal tubercle absent. Ac 0, Dc 5 – 6, Pa 2, Scts 4. Wing (Fig. 1 A). VR 1.06 – 1.13, 1.095. R 1 and R 4 + 5 bare. Brachiolum with 2 setae. Squama with 4 – 6 fringed setae. FCu distinctly distal to RM. Anal lobe moderately developed. Legs. Fore tibia with 1 seta. Mid legs with 2 tibial spurs; 13.8 µm long with 27 – 28 lateral teeth. Hind leg with 2 tibial spurs; 16 µm and 18.5 µm long with 32 teeth. Lengths and proportions of leg segments shown in table I. Hypopygium (Figs. 1 B – D). Anal tergite band Y shaped. Anal point 42 – 46, 44 µm long, apically expanded. Laterosternite IX with 2 setae on each side of anal point. Longitudinal and transverse sternapodeme 57.5 – 61, 59.25 µm and 27.5 – 30, 28.75 µm long respectively. Superior volsella 53 – 57.5, 55.25 µm long and 28 µm wide apically, somewhat racquet like, microtrichiose with bearing 13 – 16 setae and fused with inferior volsella. Gonocoxite 105 – 115, 110 µm long. Gonostylus 161 – 165, 163 µm long widest distally. HR 0.65 – 0.70, 0.675. HV 2.61 – 2.66, 2.63.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE8FFFDFF22F3E4FBADF86A.taxon	distribution	Distribution. India, West Bengal.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE8FFFDFF22F3E4FBADF86A.taxon	discussion	Remarks. Apically broadened superior volsella, distally thickened gonostylus and absence of acrostichals confirm the emplacing of the new species in the genus Beckidia Saether. The new species shows closeness with B. connexa Zorina, 2006 in having distally swollen gonostylus, nearly similar anal point and lacking of acrostichals. They differ in the shape of anal tergite band (H shaped tergite band in B. connexa while it is Y shaped in new one) and superior volsella (pad like in B. connexa while it is apically inflated in new species).	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFFBFF22F726FB74FAC7.taxon	type_taxon	Type species: Tipula pusilla Linnaeus, 1767 by original designation and monotypy.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFFBFF22F726FB74FAC7.taxon	diagnosis	Diagnosis (As in Reiss, 1988; Cranston et al., 1989; Kobayashi, 2007). The only genus in Harnischia complex having non-shortened inferior volsella covered with short setae only, lacking long apical seta.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFFBFF22F3ABFA46FBDB.taxon	discussion	Remarks. Bhattacharya et al., 1985 described the species in the genus Beckidia. Based on the original description and illustrations (presence of 8 – 10 Acs in two rows, more or less cerciform superior volsella, anal point from the apical end of tergite IX, absence of distally widest gonostylus and blunt projection on tergite IX), we tentatively place it to the genus Paracladopelma. It is also affirmed that the superior volsella is not cerciform rather it is completely pediform, microtrichiose with several apical setae, and slender gonostylus is pointed apically.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFFBFF22F3ABFA46FBDB.taxon	distribution	Distribution. India, West Bengal.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFF9FF22F628FA46F8D3.taxon	description	GenBank Accession No. OP 683429 urn: lsid: zoobank. org: act: 0 D 8 FAD 19 - 2 ADE- 4456 - 963 F- 2 E 22 C 11 D 6057	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFF9FF22F628FA46F8D3.taxon	materials_examined	Material examined. Holotype male, labelled ‘ Holotype Kloosia incurva sp. n., India, West Bengal, Purba Barddhaman [23.2393 ° N, 87.8512 ° E], 22. ix. 2021, Coll. N. Hazra’. Paratypes 3 males, labelled ‘ Paratype Kloosia incurva sp. n., India, West Bengal, Purba Barddhaman [23.2446 ° N, 87.8496 ° E], 30. iii. 2022, Coll. B. Mukherjee’.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFF9FF22F628FA46F8D3.taxon	diagnosis	Diagnostic characters. The adult male is distinguished by the following combination of characters: superior volsella with broad apex having 5 – 6 moderately long setae, elongated inferior volsella with short setae.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFF9FF22F628FA46F8D3.taxon	etymology	Etymology. The name ‘ incurva ’, originated from a Latin word ‘ incurvus ’, refers to the curved gonostylus at distal end.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFF9FF22F628FA46F8D3.taxon	description	Male (n = 4). Total length 3 – 3.3, 3.165 mm. Wing length 1.55 – 1.7, 1.62 mm. Costal length 1.45 – 1.62, 1.53 mm. Antennal length 0.92 – 0.94, 0.93 mm. Colouration. Thorax, leg and abdomen yellowish brown. Head. Head width 440 – 500, 470 µm. Temporal setae 7 – 8 (IV 4, OV 3, Po 0 – 1). Clypeal setae 6. Frontal tubercles absent. Eyes bare with dorsomedial extension of 115 – 120, 117.5 µm. AR 1.4 – 1.5, 1.45; ultimate flagellomere 620 – 630, 625 µm long. Palpomere lengths (I – V) (µm): 23: 25 – 27.5, 26.25: 66.5 – 70, 68.25: 94.5: 110.5 – 112, 111.25. CA 0.48 – 0.53, 0.505. Thorax. Scutal tubercle absent. Ac 2, Dc 6, Pa 2, Scts 4, Su 1. Wing (Fig. 2 A). VR 0.93 – 0.96, 0.945. R 1 without setae, R 4 + 5 with 3 – 4 setae. Brachiolum with 2 setae. Squama with 5 fringed setae. FCu distinctly distal to RM. Anal lobe moderately developed. Legs. Fore tibia with 2 setae. Mid legs with 2 tibial spurs; 16 – 18, 17 µm and 27.6 µm long with 16 lateral teeth. Hind leg with 2 tibial spurs; 18.4 µm and 23 µm long with 22 teeth. Lengths and proportions of leg segments shown in table II. Hypopygium (Figs. 2 B – E). Anal tergite band Y shaped. Anal point rounded apically, 57.5 – 60, 58.75 µm long and 11.5 µm wide at apex. Laterosternite IX with 2 setae on each side of anal point. Longitudinal and transverse sternapodeme 92 – 95, 93.5 µm and 27.5 µm long respectively. Superior volsella somewhat spatulate like at apex, 50 – 53, 52.5 µm long and 25.5 µm wide at apex bearing 5 – 6 long setae. Elongated inferior volsella straight and distally widened bearing 5 small setae, 90 – 92 µm long and 30 µm wide at apex. Gonocoxite 132 – 138, 135 µm long. Gonostylus with partial cleft at apex, 149.5 – 152, 150.75 µm long, fairly curved and swollen distally. HR 0.88 – 0.91, 0.895. HV 2.27 – 2.39, 2.33.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFF9FF22F628FA46F8D3.taxon	distribution	Distribution. India, West Bengal.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEEFFF9FF22F628FA46F8D3.taxon	discussion	Remarks. Characters such as weakly appeared partial cleft on apex of gonostylus, non shortened inferior volsella with small setae and distally expanded superior volsella affirm positioning of the new species in the genus Kloosia Kruseman. Somewhat similar kind of superior volsella and anal point are present in both Kloosia dorsenna (Saether, 1983) and the new species, but they differ in variable number of setae on superior volsella (5 – 6 setae on superior volsella of new species while it is 3 in number in K. dorsenna), shape of gonostylus (roughly straight gonostylus present in K. dorsenna while distally curved in the new species.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFECFFF8FF22F425FA85FEB5.taxon	type_taxon	Type species: Chironomus cryptotomus Kieffer, 1915 by subsequent designation of Townes, 1945.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFECFFF8FF22F425FA85FEB5.taxon	diagnosis	Diagnosis (As in Cranston et al., 1989; Spies et al., 1994). Anal tergite band weak V to Y-type, often separated medially, median longitudinal band long to absent; caudal region of anal tergite with or without projections; superior volsella in dorsal view generally rod-shaped, slender, straight to slightly sinuous, distal part often bending to dorsal, slightly to distinctly widened and sometime with median extension give pediform appearance, distal region have two setae arising from distinct, small and shallow, deep pits, usually one pit smaller and more median, subapical to slightly proximal position; pubescent inferior volsella without setae, rarely reaching beyond anal tergite with bluntly rounded to fairly acute median to caudal projection; gonostylus straight or curving medially and / or dorsally.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEDFFF6FF22F2DDFAB7FEB5.taxon	description	urn: lsid: zoobank. org: act: 5 C 145280 - F 27 F- 4916 - 8678 - C 55 A 2 A 150 EC 9	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEDFFF6FF22F2DDFAB7FEB5.taxon	materials_examined	Material examined. Holotype male, labelled ‘ Holotype Parachironomus salsus sp. n., India, West Bengal, Purba Barddhaman [23.2393 ° N, 87.8512 ° E], 16. viii. 2021, Coll. N. Hazra’. Paratype 1 male, same data as holotype, 07. iii. 2022, Coll. N. Hazra. Paratype 1 male, labelled ‘ Paratype Parachironomus salsus sp. n., India, West Bengal, Purba Barddhaman [23.2441 ° N, 87.8368 ° E], 30. iv. 2022, Coll. B. Mukherjee’. Diagnostic characters. The adult male can be differentiated by the presence of nearly vase shaped anal point, sharply pointed beak like superior volsella and medially swollen gonostylus.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEDFFF6FF22F2DDFAB7FEB5.taxon	etymology	Etymology. The name ‘ salsus ’, a Latin word, refers to the sharp superior volsella. Male (n = 3). Total length 2.9 – 3.1, 3 mm. Wing length 1.52 – 1.55, 1.535 mm. Costal length 1.48 – 1.50, 1.49 mm. Antennal length 0.93 – 0.96, 0.945 mm. Colouration. Yellowish thorax with dark brown marking; leg and abdomen yellowish brown. Head. Head width 520 – 530, 525. Temporal setae 8 (IV 2, OV 4, Po 2). Clypeal setae 15 – 16. Frontal tubercles absent. Eyes bare with dorsomedial extension of 108 – 112, 110 µm. AR 1.82 – 1.90, 1.86; ultimate flagellomere 610 – 620, 615 µm long. Palpomere lengths (I – V) (µm): 34.5: 27.5 – 30, 28.75: 85 – 86.5, 85.75: 120 – 124, 122: 177. CA 0.55 – 0.56, 0.555. Thorax. Scutal tubercle absent. Ac 11, Dc 7, Pa 3, Scts 10, Su 1. Wing (Fig. 3 A). VR 1.12 – 1.14, 1.13. R 1 with 32 – 34 setae, R 4 + 5 with 15 – 18 setae. Brachiolum with 2 setae. Squama with 2 setae. FCu distinctly distal to RM. Anal lobe moderately developed. Legs. Fore tibia with 2 setae. Mid legs with 2 tibial spurs; each 18.4 µm long with 21 lateral teeth. Hind leg with 2 tibial spurs; 16 µm and 11.5 µm long with 31 teeth. Lengths and proportions of leg segments shown in table III. Hypopygium (Figs. 3 B – D). Anal tergite band Y shaped. Anal point basally and apically constricted while slightly expanded medially, 65 – 69, 67 µm long and 16 – 20, 18 µm wide, with a median ridge like structure. Laterosternite IX with 3 setae on each side of anal point. Longitudinal and transverse sternapodeme 87.5 – 90, 88.75 µm and 40 µm long respectively. Superior volsella 53 – 57.5, 55.25 µm long and 28 µm wide, roughly straight with pointed beak like structure at apical end bearing 2 setae subapically, 1 of the setae (34.5 µm long) placed on a deep and wide pit and other one 16 µm long located on distinct shallow pit. Inferior volsella with a blunt projection covered with microtrichia. Gonocoxite 126.5 – 130, 128.25 µm long. Gonostylus 165.5 – 170, 167.75 µm long, bluntly rounded at apex. HR 0.76 – 0.77, 0.765. HV 2.29 – 2.38, 2.335.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEDFFF6FF22F2DDFAB7FEB5.taxon	distribution	Distribution. India, West Bengal.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFEDFFF6FF22F2DDFAB7FEB5.taxon	discussion	Remarks. Presence of distolateral beak on superior volsella with 2 setae arise from distinct pits, long anal point and variable gonostylus assert the placement of the species in the genus Parachironomus Lenz. P. vitiosus Goetghebuer, 1921 is similar with the new species in having somewhat similar shaped superior volsella but differs in the shape of gonostylus and anal point. The new species and P. matapi Spies, Fittkau & Reiss, 1994 have nearly similar shaped gonostylus but differs in the shape of superior volsella and anal point. The new species and P. elodeae (Townes, 1945) have similar type of superior volsella but they differ in the shape of gonostylus and anal point.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE3FFF3FF22F28DFA46FD0A.taxon	description	GenBank accession number. MW 012424	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE3FFF3FF22F28DFA46FD0A.taxon	materials_examined	Material examined. Holotype male labelled ‘ Holotype Cryptotendipes disparilis Pal & Hazra, 2018, India, West Bengal, Malda [24.90 ° N, 88.11 ° E], 06. ii. 2013, Coll. G. Pal’. 2 males, India, West Bengal, Purba Barddhaman, Burdwan [23.22 ° N, 87.85 ° E], 23. xii. 2019, Coll. N. Hazra; 2 males, India, West Bengal, Purba Barddhaman, Burd _ wan [23.22 ° N, 87.85 ° E], 15. i. 2020, Coll. N. Hazra; 5 males, India, West Bengal, Purba Barddhaman, Burdwan [23.22 ° N, 87.85 ° E], 16. ii. 2020, Coll. N. Hazra; 2 males, India, West Bengal, Birbhum [24.435 ° N, 87.911 ° E], 10. iv. 2022, Coll. B. Mukherjee.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE3FFF3FF22F28DFA46FD0A.taxon	diagnosis	Diagnosis. The species have V shaped anal tergite band; superior volsella distally somewhat triangular in shape bearing two apical setae and gonostylus basally expanded, gradually tapered distally with a nipple-like apex bearing single seta.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE3FFF3FF22F28DFA46FD0A.taxon	discussion	Remarks. Sasa & Hasegawa (1983) tentatively placed the species into the genus Harnischia, described by Goetghebuer, 1936 as Chironomus (Harnischia) acutus. Yan et al. (2015) tentatively transferred this species into Parachironomus. Pal and Hazra (2018) described Cryptotendipes disparilis from India. After an extensive study of specimens, we have been able to affirm its placement in the genus Cryptotendipes Beck & Beck along with synonymisation of other known species with Cryptotendipes acutus. The combination of those characters in male imagines justifying the placement and synonymy includes gonostylus with strong basal expansion and superior volsella having two apical setae not arising from conspicuous pits. Other significant characters are: total length 2.98 – 3.27, 3.13 mm; eyes with dorsomedian parallel sided extension of 150 – 180, 165 µm; AR 1.9 – 2.07, 1.99; CA 0.61 – 0.66, 0.63; Ac 12 – 14, Dc 7 – 8, Pa 3 – 5, Scts 8 – 10; vein R with 42 – 46, R 1 30 – 32, R 4 + 5 12 – 14 setae; VR 1.26 – 1.33, 1.3; T IX with 19 – 22, 20 median long setae; anal point 72.3 – 80, 75 µm long with 2 – 3 basolateral setae on each side; gonocoxite 165 – 172, 168 µm long; superior volsella 23 – 27.6, 25 µm long, 11.5 – 14, 12 µm wide; transverse sternapodeme 55 – 59, 57 µm long, lateral sternapodeme 128 – 134, 131 µm long; HR 1.2 – 1.27, 1.23.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE3FFF3FF22F28DFA46FD0A.taxon	distribution	Distribution. Egypt, Israel, Chad, Congo, Ethiopia, Kenya, Nigeria, South Africa, Japan, India.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE6FFF3FF22F051FEEBFBD5.taxon	type_taxon	Type species: Harnischia (Cladopelma) tylus Townes, 1945 by original designation. Emended diagnosis: Hypopygium: Superior volsella consists of bare dorsal lobe with or without digitiform projection and ventral lobe covered with microtrichia, bearing 1 – 2 setae; squama of wing with reduced fringe of about 7 setae.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE6FFF1FF22F738FBB1FB88.taxon	description	urn: lsid: zoobank. org: act: 60 F 40 CC 2 - CCA 8 - 44 E 2 - B 256 - 7 CB 9 E 40 BC 76 F	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE6FFF1FF22F738FBB1FB88.taxon	materials_examined	Material examined. Holotype male, labelled ‘ Holotype Saetheria circinata sp. n., India, West Bengal, Purba Barddhaman [23.2393 ° N, 87.8512 ° E], 04. ii. 2020, Coll. N. Hazra’. Paratype 1 male, same data as holotype, 25. vi. 2020, Coll. N. Hazra. Paratypes 2 males, same data as holotype, 15. ii. 2021, Coll. N. Hazra. Paratype 1 male, same data as holotype, 14. vi. 2021, Coll N. Hazra. Paratype 1 male, same data as holotype, 19. xi. 2021, Coll. N. Hazra. Diagnostic characters. The distinguished features of new species are as follows: apically slightly broadened anal point with a median ridge, Y shaped anal tergite band, anal tergite band and nearly triangular shaped ventral lobe of superior volsella.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE6FFF1FF22F738FBB1FB88.taxon	etymology	Etymology. The name ‘ circinata ’, originated from a Latin word ‘ circinatus ’, refers to circular dorsal lobe of superior volsella. Male (n = 6) Total length 2.75 – 2.8, 2.775 mm. Wing length 1.7 – 1.85, 1.77 mm. Costal length 1.65 – 1.7, 1.67 mm. Antennal length 0.82 – 0.85, 0.835 mm. Colouration. Thorax with dark brown marking; leg and abdomen yellowish brown. Head. Head width 450 – 460, 455 µm. Temporal setae 8 – 9 (IV 4 – 5, OV 3 – 4, Po 0 – 1). Clypeal setae 12. Frontal tubercles present. Eyes bare with dorsomedial extension of 130 – 140, 135 µm. AR 1.2 – 1.41, 1.30; ultimate flagellomere 460 – 480, 470 µm long. Palpomere lengths (I – V) (µm): 23 – 25, 24: 30: 80.5 – 82, 81.25: 103.5: 149.5 – 152, 150. CA 0.54 – 0.55, 0.545. Thorax. Scutal tubercle absent. Ac 8, Dc 7, Pa 3, Scts 2 – 3. Wing (Fig. 4 A). VR 1.14 – 1.16, 1.15. R 1 and R 4 + 5 bare. Brachiolum with 1 seta. Squama with 7 fringed setae. FCu distinctly distal to RM. Anal lobe moderately developed. Legs. Fore tibia with 2 setae. Mid legs with 2 tibial spurs; 16 µm and 18.4 µm long with 22 – 26 lateral teeth. Hind leg with 2 tibial spurs; 18.4 µm and 20.7 µm long with 29 – 32 teeth. Lengths and proportions of leg segments shown in table IV. Hypopygium (Figs. 4 B – D). Anal tergite band Y shaped. Anal point slightly expanded apically, 37 – 40, 38.5 µm long with a median ridge like appearance. Laterosternite IX with 3 setae on each side of anal point. Longitudinal and transverse sternapodeme 75 – 78, 76.5 µm and 73.5 µm long respectively. Superior volsella 55 – 58, 56.5 µm long and 25.3 µm wide, nearly triangular in shape, consisting of dorsal and ventral lobe, dorsal lobe having roughly circular base (23 µm long) with digitiform projection (27 – 28 µm long) while ventral lobe nearly triangular shaped bearing 2 setae (1 seta on heel and another one placed subapically). Inferior volsella with a rounded apical part covered with microtrichia. Gonocoxite 122 – 126, 124 µm long. Gonostylus 126.5 – 131, 128.75 µm long. HR 0.96 – 0.97, 0.965. HV 2.18 – 2.29, 2.235.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE6FFF1FF22F738FBB1FB88.taxon	distribution	Distribution. India, West Bengal.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE6FFF1FF22F738FBB1FB88.taxon	discussion	Remarks. Presence of roughly bilobed superior volsella having digitiform projection on dorsal lobe, Y shaped anal tergite band and spatulated anal point confirm the positioning of new species into genus Saetheria Jackson. The new species closely related with S. digitata Yan et al., 2011 in having more or less similar gonostylus and dorsal lobe of superior volsella but presence of triangular like ventral lobe of superior volsella with 1 seta at apical end and somewhat apically spatulated anal point can segregate the new species from S. digitata.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE4FFF0FF22F7DAFA47FDDD.taxon	diagnosis	Diagnostic characters. Anal point spoon-like at apex; superior volsella is somewhat flattened pad-like, composed of dorsal and ventral lobes and each lobe bearing one seta.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE4FFF0FF22F7DAFA47FDDD.taxon	discussion	Remarks. Based on the original description and illustration, following corrections have been made: anal point spoon-like at apex; superior volsella somewhat flattened pad-like, divided into dorsal and ventral lobes and each lobe bearing one seta.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE4FFF0FF22F7DAFA47FDDD.taxon	distribution	Distribution. India, West Bengal (present record).	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE5FFF0FF22F113FBCEFCF8.taxon	type_taxon	Type species. Harnischia fuscimana, Kieffer, 1921, by subsequent designation and monotypy. Emended diagnosis. Frontal tubercle present or absent. Gonostylus variable shaped (short and broad or long and narrow), anal point broad to narrow and parallel shaped to variously developed.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE5FFEEFF22F009FE2DFCC4.taxon	description	GenBank accession number. OQ 400990 urn: lsid: zoobank. org: act: 2 E 308 C 20 - 6 D 6 E- 4 FB 2 - B 3 D 5 - 323 DD 48 FD 0 F 0	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE5FFEEFF22F009FE2DFCC4.taxon	materials_examined	Material examined. Holotype male, labelled ‘ Holotype Harnischia bulbosa sp. n., India, West Bengal, Purba Barddhaman [23.2393 ° N, 87.8512 ° E], 04. ii. 2020, Coll. N. Hazra’. Paratype 1 male, labelled ‘ Paratype Harnischia bulbosa sp. n., India, West Bengal, Purba Barddhaman [23.1840 ° N, 88.0302 ° E], 14. xi. 2022, Coll. Sk. Mohim Ali’.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE5FFEEFF22F009FE2DFCC4.taxon	diagnosis	Diagnosis. Presence of spoon shaped anal point and long gonostylus are the segregating characters of this new species.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE5FFEEFF22F009FE2DFCC4.taxon	etymology	Etymology. The name ‘ bulbosa ’, a Latin word, refers to bulbous apex of the anal point.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE5FFEEFF22F009FE2DFCC4.taxon	description	Male (n = 3). Total length 2.7 – 2.85, 2.775 mm. Wing length 1.4 – 1.43, 1.415 mm. Costal length 1.30 – 1.35, 1.325 mm. Antennal length 0.70 – 0.75, 0.725 mm. Colouration. Thorax, leg and abdomen yellowish brown. Head. Head width 400 – 410, 405 µm. Temporal setae 5 – 7 (IV 0, OV 2, Po 3 – 5). Clypeal setae 12 – 14. Frontal tubercles absent. Eyes bare with dorsomedial extension of 138 – 150, 144 µm. AR 1.65 – 1.68, 1.66; ultimate flagellomere 470 – 480, 475 µm long. Palpomere lengths (I – V) (µm): 34.5: 25 – 30, 27.5: 62: 87.5: 131. CA 0.53 – 0.54, 0.535. Thorax. Scutal tubercle absent. Ac 11, Dc 7, Pa 3, Scts 2. Wing (Fig. 5 A). VR 1.12 – 1.14, 1.13. R 1 with 10 setae, R 4 + 5 bare. Brachiolum with 1 seta. Squama with 3 – 4 fringed setae. FCu distal to RM. Anal lobe moderately developed. Legs. Fore tibia with 2 setae. Mid legs with 2 tibial spurs; 13.5 µm and 16 µm long with 24 – 26 lateral teeth. Hind leg with 2 tibial spurs; 11.5 µm and 23 µm long with 36 – 40 teeth. Lengths and proportions of leg segments shown in table V. Hypopygium (Figs. 5 B – C). Anal tergite band V shaped. Anal point more or less spoon-shaped, slightly expanded sub-apically with blunt apex, 32 – 39, 35.5 µm long with a median ridge like structure and small lobe-like appearance on each side of the anal point. Anal point with 2 lateral setae on each side. Apical anal tergite with 5 setae. Laterosternite IX with 4 setae. Longitudinal and transverse sternapodeme 65 – 69, 67 µm and 22 – 25, 23.5 µm long respectively. Superior and inferior volsellae greatly reduced. Gonocoxite 92 – 96.5, 94.25 µm long. Gonostylus 131 µm long. HR 0.70 – 0.74. HV 2.96 – 3.09.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE5FFEEFF22F009FE2DFCC4.taxon	distribution	Distribution. India, West Bengal.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFE5FFEEFF22F009FE2DFCC4.taxon	discussion	Remarks. Absence of frontal tubercles and attenuated gonostylus from the junction with gonocoxite and presence of vestigial superior and inferior volsellae specify the positioning of new species into the genus Harnischia Kieffer. The new species is closely related to H. incidata (Townes, 1945) in having more or less long gonostylus. However, the presence of spoon-shaped anal point in the new species while apically pointed anal point of H. incidata can differentiate them.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFFBFFEDFF22F0DEFA46FBEB.taxon	materials_examined	Material examined. Holotype male labelled ‘ H. sibabecea Sasa, Sumita & Suzuki, 1999 (http: // www. type. kahaku. go. jp / TypeDB / diptera / 973) ’; Holotype male labelled ‘ H. minuta Dutta & Chaudhuri, 1996 (Type no. B. U. Ent. 210) ’; 2 males, India, West Bengal, Purba Barddhaman, Burdwan [23.22 ° N, 87.85 ° E], 16. ii. 2022, Coll. N. Hazra; 2 males, India, West Bengal, Purba Barddhaman, Burdwan [23.2446 ° N, 87.8496 ° E], 18. iv. 2022, Coll. B. Mukherjee; 3 males, India, West Bengal, Jalpaiguri [26.71 ° N, 88.76 ° E], 07. iv. 2022, Coll. P. Hui’.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFFBFFEDFF22F0DEFA46FBEB.taxon	discussion	Remarks. Harnischia ohmuraensis Kobayashi & Suzuki, 1999, H. sibabecea Sasa, Sumita & Suzuki, 1999, H. minuta Dutta & Chaudhuri, 1996 and H. tenuitubercula Chaudhuri & Chattopadhyay, 1990 have medially setose anal tergite, long gonostylus, broad anal point, and very small superior volsella. Kobayashi & Endo (2008) have synonymised H. sibabecea Sasa, Sumita & Suzuki, 1999 with H. ohmuraensis Kobayashi & Suzuki, 1999. Besides that, Chen et al. (2017) mentioned that H. tenuitubercula Chaudhuri & Chattopadhyay, 1990 and H. ohmuraensis Kobayashi & Suzuki, 1999 may be conspecific. After a considerate study of types, we come to conclusion that H. minuta Dutta & Chaudhuri, 1996 is also conspecific with H. sibabecea Sasa, Sumita & Suzuki, 1999. The original description of H. tenuitubercula was published in 1990 while descriptions of other species were made on 1996 and 1999 respectively. Therefore, the species H. minuta, H. ohmuraensis and H. sibabecea are junior synonyms of H. tenuitubercula.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFFBFFEDFF22F0DEFA46FBEB.taxon	description	(Modified after Chen et al., 2017)	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFF8FFEDFF22F7FEFEC6F914.taxon	materials_examined	Material examined. 3 males, labelled ‘ Microchironomus ishii Sasa, 1987, India, West Bengal, Jalpaiguri [26.71 ˚ N, 88.76 ˚ E], 07. iv. 2022, Coll. P. Hui’.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFF8FFEDFF22F7FEFEC6F914.taxon	discussion	Remarks (Figs. 6 A – C). Sasa, 1987 described this species from Japan. Sasa, 1987 described this species from Japan, collected in open light open light trap during this revisionary study. The specimen fully corroborates with M. ishii due to the presence of following diagnostic characters: anal point with small setigerous lobe at basal region, rod-like superior volsella subapically slight expanded with 3 simple setae, similar LR 2, LR 3, and wing length. Upon examining the Indian specimens, we observed that AR 2 (AR 1.6 – 1.8 in Japanese specimens), and Scts 4 (Scts 6 – 12 in Japanese specimens). Moreover, other significant characters are as follows: eyes with dorsomedial extension of 140 µm; V shaped anal tergite band; gonocoxite 105 – 110.5 µm long; gonostylus 122 – 130 µm long; anal point 70 – 73.5 µm long and widened basally with 3 setae lateral to anal point; superior volsella 48 – 50 µm long with little expansion at sub-apical end have 3 setae (1 seta at apex, 2 setae located sub-apically); HR 0.86 – 0.87; HV 2.85 – 3.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFF8FFEDFF22F7FEFEC6F914.taxon	distribution	Distribution. The species is reported first time from India, West Bengal. Previously the species was reported from Japan.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFF8FFEAFF22F466FD9EFD55.taxon	materials_examined	Material examined. 8 males, labelled ‘ Microchironomus fuscitarsus (Guha & Chaudhuri, 1979), India, West Bengal, Burdwan [23.2393 ° N, 87.8512 ° E], 05. ii. 2021, 10. v. 2021, 6. vii. 2021, 03. ii. 2022, Coll. N. Hazra’, 2 males labelled ‘ Microchironomus fuscitarsus (Guha & Chaudhuri, 1979), India, West Bengal, Birbhum (23.668 ° N, 87.682 ° E), 03. iii. 2021, Coll. B. Mukherjee’, 3 males, labelled ‘ Microchironomus fuscitarsus (Guha & Chaudhuri, 1979), India, West Bengal, Burdwan [23.2446 ° N, 87.8496 ° E], 05. iii. 2022, 10. iv. 2021, Coll. B. Mukherjee’, 3 males, ‘ Microchironomus fuscitarsus (Guha & Chaudhuri, 1979), India, West Bengal, Jalpaiguri [23.71 ° N, 88.76 ° E], 06. iv. 2022, Coll. P. Hui’.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFF8FFEAFF22F466FD9EFD55.taxon	discussion	Remarks (Figs. 7 A – C). Guha & Chaudhuri (1979) described adult male of this species as Cryptochironomus fuscitarsus while Chaudhuri & Chattopadhyay (1988) described immature stages of this species and transferred this species into the genus Microchironomus. We noticed that few characters of the original description of the adult male are erroneous. Revised characters are: anal tergite band V shaped; superior volsella tapered gradually bearing 3 setae (1 seta at apex and 2 at sub-apex), 57 – 60 µm long; anal point medially swollen, 76 – 80 µm long, parallel shaped; laterosternite IX with 3 – 4 setae on each side of anal point.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
038EF23FFFF8FFEAFF22F466FD9EFD55.taxon	distribution	Distribution. India, West Bengal.	en	Mukherjee, Bindarika, Hazra, Niladri (2023): Taxonomic studies on Harnischia complex from India (Diptera: Chironomidae). Zootaxa 5278 (2): 239-263, DOI: 10.11646/zootaxa.5278.2.2, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.2
