taxonID	type	description	language	source
038E4022B10BFFECFF6DF8805FB7E8FD.taxon	materials_examined	Material examined. TAIWAN: 1 male, Taichung City, Daxue Mt. Forest Trail 31.7 k [L ṞƜẇae 31.7 k], Heping Dist., 24.2255, 120.9764, alt. about 1950 m; 28 – 11 – 2019, leg. F. S. Hu (in dead wood) / Quwatanabius chiaw (Smetana, 1995), Det. By F. S. Hu, 2019.	en	Hu, Fang-Shuo, Bogri, Amalia (2020): Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae). Zootaxa 4743 (2): 285-288, DOI: 10.11646/zootaxa.4743.2.12
038E4022B10BFFECFF6DF8805FB7E8FD.taxon	description	Description. Male. Body (Fig. 1) matches the species description of the female (Smetana 1995). Sternite VIII (Fig. 2) with rather narrow, deep, triangular medioapical emargination. Sternite IX (Fig. 3) with rather narrow, U-shaped apical emargination, with one strong and long seta on each side of the emargination, plus two longitudinally arranged strong and long setae near the apex of each lobe of the sternite. Tergite X (Fig. 4) with a pair of short setae in the middle; with rather narrow, U-shaped apical emargination with two to three pairs of strong and long setae and a pair of rather short longitudinally arranged setae on each side of the emargination. Aedeagus (Figs. 5 – 6) relatively small; median lobe subparallel-sided in middle portion, anteriorly conically narrowed into acute apex. Paramere small, trapezoid-shaped, with a broad medioapical emargination; subtriangular-shaped in lateral view. Internal sac with two strip-like sclerites, each sclerite with two sizeable teeth and a blunt protrusion. Female. See Smetana (1995).	en	Hu, Fang-Shuo, Bogri, Amalia (2020): Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae). Zootaxa 4743 (2): 285-288, DOI: 10.11646/zootaxa.4743.2.12
038E4022B10BFFECFF6DF8805FB7E8FD.taxon	diagnosis	Diagnosis. Quwatanabius chiaw is similar to Q. flavicornis but can be distinguished by the following characters: pal- er, rufo-testaceous pronotum; relatively shorter elytra; absent pale apical seam of palisade fringe on tergite VII; broader apex of median lobe (in ventral view); simpler structure of internal sac, where lateral protrusions are not visible in ventral view. Bionomics. Smetana (1995) was not able to report the bionomics when he described this species. Additional specimens in Smetana (1996) were collected by sifting rotting, moist twigs, logs and bark along fallen trees. The specimen in this study was collected from the inside of a large and moist, strongly decomposed piece of fallen dead wood in primary, evergreen forest. According to the available locality data, the species is found on high elevations (approximately 1900 m or higher).	en	Hu, Fang-Shuo, Bogri, Amalia (2020): Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae). Zootaxa 4743 (2): 285-288, DOI: 10.11646/zootaxa.4743.2.12
038E4022B10BFFECFF6DF8805FB7E8FD.taxon	discussion	Comments. The hind wings of this species are quite short and probably non-functional. The absence of the apical seam of palisade fringe on tergite VII also suggests this species is flightless. Quwatanabius chiaw is the only species considered to be flightless in this genus at the moment. Hence, we suggest that this is an endemic species with restricted distribution in the mountainous areas of central Taiwan.	en	Hu, Fang-Shuo, Bogri, Amalia (2020): Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae). Zootaxa 4743 (2): 285-288, DOI: 10.11646/zootaxa.4743.2.12
038E4022B10BFFECFF6DF8805FB7E8FD.taxon	distribution	Distribution. The species is known from central Taiwan (Nantou County and Taichung City).	en	Hu, Fang-Shuo, Bogri, Amalia (2020): Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae). Zootaxa 4743 (2): 285-288, DOI: 10.11646/zootaxa.4743.2.12
038E4022B109FFEDFF6DFEA95A42E8D2.taxon	materials_examined	Material examined. TAIWAN: 1 male, Nantou Co., Guandao Mt. [ḆBƜ], Renai To., 24.0191, 121.0108 (alt. 1470 m), 23 – III – 2019, leg. F. S. Hu (in dead wood) / Quwatanabius flavicornis (Sharp, 1889), Det. By F. S. Hu, 2019; 1 female, same location, 01 – XI – 2019, leg. F. S. Hu (sifting pieces of dead wood).	en	Hu, Fang-Shuo, Bogri, Amalia (2020): Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae). Zootaxa 4743 (2): 285-288, DOI: 10.11646/zootaxa.4743.2.12
038E4022B109FFEDFF6DFEA95A42E8D2.taxon	description	Description. See Smetana (1995); for illustration of internal sac of aedeagus see Smetana (2002).	en	Hu, Fang-Shuo, Bogri, Amalia (2020): Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae). Zootaxa 4743 (2): 285-288, DOI: 10.11646/zootaxa.4743.2.12
038E4022B109FFEDFF6DFEA95A42E8D2.taxon	diagnosis	Diagnosis. Quwatanabius flavicornis (Fig. 7) is similar to Q. chiaw but can be distinguished from it by the following characters: pronotum much darker; elytra relatively longer; pale apical seam of palisade fringe on tergite VII present; median lobe rather narrow at apex, internal sac of more complex structure. Bionomics. The specimens in this study were collected from a large, fallen, moist, strongly decomposed piece of dead wood in primary, evergreen forest (Fig. 8), similar to the habitat where Quwatanabius chiaw was collected. The two species, however, have been collected from different mountains, and most specimens of Q. flavicornis were collected from lower altitudes (approximately 1500 m), compared to the higher altitudes where Q. chiaw has been found.	en	Hu, Fang-Shuo, Bogri, Amalia (2020): Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae). Zootaxa 4743 (2): 285-288, DOI: 10.11646/zootaxa.4743.2.12
038E4022B109FFEDFF6DFEA95A42E8D2.taxon	discussion	Comments. The bionomics of the Chinese species remains unknown, except Q. dayaoensis which we know was collected from dead wood (Hu et al. 2013). Since both species from Taiwan were collected from dead wood as well, we assume that this genus may be saproxylophilic.	en	Hu, Fang-Shuo, Bogri, Amalia (2020): Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae). Zootaxa 4743 (2): 285-288, DOI: 10.11646/zootaxa.4743.2.12
038E4022B109FFEDFF6DFEA95A42E8D2.taxon	distribution	Distribution. The species is widespread in Japan (Honshu, Kyushu, and Shikoku) and Taiwan (Nantou County and Chiayi County).	en	Hu, Fang-Shuo, Bogri, Amalia (2020): Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae). Zootaxa 4743 (2): 285-288, DOI: 10.11646/zootaxa.4743.2.12
