taxonID	type	description	language	source
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	diagnosis	Diagnosis. Eutyphlus dybasi differs from all known Eutyphlus species, except for E. spiralis, in lacking the anteroprosternal foveae, the presence of secondary sexual modifications on the legs of males (Fig. 19), the quadrifoveate elytra, and concave metaventrite of both sexes, along with the larger, broadly ovate seventh ventrite of males and the large, concave, triangular seventh ventrite of females. The aedeagus is larger than that of other species in the genus, with the exception of E. spiralis, and differs in structure, bearing an enlarged dorsal lobe, a left paramere bearing distal ctenidia, a median lobe with an extensive fimbriated process, and a flattened, ventral right paramere (Fig. 13). Eutyphlus dybasi is most similar to E. spiralis and can be separated by differences in the morphology of the aedeagus. In E. dybasi, a brush of ctenidial spines occurs on the left, dorsally located paramere, and the distal expansion of the right, ventrally located paramere curves towards the right; the left paramere is spiralshaped, and the right paramere curves towards the left in E. spiralis.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	description	Redescription. MALE. Measurements: HL 0.20 mm, HW 0.29 mm; PL 0.34 mm, PW 0.32 mm; EL 0.55 mm, EW 0.55 mm; A 1 - 6 respectively 0.02, 0.15, 0.15, 0.11, 0.11, 0.3 mm; An 1 – 11 respectively 0.09, 0.05, 0.03, 0.02, 0.02, 0.02, 0.03, 0.03, 0.04, 0.04, 0.10 mm; MP 1 – 4 respectively 0.01, 0.05, 0.01, 0.05 mm. GL 0.22 mm, GW 0.14 mm. ML 1.62 mm. Integument: Typical for genus. Head: Slightly narrower than pronotum. Tempora evenly rounded. Eyes present and well-developed with 35 facets, slightly emarginate posteriorly, ocular canthus weakly developed, projecting weakly laterally and sloping gently to gular area. Ventral surface of head flat, slightly rounded in gular area, gular sulcus well-developed anteriorly and projecting to margin of posteriorly located, paired, gular foveae. Thorax: Prothorax with basolateral margin slightly crenulate, median longitudinal sulcus present, area behind basal sulcus with thin longitudinal sulcus extending from basal bead to basal sulcus. Prosternum lacking anteroprosternal foveae. Mesotrochanter bearing tubercle, mesotibia with blunt apical spine. Metaventrite slightly concave baso-medially. Metatrochanter with tubercle. Elytra bearing 3 basal foveae as well as subhumeral foveae. Flight wings well-developed. Abdomen: Tergites unmodified. Seventh ventrite broadly oval, other ventrites unmodified. Genitalia: Aedeagus asymmetrical. Dorsal lobe broad, membranous. Left paramere expanded distally, bearing brush of ctenidial spines. Median lobe with distally fimbriated process. Right paramere more ventral, flattened, distally expanded towards right, with apical patch of scattered, flattened setae. FEMALE. Measurements: HL 0.25 mm, HW 0.30 mm; PL 0.38 mm, PW 0.32 mm; EL 0.50 mm, EW 0.60 mm; A 1 - 6 respectively 0.03, 0.17, 0.15, 0.15, 0.13, 0.05 mm. An 1 – 11 respectively 0.09, 0.05, 0.03, 0.02, 0.02, 0.02, 0.03, 0.03, 0.04, 0.04, 0.10 mm. MP 1 – 4 respectively 0.01, 0.05, 0.01, 0.05 mm. GL 0.15 mm, GW 0.21. ML 1.7 mm. Integument, head, thorax, and abdomen: Similar to male except eyes smaller with 12 facets, legs unmodified, 7 th abdominal ventrite slightly concave, broadly triangular. Genitalia: Membranous, narrowed towards distal end. Internally sclerotized portion forked at base and fused at 1 / 3 length. Variation. Considerable variation occurs among males with respect to the presence of secondary sexual characters on the legs, ranging from no 17) E. spiralis; 18) E. thoracicus. AP = accessory process; LP = left paramere; ML = median lobe; RP = right paramere. modifications to possessing the full spectrum of modifications present within the species (Fig. 19). These characters are variable within populations, sometimes among individuals from the same collection event and apparently not linked with differences in locality or habitat. No differences in genitalia correlated with these variations were observed. Within the genus, the eyes of E. dybasi females are larger and more convex. The genitalia of females vary in the degree of sclerotization of the internal y-shaped process, possibly as an artifact of clearing and preparation. Bionomics. Biological information on E. dybasi in the literature is limited to locality data in the original description (Park 1956) and a study of leaf litter Coleoptera of Great Smoky Mountains National Park (Ferro et al. 2012 b). The following information was obtained from data on collection labels of 99 specimens. Seasonality: Eutyphlus dybasi has been collected during late spring through winter (earliest date 21 March; latest 18 December). Elevation: The majority of individuals were collected at 620 – 1,800 m elevation. However, several individuals were collected from an elevation of 2,750 m. Habitat / Microhabitat: Substrates at collection sites were mixed forest and hardwood litter. Specific woody plant cover mentioned included oak, Rhododendron, tulip tree, and alder. Collecting methods: Most specimens were collected by litter sifting and extraction using a Berlese funnel. However, E. dybasi was the only species of Eutyphlus collected by both Malaise traps and pitfall traps.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	distribution	Distribution. Eutyphlus dybasi is known from Georgia (Union County), North Carolina (Macon and Swain Counties), and Tennessee (Blount, Cocke, and Sevier Counties) (Fig. 28). 2. Eutyphlus prominens Casey, 1894 (Figs. 5, 6, 14, 20 – 25, 29)	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	diagnosis	Diagnosis. Eutyphlus prominens may be distinguished from all known Eutyphlus species by the combination of a lateral excavation and vertical 30) E. schmitti; 31) E. similis; 32) E. spiralis; 33) E. thoracicus. lamina on the third male ventrite and an excavated fourth male ventrite (Figs. 20 – 25). The aedeagus bears a large, membranous projection associated broadly with the fimbriated median lobe and three distal setae on the single ventrally located paramere (Fig. 14).	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	description	Redescription. MALE. Measurements: HL 0.20 mm, HW 0.25 mm; PL 0.30 mm, PW 0.29 mm; EL 0.40 mm, EW 0.42 mm; A 1 – 6 respectively 0.02, 0.14, 0.11, 0.10, 0.10, 0.02 mm. An 1 – 11 respectively 0.08, 0.06, 0.03, 0.03, 0.02, 0.02, 0.02, 0.03, 0.04, 0.04, 0.11 mm. MP 1 – 4 respectively 0.01, 0.05, 0.01, 0.05 mm. GL 0.06 mm, GW 0.04 mm. ML 1.53 mm. Integument: Typical for genus. Head: Slightly narrower than pronotum. Eyes present and well-developed, bearing 25 facets, convex, longer than wide, ocular canthus weakly developed, projecting laterally and sloping gently to gular area. Ventral surface of head slightly rounded in gular area, gular sulcus developed anteriorly and projecting to margin of posteriorly located, paired, gular foveae. Thorax: Pronotum with margin slightly crenulate, median longitudinal sulcus present, area behind basal sulcus bearing small, median longitudinal carina that extends into and bisects sulcus. Prosternum bearing anteroprosternal foveae. Paired, median mesoventral foveae present. Metaventrite convex. Elytron bearing 2 closely approximate basal foveae, as well as subhumeral foveae. Flight wings well-developed. Abdomen: First visible abdominal tergite with oval depression wide (at least ¾ segment length); other tergites unmodified. Second ventrite with longitudinal carina interrupted medially. Third ventrite modified with shelf-like excavation laterally, setose lamina produced vertically in median of excavation, sclerotized shelf produced over excavation bearing fringe of setae projecting laterally. Fourth ventrite excavated in lateral 1 / 3, excavation unmodified. Seventh ventrite small and strap-like. Other ventrites unmodified. Genitalia: Aedeagus complex and asymmetrical. Left paramere more dorsal, comprising an oblique, sclerotized bar. Associated process immediately beneath, sclerotized, bearing 2 erect setae. Median lobe modified into internal, fimbriated process associated with extensive, membranous projection. Right paramere more ventral, bearing 3 distal setae. FEMALE. Measurements: HL 0.15 mm, HW 0.25 mm; PL 0.28 mm, PW 0.28 mm; EL 0.40 mm, EW 0.41 mm; A 1 – 6 respectively 0.03, 0.15, 0.11, 0.10, 0.10, 0.02 mm. An 1 – 11 respectively 0.08, 0.05, 0.03, 0.02, 0.02, 0.02, 0.02, 0.03, 0.03, 0.04, 0.11 mm. MP 1 – 4 respectively 0.01, 0.05, 0.01, 0.05 mm. ML 1.32 mm. Integument, head, thorax, and abdomen: Similar to male except eyes greatly reduced to 2 facets, ocular canthus well-developed. Tempora rounded and broad. Pronotum more quadrate. Third and fourth ventrites unmodified. Seventh ventrite small and transverse. Genitalia: Membranous, completely lacking sclerotization. Produced into voluminous median structure and billowy right sac. Variation. The size and shape of the eyes vary in male E. prominens, and the elytra vary from being bi- to trifoveate. Major intraspecific variation occurs in male secondary sexual characters. Specifically, the sclerotized shelf of the third ventrite that overhangs the lateral excavation on this segment ranges from a small, crescent-shaped projection to an expanded shelf that extends posteriorly over part of the excavation of the fourth ventrite. In addition, the vertical lamina of the excavation on the third ventrite and the degree of excavation of the fourth ventrite are produced to varying degrees. Examination of 27 males, including at least one male from each of the 22 localities, resulted in identification of six distinct “ types ” of ventrite modifications in E. prominens (Figs. 20 – 25), with a range of more subtle variations observed within each type. Type II (Fig. 21) was observed most frequently, with 16 of the 27 males possessing characters that fell into this category. When multiple individuals were observed from a single locality, most exhibited the same form. However, two specimens collected from the same locality in Caldwell County, North Carolina were found to exhibit different forms. Slight variations were observed in the aedeagus of E. prominens. The number of setae on the ventral paramere varies from three to four. Most possess three, but some individuals have a fourth offset slightly from the three main setae that are shorter and narrower in form. The degree of extension of the inflated membranous projection varies among individuals. This process is a component of the median lobe of the aedeagus and likely functions as the intromittent organ during mating. The apparent differences in inflation may be related to specimen preparation. Female E. prominens specimens exhibit variations in eye facet number and the shape of the ocular canthus, but are otherwise similar. Bionomics. No previously published information exists dealing with the biology of E. prominens. Seasonality: Specimens were collected during early spring and early winter (earliest date 6 February; latest 11 November). The two-month gap likely reflects lack of collecting events. Elevation: Most specimens were collected from elevations between 430 and 1,520 m. Habitat / Microhabitat: Most specimens were associated with heavily decayed and rotting wood, either in litter near wood or in wood. Collecting methods: Most specimens were collected using sifting and Berlese funnel extraction, with a small number obtained using flight intercept traps.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	distribution	Distribution. Eutyphlus prominens is known from Georgia (Rabun County), North Carolina (Buncombe, Caldwell, Haywood, Macon, and Yancy Counties), South Carolina (Oconee County), Tennessee (Blount, Cocke, Sevier, Unicoi Counties), Virginia (Smythe County), and West Virginia (Pocahontas County) (Fig. 29). 3. Eutyphlus schmitti Raffray, 1904 (Figs. 7, 8, 15, 27, 30)	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	diagnosis	Diagnosis. Eutyphlus schmitti differs from all known Eutyphlus species by the structure of the male ventrites. The apical margin of the second ventrite is excavated, the third is excavated laterally with vertical lamina and median plates that overlap the fourth, and the fourth and fifth are excavated laterally (Fig. 27). The aedeagus bears a process on either side of the parameres, and the median lobe is modified into a fimbriated, sclerotized process (Fig. 15).	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	description	Redescription. MALE. Measurements: HL 0.21 mm, HW 0.25 mm; PL 0.27 mm, PW 0.26 mm; EL 0.43 mm, EW 0.51 mm; A 1 – 6 respectively 0.04, 0.11, 0.11, 0.11, 0.09, 0.03 mm. An 1 – 11 respectively 0.07, 0.05, 0.04, 0.03, 0.02, 0.02, 0.02, 0.04, 0.04, 0.05, 0.14 mm. MP 1 – 4 respectively 0.01, 0.05, 0.01, 0.05 mm. GL 0.08 mm, GW 0.05 mm. ML 1.44 mm. Integument: Typical for genus. Capitate genal setae sparse. Head: About as wide as pronotum. Eyes small, with 12 facets, weakly convex, longer than wide, ocular canthus weakly developed, projecting laterally and sloping gently to gula. Ventral surface of head rounded at gula, gular sulcus developed weakly anteriorly and projecting to margin of posteriorly located, paired, gular foveae. Thorax: Pronotum with lateral margin crenulate, median longitudinal sulcus present, area behind basal sulcus bearing small, median longitudinal carina that extends into and bisects sulcus. Prosternum bearing anteroprosternal foveae. Paired, median mesoventral fovea present. Mesotrochanters bearing single, median spine. Metaventrite concave baso-medially. Elytra bearing 2, closely approximate basal foveae, as well as subhumeral and sutural foveae. Flight wings well-developed. Abdomen: Tergites unmodified. Second ventrite with apical margin excavated laterally to receive vertical lamina of 3 rd ventrite. Third ventrite excavate laterally, bearing lamina with numerous setae, median portion of sclerites with sclerotized plates bearing fringes of setae that extend over excavation and slightly over margin of 4 th ventrite. Fourth ventrite excavated laterally, middle portion of sclerites contiguous with 2 lateral, sclerotized projections that overhang excavated portion of 5 th ventrite. Fifth ventrite laterally excavated to about ⅔ length of segment. Seventh ventrite small and elongate teardrop-shaped. Other ventrites unmodified. Genitalia: Aedeagus asymmetrical. Left paramere more dorsal, formed as oblique, sclerotized bar with numerous small setae and a large seta about middle, distal end flattened and slightly twisted to expose ventral face. Process associated with paramere branching ventrally to left, similar in form to paramere. Median lobe comprising internal, sclerotized process bearing 2 distinct arms, left arm bearing numerous spicules and spines and right arm more membranous, bearing longer projections. Process associated with right paramere sclerotized, slightly narrowed and curved towards distal end. Right paramere more ventral, flattened and bearing 3 enlarged, thickened distal setae. FEMALE. Measurements: HL 0.20 mm, HW 0.30 mm; PL 0.30 mm, PW 0.30 mm; EL 0.41 mm, EW 0.50 mm; A 1 – 6 respectively 0.03, 0.14, 0.14, 0.13, 0.12, 0.05 mm. An 1 – 11 respectively 0.07, 0.06, 0.02, 0.01, 0.01, 0.01, 0.02, 0.02, 0.04, 0.03, 0.10 mm. MP 1 – 4 respectively 0.01, 0.04, 0.01, 0.04 mm. MP 1.73 mm. Integument, head, thorax, and abdomen: Similar to male except eyes greatly reduced, comprising 2 facets, ocular canthus well-developed. Tempora rounded and broad. Pronotal margin barely crenulate, basal sulcus on pronotum straight between basal foveae. Lacking abdominal secondary sexual characters. Seventh ventrite small and transverse. Genitalia: Membranous, completely lacking sclerotization. Produced into slightly elongate median structure and billowy right sac. Variation. Slight variation was observed in the shape and size of the sclerotized plates and excavated portions of the male ventrites (Fig. 27). This variation was less extreme than those noted for E. prominens. Among females, the number of eye facets varied 1 – 6. Length of the hind margin of the prothorax posterior to the basal sulcus was slightly different among individuals. The depth and angle of this sulcus was slightly variable as well, but never attaining the same angle as in E. prominens or E. similis. Bionomics. Seasonality: E. schmitti was collected from late spring through late fall / early winter (earliest date 16 May; latest 11 November). Based on the data available, occurrence of adults in this species is somewhat more seasonal than it is for other species, though this may be an artifact of sampling. Elevation: Two elevations were included in the label data from the available material: 274 m and 914 m, somewhat lower than most other species in the genus, but the data is limited. Habitat / Microhabitat: Specimens were collected from forest hardwood leaf litter, Rhododendron litter, beech litter, oak litter, and birch litter. Specimens were also found in Sphagnum moss. This species was represented in the study of an old growth forest habitat (“ The Bowl ”) in New Hampshire (Chandler 1987). Collecting methods: Sifting and processing by Berlese funnels was the only collection method mentioned for this species.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	distribution	Distribution. Eutyphlus schmitti is known from New Hampshire (Carroll, Coos, and Grafton Counties), New York (Cattaraugus County), Ohio (Highland County), Pennsylvania (Elk, Huntington, Jefferson, McKean, Somerset, and Westmoreland Counties), and West Virginia (Pocahontas County) (Fig. 30). 4. Eutyphlus similis LeConte, 1880 (Figs. 1, 2, 9, 10, 16, 31)	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	diagnosis	Diagnosis. Eutyphlus similis differs from all known Eutyphlus species in the form of the seventh ventrite of males (small and strap-like) (Fig. 2, VII), the absence of sexual modifications on ventrites of the males, and absence of flight wings in both sexes. The aedeagus is broadly egg-shaped with a rounded internal membrane associated with the distinctly forked, spinose median lobe, and the more ventral paramere bears four distal setae (Fig. 16).	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	description	Redescription. MALE. Measurements: HL 0.24 mm, HW 0.28 mm; PL 0.31 mm, PW 0.30 mm; EL 0.41 mm, EW 0.50 mm; A 1 – 6 respectively 0.03, 0.15, 0.12, 0.10, 0.10, 0.03 mm. An 1 – 11 respectively 0.08, 0.05, 0.03, 0.03, 0.02, 0.02, 0.02, 0.03, 0.03, 0.04, 0.11 mm. MP 1 – 4 respectively 0.01, 0.05, 0.01, 0.05 mm. GL 0.08 mm long, GW 0.05 mm. ML 1.60 mm. Integument: Typical for genus. Capitate genal setae sparse. Head: About as wide as pronotum. Eyes small, with 12 facets, weakly convex, longer than wide, ocular canthus weakly developed, projecting laterally, and sloping gently to gular area. Ventral surface of head rounded in gular area, gular sulcus developed anteriorly and projecting to margin of posteriorly located, paired, gular foveae. Thorax: Pronotum with lateral margin barely crenulate, if at all. Median longitudinal sulcus present. Area behind basal sulcus bearing small, median longitudinal carina that extends into and bisects sulcus. Prosternum bearing anteroprosternal foveae. Paired, median mesoventral fovea present. Metaventrite strongly convex, more prominently so in apical ¼. Elytra bearing 2 basal foveae, subhumeral foveae present. Flight wings absent. Abdomen: Tergites unmodified. Fifth ventrite with lateral sides flattened, quadrate. Seventh ventrite small and strap-like. Other ventrites unmodified. Genitalia: Aedeagus asymmetrical. Left paramere dorsally attached, forming an oblique, sclerotized bar with distal end curved and slightly expanded, bearing 2 large, erect setae. Median lobe modified into internal, sclerotized process bearing 2 distinct arms with numerous spicules and spines, associated with inflated, membranous internal sac. Right paramere flattened, ventrally associated with phallobase and bearing four distal setae. FEMALE. Measurements: HL 0.20 mm, HW 0.30 mm; PL 0.33 mm, PW 0.32 mm; EL 0.41 mm, EW 0.50 mm; A 1 – 6 respectively 0.04, 0.15, 0.12, 0.11, 0.11, 0.03 mm. An 1 – 11 respectively 0.08, 0.06, 0.03, 0.02, 0.02, 0.02, 0.02, 0.03, 0.04, 0.04, 0.12 mm. MP 1 – 4 respectively 0.01, 0.05, 0.01, 0.05 mm. ML 1.74 mm. Integument, head, thorax, and abdomen: Similar to male except eyes greatly reduced, comprising a single facet (this exemplar; see variation comments below), ocular canthus well-developed. Tempora rounded and broad. Pronotum more quadrate. Metasternum more flattened and evenly rounded. Fifth ventrite not flattened laterally as in males. Seventh ventrite small and transverse. Genitalia: Membranous, completely lacking sclerotization. Produced into voluminous median membrane and billowy right sac. Variations. The median longitudinal and antebasal pronotal sulci vary from relatively shallow to well-defined. The aedeagus varies slightly among individuals. The number of setae on the ventral paramere varies from three to six, with four the most common number. When more are present, they are shorter and / or narrower in form and set off slightly from the four main setae. The median membranous projection gives the aedeagus of E. similis its characteristic “ eggshape. ” It appears to vary in degree of inflation, but this may be an artifact of preparation. Number of eye facets in females varies greatly from one to over 20, and the ocular canthus covaries in development.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	discussion	Comments. Several collecting events produced over 40 females and fewer than 10 males. Other samples included only females, making the accurate identification of females important. However, the process of distinguishing E. similis females from the females of E. prominens is problematic and complicates efforts to determine accurate species placement from sites producing only females. In his revision, Park (1956) suggested differences in head and pronotal widths as a reliable way to discriminate between females of the two species. To test this and to investigate other possible characters useful for identification, we measured body parts for 111 E. similis females and 13 E. prominens females. A single individual was taken from each locality represented in this study. Eutyphlus similis females exhibited a slightly wider range of sizes, but the two species overlapped broadly in these measurements (Table 3). This overlap suggests that they cannot be distinguished based on visible characters, and association with males is the only way to identify females. Bionomics. No previously published information on the biology of E. similis exists. Seasonality: The majority of E. similis specimens were collected during April, May, June, and July (earliest date 19 February; latest 6 December). Adults are probably present year-round, and at least one sample from snow- and ice-covered litter yielded specimens. Elevation: Specimens were collected from elevations between 460 m and 2,024 m. Habitat / Microhabitat: Specimens were collected from a wide range of forest litter habitats, including rotted leaves, oak litter, beech litter, Rhododendron, old-growth and non-old growth hardwood litter, and conifer litter (pine, spruce, and fir). The occurrence in conifer litter differs from the habitats that the other species are known from, but this may simply be due to the much larger sample size (approximately 700) for this species. This was the only Eutyphlus species collected from tree hole debris, gilled fungi, and cave litter. In a study of the Coleoptera community of the woody debris of Great Smoky Mountains National Park, a single male specimen of E. similis was collected from a dead-wood emergence chamber (Ferro et al. 2012 a). Collecting methods: Most individuals were collected by hand-collecting or sifting the substrate and then processing the material in a Berlese funnel. Some individuals were collected via flight intercept traps.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFAA5809FE9C3FB9FEE1FCD0.taxon	distribution	Distribution. This species is known from Arkansas (Washington County), Georgia (Dade and White Counties), Kentucky (Edmonson County and Mammoth Cave), Maryland (Calvary County), North Carolina (Avery, Buncombe, Caldwell, Haywood, Mecklenburg, Swain, Transylvania, Watauga, Wilkes, and Yancy Counties), Ohio (Fairfield and Lawrence Counties), Pennsylvania (Somerset and Westmoreland Counties), South Carolina (Greenville and Pickens Counties), Tennessee (Blount, Carter, Cocke, and Sevier Counties), Virginia (Giles, Madison, Page, Smythe, and Washington Counties) and West Virginia (Fayette, Mercer, and Pocahontas Counties) (Fig. 31). The single Illinois record is a checklist record (Chandler 1997) and requires confirmation. The Washington Co., Arkansas record is unusual in that it is the only known locality west of the Mississippi River where any Eutyphlus specimens have been collected. Only E. similis females (16 individuals) are known from this locality, so the identification is considered provisional until additional specimens, including males, are found to confirm the identity of the species that lives in this disjunct area.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB5580EFEA83A29FB2DF991.taxon	materials_examined	Material Examined. (n = 99) * USA Tenn. Cocke Co. GSMNP Cosly Creek trail / Lot # 76 - 105 Oct. 15, 1976 Berlesate R. Chenowith & R. T. Allen (LSAM) (slide mounted M) (description of male based on this specimen). * USA: TN: Cocke Co. GSMNP Snake Den Ridge Trail btw 299077 3957780 & 299657 3957885 CWright June 2005 (LSAM) (slide mounted F) (description of female based on this specimen). * # 5 / SMOKIES, TENN DYBAS 1953 / male symbol / Field Mus. Nat. Hist Orlando Park Pselaphidae Colln / (orange paper) HOLOTYPE / (orange paper) Eutyphlus dybasi Park (FMNH) (1 M; HOLOTYPE). * Brasstown Bald, GA. Union Co. 8 - IX- 63 El. 2750 ′ / Forest floor debris nr. Rotten wood / H. R. Steeves, Jr. J. D. Patrick, Jr. Collectors / H. R. Steeves, Jr. Collection (FMNH) (7 F; 6 M). * USA: NC. Macon Co. 10 mi SW Franklin, Back country info center, v- 22 - 1991, berlese oak & Rhododendron, S. O’ Keefe (DSC) (3 F; 4 M). * N CAROLINA: Swain Co. GSMNP, App. Tr., Buckeye Gap. 35 ° 43.6 ′ N 83 ° 36.1 ′ W 1480 m. Forest litter. 6 June 2005. A. K. Tishechkin (LSAM) (1 F). * U. S. A., NC, Swain Co. GSMNP, Beech Gap Trail 83 ° 12 ′ 42 ″ W, 35 ° 37 ′ 39 ″ N elv. 1000 m, 20 Oct. 2001, berlese Carlton, Cline (LSAM) (3 F; 2 M). * N CAROLINA: Swain Co. GSMNP, Jenkins Ridge Tr. ∼ 1 mi from Appalachian Tr. 35 ° 33.75 ′ N 83 ° 43.2 ′ W 1540 m. Forest litter. 30 Jul; 2004. A. K. Tishechkin (LSAM) (2 F). * NORTH CAROLINA: Swain Co., GSMNP, Lakeshore Tr. 0.5 mi above Forney Creek 15 April 2004. Tulip tree, oak. A. J. Mayor collr. (LSAM) (1 M). * N CAROLINA: Swain Co. GSMNP, nr. Pecks Corner Shelter. 35 ° 39.1 ′ N 83 ° 18.5 ′ W 1620 m. Forest litter. 13 Jun 2005. A. K. Tishechkin (LSAM) (1 M). * N CAROLINA: Swain Co GSMNP, Twentymile Tr. Proctor Field Gap. 35 ° 29.1 ′ N 83 ° 50.3 ′ W. 740 m. Forest litter. 9 May 2004. W. Merritt (LSAM) (1 M). * N CAROLINA: Swain Co, GSMNP, upper Balsam Mt. Tr. ∼ 2 km from App. Tr. @ 35 ° 41.59 ′ N 83 ° 14.59 ′ W 1800 m Forest litter. 13 Jun 2005. A. K. Tishechkin (LSAM) (1 M). * USA: NC: Swain Co. GSMNP Upper Deep Creek Trail UTM 280352 E 3942040 N moist upland berlese 22 July 2002 CEC (LSAM) (1 M). * USA: NC / TN boundary; GSMNP AT btw. Silers; Bald & Miry Ridge Tr.; 265375, 3938911 1570 m; 6 Jun 2005 A. Tishechkin (LSAM) (1 F). * TENNESSEE: Blount Co., GSMNP, App. Tr. at Beechnut Gap. 35 ° 34.16 ′ N 83 ° 41.51 ′ W 1510 m. Forest litter sifting 13 April 2006. A. K. Tishechkin (LSAM) (1 F; 1 M). * U. S. A., TN, Blount Co., GSMNP, Gregory Ridge Trail 83 ° 50 ′ 6 ″ W, 35 ° 32 ′ 25 ″ N / elv. 900 m, forest litter berlese 22 March 2002 CECarlton, VLMoseley (LSAM) (2 F). * TENNESSEE: Blount Co. GSMNP, Indian Grave Gap Tr. @ 35 ° 37.6 ′ N 83 ° 47.4 ′ W 1080 m. Forest litter 3 June 2005. R. Ward (LSAM) (1 F). * TENNESSEE: Blount Co., GSMNP, upper Gregory Ridge Tr. At 35 ° 31.61 ′ N 83 ° 51.18 ′ W. 1315 m. For. Litter 12 April 2006. A. K. Tishechkin (LSAM) (2 F; 1 M). * TENNESSEE: Blount Co., GSMNP, upper Long Hungry Ridge Tr. At 35 ° 30.89 N 83 ° 51.00 ′ W. 1390 m. For. Litter 12 April 2006. A. K. Tishechkin (LSAM) (1 F; 2 M). * TN: Blount Co., GSMNP, White Oak Sink, 35 ° 38 ′ 07 ″ N 83 ° 44 ′ 49 ″ W, 620 m, mixed forest, litter hand sifting Sokolov I. M. 16.06.2006 (LSAM) (3 F; 1 M). * TENNESSEE: Cocke Co., GSMNP, Albright Grove Loop Tr. 980 m. Forest litter sifting. 13 April; 2006. C. E. Carlton (LSAM) (1 M). * U. S. A., TN, Cocke Co. GSMNP, Albright Grove Trail 83 ° 16 ′ 45 ″ W, 35 ° 44 ′ 10 ″ N / elv. 1000 m, decid. Berlese, 19 October 2001. C. Carlton, A. Cline, A. Tishechkin (LSAM) (8 F; 14 M). * USA Tenn. Cocke Co. GSMNP Cosly Creek Trail / Lot # 76 - 105 Oct. 15, 1976 Berlesate R. Chenowith & R. T. Allen (LSAM) (1 F; 7 M). * USA: TN: Cocke Co. GSMNP Snake Den Ridge Trail / btw 299077 3957780 & 299657 3957885 Cwright June 2005 (LSAM) (2 F). * U. S. A., TN, Sevier Co. Appalachian Trail at Beech Gap on Clingman’ s Dome Rd 83 ° 26 ′ 50 ″ W, 35 ° 36 ′ 36 ″ N / elv. 1750 m, forest litter berlese 28 June 2001, C. Carlton, A. Tishechkin, V. Moseley (LSAM) (2 F; 3 M). * TN: Sevier Co. GSMNP ATBI Plot. Goshen Prong Pitfall 106 83 32 34 35 36 38 Parker, Stocks, Peterson 28 Nov – 11 Dec 2000 (DSC) (1 F). * TENNESSEE: Sevier Co.; GSMNP, Goshen Prong; ATBI Plot. Malaise trap; MT- 2120022218. 5 – 18 Dec; 2001. C. R. Parker (LSAM) (2 M). * TENNESSEE: Sevier Co. GSMNP, Indian Head Tr. 35.60944 ° N 83.44659 ° W Sift litter. 5290 ′ 20 July 2003. S. O’ Keefe (LSAM) (1 F). * USA Tenn. - N. Car. Sevier Co, GSMNP, Newfound Gap to Clingman’ s Dome / Lot # 76 - 107 Oct. 11, 1976 Berlesate; R. Chenewith & R. T. Allen (LSAM) (1 F). * U. S. A., TN, Sevier Co., GSMNP Ramsey Cascade Trail (middle) 83 ° 18 ′ 24 ″ W, 35 ° 42 ′ 34 ″ N / elv. 1300 m, forest litter berlese 21 March 2002 CECarlton, VLMoseley (LSAM) (1 F). * TENNESSEE: Sevier Co., GSMNP, Road Prong Tr. at 35 ° 36 ′ 36 ″ N 83 ° 27 ′ 3 ″ W 1580 m. Leaf / moss mat’ litter. 20 July 2003; A. Tishechkin (LSAM) (1 F; 1 M). * USA: TN: Sevier Co. 10 mi S Gatlinburg 3800, v- 24 - 1977 Anewton, Mthayer / berlese alder & rhododendron litter by stream (DSC) (1 F; 2 M). * TENNESSEE: Sevier Co., GSMNP, Twin Creek ATBI Plot. Malaise Trap MT- 0120011205. 5 Nov – 5 Dec 2001. I. C. Stocks (LSAM) (1 F; 2 M).	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFA6581FFCA93BEFFE38FA60.taxon	description	(Figs. 1, 2) Redescription. Possessing characters of the tribe Trichonychini (sensu Reitter 1882) (Figs. 1, 2). MALE. Integument: Body and appendages uniformly light brown, evenly covered by sparse, short, suberect pubescence. Capitate genal setae present. Head: Vertexal foveae present, large, setose, located posterior to shallow, U-shaped vertexal sulcus. Antennal acetabula remote, antennomeres 1 – 10 unmodified, antennomere 11 with lateral, shelf-like excavation bearing dense cluster of setae. Labrum wider than long, slightly expanded distally, apical margin gently concave, bearing Character Number 00000000011111111112222222222333333333344444444445555 Taxa 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 Arthmius bulbifer 1 0 0 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 Hendecameros panamense 1 0 0 0 0 1 0 0 0 0 0 1 2 1 0 3 1 0 0 1 0 1 1 1 1 1 1 2 2 0 1 1 1 1 1 1 1 1 1 0 0 0 1 0 0 2 0 0 1 1 0 0 1 Barroeuplectoides sp. 1 1 0 0 0 0 0 1 0 0 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 1 0 0 1 1 0 0 0 0 0 1 Euplectus confluens 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 2 1 0 0 0 0 1 0 0 1 0 0 1 1 0 0 1 Leptoplectus pertenuis 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 1 1 1 0 1 0 1 2 1 0 0 0 0 1 0 0 1 0 0 0 1 0 0 1 Pycnoplectus linearis 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 2 1 0 0 0 0 1 0 0 1 0 0 1 1 0 0 1 Jubus sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 2 1 1 2 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 1 0 0 0 0 0 1 1 1 0 1 1 0 0 1 Sebaga sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 2 1 1 2 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 1 0 0 0 0 0 1 1 0 0 1 1 0 1 1 Mayetia domestica 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 0 0 0 0 1 0 3 1 1 1 1 1 2 2 1 1 1 0 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 1 Metopias sp. 1 0 0 0 0 1 1 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 1 1 2 1 0 0 0 0 0 0 0 1 0 0 0 0 0 0 1 Foveoscapha terracola 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 1 0 1 1 0 0 1 1 0 1 1 Bibloplectus cherokee 1 1 0 3 0 0 0 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 0 0 0 1 0 0 0 1 0 0 1 Eutyphlus dybasi 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 0 1 1 0 0 0 1 1 0 1 Eutyphlus spiralis 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 0 1 1 0 0 0 1 1 0 1 Eutyphlus prominens 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 1 0 1 0 0 0 1 1 0 1 Eutyphlus schmitti 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 1 0 1 0 0 0 1 1 0 1 Eutyphlus similis 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 0 1 1 0 0 0 1 1 0 1 Eutyphlus thoracicus 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 0 1 0 0 0 1 1 0 1 Pseudactium carolinae 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 0 1 1 0 0 1 1 0 0 1 Rhinoscepsis bistriatus 1 1 0 0 0 1 1 0 0 0 0 0 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 1 1 0 0 1 0 1 1 1 0 1 1 0 0 1 Thesiastes sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 1 0 0 1 0 0 1 1 0 0 1 Thesium sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 1 1 0 0 1 0 1 1 0 0 1 1 0 0 1 Trimioplectus australis 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 1 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 0 0 1 0 0 0 0 1 0 1 1 Actiastes globiferum 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 0 0 0 1 0 0 1 1 1 1 1 Dalmosanus quercavum 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 0 0 1 0 0 0 0 1 0 0 1 Dalmosella tenuis 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 0 1 0 0 2 1 0 0 1 Hanfordia sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 1 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 Melba clypeata 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 1 0 0 0 0 1 0 0 0 Trimiomelba dubia 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 Trimium brevicorne 1 1 0 0 0 0 0 0 0 1 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 1 0 0 0 0 1 0 0 0 Quadrelba sp. 1 1 0 0 0 0 0 0 0 0 0 1 0 1 0 1 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 Conoplectus acornus 1 1 0 0 0 0 0 0 0 2 0 1 0 1 0 1 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 0 1 0 0 0 0 1 0 1 1 0 0 0 1 0 1 1 Rhexius sp. 1 1 0 0 0 0 0 0 0 2 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 1 1 0 0 0 2 1 0 1 1 1 0 2 1 0 1 1 Sonoma baylessae 1 0 1 3 1 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0 0 1 1 1 1 1 1 1 2 0 1 1 0 1 3 1 1 1 2 2 0 0 0 0 0 0 0 0 0 0 0 0 1 Reichenbachia howardi 1 0 0 3 0 0 0 0 0 0 0 1 0 1 0 3 1 0 0 1 0 1 1 1 1 1 1 1 2 0 1 1 0 1 0 1 2 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 Pselaptrichus perfidus 1 1 0 0 0 1 1 0 1 0 1 1 0 1 0 0 1 0 0 1 0 2 4 1 4 1 1 1 3 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 Lucifotychus cognatus 1 1 0 0 0 1 1 0 0 0 0 1 0 1 0 0 1 0 0 2 0 2 1 2 1 2 3 2 3 1 1 1 0 1 0 1 2 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 1 Character Number 55555566666666667777777777888888888899999999991111111 Taxa 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 Arthmius bulbifer 0 0 1 1 1 0 0 1 0 1 1 1 1 0 0 0 0 0 1 0 0 0 1 1 0 1 0 0 0 0 1 1 0 1 0 0 0 1 1 1 0 0 0 0 0 3 0 0 0 0 0 1 1 Hendecameros panamense 0 0 1 0 1 0 1 1 0 1 1 1 1 1 0 0 1 0 0 0 1 1 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 0 1 0 0 0 0 0 0 1 0 0 1 0 0 2 1 Barroeuplectoides sp. 1 0 0 1 1 0 1 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 1 0 0 1 0 0 0 0 0 0 0 4 0 0 1 0 0 2 1 Euplectus confluens 1 0 0 2 1 0 0 1 0 0 1 1 1 1 0 1 0 0 1 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 1 1 0 1 0 1 4 0 0 0 0 0 1 1 Leptoplectus pertenuis 1 0 0 3 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 1 0 0 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 4 0 0 0 0 0 1 1 Pycnoplectus linearis 0 0 0 1 1 0 0 1 0 0 1 1 1 1 0 1 0 0 1 0 0 0 1 1 0 0 0 0 1 0 0 1 0 0 0 0 0 0 1 1 0 0 1 1 1 4 0 0 0 0 0 1 1 Jubus sp. 0 0 0 3 2 0 0 1 0 0 1 1 1 1 0 0 1 0 0 0 0 0 1 0 0 0 0 0 1 1 1 2 0 0 1 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Sebaga sp. 0 0 0 3 2 0 0 1 0 0 1 1 2 1 0 0 1 0 0 0 0 0 1 0 0 0 0 0 1 1 1 2 0 0 1 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Mayetia domestica 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1 1 0 0 0 0 0 1 1 0 2 0 1 0 0 0 0 0 0 0 0 0 0 0 4 0 0 1 0 0 2 1 Metopias sp. 0 0 0 0 1 0 0 1 1 1 1 1 1 0 0 1 0 0 0 0 1 0 1 1 1 0 0 0 0 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 Foveoscapha terracola 0 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 1 1 Bibloplectus cherokee 0 0 0 0 1 0 0 1 0 1 1 1 1 1 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 1 1 0 0 0 0 0 1 0 2 0 0 0 0 0 0 0 0 0 0 0 1 1 Eutyphlus dybasi 0 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 1 1 1 0 0 0 1 1 1 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Eutyphlus spiralis 0 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 1 1 0 0 0 0 1 1 1 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Eutyphlus prominens 1 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 1 0 0 1 0 1 1 1 1 0 1 3 0 0 0 0 0 1 1 Eutyphlus schmitti 1 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 1 0 0 1 0 1 1 0 1 1 1 3 0 0 0 0 0 1 1 Eutyphlus similis 1 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 1 0 0 0 0 0 3 0 0 0 0 0 1 1 Eutyphlus thoracicus 1 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 1 0 0 1 0 1 0 0 0 0 0 3 0 0 0 0 0 1 1 Pseudactium carolinae 0 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 1 0 1 0 0 0 1 1 1 0 0 0 1 1 1 2 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Rhinoscepsis bistriatus 0 0 0 0 1 0 0 1 1 1 1 1 2 1 0 2 0 0 1 0 0 0 1 1 0 0 0 0 1 1 1 2 0 0 1 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Thesiastes sp. 0 0 0 1 1 0 0 1 0 1 1 1 1 1 0 1 1 0 1 0 0 0 1 1 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0 1 0 0 1 0 1 0 0 0 0 0 0 1 1 Thesium sp. 1 0 0 3 1 0 1 1 0 1 1 1 1 1 0 1 1 2 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 Trimioplectus australis 0 0 0 2 2 0 0 1 0 0 1 1 2 0 0 1 1 0 1 0 0 0 1 1 0 1 1 0 0 0 1 1 0 0 0 0 0 1 0 0 1 1 1 1 0 0 0 0 0 0 0 1 1 Actiastes globiferum 0 0 0 1 1 0 0 1 0 0 1 1 1 1 0 1 1 0 1 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 1 0 0 1 0 2 0 0 0 0 0 0 0 0 0 0 0 1 1 Dalmosanus quercavum 0 0 0 3 1 0 0 1 0 1 1 1 2 1 0 0 0 0 1 0 0 0 1 0 0 0 0 0 0 1 1 1 0 1 0 0 0 1 0 2 0 0 0 0 1 0 0 0 0 0 0 1 1 Dalmosella tenuis 0 0 0 0 1 0 0 1 0 0 1 1 2 1 0 0 1 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 2 0 0 0 0 0 0 0 0 0 0 0 1 1 Hanfordia sp. 0 0 0 3 2 0 0 0 0 0 1 1 2 0 0 0 0 0 1 0 0 0 1 1 1 1 0 0 1 1 1 2 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 Melba clypeata 0 0 0 0 2 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 0 0 1 1 0 1 0 0 0 1 1 1 0 0 0 0 0 1 0 1 1 1 1 0 1 0 0 0 0 0 0 1 1 Trimiomelba dubia 0 0 0 3 2 0 0 0 0 1 1 1 2 0 0 0 0 0 1 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 Trimium brevicorne 0 0 0 3 2 0 1 1 0 0 1 1 2 0 0 0 1 0 0 0 0 0 1 0 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 Quadrelba sp. 0 0 0 3 2 0 0 1 0 1 1 1 2 1 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Conoplectus acornus 0 0 0 1 2 0 0 1 0 1 1 1 1 1 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Rhexius sp. 0 0 0 1 2 0 0 1 0 0 1 1 1 1 0 1 0 0 0 0 0 0 1 1 0 0 0 0 0 0 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 1 Sonoma baylessae 0 1 0 0 1 1 1 1 1 1 1 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 1 Reichenbachia howardi 0 0 0 3 2 0 0 1 0 1 1 1 1 0 0 2 0 0 2 0 0 0 1 1 0 0 0 0 0 0 1 1 0 1 0 0 0 1 1 1 0 0 0 0 0 3 0 0 0 0 0 0 0 Pselaptrichus perfidus 0 0 0 3 1 0 0 1 0 1 1 1 1 1 0 0 0 1 1 1 0 0 1 0 0 0 0 0 0 1 1 1 0 3 0 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 Lucifotychus cognatus 0 0 0 1 2 0 0 1 0 1 1 1 1 0 0 1 0 1 1 1 0 0 1 0 1 0 0 0 0 0 1 1 0 3 1 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 2 simple sensory pegs, lateral angles rounded. Mandibles sickle-shaped with 6 teeth on incisor edge. Labial palpi small, 2 - segmented. Maxillary palpi unmodified, MP 1 minute and obliquely joined to MP 2, MP 2 pedunculate, narrower in basal 1 / 3, then slightly expanded and rounded distally, MP 3 subtriangular, MP 4 widest, unmodified, bearing apical palpal cone. Paired gular fovea present. Thorax: Prothorax moderately convex. Basal sulcus present, margin delimited by basolateral foveae. Mediobasal foveae absent. Prosternum bearing robust prosternal median carina, procoxal foveae present on either side of carina. Lateral mesoventral and mesocoxal foveae both present. Mesocoxae contiguous. Metacoxae conjunct, not separated by short intercoxal process of ventrite 1. Lateral metaventral foveae present, paired, metaventrite otherwise afoveate. Elytra narrower basally, then gradually wider posteriorly, bearing sutural striae with fovea. Lateroapical cleft present. Abdomen: First visible abdominal tergite bearing laterobasal foveae with weak, oval depression, margined anteriorly. Second tergite bearing similar, weaker depression and foveae. Other tergites unmodified. Second ventrite with basal carina extending between basolateral foveae, continuous. Third ventrite bearing small basolateral foveae. Other ventrites variously modified in some species as male secondary sexual characters. Male genitalia: Asymmetrical, somewhat laterally compressed. Dorsal diaphragm and internal sac visible. Parameres fixed, rotated to dorsal and ventral orientation. Median lobe sclerotized with varying combinations of spines and spicules. FEMALE. Generally more heavy-bodied and dorso-ventrally flattened than males. Ocular canthus more prominent. Eyes usually reduced to absent (with the exception of E. dybasi). Female genitalia: Largely membranous, comprising internalized ventrite 7 (visible as a transverse triangular strap in some specimens) and 2 internal sclerites that often appear sac-like.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB5580FFCAB3D6AFC13FD6B.taxon	materials_examined	Material Examined. (n = 68) * TENNESSEE: Great Smoky Mtns. NP, Newfound Gap. F. I. T. # 2, 26 June – 1 July 2001. 35 ° 36.7 ′ N 83 ° 25.30 ′ W. C. Carlton, V. Moseley, A. Tishechkin (LSAM) (slide mounted M) (description of male based on this specimen). * TENNESSEE: Sevier Co. GSMNP, Trillium gap Tr. @ 35 ° 39.9 ′ N 83 ° 26.2 ′ W 1400 m. Forest litter. 29 July 2004. A. Tishechkin (LSAM) (slide mounted F) (description of female based on this specimen). * Rabun Bald, GA. Rabun Co. El. 3630 30. v. 64 / Forest floor debris nr. Dead wood / H. R. Steeves Jr. Collector / H. R. Steeves Jr. Collection (FMNH) (1 F; 2 M). * Clingmans Peak Buncombe Co., N. C. 3. VII. 60 Rhododendron duff / H. R. Steeves Jr. Collector / H. R. Steeves Jr. Collection / CNHM 1963 H. R. Steeves, Jr. Pselaphidae Colln. Acc. Z- 13, 288 / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (15 F; 5 M). * USA: NC. Caldwell Co. 1 miS Julian Price Mem Park v- 20 - 1991 sift oak litter S. O’ Keefe (DSC) (4 F; 2 M); (FSAC) (3 F). * N CAROLINA: Caldwell Co, Pisgah Natl. For., nr. Mortimer Cmg. 35 ° 59.54 ′ N 81 ° 45.64 ′ W 430 m. Litter sifting. 30 April 2006. A. K. Tishechkinv (LSAM) (1 M). * N. C. / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. / Compared with type specimen O. Park XI. II. 41 (FMNH) (1 F). * NC: Haywood Co. GSMNP. Purchase Knob, 35 ° 35 ′ 05 ″ N 83 ° 03 ′ 45 ″ W, mixed forest, 1520 m, litter hand sifting Sokolov, LM. 22.06.2006 (LSAM) (1 M). * USA: NC: Macon Co. 3 mi NW highlands v- 29 - 1983 DSChandler, sift; Rhododendron duff (DSC) (1 M). * USA: NC, Yancy Co. B. R. Pkw. 37.4 mi. SW intersection US 221 elv. 4600 ′, log litter 2. vi. 1991 C. E. Carlton (LSAM) (1 M). * USA: SC: Oconee Co 7 mi S NC state line on Hwy. 107 / v- 29 - 1983 DSChandler, sift forest litter (DSC) (1 F; 1 M). * TENNESSEE: Blount Co.; GSMNP, lower Gregory Ridge Tr. @ 35 ° 33.5 ′ N; 83 ° 50.5 ′ W. 630 m. For. Litter; 28 July 2004. A. K. Tishechkin (LSAM) (1 M). * USA: TN: Blount Co. GSMNP White Oak Sink: F. I. T / 251399 3946695 920 m 11 Jun 2005 A. Tishechkin (LSAM) (1 M). * U. S. A., TN, Cocke Co. GSMNP, Albright Grove Trail 83 ° 16 ′ 45 ″ W, 35 ° 44 ′ 10 ″ N elv. 1000 m, old growth Berlese 29 June 2001, C. Carlton, A. Tishechkin, V. Moseley (LSAM) (1 M). * USA: TN: Cocke Co., 6 mi SE Cosby v- 31 - 1983 DSChandler, sift forest litter (DSC) (1 F; 4 M). * TENNESSEE: Great Smoky Mtns. NP, Newfound Gap. F. I. T. # 2, 26 June – 1 July 2001. 35 ° 36.7 ′ N 83 ° 25.30 ′ W. C. Carlton, V. Moseley, A. Tishechkin (LSAM) (1 M). * USA: TN: Sevier Co. GSMNP: AT 2 km fr. Derrick Knob Shelter 258686, 3939145, 1440 m Berlese 7 Jun 2005 A. Tishechkin (LSAM) (1 M). * TENNESSEE: Sevier Co. GSMNP, nr. Derrick Knob Shelter. 35 ° 33.87 ′ N 83 ° 38.53 ′ W. 1465 m. beating 6 Jun 2005. S. Kazantsev (LSAM) (3 M). * TENNESSEE: Sevier Co. GSMNP, Porters Creek Tr. Sift litter near rotten logs Beetle Blitz - 2003 18 July 2003. S. O’ Keefe (LSAM) (1 M). * TN: Sevier Co., GSMNP Porter’ s Creek Trail 9617 El. 738 m 18 – 19 June 1996 Coyle, Brooks, Aiken, Davis (LSAM) (1 M). * TENNESSEE: Sevier Co. GSMNP, Trillium gap Tr. @ 35 ° 39.9 ′ N 83 ° 26.2 ′ W 1400 m. Forest litter. 29 July 2004. A. Tishechkin (LSAM) (3 F; 3 M). * USA: TN: Sevier Co. upper miry ridge tr: litter / 264327 3941078 1520 m 6 Jun 2005 A. Tishechkin (LSAM) (1 M). * Unicoi Co. Tenn. 3500 ′ VII 5, 53 / Field Mus. Nat. Hist, Orlando Park Pselaphidae Colln. (FMNH) (3 F; 1 M). * Unicoi Co, Tenn Una (illegible) Mtn. July 5, 1953 / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln (FMNH) (1 F). * USA, VA. Smythe Co. 7.5 mi SE Marion, Racoon; Branch cmpgrnd, v- 10 - 91 sift pine & oak litter S. O’ Keefe collector (DSC) (1 M). * USA: WV. Pocahontas Co. 6 mi SE Marlinton forest trail 407 off Hwy 92, v- 16 - 1991 sift oak S. O’ Keefe (DSC) (1 F).	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB4580FFCC439EDFB11F914.taxon	materials_examined	Material Examined. (n = 31) * Ohio, Highland Co, Mohican St. For. V- 31 - 1976 / LEWatrous Berlese forest litter (DSC) (slide mounted male) description of male based on this specimen. * USA: PA: Hunt. Co. Rothrock St. For. Seeger Nat. Area 900 ′, V- 30 - 1985 / DSChandler, sift Rhododendron leaf litter (DSC) (slide mounted female) (description of female based on this specimen). * USA: NH: Carr. Co., The Bowl, 2,5 mi; NW Wonalancet VII- 17 - 1985 / DSChandler sift birch / beech leaf litter (DSC) (1 M). * Jefferson Notch, N. Hamp., Coos Co. 3000 ′ IX- 20 - 1974 ANewton / Berlese forest litter (FMNH) (6 F). * USA: NH. Grafton Co Waterville valley viii- 30 - 1990; sift litter; S. O’ Keefe (FSAC) (4 F). * NY: Cattaraugus Co. Allegany St. Park VI- 27 - 1971 berlesed deciduous litter W. B. Muchmore coll. (DSC) (1 M). * Ohio, Highland Co. Mohican St. For. V- 31 - 1976 / LEWatrous; berlese forest litter (DSC) (2 M). * USA: PA: Elk Co., Allegheny Nat. For. 3 mi NW Beltown Millstone Creek, VI- 7 - 1997, DSChandler hardwood leaf litt. (DSC) (2 M). * USA: PA: Hunt. Co. Rothrock St. For. Seeger Nat. Area 900 ′, V- 30 - 1985 / DSChandler, sift Rhododendron leaf litter (DSC) (3 F; 2 M). * USA: PA: Jeffer. Co., Cook State Forest, 0.4 mi S Cooksburg; Mohawk Trail, V- 6 - 1997, DSChandler hardwood leaf litt. (DSC) (1 F; 1 M). * USA: PA: Jeffer. Co., Cook State Forest, 0.4 mi S Cooksburg Mohawk Trail, V- 6; - 1997, DSChandler hardwood leaf litt. (DSC) (3 M). * USA: PA: McKean Co, Allegheny N. For. Tionesta Scenic / Area, V- 29 - 1985 DSChandler, sift beech leaf litter (DSC) (3 M). * Penn., Somer Co., 1 mi S Kentner VI- 10 - 1976 / LEWatrous berlese forest litter (DSC) (1 F). * USA: WV. Pocahontas Co. 6 mi SE Marlinton forest trail 407 off Hwy 92, V- 16 - 1991 sift oak, S. O’ Keefe (DSC) (1 M). *	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFBB5803FEA83BDFFECAFD0D.taxon	materials_examined	Material Examined. (n = 737) * TENNESSEE: Sevier Co.; GSMNP, Indian Head Tr.; 35.60944 ° N 83.44659 ° W; Sift litter. 5290 ′; 20 July 2003. S. O’ Keefe / (LSAM) (slide mounted male) (description of male based on this specimen). * VIRGINIA: Page Co., Shennandoah Natl. Park, Little Stony Man at 38 ° 36.24 ′ N 78 ° 21.99 ′ W. 1024 m. For. litter 3 May 2006. A. Tishechkin (LSAM) (slide mounted female) (description of female based on this specimen). * USA, AR, Washington Co. Ozark Nat. Forest Wedington 36 ° 06.312 N 94 ° 23.390 W Litter between rocks 25 Jul. 2010. MJ Skvarla APGD 10 - 0726 - 004 (LSAM) (16 F). * Cloudland Canyon S. Pk. Dade Co., GA. 17. VI. 62 Forest floor debris / H. R. Steeves Jr. collector / H. R. Steeves Jr. Collection / CNHM 1963 H. R. Steeves, Jr. Pselaphidae Colln. Acc. Z- 13, 288 (FMNH) (2 F; 1 M). * USA: GEORGIA, Dade Co., Cloudland Canyon State Pk. 34 ° 48.88 ′ N 85 ° 29.10 ′ W 510 m. 17 Sept. 2006. Forest litter sifting. I. M. Sokolov (LSAM) (6 F). * GA: White Co. 8 August 1984 coll. R. J. Beshear “ Eutyphlus ” / det. DSChandler (GMNH). * KY. Edmonson Co. Mammoth Cave Nat. Park Ugly Creek 8.5 mi NE Rhoda Berlese sample of rotted leaves 22 - NOV- 1992, E. A. Lisowski (LSAM) (4 F). * Mammoth Cave KY. V- 3 - 47 O. Park / Field. Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (2 F). * USA: MD: Calv. Co. 4 mi S Prince Frederick, IV- 16 / V- 7 - 1987, L Masner FIT, forest by crk (DSC) (2 M). * N CAROLINA: Avery Co. Blue Ridge Pkwy., Grandfather Mt. at 36 ° 5.70 ′ N 81 ° 49.62 ′ W 1580 M. Litter sifting 1 May; 2006. A. K. Tishechkin (LSAM) (19 F). * Clingman’ s Peak Buncoabe Co., N. C. 3 - VII- 60 spruce grove debris / H. R. Steeves Jr. Collector / H. R. Steeves Jr. Collection / CNHM 1963 H. R. Steeves, Jr. Pselaphidae Colln. Acc. Z- 13, 288 / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (1 F). * NCBuncoabeCo.; MtPisgah 5600 26 Ap 74 PDebs soiluRocks (LSAM) (2 F). * USANCCALDWELLCO Nr Cragg 36 ° 4 ′ 31 ″ N 81 ° 46 ′ 52 ″ W el 2930 ′ 14 Nov 2009 J & SCornell ExMoss & Rhododendron litter RoadsideDitch (LSAM) (1 F). * Grandfather Mt. N. C. 14. X. 50 T. Daggy / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (1 F). * USA: NC: Haywood GSMNP BRPW nr. Cove Field Ridge Overlook: litter / 35 ° 25.84 ′ N 83 ° 21.15 ′ W’ 1420 m 21 Sep 2005 ATishechkin (LSAM) (5 F). * NCAROLINA: Haywood Co. GSMNP, Caldwell Fork Tr. At UTM 30897 E; 3940883 N. Moist forest Berlese. 3 August 2002. C. Carlton & N. Lowe (LSAM) (6 F; 1 M). * U. S. A, NC Haywood Co. GSMNP, Balsam Mt. Trail 83 ° 10 ′ 57 ″ W, 35 ° 38 ′ 12 ″ N / 20 Oct 2001, A. Tishechkin elv. 1500 m, hdwd. Berlese (LSAM) (3 F). * U. S. A., NC, Haywood Co. GSMNP, Baxter Creek Tr. 83 ° 06 ′ 36 ″ W, 35 ° 43 ′ 24 ″ N / elv. 1350 m. forest litter, Berlese 23 March 2002, C. Carlton V. Moseley / LSAM 0014787 (LSAM) (1 F; 2 M). * USA: NC: Haywood lower Baxter Cr. Tr / 308798 3957165 5 Jun 2005 CWright / LSAM 0092285 (LSAM) (8 F). * USA: NC: Haywood Co. GSMNP Catalooche Divide Tr. Near Purchase / 311819 E 3940339 N 17 Jul 2002 CCarlton (LSAM) (2 F). * USA: NC: Haywood Co. GSMNP Cataloochee Rough Ridge Tr., lower / 306360 E 3940881 N moist berlese 29 Jul 2002 CCarlton (LSAM) (2 F). * U. S. A., NC, Haywood Co. GSMNP, Chestnut Branch Trail 83 ° 07 ′ 24 ″ W, 35 ° 45 ′ 34 ″ N elv. 740 m, leaf litter Berlese 1 August 2001, A. Tishechkin (LSAM) (4 F). * USA: NC: Haywood Co. GSMNP McKee Branch Tr. / 310220 E 3940349 N, most upland berlese 16 Jul 2002 CCarlton (LSAM) (1 F). * USA: NC: Haywood Co.; GSMNP Purchase; Knob, berlese / 313124 E; 3939640 N 5089 ft; 20 July 2002; CCarlton (LSAM) (1 F). * MECKLENBURG Co., N. C. iv- 22 - 57 T. Daggy (NCSUIM) (2 F; 1 M). * NORTH CAROLINA: Swain Co., GSMNP, admin. Road W of Fontana Dam, Lewellyn Branch UTM 0250085 E 392971 N 12 Apr 2004. A. J. Mayor (LSAM) (1 F). * N CAROLINA: Swain Co. GSMNP, Appalachian Tr. ∼ 1.5 mi E Newfound Gap 35 ° 36.96 ′ N 83 ° 24.66 ′ W 1730 m. Forest litter 18 July 2002. C. Carlton (LSAM) (10 F). * USA: NC: Swain Co GSMNP Appalachian Tr. 1.5 mi NE Newfound Gap / 281639 E 3944038 N ridgetop berlese 18 July 2002 CCarlton (LSAM) (3 F; 1 M). * USA: NC: Swain Co. GSMNP Appalachian Tr. at Sweat Heifer Cr. Tr. Ridgetop berlese / 282160 E 3944324 N 18 July 2002 CCarlton (LSAM) (2 F; 1 M). * N CAROLINA: Swain Co. GSMNP, E branch of Balsam Mt. Tr. Nr. Balsam Mt. Rd; 35 ° 37.5 ′ N 83 ° 12.7 ′ W. 980 m Forest litter. 26 July 2004 J. Ciegler & M. Fitzgerald (LSAM) (1 F). * N CAROLINA: Swain Co. GSMNP, W branch of Beach Gap Tr. @ 35 ° 37.3 ′ N; 83 ° 12.8 ′ W. 975 m. Forest litter. 9. vi. 2005. A. Tishechkin (LSAM) (1 F). * U. S. A., NC Swain Co. GSMNP, Clingman’ s Dome 83 ° 29 ′ 50 ″ W, 35 ° 33 ′ 34 ″ N / elv. 2024 m, upper Berlese 30 June 2001, Ccarlton, A. Tishechkin, V. Moseley (LSAM) (3 F). * USA: NC: Swain Co. GSMNP Upper Deep Creek Trail UTM 280352 E 3942040 N moist upland berlese 22 July 2002 CEC / LSAM 0077201 (LSAM) (7 F; 2 M). * USA: NC: Swain Co. GSMNP, Deep Creek area, Loop Tr. at Indian Creek Trail, UTM 280689 E 3928371 N, moist forest berlese C. Carlton 26 July 2002 (LSAM) (1 F). * U. S. A., NC, Swain Co. GSMNP, Flat Creek Trail 83 ° 10 ′ 21 ″ W, 35 ° 31 ′ 1 ″ N / elv. 1500 m, leaf litter Berlese 31 July 2001 A. Tishechkin (LSAM) (1 F). * same locality / LSAM 0002509 (LSAM) (1 F). * same locality / LSAM 0002510 (LSAM) (2 F). * USA: NC: Swain Co. 5.8 mi. S. Heintooga Overlook, Great Smoky Mt. N. P. 1. vi. 1991 C. E. Carlton (LSAM) (1 F). * USA N. Car. Swain Co. Dirt Rd. from Heintooga Overlook to Cherokee / Lot # 76 - 102 Oct. 14, 1976 Berlesate 5000 ′ – 4500 ′ R. Chenowith & R. T. Allen (LSAM) (46 F; 3 M). * N CAROLINA: Swain Co.; GSMNP, Jenkins Ridge Tr. ∼ 1 mi from Appalachian Tr. 35 ° 33.75 ′ N 83 ° 43.2 ′ W 1540 m. Forest litter. 31 Jul; 2004. A. K. Ticheshkin (LSAM) (1 F). * same locality / LSAM 0094941 (LSAM) (1 F). * same locality / LSAM 0094945 (LSAM) (3 F). * N CAROLINA: Swain Co. GSMNP, upper Jenkins Ridge Tr. At Gunna Cr. 35 ° 28.3 ′ N; 83 ° 42.65 ′ W 1400 m. Forest litter. 29 July 2004. A. Tishechkin (LSAM) (1 M). * N CAROLINA: Swain Co. GSMNP, Kephart Prong Tr. at 35 ° 35 ′ 36 ″ N 83 ° 21 ′ 55 ″ W 930 m. Forest litter 20 July 2003. A. Tishechkin (LSAM) (1 F). * N CAROLINA: Swain Co. GSMNP, Lakeshore Tr. at 35 ° 28 ′ 20 ″ N 83 ° 27 ′ 3 ″ W 1580 m. Forest litter 18 July 2003. A. Tishechkin (LSAM) (6 F). * N CAROLINA: Swain Co. GSMNP, Lost Cove Tr. @ 35 ° 29.45 ′ N 83 ° 48.10 ′ W 685 m. Forest litter. 8 May 2004. W. D. Merritt (LSAM) (13 F; 3 M). * N CAROLINA: Swain co. GSMNP, McGee Spring Cmpgr., 5 site. 35 ° 38.31 ′ N 83 ° 14.36 ′ W. 1535 m. Forest litter. 2 Oct 2004. W. Merritt (LSAM) (1 F). * N CAROLINA: Swain Co. GSMNP, ∼ 250 m W Mingus Mill Parkgr. 650 m. 1 m 2 litter 28 Apr 1996. Coyle collr. (LSAM) (2 F). * USA: NC, SwainCo. 2 mi. Neintooga overlook. Beech-fir berlesate 1. vi. 1990 C. E. Carlton (LSAM) (4 F; 1 M). * N CAROLINA: Swain Co. GSMNP, Noland Divide Tr. at 35 ° 33 ′ 57 ″ N 83 ° 28 ′ 36 ″ W 1770 m. Leaf / moss mat litter 19 July 2003. A. Tishechkin (LSAM) (18 F; 4 M). * USANCTRANSYLVANIACONr Brevard PisgahNFPinkBedsPicnic Area N 35 ° 21 ′ 11 ″ W 82 ° 43. 557 ′ El 3208 ′ 5 Aug 09. J. F. & TADCornell AspirExGillFungi (LSAM) (3 F). * USA: NC, Watauga Co. Rough Ridge Trailhead 8 mi. SW Blowing Rock elv. 4200 ′, 2. vi. 1991 (LSAM) (2 F). * USA: NC: Wilkes Co. Stone Mt. State Park, nr. Hutchinson’ s Homestead 36 ° 23.37 ′ N 81 ° 2.75 ′ W. 485 m. Litter sifting. 2 May 2006 A. K. Tishechkin (LSAM) (2 F; 1 M). * USA: NC: Yancy Co. B. R. Pkw. 37.4 mi. SW intersection US 221 elv. 4000 ′ log litter 2. vi. 1991 C. E. Carlton / (LSAM) (1 F; 2 M). * Clear Creek 10 mi S Lancaster, Fair. Co., Ohio v 010 - 1974 / DSChandler sifting rotten logs (DSC) (1 M). * USA, OH, Fairfield Co. Barneby Center 11 May 1989 P. Kovarik (LSAM) (3 F). * OHIO: Lawrence Co. Vinton Furnace Experim. Forest, nr. Forest Serv. Stn. Pu. Carrionbaited pitfall 19 Aug 1995. P. W. Kovarik (LSAM) (1 M). * USA, PA, Somerset Co. Mt. Davis, elv. 3200 ft. Hdwd forest berlesate 16 Aug. 1989, Carlton (LSAM) (4 F). * St. Vinc. Penn. / 12 / C. N. H. M. Colln. (F. Psota Colln. Ex H. F. Wickham Colln.) / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln / Eutyphlus similis 12 Lee (FMNH) (1 F). * SCGreenvilleCo 5 miCaesar’ s HeadStPk 5 Aug 95 J & Scornell Ex ForestLitter & rotFungi; 995 VIII- 5 - 2 (LSAM) (4 F). * USA SC PICKENS CO RockyBottomN 35 ° 3 ′ 32 ″ W 82 ° 47 ′ 52 ″ E 12027 ′ 15 Aug O 9 J & Scornellsifte & Berl- 20 kglitteronRockyBottom Creek (LSAM) (6 F). * USA SC PICKENS CO SassafrasMtN 35 ° 3 ′ 49 ″ N 82 ° 46 ′ 42 ″ W 15 Aug 2009 J & SCornellExAlba trelius CristatusinConiFor (LSAM) (2 F). * # 8 / SMOKIES, TENN. DYBAS 1953 / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (13 F). * USA: TN, Blount Co. Great Smoky Mt. N. P. S. of entrance. Elv. 3000 ′ leaf / log litter 1. vi. 1991 C. E. Carlton (LSAM) (6 F). * TENNESSEE: Blount Co. GSMNP, Ace Gap Tr. @ 35 ° 38.7 ′ N 83 ° 48.3 ′ W. 600 m Forest litter. 2 Aug 2004 V. Bayless & S. Gil (LSAM) (2 F). * TENNESSEE: Blount Co. GSMNP, App. Tr. At Beechnut Gap. 35 ° 34.16 ′ N 83 ° 41.51 ′ W 1510 m. Forest litter sifting 13 April 2006. A. K. Tishechkin (LSAM) (1 F; 2 M). * TENNESSEE: Blount Co. GSMNP, NE corner of Cades Cove at 35 ° 36 ′ 33 ″ N 83 ° 47 ′ 12 ″ W. 640 m Forest litter. 18 July 2003. A. Tishechkin (LSAM) (2 F). * TENNESSEE: Blount Co. GSMNP, Little Bottoms Tr. @ 35 ° 37.125 ′ N 83 ° 55.397 ′ W 520 m. Forest; litter. 31 Jul 2004. J. Ciegler (LSAM) (14 F). * TENNESSEE: Blount Co. GSMNP, lower Gregory Ridge Tr. @ 35 ° 33.5 ′ N 83 ° 50.5 ′ W. 630 m. For. Litter 28 July 2004. A. K. Tishechkin (LSAM) (43 F). * TENNESSEE: Blount Co. GSMNP, Upper Long Hungry Ridge Tr. At 35 ° 30.89 ′ N; 83 ° 51.00 ′ W 1390 m. For. Litter 12 April 2006. A. K. Tishechkin (LSAM) (1 F). * TENNESSEE: Blount Co. GSMNP, Mt. Thunderhead nr. Summit @ 35 ° 34.1 ′ N 83 ° 42.5 ′ W. 1650 m. Litter 30 July 2004. A. K. Tishechkin (LSAM) (2 F; 1 M). * TENNESSEE: Blount Co. GSMNP, West Prong Campgr. 35 ° 37.7 ′ N 83 ° 42.3 ° W. 520 m. Forest litter. 30 Jul 2004. J Ciegler (LSAM) (32 F). * Carter Co. Tenn. # 228 Copeland 12. VII. 53 / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (1 F). * TENNESSEE: Cocke Co. GSMNP, Albright Grove @ 35 ° 44.11 ′ N 83 ° 16.78 ′ W 970 m Forest litter. 1 Aug; 2004. J. Ciegler, A. Tishechkin (LSAM) (6 F). * TENNESSEE: Cocke Co. GSMNP, Albright Grove Loop Tr. Ca. 950 m. Berlese near rotten longs 19 Jul 2003. S. O’ Keefe (LSAM) (10 F). * U. S. A., TN, Cocke Co. GSMNP, Albright Grove Trail 83 ° 16 ′ 45 ″ W, 35 ° 44 ′ 10 ″ N / elv. 1000 m, old growth Berlese 29 June 2001, C. Carlton, A. Tishechkin, V. Moseley (LSAM) (10 F; 1 M). * TN: Cocke Co GSMNP Albright Grove Trail loop / 35 ° 39 ′ 42 ″ N 83 ° 27 ′ 46 ″ W 1500 m Carlton Mayor 6 Dec 2003 (LSAM) (16 F). * USA Tenn. Cocke Co. GSMNP Cosby Creek Trail / Lot # 76 - 106 Oct. 15, 1976 Berlesate R. Chenowith & R. T. Allen (LSAM) (19 F; 1 M). * USA: TN: Cocke Co. GSMNP Cosby House 35 ° 46.67 N 83 ° 12.83 W 7 Oct 2005 M. Gimmel sifting litter (LSAM) (5 F). * U. S. A., TN, Cocke Co. GSMNP, Cosby Ranger Station 83 ° 12 ′ 25 ″ W, 35 ° 46 ′ 12 ″ N elv. 622 m, leaf litter Berlese 31 July 2001, A. Tishechkin (LSAM) (2 F). * USA: TN: Cocke Co. GSMNP Gabes Mnt. Tr @ Hen Wallow Fall, moist forest berlese / 297649 E 3959487 N 19 Jul 2002 CCarlton, NLowe (LSAM) (2 F). * U. S. A., TN, Cocke Co. GSMNP, Hen Wallow Falls 83 ° 14 ′ 27 ″ W, 35 ° 46 ′ 22 ″ N elv. 580 m, leaf litter Berlese 31 July 2001, A. Tishechkin (LSAM) (5 F). * USA TN CockeCo GSMNP MaddronBaldTrail 17 - 2916 31 E 3959236 NA 027 2320 8 Aug 06 JCornell & Sranger BerlExSiftLitter Of 30 ″ DiamHem RockStumpJC 006 - VII- 0 - 40 (LSAM) (3 F). * TENNESSEE: Cocke Co. GSMNP, Maddron Bald Tr. @ lower end of the Albright Grove Loop. ∼ 970 m. Forest litter. 19 Feb 2004. A. Mayor (LSAM) (7 F). * USA: TN: Cocke Co. GSMNP Snake Den Ridge Trail / btw 299077 3957780 & 299657 3957885 Cwright June 2005 (LSAM) (1 F). * TENNESSEE: Sevier Co. GSMNP, App. Tr. ∼ 1 mi E Newfound Gap. 35 ° 36.96 ′ N; 83 ° 24.88 ′ W. 1650 m. Forest; litter. 7 Oct 2005 D. Chouljenko & M. Gimmel (LSAM) (1 F). * TENNESSEE: Sevier Co. GSMNP, App. Tr. At Sweat Heifer Cr. Tr. 35 ° 37.29 ′ N 83 ° 24.25 ′ W. 1725 m. Forest litter. 23 Aug 2006. V. Bayless (LSAM) (8 F; 1 M). * TENNESSEE: Sevier Co. GSMNP, App. Tr. Just W of Sweat Heifer Cr. Trhd. 1720 m 35 ° 37.3 ′ N 83 ° 24.35 ′ W. Forest litter. 14 April 2006. C. Carlton (LSAM) (3 F; 1 M). * U. S. A., TN Sevier Co. GSMNP, Chimneys Picnic Area Nature Trail, 83 ° 29 ′ 45 ″ W, 35 ° 38 ′ 6 ″ N, elv. 891 m / soil litter berlese 30 June 2001, C. Carlton A. Tishechkin, V. Moseley (LSAM) (2 F). * Sevier Co. Tenn. Smoky Mt. Pk. Summer 1953 T. P. Copeland / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (1 F). * Cove Forests S. & E. Gatlinburg, TENN. Sevier Co. V- 16 - 23 1972 2500 - 3500 ′ / Eutyphlus similis LeConte det. DSChandler ’ 95 (FMNH) (1 F). * Rocky Creek Gap Smoky Mts. Nat. Pk. Tenn. IX: 14: 53, SA-HD / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (3 F). * Gt. Smoky Mts. Nat. Pk. Elkmont Sevier Co., Tenn. 8 - VI- 60 Allt. 2250 ′ Leaf Duff / H. Buter J. Wagner Collectors / H. R. Steeves, Jr. Collection / CNHM 1963 H. R. Steeves, Jr. Pselaphidae Colln. Acc. Z- 13, 288 / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (2 F) (2 M). * Greenbrier Cove Smoky Mts. Nat. Pk. Sevier Co., TENN. / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (8 F). * USA: TN: Sevier Co. GSMNP Greenbrier Ridge Tr. 26014. 3941340. 1260 m Berlese 7 Jun 2005 A. Tishechkin (LSAM) (2 F). * TENNESSEE: Sevier Co. GSMNP, Indian Head Tr. 35.60944 ° N 83.44659 ° W sift litter. 5290 ′ 20 July 2003. S. O’ Keefe (LSAM) (18 F; 9 M). * U. S. A., TN, Sevier Co. GSMNP, Laurel Falls Trail 83 ° 35 ′ 36 ″ W, 35 ° 40 ′ 19 ″ N / elv. 747 m, leaf litter berlese 1 July 2001, C. Carlton, V. Moseley, A. Ticheshkin (LSAM) (3 F). * TENNESSEE: Sevier Co., GSMNP Middle Prong Tr. at 35 ° 36.9 ′ N 83 ° 39.4 ′ W 30 July 2004. J. Ciegler (LSAM) (3 F). * TENNESSEE: Sevier Co. GSMNP, lower Middle Prong Tr. @ 35 ° 36.9 ′ N 83 ° 39.4 ′ W. 700 m. Forest litter. 30 Jul 2004. J. Ciegler (LSAM) (8 F). * TENNESSEE: Sevier Co. GSMNP, Mt LeConte nr. Cliff Top Cabins 35 ° 39 ′ 20 ″ N 83 ° 26 ′ 40 ″ W. 1980 m. Leaf moss litter. 18. VII. 2003 A. Mayor & P. Skelly (LSAM) (3 F; 2 M). * Newfound Gap 16. IX. 53 HSD / Field Mus. Nat Hist. Orlando Park Pselaphidae Colln (FMNH) (2 F) (1 M). * USA Tenn. - N. Car.; Sevier Co.; GSMNP Newfound; Gap to Clingmans Dome / Lot # 76 - 107; Oct. 11, 1976; Berlesate; R. Chenowith & R. T. Allen (LSAM) (5 F). * U. S. A., TN, Sevier Co. GSMNP, 0.5 km NE Newfound Gap, elv. 1600 m 83 ° 24 ′ 46 ″ W, 35 ° 38 ′ 9 ″ N / forest litter Berlese 26 June 2001 C. Carlton, V. Moseley, A. Tishechkin (LSAM) (1 F). * TENNESSEE: Sevier Co. GSMNP, W Old Settlers Trhd. 35 ° 42.46 ′ N 83 ° 22.9 ′ W Forest litter. 6 Oct 2005 D. Chouljenko, M. Gimmel & A. Hepperman (LSAM) (2 F). * USA:, TN, Sevier Co. GSMNP, Porters Crk. Tr. 83 ° 23 ′ 52 ″ W, 35 ° 40 ′ 13 ″ N / elv. 870 m, 18 Oct. 2001 old growth berlese A. Tishechkin, A. Cline (LSAM) (6 F). * TENNESSEE: Sevier Co;. GSMNP, Porters Creek Tr. @ 35 ° 40.1 ′ N 83 ° 23.6 ′ W 850 m. Forest litter. 31 July 2004. C. E. Carlton & N. Lowe (LSAM) (3 F). * TENNESSEE: Sevier Co. GSMNP, Porters Creek Tr. Sift litter near rotten logs Beetle Blitz- 2003 18 July 2003. S. O’ Keefe (LSAM) (44 F). * USA: TN: Sevier Co. GSMNP Rainbow Falls snowy litter berlese / 35 ° 39 ′ 42 ″ N / 83 ° 27 ′ 46 ″ W 1500 m Carlton Mayer 6 Dec 2003 (LSAM) (1 F). * TENNESSEE: Sevier Co. GSMNP Ramsey Cascades Tr. @ 38 ° 42.53 ′ N 83 ° 19.21 ′ W 1165 m. Forest litter. 10 May 2002. W. D. Merritt (LSAM) (3 F). * U. S. A, TN, Sevier Co., GSMNP Ramsey Cascade Trail (middle) 83 ° 18 ′ 24 ″ W, 35 ° 42 ′ 34 ″ N / elv. 1300 m. Forest litter berlese 21 March 2002 CECarlton, VLMoseley (LSAM) (2 F). * U. S. A, TN, Sevier Co., GSMNP Ramsey Cascade Trail (upper) 83 ° 18 ′ 7 ″ W, 35 ° 42 ′ 30 ″ N / elv. 1375 m. Forest litter berlese 21 March 2002 CECarlton, VLMoseley (LSAM) (3 F). * TENNESSEE: Sevier Co. GSMNP, Road Prong Tr. at 35 ° 36 ′ 36 ″ N 83 ° 27 ′ 3 ″ W 1580 m. Leaf / moss mat litter. 20 July 2003 A. Ticheshkin (LSAM) (20 F; 1 M). * USA: TN: Sevier Co. GSMNP, Roaring Fork Area, Rainbow Falls Tr. / 1 Aug 2002 moist cave Berlese UTM 275790 E 3949640 N C. Carlton (LSAM) (21 F). * TENNESSEE: Sevier Co. GSMNP Trillium Gap Tr. @ 35 ° 39.9 ′ N 83 ° 26.2 ′ W 1400 m. Forest litter. 29 July 2004. A. Tishechkin (LSAM) (38 F; 7 M). TENNESSEE: Sevier Co. GSMNP, jct. Trillium Gap & Brushy Mt. Tris. 35 ° 40.4 ′ N 83 ° 26.0 ′ W. 1430 m. Forest litter. 29 July 2004 C. E. Carlton & J. Ciegler (LSAM) (16 F). * VA., Giles Co. Jefferson Nat’ l. Forest Mt. Lake Tech. Area 4. VII. 67 / Rhododendron duff / H. R. Steeves, Jr. H. R. Steeves III Collectors / H. R. Steeves, Jr. Collection (FMNH) (1 F; 1 M). * VIRGINIA: Madison Co., Shenandoah NP, nr. Upper Hawksbill Parking. 38 ° 32.73 ′ N; 78 ° 23.64 ′ W. 1115 m. Litter sifting. 3. v. 2006. A. Tishechkin (LSAM) (2 F). * Mountain Lake, VA. / VII- 23 - 47 J. H. Davis / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. (FMNH) (1 F). * VIRGINIA: Smyth Co., Lewis Fork Wildrns., Mt. Rogers Tr. at 36 ° 39.63 ′ N 81 ° 33.11 ′ W 1540 m. Litter sifting. 30 April 2006. A. K. Tishechkin (LSAM) (13 F; 1 M). * USA, VA. Smythe Co. 7.5; mi SE Marion, Raccoon; Branch cmpgrnd, v- 29 - 91; sift pine & oak litter; S. O’ Keefe collection (DSC) (1 M). * USA: VA, Wash. Co. E of Damascus leaf litter 3. vi. 1991 C. E. Carlton (LSAM) (3 F). * USA WV Fayette Co. 35 mi N Beckley 18 - III- 83 JFcornell ExConiferlitter JFC 983 - III- 18 - 4 (LSAM) (1 M). * W. Virginia: Mercer Co. Camp Ck. St. Forest. IV: 8: 1971. Leg: S. Bird. FM (HD) # 71 - 72 / Eutyphlus similis det. DSChandler (FMNH) (1 F; 1 M). * WVPocahontasCo Snedegar’ sCave (illegible) 10 - vi- 1989 JPCornall (illegible) forest litter (NCSUIM) (1 F; 6 M).	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB2580DFF51382DFE23FD55.taxon	description	(Figs. 17, 19 A, 19 C, 26, 32)	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB2580DFF51382DFE23FD55.taxon	materials_examined	Holotype. Male. USA: NC. Macon Co. 10 mi SW Franklin; Back Country info center; V- 22 - 1991; Berlese oak & Rhododendron; S. O’ Keefe (FMNH) (dissected and slide-mounted in Euparal).	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB2580DFF51382DFE23FD55.taxon	diagnosis	Diagnosis. Eutyphlus spiralis is similar to E. dybasi based on the absence of anteroprosternal foveae, presence of secondary sexual characters on the legs of males (Figs. 19 A, C, 26), quadrifoveate elytra, concave metaventrite, and larger, broadly ovate seventh ventrite of males. In E. spiralis, the right paramere of the aedeagus is ventrally located and expanded distally to the left (the right paramere of E. dybasi is expanded distally to the right), and the left paramere is dorsally located, twisted counterclockwise, and bears a series of spines along both sides of the process (Fig. 17) (the left paramere of E. dybasi is expanded distally into a process that bears ctenidial spines along one face).	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB2580DFF51382DFE23FD55.taxon	description	Description. MALE. Measurements: HL 0.20 mm, HW 0.29 mm; PL 0.35 mm, PW 0.33 mm; EL 0.56 mm, EW 0.56 mm; A 1 - 6 respectively 0.02, 0.16, 0.15, 0.12, 0.11, 0.3 mm. An 1 – 11 respectively 0.10, 0.05, 0.03, 0.02, 0.02, 0.02, 0.03, 0.03, 0.04, 0.04, 0.11 mm. MP 1 – 4, respectively, 0.01, 0.05, 0.01, 0.05 mm. GL 0.23 mm, GW 0.14 mm. ML 1.63 mm. Integument: Typical for genus. Head: Head slightly narrower than pronotum. Eyes present and well-developed with 27 facets, slightly emarginate posteriorly, ocular canthus weakly developed, projecting weakly laterally and sloping to gula. Ventral surface flat to slightly rounded at gula, gular sulcus well-developed anteriorly and projecting to margin of posteriorly located, paired, gular foveae. Thorax: Prothorax with basolateral margin slightly crenulate, median longitudinal sulcus present, area behind basal sulcus with thin, longitudinal sulcus extending from basal bead to basal sulcus. Prosternum lacking anteroprosternal foveae. Protibia with apical spine. Mesotrochanter with tubercle, mesotibia with apical flange. Metaventrite slightly concave baso-medially. Elytra bearing 3 basal foveae, as well as subhumeral foveae. Flight wings well-developed. Abdomen: Tergites unmodified. Seventh ventrite broadly oval. Other ventrites unmodified. Genitalia: Aedeagus asymmetrical. Broad, membranous lobe dorsal and associated with left paramere that is twisted counter-clockwise distally and bears a row of ctenidial spines on both sides. Median lobe with distal fimbriated process. Right paramere more ventral, flattened, distally expanded to left, with apical patch of very sparse, scattered, short, suberect setae. FEMALE. Unknown. Variation. Unknown. Bionomics. Seasonality: The type was collected during May. Elevation: The elevation at the type locality is 646 m. Habitat / Microhabitat: The holotype was collected from oak and rhododendron litter. Collecting methods: The holotype was collected by sifting forest litter and extracting via a Berlese funnel.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB2580DFF51382DFE23FD55.taxon	distribution	Distribution. This species is known from Macon County, North Carolina (Fig. 32). 6. Eutyphlus thoracicus Park, 1956 (Figs. 11, 12, 18, 33)	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB2580DFF51382DFE23FD55.taxon	diagnosis	Diagnosis. Eutyphlus thoracicus is distinguished from all known Eutyphlus species by the smaller body size, absence of a longitudinal pronotal sulcus, the left paramere of the aedeagus forming a sclerotized bar that bears small spicules and setae, and the ventrally located flattened right paramere (Fig. 18).	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB2580DFF51382DFE23FD55.taxon	description	Redescription. MALE. Measurements: HL 0.14 mm, HW 0.20 mm; PL 0.21 mm, PW 0.20 mm; EL 0.34 mm, EW 0.38 mm; A 1 – 6 respectively 0.03, 0.11, 0.10, 0.09, 0.09, 0.02 mm. An 1 – 11 respectively 0.05, 0.03, 0.01, 0.01, 0.01, 0.01, 0.02, 0.02, 0.02, 0.03, 0.09 mm. MP 1 – 4 respectively 0.01, 0.03, 0.01, 0.03 mm. GL 0.05 mm, GW 0.04 mm. ML 1.20 mm. Integument: Typical for genus. Capitate genal setae sparse. Head: About as wide as pronotum. Vertex bearing small, longitudinal depression extending dorsally from posterior margin of head to about the margin of the eye. Eyes present, well-developed with 27 facets, convex, about as long as wide. Ocular canthus weakly developed, projecting laterally and sloping to gula. Ventral surface of head rounded at gula, gular carina or sulcus absent or weakly developed anteriorly, posteriorly located paired, gular foveae present. Thorax: Pronotum with lateral margin smooth, median longitudinal sulcus absent, area behind basal sulcus smooth. Prosternum bearing anteroprosternal foveae. Paired median mesoventral fovea present. Metaventrite flat. Elytra bearing 2 basal foveae as well as subhumeral foveae. Flight wings well-developed. Abdomen: Tergites unmodified. Fourth ventrite bearing small basolateral foveae. Seventh ventrite small and strap-like. Other ventrites unmodified. Genitalia: Aedeagus asymmetrical. Left paramere most dorsal, oblique, forming a sclerotized bar bearing small spicules and setae, and possessing a single, elongate, median seta. Median lobe modified into internal, fimbriated process associated with internal, membranous sac. Right paramere attached ventrally, flattened, and bearing 4 distal setae. FEMALE. Measurements: HL 0.16 mm, HW 0.16 mm; PL 0.23 mm, PW 0.21 mm; EL 0.30 mm, EW 0.29 mm; A 1 – 6 respectively 0.02, 0.11, 0.10, 0.10, 0.08, 0.04 mm. An 1 – 11 respectively 0.04, 0.02, 0.01, 0.01, 0.01, 0.01, 0.01, 0.01, 0.02, 0.03, 0.07 mm. MP 1 – 4 respectively 0.01, 0.03, 0.01, 0.03 mm. ML 1.20 mm. Integument, head, thorax, and abdomen: Similar to male except eyes completely absent, ocular canthus weakly developed. Tempora rounded and nearly parallel. Pronotum more quadrate. Seventh ventrite small and transverse. Genitalia: Completely membranous. Produced into somewhat elongate, median membranous structure and billowy right sac. Variation. Although generally smaller than members of other species, size is variable (± approximately 0.1 mm total length). The degree of pronotal convexity varies slightly. The aedeagus of E. thoracicus varied little among the individuals examined, but three distal setae were present on the right, ventrally located paramere of a few specimens instead of the usual four. Bionomics. No previously published information on the biology of E. thoracicus is available. Seasonality: Specimens of E. thoracicus were all collected from late winter / early spring to early fall (earliest date 21 February; latest 20 October). Elevation: Specimens were obtained from between 870 m and 1,920 m elevation. Habitat / Microhabitat: Specimens were collected from hardwood forest litter, Rhododendron litter, oak litter, spruce litter, fir litter, and old-growth forest mixed litter. Several individuals were also collected from moss mats. Collecting methods: The majority of individuals were collected via litter sifting and Berlese extraction. Several individuals were collected in flight intercept traps.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB2580DFF51382DFE23FD55.taxon	distribution	Distribution. The species is known from Georgia (Dade County), North Carolina (Haywood, Macon, Sway, and Yancy Counties), Tennessee (Blount, Cocke, and Sevier Counties), Virginia (Washington County), and West Virginia (Pocahontas County) (Fig. 33). PHYLOGENETIC ANALYSIS Results. Cladistic analysis of the data matrix resulted in 26 equally parsimonious trees (length = 347, CI = 0.37, RI = 0.52). The preferred tree selected for illustration (Fig. 34) was rooted with Sonoma baylessae Ferro and Carlton (tribe Faronini). A clade comprising one metopiasine genus, Rhinoscepsis LeConte, and one panaphantine genus, Trimioplectus Brendel, were recovered as sister to the remaining taxa. Remaining taxa were supported by five synapomorphies (14 - 1, 17 - 1, 20 - 1, 68 - 0, 71 - 0) and one homoplasious character (50 - 1). Hendecameros Comellini, the single representative of the tribe Bythinoplectini, diverged based on a number of homoplasies and a single autapomorphy (13 - 2). The clade containing the remaining taxa was supported by five synapomorphies (2 - 1, 12 - 1, 35 - 0, 60 - 0, 91 - 1). Two homoplasies (39 - 0, 49 - 0) supported a clade comprising all representatives of the Trimiina, plus members of the genus Bibloplectus Reitter (subtribe Panaphantina) and Mayetiini. Bibloplectus was sister to Trimiina, and Mayetia Mulsant and Rey was nested within the trimiine taxa close to the genus Hanfordia Park. The second branch, linked by the homoplasies (45 - 1, 57 - 3), contained Eutyphlus and a number of trichonychine genera and genera from the remaining Euplectitae tribes and supertribes, with the exception of the basal Faronitae. The two genera of the tribe Jubini were recovered as sister taxa based on a number of synapomorphies and homoplasies. Thesium Casey was not recovered with other panaphantines. Actiastes Casey was not recovered with the other trimiine genera but was paired with Pseudactium Casey. Eutyphlus was recovered as a monophyletic group alongside other members of the Panaphantina, excluding Thesium, based on two homoplasies. Within the genus, E. dybasi and E. spiralis formed a sister clade to the other four species based on a single character (78 - 1). The remaining species were grouped as E. similis sister to E. thoracicus and E. schmitti sister to E. prominens. Other clades included a mixture of trichonychine genera on trees that differed in topology among various equally parsimonious trees. Non-trichonychine genera generally segregated on the trees with batrisite and goniacerite genera as sister taxa, and the two representatives of the goniacerite tribe Tychini recovered as sister taxa. Bootstrap support was low for many of the internal branches of the tree (Fig. 35). Four branches on the tree had a branch support above 90 %. The branch with Sonoma Casey diverging as sister to all other taxa, the Rhinoscepsis - Trimioplectus clade, the Jubini clade, and the branch linking E. prominens and E. schmitti had branch supports of 1.0, 1.0, 0.99, and 0.94, respectively. The next highest branch support was 0.71 for the goniacerite clade containing Pselaptrichus Brendel and Lucifotychus Park and Wagner. The branch containing Arthmius LeConte and Reichenbachia Leach had support of 0.55. The rest of the branches in the tree had support below 0.50.	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB2580DFF51382DFE23FD55.taxon	discussion	Discussion. The main objective of the phylogenetic analysis was to test for the monophyly of Eutyphlus. Although branch support was low, monophyly of the genus was supported by several homoplasious character states. These results, in addition to the relatively straightforward diagnosis of the genus using easily observable external characters, argue for retaining genus status for this group of species. Several tribes and subtribes were recovered as paraphyletic among different branches of the tree. Only the Bythinoplectini, Jubini, Trogastrini, and Dimerini formed monophyletic groups, and only the Bythinoplectini and Jubini were not nested within other tribes. Limited taxon sampling of these genus groups in this analysis makes assertions regarding monophyly problematic. The supertribe Euplectitae is a collection of tribes and subtribes that have been taxonomically defined on the basis of regional treatments, with previously untreated genera sometimes placed into these taxa arbitrarily (Newton and Chandler 1989). The subjective nature of these higher classifications has resulted in downstream issues for the classification of large genus groups that share a number of dis- tinct morphological features, yet cannot be separated in a consistent manner under the current system. In addition, a number of characters exhibited by the Euplectitae, such as foveae and sulci, have proven to be highly variable within tribes and supertribes, sometimes differing among species within the same genus. This results in a large number of homoplasious characters and requires inclusion of a large number of outgroups within the Euplectitae and other supertribes to polarize characters for tree rooting. Homoplasy is a particularly vexing problem with the external characters available for use in morphological phylogenetic analysis, and this may reflect conserved developmental genes that may be turned on and off in phylogenetically remote taxa. Thus, these characters are poor indicators of relationship at genus group nodes (tribe and subtribe), but may provide useful data for defining phylogenetically shallow nodes within genera, as required in the present study for testing monophyly of Eutyphlus. The low support values at the majority of nodes do not inspire confidence in the phylogenetic hypothesis proposed, but they are consistent with results of other species-level pselaphine phylogenies (e. g., Carlton 2003 that featured a 16 - species polytomy of Reichenbachia species).	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
038FA12DFFB85805FEAB3817FB1AF914.taxon	materials_examined	Material Examined. (n = 150) * TENNESSEE: Sevier Co. GSMNP, Trillium Gap Tr. @ 35 ° 39.9 ′ N 83 ° 26.2 ′ W 1400 m. Forest litter. 29 July 2004. A. Tishechkin / LSAM 0080787 (LSAM) (slide mounted male) (description of male based on this specimen). * TENNESSEE: Sevier Co.; GSMNP, Trillium Gap Tr. @; 35 ° 39.9 ′ N 83 ° 26.2 ′ W; 1400 m. Forest litter. 29 July; 2004. A. Tishechkin (LSAM) (slide mounted female) (description of female based on this specimen). * Rocky Creek Gap Smoky Mts. Nat. Pk. Tenn. 1 X: 14: 53, SA-HD / Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln. / (orange label) HOLOTYPE / Eutyphlus thoracicus. Park, nsp. (FMNH) (1 M; HOLOTYPE). * same locality / (orange label) PARATYPE. (FMNH) (2 F). * Chimney Cp. Gr Smoky Mts. N. P. Tenn. 16: IX: 53 Auerbach # 5 / Field Mus. Nat. Hist Orlando Park Pselaphidae Colln. / (orange label) PARATYPE (FMNH) (1 F). * Chimney Cp. Gr Smoky Mts. N. P. Tenn. 16: IX: 53 Auerbach # 5 / Field Mus. Nat. Hist Orlando Park Pselaphidae Colln. / (orange label) PARATYPE (FMNH) (1 F). * Cloudland Canyon S. Pk. Dade Co., GA. 14. VI. 62 Forest floor debris / H. R. Steeves, Jr. J. D. Patrick Jr. Collectors / H. R. Steeves Jr. Collection (FMNH) (5 F). * N CAROLINA: Haywood Co. GSMNP, Caldwell Fork Tr. At UTM 30897 E 3940883 N. Moist forest Berlese. 3 August 2002. C. Carlton & N. Lowe (LSAM) (18 F; 2 M) * USA: NC Haywood Co. GSMNP McKee Branch Tr. 35 ° 35 ′ 6 ″ N 83 ° 05 ′ W / 1370 m 16 Jul – 12 Aug 2002 FIT C. Carlton & S. Gil (LSAM) (1 M). * N CAROLINA: Haywood Co. GSMNP, Mt. Sterling Gap. 35 ° 42.0 ′ N 83 ° 5.9 ′ W 1190 m. Forest litter. 29 July 2004. R. T. Allen / LSAM 0095711 (LSAM) (1 F). * same locality (LSAM) (16 F; 1 M). * USA: NC: Haywood Co. GSMNP Purchase Knob, berlese / 313124 E 3939640 N 5089 ft. 20 Jul 2002 CCarlton (3 F; 1 M). * USA: NC: Macon Co. 3 mi NW Highlands v- 29 - 1983, sift rhododendron litter, DSChandler (DSC) (7 F). * USA: NC: Swain Co GSMNP Appalachian Tr. 1.5 mi NE Newfound Gap / 281639 E 3944038 N ridgetop berlese 18 Jul 2002 CCarlton (LSAM) (3 F). * N CAROLINA: Swain Co. Appalachian Tr. ∼ 1.5 mi E Newfound Gap. 35 ° 36.96 ′ N 83 ° 24.66 ′ W 1730 m. Forest litter; 18 July 2002. C. Carlton (LSAM) (1 F). * U. S. A., NC, Swain Co.; GSMNP, Beech Gap Tr.; 83 ° 12 ′ 32 ″ W, 35 ° 38 ′ 23 ″ N / elv. 1400 m, 20, Oct. 2001; N. hardwood berlese; LSAM, C. Carlton (LSAM) (2 F). * N CAROLINA: Swain Co. GSMNP, jct. of Beech Gap & Hyatt Ridge Trails 35 ° 37.82 ′ N 83 ° 42.07 ′ W 1495 m. Forest litter. 2 Oct 2005. W. D. Merritt (LSAM) (3 F). * USA N. C. Swain Co. Dirt Ri. From Heintooga Overlook to Cherokee / Lot # 76 - 102 Oct. 14, 1976 Berlesate 5000 ′ - 4500 ′ R. Chenowith & R. Tallen (LSAM) (7 F). * U. S. A., NC, Swain Co. GSMNP, Flat Creek Trail 83 ° 10 ′ 21 ″ W, 35 ° 33 ′ 1 ″ N / elv. 1500 m, leaf litter Berlese, 31 July 2001. A. Tishechkin (LSAM) (1 M). * N CAROLINA: Swain Co. GSMNP, Jenkins Ridge Tr. ∼ 1 mi from Appalachian Tr. 35 ° 33.75 ′ N 83 ° 43.2 ′ W 1540 m. Forest litter. 30 Jul 2004. A. K. Tishechkin (LSAM) (3 F). * N CAROLINA: Swain Co. GSMNP, upper Jenkins Ridge Tr. At 35 ° 33.75 ′ N 83 ° 43.2 ′ W. 1540 m. Leaf litter 31 Jul 2004. A. K. Tishechkin (LSAM) (1 F). * N. CAR. Swain Co. Mt. Collins 5900 ′ v. 21.1977 A. Newton, M. Thayer (DENH) (2 F). * N CAROLINA: Swain Co. GSMNP, Noland Divide Tr. at 35 ° 33 ′ 57 ″ N 83 ° 28 ′ 36 ″ W 1770 m. Leaf / moss mat litter 19 July 2003. A. Tishechkin (LSAM) (2 F). * USA: NC. Yancey Co. Black Mtn. info cntr v- 21 - 1991 sift fir-oak S. O’ Keefe collr (DSC) (3 M). * USA: NC, Yancy Co. B. R. Pkwy. 37.4 mi. SW intersection US 221 elv. 4600 ′ log litter 2. vi. 1991 C. E. Carlton (LSAM) (1 F). * SC: Oconee Co. 6.7 mi NNW Walhalla 1. vii 1983 1760 ′ hdwd forest litter J. Pakaluk # 148 (DSC) (1 M). * TENNESSEE: Blount Co. GSMNP, App. Tr. ∼ 0.5 mi E Thunderhead Mt. summit 35 ° 34.05 ′ N 83 ° 43.05 ′ W 1510 m. Forest litter. 30 Jul 2004 A. K. Tishechkin (LSAM) (5 F). * TENNESSEE: Blount Co. GSMNP, App. Tr. ∼ 0.5 km W Mt. Thunderhead summit at 35 ° 34.11 ′ N 83 ° 42.00 ′ W 1585 m. Forest litter sifting 13 April 2006. A. K. Tishechkin (LSAM) (4 F). * USA: TN, Blount Co. 5.8 mi. ECling. Dome 1. vi. 1991 C. E. Carlton (LSAM) (2 F). * TENNESSEE: Blount Co. GSMNP, Gregory Cave entrance. 35 ° 33.74 ′ N 83 ° 51.624 ′ W. Litter. 28 July 2004. V. Bayless, C. Carlton & A. Tishechkin (LSAM) (1 F). * TENNESSEE: Blount Co. GSMNP, lower Gregory Ridge Tr. @ 35 ° 33.5 ′ N 83 ° 30.5 ′ W. 630 m. For. Litter 28 Jul 2004. A. K. Tishechkin (LSAM) (3 F). * TENNESSEE: Cocke Co. GSMNP, Albright Grove Loop Tr. Ca. 950 m. Berlese near rotten longs 19 July 2003. S. O’ Keefe (LSAM) (2 F). * USA Tenn Cocke Co. GSMNP Cosly Creek Trail / Lot # 76 - 105 Oct. 15, 1976 Berlesate R. Chenowith & R. T. Allen (LSAM) (4 F). * TENNESSEE: Sevier Co. GSMNP, App. Tr. Just W of Sweat Heifer Cr. Trhd. 1720 m 35 ° 37.3 ′ N 83 ° 24.35 ′ W. Forest litter. 14 April 2006. C. Carlton (LSAM) (1 F). * TENNESSEE: Sevier Co. GSMNP, Indian Head Tr. 35.60944 ° N 83.44659 ° W Sift litter. 5290 ′ 20 July 2003. S. O’ Keefe (LSAM) (2 F; 2 M). * USA-TN: Sevier Co. GSMNP. Mount LeConte, Alum Cave Tr. at 35 ° 39.33 ′ N 83 ° 26.75 ′ W, 1920 m, spruce-fir litter berlese Sokolov I. M 6 Oct 2008 (LSAM) (3 F; 3 M). * TENNESSEE: Sevier Co. GSMNP, Porters Creek Tr. Sift litter near rotten logs Beetle Blitz- 2003 18 July 2003. S. O’ Keefe (LSAM) (9 F). * U. S. A., TN, Sevier Co. GSMNP, Porters Crk. Tr. 83 ° 23 ′ 52 ″ W, 35 ° 40 ′ 13 ″ N / elv. 870 m, 18 Oct. 2001 old growth berlese A. Tishechkin, A. Cline (LSAM) (2 F; 4 M). * TENNESSEE: Sevier Co. GSMNP, Road Prong Trail 35 ° 36 ′ 36 ″ N 83 ° 27 ′ 3 ″ W 1580 m. Leaf / moss mat litter. 20 July 2003 A. Tishechkin (LSAM) (3 F; 1 M). * USA: TN: Sevier Co. GSMNP, Roaring Fork Area, Rainbow Falls Tr UTM 275540 E 3949710 N, Moist forest Berlese 1 Aug 2002 CEC (LSAM) (6 F). * TENNESSEE: Sevier Co. Trillium Gap Tr. @ 35 ° 39.9 ′ N 83 ° 26.2 ′ W 1400 m. Forest litter. 29 July; 2004. A. Tishechkin (LSAM) (1 F). * USA Tenn. - N. Car. Sevier Co. GSMNP, Newfound Gap to Clingman’ s Dome / Lot # 76 - 107 Oct. 11, 1976 Berlesate R. Chenowith & R. T. Allen (LSAM) (2 F). * USA: VA, Wash. Co. E. of Damascus leaf litter 3. vi. 1991 C. E. Carlton (LSAM) (1 M). * USA: WV. Pocahontas Co. 6 mi SE Marltinon Co. forest trail # 407, off Hwy 92, v- 16 - 1991, sift oak litter, S. O’ Keefe (DSC) (1 F). APPENDIX 2 List of taxa (with specimen label data) included in the phylogenetic analysis. SUPERTRIBE BATRISITAE TRIBE BATRISINI Reitter Arthmius bulbifer Casey, 1983: USA: AR, Montgomery Co. approx. 5 mi. W. of Little Mo. Falls rec. Area MVLT 25. vii. 1991 C. E. Carlton SUPERTRIBE EUPLECTITAE TRIBE BYTHINOPLECTINI Schaufuss Hendecameros panamense Comellini, 1985: PANAMA: Colon Pr., San Lorenzo Forest. 9 ° 17 ′ N 79 ° 58 ′ W. Flight intercept FIT-C 1 - 5. 27 - 28. ix. 2003 A. Tishechkin – IBISCA’ 03 TRIBE DIMERINI Raffray Barroeuplectoides sp. Park, 1942: BELIZE: Orange Walk Dist. Rio Bravo Conservation Area La Milpa Ruins 17. IV. 1995 P. W. Kovarik rot. Log. Berlese det. C. Carlton 1995 TRIBE EUPLECTINI Streubel Euplectus confluens LeConte, 1849: USA, AR, Newton Co. Buf. Nat. Riv. Kyles Landing Hdwd. Berlesate 10. Mar. 1988 C. Carlton Leptoplectus pertenuis (Casey, 1884): USA, Ark, Polk Co, Shady Lake Rec. Area, 1200 ft. Hdwd-pine berlese. 13 Oct. 1974 A. Newton Pycnoplectus linearis (LeConte, 1849): FLORIDA: Alachua Co. 29 ° 34 ½ ′ N, 82 ° 29 ′ W 18 - X- 1994 Randall W. Lundgren Flightbarrier trap in hardwood hammock det. C. Carlton 1996 TRIBE JUBINI Raffray Jubus sp. Schaufuss, 1872: ECUADOR, Azuay Pr. 45 km NE Cuenca, 195 elv. 2850 m, 1 Jan 1992 cloud forest Berlese C. Carlton, R. Leschen Sebaga sp. Raffray, 1891: BELIZE: Orange Walk Dist. Rio Bravo Cons. Area Research Station 15 - 20 IV 1995, F. I. T. P. W. Kovarik TRIBE MAYETIINI Winkler Mayetia domestica Schuster, Marsh, and Park, 1959: USA: MO: Taney Co., Ozark Underground Lab 36.5789 ° N, 92.8641 ° W. 19. X. 2007, soil under fungi, # 998. L. F. Watrous TRIBE METOPIASINI Raffray Metopias sp. Gory, 1832: ECUADOR: Depto. Orellana, Res. Ethnica Waorani, 1 km S Onkone Gare Camp, Trans. Ent. 0 ° 39 ′ 10 ″ S 76 ° 26 ′ W. 220 m. 5 February 1996 T. L. Erwin et al. collectors Insecticidal fogging of mostly bare green leaves, some with covering of lichenous or bryophytic plants in terra firme forest. Project MAXUS Lot 1427 Trans. 3 Sta. 7 Subtribe Rhinoscepsina Bowman 1934 Rhinoscepsis bistriatus LeConte, 1878: Crescent City, Fla. Coll Hubbard & Schwarz TRIBE TRICHONYCHINI Reitter Subtribe Trichonychina Reitter Foveoscapha terracola Park and Wagner, 1962: USA: Washington, Pierce Co. Mt Rainier NP, 1000 ft N Green Lake, 12.3 mi (19.8 km) SE Wilkeson UTM: 10 Z, 586840 mE, 5203500 mN elev: 3360 ft 27 Oct 1996 E. A. Lisowski Berlese of rotting wood and forest litter # 105 - 01 Subtribe Panaphantina Jeannel Bibloplectus cherokee Chandler, 1990: USA: LA: Bossier Par Bodcau WMA. UVLT 22 May 1996 C. Carlton / V. Moseley. Det. CE CARLTON 2002 Eutyphlus dybasi Park, 1956: USA: TN: Cocke Co. GSMNP Snake Den Ridge Trail btw 299077 3957780 & 299657 3957885 CWright June 2005 Eutyphlus spiralis Owens and Carlton, new species: USA: NC: Macon Co. 10 mi SW Franklin; Back country info center; V- 22 - 1991; Berlese oak & Rhododendron; S. O’ Keefe Eutyphlus prominens Park, 1956: TENNESSEE: Sevier Co. GSMNP, Trillium Gap Tr. @ 35 ° 39.9 ′ N 83 ° 26.2 ′ W 1400 m. Forest litter. 29 July 2004. A. Tishechkin Eutyphlus schmitti Raffray, 1904: USA: PA: Jeffer. Co., Cook State Forest 0.4 mi S Cooksburg Mohawk Trail. VI- 6 - 1997, DSChandler hardwood leaf litt. Eutyphlus similis LeConte, 1880: USA: Tenn. Cocke Co. GSMNP Colby Creek Trail Lot # 76 - 106 Oct 15, 1976 Berlesate R. Chenowith & R. T. Allen Eutyphlus thoracicus Park, 1956: N CAROLINA: Haywood Co. GSMNP, Caldwell Fork Tr. At UTM 30897 E 3940883 N. Moist forest Berlese. 3 August 2002. C. Carlton & N. Lowe Pseudactium carolinae Casey, 1908: USA, TX, Sabine Co. 9 mi. E Hemphill Beech Bottom 11 May 1988 beech, magnolia sifter R. Anderson Thesiastes sp. Casey, 1894: Falls Church 6: iv: 1919 Va EA Chapin Collector Thesium sp. Casey, 1884: USA Ark. Scot Co. 6.7 Mi. S. Inters Hwys. 270 & 71 on 71 Lot # 77 - 103 July 25, 1977 Berlesate Hardwood-Pine Chenowith, Carlton, Heiss Trimioplectus australis Chandler, 1990: USA, AR, Polk Co. Rich Mt., Eagleton Overlook Berlesate R. T. Allen, 17 Feb. 1988 Subtribe Trimiina Bowman Actiastes globiferum (LeConte, 1849): Nashville Tenn H Soltau Collection Dalmosanus quercavum (Chandler, 1990): USA Ark. St. Francis Co. 6.5 Mi. NW. Inters Hwys 79 & 38 on 38 Lot # 77 - 23 March 10, 1977 Berlesate Oak R Chenowith Dalmosella tenuis Casey, 1878: USA, AR, Cross Co., Village Crk. St. Pk. Camp loop A Berlesate 7 Feb. 1989 C. E. Carlton Hanfordia sp. Park, 1960: BELIZE: Orange Walk Dist. Rio Bravo Conservation Area La Milpa ruins, 18. IV. 1995 Atta debris pile, P. W. Kovarik det. C. Carlton 1995 Melba clypeata (Reitter, 1883): Virgin IS: St Croix Est. Rust-up-Twist North Shore Rd. 01 Oct 1987, M. A. Ivie under bark Trimiomelba dubia (LeConte, 1849): USA: TX Brewster Co. Big Bend N. P., Cattail Falls, elv. 1310 m 6 Sept 1988 hdwd. Berlesate R. Anderson Trimium brevicorne (Reichenbach, 1816): PQ # A 20402 DC 9. XII. 32 in moss, taking from Czechoslovzk USNM Quadrelba sp. Park, 1942: VIRGIN IS: St. John Bordeaux Mt. Elv. 1150 ft. 13 - AUG 1980 M A Ivie Colr. Rotten Log TRIBE TROGASTRINI Jeannel Subtribe Trogastrina Jeannel Conoplectus acornus Carlton, 1983: USA Ark. Fulton Co. 1.4 Mi. N. Inters. Hwys. 62 & 289 on 289 Lot # 77 - 186 Oct. 18, 1977 Berlesate Hardwoods R. Chenowith Subtribe Rhexiina Park Rhexius sp. LeConte, 1849: USA, Ark., Desha Co. 1.9 mi. W. intersec. Hwy. 65 + I 59 on I 59. Bottomland Hdwd. Mar. 8, 1977 Chenowith 77 - 19 SUPERTRIBE FARONITAE Sonoma baylessae Ferro and Carlton, 2010: USA Tenn-N. Car. Sevier Co. GSMNP, Newfound Gap to Mt. Le Conte Lot # 77 - 74 June 22, 1977 Berlesate 5000 ′ – 6300 ′ R. Chenowith & J. Heiss SUPERTRIBE GONIACERITAE TRIBE BRACHYGLUTINI Raffray Reichenbachia howardi Park, 1958: Gueydan La Aug 7 1925 EKalmbach At lights TRIBE BYTHININI Raffray Pselaptrichus perfidus Schuster and Marsh, 1956: USA: Washington, King Co. Mt. Baker- Snoqualmie Nat. Forest. 0.8 mi SSW Mount Defiance summit 11.3 mi ESE North Bend UTM: 10 Z, 607800 mE, 5253180 mN Elev: 3200 ft, 1 - ix- 1996 E. A. Lisowski # 523 - 01 berlese of moss and liverworts on wood TRIBE TYCHINI Raffray Lucifotychus cognatus (LeConte, 1874): USA: WASHINGTON, Yakima Co. # 567 - 1 ravine north of Wild rose Campground 3.8 mi (6.1 km) ENERimrock UTM: 10 Z, 649460 mE, 5170760 mN Elev: 2650 ft, 780 m 15 - II- 1998, EA Lisowski Berlese creek litter	en	Owens, Brittany E., Carlton, Christopher E. (2016): Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini. The Coleopterists Bulletin 70 (1): 1-29, DOI: 10.1649/072.070.0102, URL: http://dx.doi.org/10.1649/072.070.0102
