identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038FEF5F2F10FF9B8ED1FC70FA882ED6.text	038FEF5F2F10FF9B8ED1FC70FA882ED6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetilla pentatriaena	<div><p>Tetilla pentatriaena sp. nov.</p><p>(Figs 2, 3; Tables 1, 3)</p><p>Holotype. UFBA 2048-POR, Camaçari, north of Bahia State, Brazil (12o45’41.61”S – 38o06’12.77”W), 26 m depth, ii.1999, coll. Walter Andrade.</p><p>Paratypes. (1) Camaçari, north littoral of Bahia State, Brazil (1247’05.0”S 3806’38.4”W), (same collector and depth as holotype), UFBA 2049-POR, vii.2003; UFBA 2051-POR, UFBA 2052-POR, UFBA 2053-POR, UFBA 2054-POR and UFBA 2055-POR, vii.2005; UFBA 2057-POR, UFBA 2058-POR, UFBA 2059-POR and UFBA 2060-POR, ii.2006; UFBA 2373-POR, UFBA 2374-POR, UFBA 2375-POR, UFBA 2376-POR and UFBA 2377- POR, vii.2007; UFBA 3272-POR, i.2010. Camaçari, north littoral of Bahia State, Brazil (1244’59.7”S 3804’05.5”W), UFBA 2050-POR, 28 m depth, ii.2004, coll. (same collector as holotype). Camaçari, north littoral of Bahia State, Brazil (1247’05.0”S 3807’58.1”W), UFBA 2056-POR, 23 m depth, ii.2006, coll. (same collector as holotype). (2) Salvador, Bahia State, Brasil (12o58’50.6”S 38o22’44.3”W), UFBA 2904-POR, 35 m depth, i.2009, coll. (same collector as holotype). (3) Todos os Santos Bay, Salvador, Bahia State, Brazil (1256’46”S 3830’38”W), UFBA 3877-POR, 19 m depth, iii.2010, coll. (same collector as holotype). Todos os Santos Bay, Salvador, Bahia State, Brazil (1257’12”S 3830’26”W), UFBA 3878-POR, 10 m depth, iii.2010, coll. (same as holotype). Todos os Santos Bay, Salvador, Bahia State, Brazil (1257’43”S 3830’34”W), UFBA 3879-POR (12 specimens), 12 m depth, iii.2010, coll. (same as holotype). (4) Ilha Grande, Camamu Bay, Mara, south of Bahia (13o55’08.8”S 38o59’59.3”W), UFBA 2112-POR and UFBA 2400-POR, 16 m depth, viii.2004, coll. (same as holotype).</p><p>Diagnosis. Tetilla pentatriaena sp. nov. is the only species possessing five categories of triaenes, two of choanosomal protriaenes, two of choanosomal anatriaenes, and one of exclusively rhizoidal and quite long anatriaenes (average of 7000 µm).</p><p>Description. The shape can vary from spherical to oval, with a single oscule on the apical side of the sponge (Figs 2 A and C). The body height varies from 3 to 6 mm and the diameter from 1 to 6 mm. On the underside there is a slender rhizoid, generally longer than the sponge body. This structure consists of long spicules surrounded by spongin-like fibres. The consistency of the body is hard, but compressible. The entire surface is quite hispid, with bundles of spicules protruding about 2 mm above, arranged radially and slightly spirally. They feature a light beige color in 80% ethanol. Their color in vivo is unknown.</p><p>Spicules. Megascleres: Oxeas I (Fig. 3A), abundant, smooth, straight, fusiform, with both ends slender and gradually pointed: 700–1400–2100 μm / 10–16.0–28 μm. Oxeas II (Fig. 3B), usually smaller than oxeas I, and sometimes a little thicker. Smooth, slightly curved near the middle region; commonly slightly aniso: 235–650–940 μm / 12–15.0–25 μm. Oxeas III (Fig. 3C), smooth, straight, with unequal ends, one slightly conical and the other very thin: 350–590–900 μm / 3.5–6.0–10.6 μm. Protriaenes I (Fig. 3D), large, also as prodiaenes. Rhabdome: 1080–1435.5–2880 μm / 6.5–8.0–11.5 μm. Cladi are stout, gradually pointed, with equal lengths: 37– 70.5–151 μm / 1.8 4.6 –10.8 μm. Protriaenes II (Fig. 3E), also as prodiaenes. Thin rhabdome: 168–390–560 μm / 0.5–2.5–5.0 μm. Cladi are small, pointed, slender and generally with equally long ends, but sometimes one clade may be larger than the others (usually 1.5 x the smaller ones): 10–20–38 μm / 0.5–1.0–2.5 μm. Anatriaenes I (Fig. 3F and J), are choanosomal, long rhabdome: 1260–2108 –3528 μm, with variable thicknesses along it: in the insertion point of the cladome, 5.5–8.5–10.8 μm; 70 μm distant from the cladome, 1.5–3.2–7.2 μm; and a third measure further down the rhabdome, 3.0–4.5–7.2 μm. Cladi are stout, pointed, forming an angle of 45–60 with the rhabdome: 11–34.5–54 μm / 3.6–8.0–11 μm. Anatriaenes II (Fig. 3G and J), are choanosomal, smaller than anatriaenes I, with slender rhabdomes, becoming gradually thinner from the point of insertion of the cladome: 280–526.5–1300 μm / 3.0–4.0– 7.2 μm. Cladi are also more slender in relation to the first category, forming an angle of nearly 90° with the rhabdome, but with their apices bent further down: 11–14.5–36 μm / 1.5–3.5–7.2 μm. Anatriaenes III (Fig. H), are exclusively rhizoidal, presenting cladomes with similar size and morphology as the anatriaenes I, but differing from those by the length of the rhabdome which is over 7000 μm long. Microscleres: small and distorted sigmaspires with microspined surface (Fig. I): 7–10–16 μm.</p><p>Skeleton. The skeleton is composed of main tracts of radially and slightly spirally oriented spicules, which develop from the center of the sponge body and reach the ectosome, usually piercing it (Fig. 2 B). These tracts are spaced by about 40 μm on the inner portion of the choanosome, becoming denser and more closely spaced toward the center of the choanosome and surface, due to a slight expansion in their width. These main tracts are composed mainly by oxeas I and also by anatriaenes I. The latter are abundant and well attached to the tracts. The cladomes of anatriaenes I are clearly seen one above the other, arranged from the inner portion of the choanosome to the surface of the sponge (Fig. 2 D), with their rhabdomes pointing inward. Anatriaenes II exhibit a similar arrangement as the first category, but are located from the center of the sponge until only the middle portion of the choanosome. Closer to the surface protriaenes I mix up with the main tracts, which are usually abundant and may protrude their cladomes 500 μm beyond the surface. Protriaenes II are abundant and visible on the surface, interspersed to the main tracts. Oxeas II and III are located in the subectosomal region, scattered without an apparent arrangement and located also between the main tracts in the choanosomal region (Fig 2 E). Sigmaspires are abundant and distributed throughout the sponge body. It is common to find foreign material adhered to the surface, like sand-grains. Also common is the presence of foreign material in the choanosome, but in smaller quantities. The rhizoids are formed mostly by anatriaenes III, with their cladomes in the sediment, thus anchoring the sponge in the soft substrate. The rhabdome of the anatriaenes may be coiled inside the choanosome. There are also small amounts of oxeas I, II and III reinforcing the rhizoids.</p><p>Ecology. The collection points on the north coast of Bahia, as well as one located off Salvador, outside Todos os Santos Bay, are in the proximity of sewage outfalls that release organic effluents. In addition to these, another outfall throws inorganic acids and heavy metals nearby; i.e. Millenium submarine out fall. All samples of Tetilla pentatriaena sp. nov. obtained on the north coast originated from the area under the influence of organic waste. The absence of this species in the stretch under the influence of inorganic effluents could not be explained by the nature of substrate, as it is quite similar in both areas, composed by a mix of biodetritic debris with gravel and sand.</p><p>In contrast, the collection point in Camamu Bay was located in an estuary, with a considerable supply of freshwater from nearby rivers. However, due to its depth of 16 m, and location near the exit of the bay, it is likely that salt water predominates. Substrate in this area was a mixture of mud and sand. The substrate at Todos os Santos Bay was where found this species similar to that of the north coast.</p><p>Etymology. The name pentatriaena refers to the characteristic and unique presence of five categories of triaenes.</p><p>Remarks. Tetilla pentatriaena sp. nov. is the only species of Tetilla with five categories of triaenes, two choanosomal protriaenes, two choanosomal anatriaenes, and one exclusively rhizoidal anatriaenes. No species of Tetilla occurring in the Atlantic has more than two categories of oxeas together with two categories of protriaenes or anatriaenes, and one category of sigmaspires. Two of these are Tetilla bonaventura Kirkpatrick, 1902 and T. capillosa Levi, 1967, both possessing two categories of oxeas and two of protriaenes, but with only one category of anatriaenes. Tetilla nimia (Topsent, 1927), also recorded in the Atlantic Ocean, has two categories of oxeas and two of anatriaenes, but only one category of protriaenes. Tetilla euplocamos and T. radiata are recorded from southeastern Brazil, but are both devoid of sigmaspires and therefore differ from Tetilla pentatriaena sp. nov. in this important aspect. Besides this difference, the new species can be distinguished from T. euplocamos and T. radiata by its possession of three categories of oxeas, two of protriaenes and three of anatriaenes, compared to only one category in each spicule type in both previously known species. In comparison with other species of Tetilla in the world, the new species is closer to Tetilla disigmata Levi, 1964, reported from Indonesia, by sharing three categories of oxeas and two of protriaenes, but it can still be easily distinguished in having three categories of anatriaenes and only one of sigmaspires, in contrast to one category of anatriaenes and two of sigmaspires in the Indonesian species.</p></div>	https://treatment.plazi.org/id/038FEF5F2F10FF9B8ED1FC70FA882ED6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fernandez, Julio C. C.;Peixinho, Solange;Pinheiro, Ulisses S.;Menegola, Carla	Fernandez, Julio C. C., Peixinho, Solange, Pinheiro, Ulisses S., Menegola, Carla (2011): Three new species of Tetilla Schmidt, 1868 (Tetillidae, Spirophorida, Demospongiae) from Bahia, northeastern Brazil. Zootaxa 2978: 51-67, DOI: 10.5281/zenodo.206930
038FEF5F2F14FF918ED1F969FCBC2D43.text	038FEF5F2F14FF918ED1F969FCBC2D43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetilla muricyi	<div><p>Tetilla muricyi sp. nov.</p><p>(Figs 4, 5; Tables 2, 3)</p><p>Holotype. UFBA 2568-POR, estuary of Acara river, Camamu Bay, Mara, south of Bahia State, Brazil (13o56’24.0”S – 39o05’04.0”W), 5 m depth, 25.ix.2004, coll. U. S. Pinheiro.</p><p>Paratypes. Estuary of Acara river, Camamu Bay, Mara, south of Bahia State, Brazil (13o56’24.0”S – 39o05’04.0”W), (same collector and depth as holotype), UFBA 2569-POR, 25.ix.2004; UFBA 2570-POR, 19.xii.2004; and UFBA 2571-POR, 07.viii.2005.</p><p>Diagnosis. Tetilla muricyi sp. nov. is devoid of microscleres and has a secondary skeleton composed of oxeas on average 500 μm long and 15 μm thick, predominantly reticulate in the subectosomal and choanosomal regions.</p><p>Description. Spherical and massive sponges with a thick rhizoid at the underside (Fig. 4 A). The body of the sponge is usually slightly flattened perpendicularly to its main axis on opposite sides, thereby having two diameters. Largest diameter is on average 29 mm and the smallest 23 mm. The average height of the sponge body is 28 mm. Variably conspicuous external longitudinal grooves are present, usually 6–10, which are distributed from the base of an apical oscule to the basal portion of the sponge body, near the rhizoids. These grooves are usually shallower closest to the oscule, and slightly deeper near the rhizoids. The apical oscule protrudes slightly above the sponge surface (Fig. 4 B) and bears spicules piercing its outer rim, albeit not in a regular, palisade-like manner. On the underside there is a fairly thick rhizoid for settlement, and among the specimens, the width at the base varied from 16 mm to 26 mm, and the overall length from 15 mm to 65 mm. The surface is slightly rough and hispid. There are some conules formed mainly near the base of the sponge. The consistency is very hard and almost incompressible. The color of the species in vivo is entirely cream, or be cream with dark green parts. Preserved specimens are cream-colored.</p><p>Spicules. Megascleres: Oxeas I (Fig. 5A), smooth, straight, fusiform, provided with equally long and thin ends: 1116–1990 –2772 μm / 11–23–33 μm. Oxeas II (Fig. 5 B and G), smooth, varying from straight to slightly curved in one or two points along their length, usually close to the edges. The ends are thin and usually mucronate: 288–546–864 μm / 7.2–17.5–21.6 μm. Prodiaenes I (Fig. 5 C), rare like-protriaenes, with long rhabdomes uniformly thick: 1584–2650 –4140 μm / 3.5–6.0–9.0 μm. Cladi may have slightly different sizes; major clade: 25–5– 90 μm / 1.8–3.2–5.5 μm. Prodiaenes II (Fig. 5 D), smaller than prodiaenes I, also rare like-protriaenes: 616–1170– 1960 μm / 1.0–2.5–4.0 μm. Thin cladi: 15–19.0–29 μm / 0.7–1.2–1.8 μm. Anatriaenes I (Fig. 5 E), are choanosomal, rhabdomes long and gradually pointed; length 1260–1895 –3060 μm. The rhabdome is thicker immediately below the insertion of the cladome; 5.4–6.8–9.0 μm, tapering abruptly to 3.5–3.8–7.2 μm, about 150 μm below that point, and then becoming slightly thicker again (ca. 6 μm) where tapering gradually to the tip. Cladomes with curved cladi bearing very thin ends: 18–39–58 μm / 3.6–5.0–7.2 μm. Anatriaenes II (Fig. 5 F), exclusively rhizoidal, similar to anatriaenes I in overall morphology, but distinguished by the length of the rhabdomes (over 6000 μm).</p><p>Skeleton. Radial skeleton composed of bundles of oxeas I mixed with anatriaenes I extending from the center of the chaonosome toward the ectosome (Fig. 4 C), where oxeas can pierce the surface up to about 50 μm. The cladomes of the anatriaenes reach the subectosomal region, but only few of these trespass it (Fig. 4 D). The prodiaenes I and II are mixed with the main tracts in the ectosome, with their cladomes normally protruding up to 200 μm beyond the surface (Fig. 4 D – F). Prodiaenes II may form tracts among the main tracts in the ectosomal region. Oxeas II are disposed throughout the choanosome and subectosomal region, exhibiting a reticulate or sometimes disorganized arrangement. These oxeas form a halichondroid secondary skeleton (Fig. 4 C) and as well as the first category, they can also reach the surface (Fig. 4 D and E). Foreign particles such as sand-grains can be found in the choanosome as well as adhered to the ectosome. There are channels throughout the body of the sponge, showing a wide variation in diameter (ca. 100 to over 1000 μm). Oxeas II surround these channels, both in the choanosome and the subectosomal region. The rhizoids are formed mostly by anatriaenes II and to a lesser extent by oxeas II, both packed by spongin. The cladomes of these anatriaenes were inserted in the sediment, thus anchoring the sponge to the soft substrate. Microscleres were not found in either the body or the rhizoids.</p><p>Ecology. Tetilla muricyi sp. nov. occurs at several localities along river Acara, up to approximately 1.5 km from its mouth. This species is exposed to large fluctuations of salinity (0.8 at low tide to 32 at high tide—data obtained by U.S. Pinheiro and M.C. Guerrazzi) and we can therefore classify it as euryhaline. The new species occurs in extensive sympatry with Craniella quirimure in Camamu Bay.</p><p>Etymology. The species name is in honour of Professor Guilherme Ramos da Silva Muricy, who greatly contributed to an expanded knowledge of the Brazilian sponge fauna.</p><p>Remarks. Prodiaenes markedly predominate over protriaenes in Tetilla muricyi sp. nov. . Therefore, we consider the former a diagnostic feature of the new species. Its secondary halichondroid skeleton is unique in the genus, and thus of considerable diagnostic value. Tetilla muricyi sp. nov. is differentiated from T. pentatriaena sp. nov. by presenting one less category of oxeas and anatriaenes and, in addition, a category of prodiaenes and a unique secondary halicondroid skeleton. Worldwide there are seven species of Tetilla devoid of sigmaspires, three of which occurring in the Tropical Atlantic: Tetilla euplocamos, T. radiata and T. truncata Topsent, 1890 . The first two of these have only one category of oxeas and protriaenes, plus a category of anatriaenes in T. radiata . The third species has only one category of styles and anatriaenes. Tetilla muricyi sp. nov. differs from these three species by its possession of two categories of oxeas and two of anatriaenes, besides its prodiaenes and secondary halichondroid skeleton. Other species lacking sigmaspires were reported from Indonesia ( Tetilla enoi Brøndsted, 1934, T. pedifera Sollas, 1886 and T. schulzei Kieschnick, 1898) and South Australia ( T. globosa (Bear, 1906)) . Tetilla muricyi sp. nov. differs from these as they all possess only one category of oxeas, protriaenes and anatriaenes, but also by their quite unrelated biogeography and non-mangrove habitat.</p></div>	https://treatment.plazi.org/id/038FEF5F2F14FF918ED1F969FCBC2D43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fernandez, Julio C. C.;Peixinho, Solange;Pinheiro, Ulisses S.;Menegola, Carla	Fernandez, Julio C. C., Peixinho, Solange, Pinheiro, Ulisses S., Menegola, Carla (2011): Three new species of Tetilla Schmidt, 1868 (Tetillidae, Spirophorida, Demospongiae) from Bahia, northeastern Brazil. Zootaxa 2978: 51-67, DOI: 10.5281/zenodo.206930
038FEF5F2F1EFF928ED1FAE9FC5A2970.text	038FEF5F2F1EFF928ED1FAE9FC5A2970.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetilla rodriguesi	<div><p>Tetilla rodriguesi sp. nov.</p><p>(Figs 6, 7; Table 3)</p><p>Holotype. UFBA 2046-POR, Goi Island, Camamu Bay, Mara, south of Bahia State, Brazil, (13o56’01.5”S – 38o59’31.6”W), &lt;5 m depth, viii.2004, coll. W. Andrade.</p><p>Diagnosis. Tetilla rodriguesi sp. nov. is devoid of microscleres, has three categories of oxeas, two of trichoidal protriaenes and one of rhizoidal anatriaenes.</p><p>Description. The sponge is pear-shaped, 4 mm in diameter and 9 mm in height. At the posterior end there is a rhizoid for attachment of the sponge (Fig. 6 A). This structure is long and slender, with a length of 13 mm and diameter of 0.5 mm. The consistency of the sponge body is quite soft. Its surface is slightly hispid, except the region near the rhizoids. There are small pores arranged on the side of the sponge, with an average diameter of 150 μm. No oscule was observed. The color of the species was not registered in vivo; and is dark brown after preservation in ethanol.</p><p>Spicules. Megascleres, Oxeas I (Fig. 7 A), straight, smooth, fusiform, with thin ends, larger and thicker than the oxeas II, 648– 1097 –2016 μm / 4.7–8.8–10.8 μm. Oxeas II (Fig. 7 B), straight and smooth, with one end slightly conical and the other long and quite thin, the latter is flexuous, 294–452–672 μm / 3–3.5–4 μm. Oxeas III (Fig. 7 C), isoactinal, slight to moderately curved in the middle portion, 130–417–560 μm / 1.8–5.0–7.2 μm. Trichoidal protriaenes I (Fig. 7 D), larger and thicker than the second category, long rhabdomes tapering gradually from the insertion point of the cladome, 1000–1860–2520 μm / 3.6–4.0–5.5 μm; cladi with similar dimensions, sharp and thinner than rhabdomes, 50–78.8–105 μm / 1.5–1.8–2.0 μm. Trichoidal protriaenes II (Fig. 7 E), rhabdomes tapering gradually to their points, 154–279–532 μm / 0.5–0.7–1.0 μm; cladomes with thin cladi of similar or different lengths (larger up to 2 x the length of the smaller, sometimes three different lengths); cladi, when these are of equal dimensions, 14.5–36.5–90 μm / 0.4–0.5–0.7 μm. Anatriaenes (Fig. 7 F), exclusively rhizoidal; rhabdome relatively thin and long, over 6000 μm, provided with three main diameters; the largest one immediately under the insertion of the cladome, 4 μm on average; about 60 μm further along its extension, it quickly thins out to 1.7 μm on average; then reexpanding back to about 3.5 μm; wherefrom it tapers gradually to the point; stout cladome, often strongly curved, cladi with very fine tips, 25–30– 45 μm / 3.0–3.5 – 5.5 μm.</p><p>Skeleton. The skeleton is composed of main radial tracts of oxeas I, starting from the center of the sponge toward the ectosome, and often reaching the surface, piercing it 150–200 μm on average (Fig. 6 B). Oxeas II are intermingled with these bundles close to the ectosome, thus rendering this area denser (Fig. 6 C). Protriaenes I and II are mixed with the main tracts in the ectosome, usually with their cladomes piercing the surface slightly. Oxeas III occur in the portion between the choanosome and the subectosomal region, do not have a visible arrangement being interspersed in a criss-cross manner with the radial bundles. The choanosome is crossed by canals with a diameter of 100–200 μm. Some foreign particles, such as sand grains, may be found in the choanosome. Microscleres of any kind are absent both in the body and in the rhizoids of the sponge.</p><p>Ecology. Tetilla rodriguesi sp. nov. was collected at the northwest side of Goi Island, southwest of Campinho Island, Mara Peninsula. This area is a typical mangrove with muddy substrate, but sandy bottoms are also present in the area. The waters around Goi Island are exposed to large amounts of nutrients as well as a considerable fluctuation of salinity, as a consequence of their proximity to river Mara and other smaller rivers in the area. Although the salinity in this collection point has not been measured, the new species appears to be euryhaline.</p><p>Etymology. The name of the new species is a tribute to Professor Sergio de Almeida Rodrigues, who contributed significantly to the science of marine biology in Brazil.</p><p>Remarks. Tetilla rodriguesi sp. nov. is devoid of sigmaspires, similar to Tetilla muricyi sp. nov. and the seven other species of the genus mentioned before. This new species shares with Tetilla muricyi sp. nov. the mangrove habitat, but can be distinguished through a series of morphologic traits: the possession by the former of three categories of oxeas, only one of anatriaenes and a strictly radial skeleton. Tetilla muricyi sp. nov. has two categories of oxeas and of anatriaenes, a single category of prodiaenes and a criss-crossed secondary skeleton. The new species differs from Tetilla pentatriaena sp. nov. by presenting a single category of anatriaenes, and by its lack of sigmaspires. Regarding the species that are devoid of microscleres ( Tetilla euplocamos, T. globosa, T. radiata, T. truncata, T. enoi, T. pedifera and T. schulzei) Tetilla rodriguesi sp. nov. presents a distinct spiculation, the main differences being its three categories of oxeas and mangrove habitat.</p></div>	https://treatment.plazi.org/id/038FEF5F2F1EFF928ED1FAE9FC5A2970	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fernandez, Julio C. C.;Peixinho, Solange;Pinheiro, Ulisses S.;Menegola, Carla	Fernandez, Julio C. C., Peixinho, Solange, Pinheiro, Ulisses S., Menegola, Carla (2011): Three new species of Tetilla Schmidt, 1868 (Tetillidae, Spirophorida, Demospongiae) from Bahia, northeastern Brazil. Zootaxa 2978: 51-67, DOI: 10.5281/zenodo.206930
038FEF5F2F1DFF928ED1FE25FA562DF8.text	038FEF5F2F1DFF928ED1FE25FA562DF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetilla	<div><p>Key to the Tetilla species occurring in the Atlantic Ocean</p><p>1a. Oxeas present....................................................................................... 2</p><p>1b. Oxeas absent; one size of protriaenes and anatriaenes; microscleres absent................................ T. truncata</p><p>2a. One category of oxeas.................................................................................. 3</p><p>2b. Two or more categories of oxeas.......................................................................... 9</p><p>3a. Protriaenes present................................................................................... 4</p><p>3b. Protriaenes absent; a single category of oxeas, anatriaenes and sigmaspires............................. T. sigmophora</p><p>4a. One category of protriaenes............................................................................. 5</p><p>4b. Two categories of protriaenes; sigmaspires centrotylote.............................................. T. geniculata</p><p>5a. Regular sigmaspires present............................................................................ 6</p><p>5b. Regular Sigmaspires absent............................................................................. 7</p><p>6a. Anatriaenes present.................................................................................... 8</p><p>6b. Anatrienes absent; oxeas of 2326 / 24 µm; protriaenes with 197 / 5 µm (larger clade) and 51/ 4 µm (smaller clade); sigmaspires 24–28 µm................................................................................... T. sandalina</p><p>7a. Oxeas (medium length/ medium thickness): 1.100 µm / 7.6 µm; triaenes (rhabdome / cladome): protriaenes&gt; 3.500 / 49–194 µm; anatriaenes&gt; 5.600 µm/ 29–50................................................................ T. radiata</p><p>7b. Oxeas (medium length/ medium thickness): 2.000 µm/ 20 µm; protriaenes not measured; anatriaenes absent.. T. euplocamos</p><p>8a. Oxeas less than 1.500 µm; triaenes (rhabdome, cladome length/thickness): protriaenes 1800–2000/ 4 µm, 35–70/ 3 µm; anatri- aenes 2500–3.000/ 5 µm, 30–40 µm; sigmaspires 10 µm............................................ T. pedonculata</p><p>8b. Oxeas greater than 3.000 µm; protriaenes 2.000–3.000 µm; sigmaspires 12 µm............................. T. africana</p><p>9a. Two categories of oxeas............................................................................... 10</p><p>9b. Three categories of oxeas.............................................................................. 13</p><p>10a. Three categories of triaenes; sigmaspires present............................................................ 11</p><p>10b. Four categories of triaenes (two of protriaenes and two of anatriaenes); sigmaspires absent and oxeas form a halichondroid sec- ondary skeleton......................................................................... T. muricyi sp. nov.</p><p>11a. Two categories of protriaenes and one single of anatriaenes................................................... 12</p><p>11b. A single category of protriaenes (rhabd. 5.300 µm, larger clade&gt; 140 µm, smaller clades 80–95 µm) and two of anatriaenes (I: rhabd 6.000, larger cladi 225–240/20 µm; II: 1.150 µm, 55–110 µm); sigmaspires 34–40 / 4 µm................. T. nimia</p><p>12a. Oxeas I with a maximum length of 5.000 µm; protriaenes I with length up to 2.000 µm and protriaenes II with up to 300 µm; anatriaenes of 5.000–8.000 µm in length by 10–15 µm thick......................................... T. bonaventura</p><p>12b. Oxeas I greather than 5.500 µm;; protriaenes I with length greater than 5.000 µm (up to 10.000 µm) and protriaenes II with up to 1.000 µm; anatriaenes of 12.000 µm in length by 35–40 µm thick..................................... T. capillosa</p><p>13a. Five categories of triaenes (2 types of protriaenes and three of anatriaenes); one single category of sigmaspires............................................................................................. T. pentatriaena sp. nov.</p><p>13b. Three categories of triaenes (2 of protriaenes and one of anatriaenes); sigmaspires absent............ T. rodriguesi sp. nov.</p></div>	https://treatment.plazi.org/id/038FEF5F2F1DFF928ED1FE25FA562DF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fernandez, Julio C. C.;Peixinho, Solange;Pinheiro, Ulisses S.;Menegola, Carla	Fernandez, Julio C. C., Peixinho, Solange, Pinheiro, Ulisses S., Menegola, Carla (2011): Three new species of Tetilla Schmidt, 1868 (Tetillidae, Spirophorida, Demospongiae) from Bahia, northeastern Brazil. Zootaxa 2978: 51-67, DOI: 10.5281/zenodo.206930
