taxonID	type	description	language	source
038C87B5FFB7B11253ECFD03FBBB395D.taxon	description	TYpe species: Pachylaelaps (Longipachylaelaps) anatolicus Özbek, 2015.	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFB7B11253ECFD03FBBB395D.taxon	diagnosis	Diagnosis (adults). Dorsal idiosoma (Fig. 1). Dorsal shield suboval, elongate, with almost parallel lateral margins and 30 pairs of smooth and needle-like setae; setae mostly similar in length; dorsocentral setae J 2 in normal posterior position to setae J 1; J 5 relatively short but normally developed. Opening of poroids gdS 4 minute, slit-like (poroids gdZ 1 with normal circular opening). Ventral idiosoma (Fig. 2). Presternal platelets absent. Male with holoventral shield bearing nine pairs of setae (plus three circum-anal setae close to anus). In female, sternal shield fused to metasternal and endopodal shields, with four pairs of setae; genitoventral shield oblong, subpentagonal, with two pairs of setae (st 5 and Jv 1); anal shield free, subtriangular, closely abutting genitoventral shield. Peritremes and peritrematal shields welldeveloped, peritrematal shields fused to exopodal shields, produced posteriorly well behind coxae IV in female. Metapodal platelets free on soft integument (in female) or fused to holoventral shield (in male). Lateral and opisthogastric soft integument with 15 pairs of setae: seven pairs of marginal setae (r-R) and eight pairs of opisthogastric setae (JV 2 ‒ JV 5 and ZV 2 ‒ ZV 5) present.	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFB7B11253ECFD03FBBB395D.taxon	description	Spermathecal apparatus. Sperm induction system of female associated with coxae IV; paired tubular components normally developed, flattened and weakly sclerotised for whole length; inner sperm ductus thin and usually in a form of wavy line (Fig. 20). Legs (Figs 4, 12 ‒ 14, 18, 19). Legs normal in length, shorter than idiosoma. Female tarsus II (Fig. 18) with two spur-like distal setae (pl 1 and pl 2), male tarsus II with only one distal seta (pl 1) modified into spur (Figs 13, 19); tarsus IV in both adults with seta pl 2 shortened and thickened, spur-like (Fig. 14). Female femur II with ventral spur-like projection associated with a seta (Fig. 4). Genu I with 13 setae (2 - 6 / 3 - 2). Male legs II spurred (Figs 12, 19). Gnathosomal structures (Figs 7 ‒ 11, 15 ‒ 17, 21 ‒ 23). Corniculi long, horn-like (Fig. 7). Epistome with truncate or laterally concave base, and smooth, narrow and long central projection; anterior margin of base finely denticulate; projection lanceolate, slightly (in female) or markedly (in male) widened basally, with pointed, bifid or divided apex (Figs 3, 9, 21 ‒ 23). In female, chelicerae pincette-like; cheliceral digits narrow, elongate, polydentate, each with strong and sharp terminal hook, and additional lateral row of teeth close to most robust distal tooth (Figs 5, 16). In male, cheliceral digits slender, less elongate than female, terminally strongly hooked, each with one robust distal tooth (Figs 8, 15); male spermatodactyl narrow, long, regularly tapered and slightly curved, sabreshaped. In male, palptibia with three petal-like projections on ventral and outer lateroventral surface (Figs 10, 17), and palpfemur with rounded tubercle on inner proximal laterodorsal surface and small pointed tubercle on inner distal lateroventral surface (Fig. 11).	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFB7B11253ECFD03FBBB395D.taxon	etymology	Etymology. The name of the new subgenus is derived from the Latin longus, referring to the elongate idiosoma, in combination with the modified suffix - pachys, as used in the names of other pachylaelapine taxa. Gender masculine. Notes on the subgenus. The subgenera Longipachys and Longipachylaelaps are reliably distinguished from the other subgenus Pachylaelaps mainly by the presence of normal needle-like dorsal setae J 5, and only one pair of slit-like poroid structures (gdS 4) placed on the posterior lateral margins of the dorsal shield. In the subgenus Pachylaelaps, setae J 5 are vestigial, and the posterolateral dorsal shield margin bears two pairs of slit-like poroids, gdZ 1 and gdS 4. Despite their striking similarities in general appearance, idiosomal and leg chaetotaxy and some expressions of sexual dimorphism (see some features on legs, palps and chelicerae), Longipachys and Longipachylaelaps may be easily distinguished by the following characters — in the new subgenus, posterolateral seta pl 2 on tarsus IV is modified into a spur (uniform with other setae in Longipachylaelaps); female chelicera is polydentate, each digit with 10 ‒ 20 teeth and a prominent terminal hook (mono- or bidentate in Longipachylaelaps, with terminal hooks inconspicuous, often split and never curved backward); central projection of epistome is regularly narrowed toward its apex, lanceolate (with subdistal neck and widened anterior margin in Longipachylaelaps); and opisthogastric setae ZV 1 are absent on lateroventral soft integument (present in Longipachylaelaps).	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFB5B11453ECF8BDFCFB3EFD.taxon	materials_examined	New specimens examined. E Turkey, Tunceli Province, Pülümür District, Uzuntarla Village, leaf litter under Quercus sp., 27 April 2016 (two females and seven males), 25 May 2017 (three females and one male); Alacık Village, leaf litter under Quercus sp., 27 April 2016 (three females and two males). The specimens are deposited at the Acarology Laboratory of Erzincan University, Turkey; except two specimens (female and male) sent to the mite collection of the Institute of Zoology, Slovak Academy of Sciences, Bratislava. Amendments to original description. Female. The female was adequately described in detail by Özbek (2015), and his original description needs only additional metric data, with some marginal changes to the detected variation, because some of our newly collected specimens are smaller and more variable in some characters than stated in the original description. Dorsal idiosoma (Fig. 1). Dorsal shield 685 ‒ 810 µm long and 370 ‒ 445 µm wide (length / width 1.63 ‒ 1.89), reticulate on surface; net-like pattern usually with additional punctation (punctation sometimes very weakly indicated, or absent in some specimens). Revised lengths of some setae as follows: j 1 38 ‒ 48 µm, j 2 58 ‒ 76 µm, j 3 70 ‒ 90 µm, J 4 75 ‒ 96 µm, J 5 21 ‒ 31 µm (J 4 / J 5 2.88 ‒ 4), z 2 65 ‒ 90 µm, z 4 70 ‒ 90 µm, z 5 46 ‒ 63 µm, s 2 52 ‒ 80 µm, s 4 70 ‒ 90 µm, s 6 60 ‒ 80 µm, S 1, S 3 and S 4 50 ‒ 80 µm, S 5 60 ‒ 83 µm, and r 5 46 ‒ 80 µm.	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFB5B11453ECF8BDFCFB3EFD.taxon	description	Ventral idiosoma (Fig. 2). Revised measurements of some structures as follows: sternal shield length 220 ‒ 280 µm, genitiventral shield width 214 ‒ 255 µm, L / W of genitiventral shield 1.12 ‒ 1.23, anal shield length 99 ‒ 132 µm, anal shield width 150 ‒ 185 µm, L / W of anal shield 0.6 ‒ 0.78. Lengths of some selected setae on venter as follows: st 1 70 ‒ 83 µm, st 2 and st 3 50 ‒ 70 µm, st 5 60 ‒ 70 µm, and JV 1 75 ‒ 87 µm. Punctation of net-like meshes on genitiventral shield sometimes very weakly indicated or absent. Lateral and ventral soft integument with 15 pairs of setae (r 6, R 1, R 3 ‒ R 7, ZV 2 ‒ ZV 5, JV 2 ‒ JV 5), setae ZV 1 absent. Gnathosomal structures (Figs 3, 5, 16, 21, 22). Central projection of epistome lanceolate, apically with simple point or furcate. Movable digit 150 ‒ 170 µm long and 54 ‒ 59 % of the length of medial segment (270 ‒ 295 µm); fixed digit 135 ‒ 145 µm long (from dorsal seta to apex). For some other features of epistome and chelicerae see the subgeneric diagnosis above. Legs (Figs 4, 18). Femur II with small spur-like projection on ventral surface, as shown in Fig. 4. Male. Dorsal idiosoma. Dorsal shield 610 ‒ 660 µm long, 360 ‒ 415 µm wide, oblong and ornamented with netlike pattern. Chaetotaxy and other characters on dorsum as in female (Özbek 2015). Ventral idiosoma (Fig. 6). Holoventral shield present, 540 ‒ 620 µm long, regularly reticulate on whole surface, and bearing nine pairs of setae (in addition to three circum-anal setae), namely st 1 ‒ st 5, JV 1 ‒ JV 3, and ZV 2. Parapodal portions of peritrematal shields fully fused to lateral margins of holoventral shield. Lateral and opisthogastric soft integument with 12 pairs of setae (seven pairs of marginals and five pairs of opisthogastric setae). Free metapodal platelets absent, completely incorporated into holoventral shield. Peritremes with anterior ends reaching close to setae z 1. Gnathosomal structures (Figs 7 ‒ 11, 15, 17, 23). Ventral surface of hypostome as in Fig. 7. Hypostomal setae h 1 longest, h 2 similar in length to palpcoxal setae (pc), both shorter than h 3; corniculi horn-like and lanceolate; deutosternal groove with six transverse rows of denticles. Epistome with base and central projection as in female, but central projection shorter, with more widened serrate subtriangular base, thicker apical part and furcate apex (Figs 9, 23). Cheliceral digits slender but not so markedly elongate as in female, monodentate; movable digit around 85 µm long, with prominent distal tooth, curved distal hook and sabre-like spermatodactyl; spermatodactyl 158 ‒ 163 µm in length, almost two times longer than movable digit (1.85 ‒ 1.91); fixed digit with dentition similar to that in movable digit; pilus dentilis setiform, slim and minute (Figs 8, 15). Palps with three-tined apotele on tarsi and three petal-like projections on tibiae (Figs 10, 17), and rounded tubercle on femora (Fig. 11). Legs (Figs 12 ‒ 14, 19). Chaetotaxy as in female. Segments of legs II ventrally spurred, as shown in Figs 12 and 19; tarsi II with only one spur-like seta, pl 1 (Figs 13, 19).	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFB3B11853ECF8AAFD3D3A8D.taxon	materials_examined	Type specimens examined. Holotype female — NE Turkey, Erzincan Province, Geyikli Village (39 ° 43 ' N, 39 ° 40 ' E), altitude 1430 m, leaf litter under Salix sp., 19 May 2012. Paratypes: one female (deposited in IZ SAV) — with the same data as in holotype; two females — with the same data as in holotype, 30 May 2017; five females (of which one deposited in IZ SAV) — E Turkey, Tunceli Province, Pülümür District, Uzuntarla Village, leaf litter under Quercus sp., 25 May 2017. The holotype and paratypes are deposited at the Acarology Laboratory of Erzincan University, Turkey; two paratypes are deposited at the Institute of Zoology, Slovak Academy of Sciences, Bratislava (indicated as IZ SAV).	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFB3B11853ECF8AAFD3D3A8D.taxon	diagnosis	Diagnosis. Dorsal setae relatively short: dorsocentral j- and J-setae with their tips not exceeding bases of following seta. Openings of slit-like glandular poroids gdZ 1 and gdS 4 with separate position on posterolateral margins of dorsal shield. Prestigmatic section of peritremes long, with anterior tips reaching dorsal surface between setae z 1 and z 2. Soft integument with 15 pairs of setae. Cheliceral digits polydentate. Trochanter III with large posterolateral projection. Sperm induction system with tubular components well developed: tubes hyaline, flattened, progressively narrowed apically and usually curved distally.	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFB3B11853ECF8AAFD3D3A8D.taxon	description	Description. Female. Dorsal idiosoma (Fig. 25). Dorsal shield 715 ‒ 815 µm long and 405 ‒ 455 µm wide, suboval and oblong (length / width 1.7 ‒ 1.96), delicately and evenly reticulate on whole surface, and bearing 30 pairs of smooth and needleshaped dorsal setae. Setae J 5 strongly reduced in length, vestigial microsetae; setae z 1 short, 20 ‒ 26 µm long; other setae relatively well developed, similar in length and uniform. Length and spacing of some selected dorsal shield setae as follows: j 1 45 ‒ 53 µm, j 4 ‒ j 6, and z 5 50 ‒ 60 µm, other setae 60 ‒ 80 µm, length of j 5 similar or shorter than distance between j 5 - z 5, and length of J 1 similar or shorter than distance between J 1 - J 2. Two posterolateral poroid structures (gdZ 1 and gdS 4) with their openings slit-like, separate and placed approximately between setae Z 2 and S 4. Ventral idiosoma (Fig. 26). Sternal shield 277 ‒ 290 µm long, proportionally 1.02 ‒ 1.15 longer than genitiventral shield, with slightly concave anterior margin and two small projections close to bases of sternal setae st 1. Genitiventral shield slightly longer than wide or subequal in length and width (length 235 ‒ 278 µm, width 230 ‒ 277 µm, length / width 1 ‒ 1.07). Anal shield subtriangular, 111 ‒ 128 µm long and 138 ‒ 172 µm wide (length / width 0.65 ‒ 0.80); anus with three circum-anal setae situated close to posterior margin of shield. Peritremes normally developed, relatively long, with anterior end reaching dorsal surface between setae z 1 and z 2. Peritrematal shields with weak longitudinal sculptural lines, other ventral shields distinctly and evenly reticulate and usually delicately punctate on surface. Metapodal platelets minuscule, free and well separate from peritrematal shields. Lateral soft integument with seven pairs of marginal setae (r 6, R 1, R 3 ‒ R 7). Ventral soft integument with eight pairs of opisthogastric setae (JV 2 ‒ JV 5, ZV 2 ‒ ZV 5). Ventrally placed setae similar to those on dorsal idiosoma. Spermathecal apparatus (Figs 28, 37, 38). Tubes of sperm induction system relatively well developed but weakly sclerotised, hyaline, flattened, with broader base and narrowed distal part; basal section widely abutting to inner anterolateral margin of coxa IV; distal part usually curved and irregularly formed, sometimes straightened inward. Gnathosomal structures (Figs 24, 27, 29, 30, 34). Corniculi elongated and horn-like; laciniae densely pilose, lanceolate, longer than corniculi; deutosternum slightly widened anteriorly, with six transverse rows of denticles; subcapitular setae smooth and needle-shaped: setae h 1 longest, h 2 similar in length to pc, and shorter than h 3 (Fig. 29). Epistome with wide, subtriangular and marginally serrate base, and central projection; projection long and narrow, smooth, with inconspicuous neck and bifurcate brush-like apex; each furca delicately serrate apically, with a row of 1 ‒ 5 minute prongs (Fig. 30). Chelicerae pincer-like, with relatively short and robust polydentate digits; fixed digit 94 ‒ 100 µm long, having bidentate terminal hook, one robust distal tooth associated with minute pilus dentilis, and 10 ‒ 15 small and sharp teeth on masticatory area; movable digit 92 ‒ 105 µm long, 42 ‒ 48 % of the length of medial segment (210 ‒ 220 µm), armed with simple hook and 7 ‒ 9 sharp teeth, including the biggest one with distal position (Figs 24, 27, 34). Legs (Figs 31 ‒ 33, 35, 36). Leg setation normal for genus (Mašán, 2007). Tarsus II with two spur-like distal setae, pl 1 and pl 2 (Figs 32, 35). Femur II with small spine-like process and associated seta on distal ventral surface (Figs 33, 35). Trochanter III with conspicuous bump-shaped projection directed backward (Figs 31, 36).	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFB3B11853ECF8AAFD3D3A8D.taxon	etymology	Etymology. The specific name of the new species is derived from the Latin words " arma " (instrument of war, arm) and " gerō " (carry or bear), and it alludes to the fact that the chelicerae of these mites are heavily armed with sharp teeth. Taxonomic notes. Mašán (2007) divided the European members of the subgenus Pachylaelaps into five clusters of species. In his classification scheme, Pachylaelaps (Pachylaelaps) armiger could be classified as a species of the bellicosus group, which is representated by Pachylaelaps bellicosus Berlese, 1920 and Pachylaelaps multidentatus Evans & Hyatt, 1956. The bellicosus group is characterised by the separate position of slit-like poroid structures gdZ 1 and gdS 4 on the dorsal shield, multidentate cheliceral digits, weakly sclerotised and hyaline tubiform structures of sperm induction system, and males apparently absent. The new species can be easily distinguished from both above mentioned congeners especially by the presence of large bump-like projection on posterior surface of trochanter III. This conspicuous structure is easily detectable and not found in its relatives in the species group, including Pachylaelaps mandibularis Moraza & Peña, 2005 previously synonymised with P. bellicosus by Mašán in 2007 (Moraza, pers. comm).	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFBFB11853ECFCBAFC013DEE.taxon	description	Pachylaelaps evansi COsta, 1971: 255. Pachylaelaps (Pachylaelaps) evansi. — Mašán & HallidaY, 2014: 35. Specimens examined. E Turkey, Tunceli Province, Pülümür District, Uzuntarla Village, leaf litter under Quercus sp., 25 May 2017 (two females). The specimens are deposited at the Institute of Zoology, Slovak Academy of Sciences, Bratislava.	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
038C87B5FFBFB11853ECFCBAFC013DEE.taxon	discussion	Remarks. The species shows two atypical characters for Pachylaelaps, namely: (1) posterolateral areas of dorsal shield only with poroids gdS 4 modified into a widely slit-like structures (gdZ 1 uniform with other poroids on dorsum), (2) genu of legs I with only 12 instead of 13 setae (pd 3 absent). Apart from these character states, the original description and illustrations given for this species by Costa (1971) do not require special amendment, except for seven pairs of r / R-setae on soft lateromarginal integument (in addition to eight pairs of opisthogastric setae on ventral soft integument) which were not specified and illustrated in his descriptive paper. Also tubular structures of the sperm induction system were not mentioned in the description but they are probably unsclerotised, rudimentary or absent because we were unable to detect them in our specimens. Pachylaelaps evansi is quite well recognisable species, described on the basis of both adult stages, and it has not been recorded for almost 50 years from its discovery in Israel. This is the first record of the species in Turkey.	en	Özbek, Hasan Hüseyin, Mašán, Peter (2018): A new species and subgenus of Pachylaelaps Berlese from Turkey (Acari: Pachylaelapidae). Zootaxa 4418 (5): 481-492, DOI: 10.11646/zootaxa.4418.5.5
