taxonID	type	description	language	source
038C87DFFFD6FFCC85F064D5FD8DEE48.taxon	etymology	Etymology: — The specific epithet “ himalayana ” refers to Himalayas of China and Azad Jammu & Kashmir. Diagnosis: — Characterized by subumbonate pileus (7 – 35 mm), smaller fusiform to utriform necropigmented cheilocystidia and larger basidiospores (av. Q = 1.69). Holotype: — PAKISTAN, Azad Jammu and Kashmir, Bagh, Saliyan, Nari, at 1 950 m a. s. l., under mixed forest of Pinus wallichiana A. B. Jacks. (1938: 85), Cedrus deodara (Roxb. Ex D. Don) G. Don (1830: 23637) and Abies pindrow Spach (1841: 423), on soil, solitary, 01 Sept. 2021, Rubab Khurshid, holotype Herb. LAH 37697; GenBank accessions ITS: OQ 449902, LSU: OQ 448897; mtSSU: OQ 448900. Description: — Basidiomata small to medium sized. Pileus 7 – 35 mm diam., conical when very young, then convex or plano-convex when mature, becoming flat with age, with an indistinct umbo or without when young, some cases subumbonate to plane with age; margin incurved when young, becoming decurved with age, longer than lamellae; surface dry, dull, covered with small scales arranged in a dense and radial pattern, floccose, finely felt pileus; orangish brown (5.1 YR 6 / 6.1) with orange red (9.3 R 3.9 / 9.3) sub-umbonate center when young, becoming brown (9.5 R 7.7 / 3.3), darker towards the disc and fades outwards; filamentous veil remnants present when young. Lamellae 1 – 2 mm wide, adnexed to emarginate, subcrowded, with three or four tiers of lamellulae, yellowish white (4 A 2) to brown (6 D 4), edges concolor, broad and indistinctly fimbriate. Stipe 12 – 30 × 2 – 3 mm, solid, equal, cylindrical to subcylindrical, widen upward, surface dry, covered with minute, dense, appressed and longitudinally silky-fibrillose, pale yellowish brown (5 B 3) when young, becoming yellowish brown (5 b 5); white at base, partial veil present when very young. Context fresh (5 A 2) in pileus, dirty white (5 A 1) to cream, 0.5 – 1 mm thick under the umbo, 2.5 – 4 mm thick at midradius. Odor unpleasant, somewhat bitter. Basidiospores [120 / 4 / 4] (8.0 –) 8.3 – 10.3 (– 11.3) × (4.4 –) 4.7 – 6.2 (– 6.9) µm, avL × avW = 9.47 × 5.6 µm, Q = 1.4 – 1.9 (2.3), avQ = 1.69, light brownish to yellowish in 5 % KOH, oval-oblong or obovate, ellipsoidal, slightly thick-walled, smooth, apiculus small and indistinct. Basidia (30.2 –) 33.1 – 38.3 (– 42.8) × (7.1 –) 8.4 – 10.2 (– 10.9) µm, avL × avW = 36.2 × 9.4 µm, narrowly clavate to clavate, thin-walled, primarily 4 - sterigmate, less often 2 - sterigmate, acute, 3 – 6 µm in length, hyaline in 5 % KOH, non-guttulate, becoming necropigmented when mature. Pleurocystidia absent. Lamellae edge sterile. Cheilocystidia (13.8 –) 14.9 – 19.8 (– 21.7) × (4.1 –) 4.3 – 7.1 (– 7.4) µm, avL × avW = 17.7 × 5.9 µm, thick-walled, fusiform or utriform; walls smooth, hyaline in 5 % KOH. Pileipellis a cutis of elongate, apically cylindrical terminal elements with some short less clavate elements interspersed, hyphae (5.1 –) 6.2 – 7.7 (– 8.8) µm in diameter, light brownish in 5 % KOH, frequently septate, crowded, regular, unbranched, thin-walled, clamp connections frequently present, pileocystidia absent. Stipitipellis a cutis, composed of parallel hyphae, (6.7 –) 7.3 – 9.3 (– 11.2) µm in diameter, yellowish in 5 % KOH, frequently septate, unbranched, regular, with narrowly cylindrical terminal ends, clamp connection present frequently, Caulocystidia (15.4 –) 17.7 – 25.4 (– 18.9) × (8.3 –) 9.3 – 10.7 (– 11.7) µm, present only at stipe apex, cylindrical to fusiform, broader than lower hyphae, smooth, empty, less frequent, hyaline in 5 % KOH. Habit and Habitat: Found on soil under conifers.	en	Naseer, Arooj, Khurshid, Rubab, Fan, Yu-Guang, Khalid, Abdul Nasir (2024): Two new species of Mallocybe from Himalayan forests of Asia. Phytotaxa 640 (3): 255-264, DOI: 10.11646/phytotaxa.640.3.3, URL: http://dx.doi.org/10.11646/phytotaxa.640.3.3
038C87DFFFD6FFCC85F064D5FD8DEE48.taxon	distribution	Distribution: It is collected from Yunnan, China and Bagh, Azad Jammu & Kashmir, Pakistan. Additional materials examined: CHINA, Yunnan, Dali Bai Autonomous Prefecture, Jianchuan County, 26 ° 27 ʹ 46 ʹʹN, 99 ° 55 ʹ 09 ʹʹE, 2199.8 m asl., under Pinus and fagaceous trees, 15 Sept. 2015, Y. - G. Fan & W. - J. Yu, FYG 2015207 (FCAS 3636), ITS-GenBank: OP 204669. PAKISTAN, Azad Jammu and Kashmir, Bagh, Saliyan, Nari, at 1,950 m a. s. l., under mixed forest of Pinus wallichiana, Cedrus deodara and Abies pindrow, on soil, solitary, 01 Sept. 2021, Rubab Khurshid LAH 37696.	en	Naseer, Arooj, Khurshid, Rubab, Fan, Yu-Guang, Khalid, Abdul Nasir (2024): Two new species of Mallocybe from Himalayan forests of Asia. Phytotaxa 640 (3): 255-264, DOI: 10.11646/phytotaxa.640.3.3, URL: http://dx.doi.org/10.11646/phytotaxa.640.3.3
038C87DFFFD1FFCE85F062EDFE60EECF.taxon	etymology	Etymology: — The specific epithet kashmirana refers to type locality Kashmir, Pakistan. Diagnosis: — characterized by yellowish brown pileus without umbo with red scales, yellowish brown concolorous lamellae, larger stipe (30 – 35 mm) and larger basidiospores (8.6 µm). Holotype: — Pakistan, Azad Kashmir, Bagh, Saliyan, Nari, at 1,950 m a. s. l., under mixed forest of Pinus wallichiana, Cedrus deodara and Abies pindrow, on soil, solitary, 01 September 2021, Rubab Khurshid, N- 213 (LAH 37695). GenBank accession ITS: OQ 448898, LSU: OQ 458716, mtSSU: OQ 448896. Description: — Basidiomata small to medium sized. Pileus 27 – 35 mm diam., conical when very young, then convex or plano-convex when mature, not umbonate, flat, margin inrolled initially, later incurved, at time undulate, not even; surface dry, rough, covered with minute scruffy appressed scales, yellowish brown (3.3 YR 7.1 / 5.2) with red scale when young, becoming yellowish red (1.7 YR 6.5 / 6) with red (6.9 R 3.4 / 7.6) scales, concentrated in central disk. Lamellae 1 – 2 mm wide, adnexed, close with three to four tiers of lamellulae, yellowish brown (3.3 YR 7.1 / 5.2), edges concolor, serrate. Stipe 30 – 35 × 3 – 5 mm, 5 mm at apex, solid, equal, central, cylindrical to subcylindrical, equal, surface dry, finally fibrillose throughout with scattered loose fibrils or prominent with age, partial veil present when very young. Context fresh (5 A 2) in pileus, dirty white (5 A 1) to cream, 0.5 – 1 mm thick. Central brown (7 R 8.2 / 4.2) to reddish brown (9.6 R 3.5 / 7.2) in stipe context when mature, darker towards lower portion, with basal white tomentose, closely adhered soil particle Odor unpleasant. Basidiospores [60 / 3 / 2] (7.6 –) 7.9 – 9.2 (– 10.3) × (4.6 –) 5.2 – 6.1 (– 6.8) µm, avL × avW = 8.6 × 5.7 µm, Q = 1.3 – 1.74 (– 1.86), avQ = 1.53, light brownish in 5 % KOH, ellipsoidal, thick-walled, smooth, apiculus absent or not prominent enough to observe. Basidia (21.8 –) 29.5 – 35.3 (– 39.4) × (7.6 –) 8.4 – 9.7 (– 9.9) µm, avL × avW = 39.1 × 9.78 µm, narrowly clavate to clavate, thin-walled, primarily 4 - sterigmate, less often 2 - sterigmate, hyaline in 5 % KOH, non-guttulate. Cheilocystidia (13.0 –) 14.2 – 19.1 (– 21.8) × (5.1 –) 6.3 – 7.9 (– 9.5) µm, avL × avW = 18.5 × 7.9 µm, thick-walled, fusiform or utriform, smooth walls, hyaline in 5 % KOH, non-guttulate. Pleurocystidia absent. Pileipellis a cutis of elongate, apically cylindrical terminal elements with some short less clavate elements interspersed, hyphae (5.2 –) 6.1 – 10.7 (– 13.6) µm in diameter, hyaline in 5 % KOH, frequently septate, crowded, regular, un branched, thin-walled, clamp connections frequently present. Pileocystidia absent. Stipitipellis a cutis, composed of parallel hyphae, (7.5 –) 10.7 – 13.8 (– 17.9) µm in diameter, light brownish in 5 % KOH, frequently septate, unbranched, regular, with narrowly cylindrical terminal ends, clamp connection present frequently. Caulocystidia (17.4 –) 18.1 – 29.4 (– 34.9) × (10.3 –) 12.3 – 14.7 (– 16.7) µm, cylindrical to fusiform, broader than lower hyphae, smooth, empty, less frequent, hyaline to light brownish in 5 % KOH. Habit and habitat: Found on soil under conifers.	en	Naseer, Arooj, Khurshid, Rubab, Fan, Yu-Guang, Khalid, Abdul Nasir (2024): Two new species of Mallocybe from Himalayan forests of Asia. Phytotaxa 640 (3): 255-264, DOI: 10.11646/phytotaxa.640.3.3, URL: http://dx.doi.org/10.11646/phytotaxa.640.3.3
038C87DFFFD1FFCE85F062EDFE60EECF.taxon	distribution	Distribution: It is first time collected from Bagh, Azad Jammu and Kashmir, Pakistan. Additional material examined: PAKISTAN, Azad Jammu and Kashmir, Bagh, Saliyan, at 2,625 m a. s. l., under mixed forest of Pinus wallichiana, Cedrus deodara and Abies pindrow, on soil, solitary, 20 August 2020, Rubab Khurshid, Herb. LAH- 258.	en	Naseer, Arooj, Khurshid, Rubab, Fan, Yu-Guang, Khalid, Abdul Nasir (2024): Two new species of Mallocybe from Himalayan forests of Asia. Phytotaxa 640 (3): 255-264, DOI: 10.11646/phytotaxa.640.3.3, URL: http://dx.doi.org/10.11646/phytotaxa.640.3.3
