taxonID	type	description	language	source
038C133CD21ED43618FBB3C1FEA97AC1.taxon	description	(Figs. 1 – 3)	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
038C133CD21ED43618FBB3C1FEA97AC1.taxon	diagnosis	Diagnosis. Frontoclypeal region strongly produced forwards, apically narrowed and emarginate; mandible robust, relatively short and wide, distinctly curved; eyes medium-sized, their minimal interocular distance 1.85 – 1.95 eye diameter; antenna pectinate; pronotum 1.30 – 1.35 times as wide as long, with lateral sides almost straight, lateral carina incomplete, distinct at basal half, then only vaguely defined and missing after 3 / 4 of prothorax length; prosternal process reduced; mesoventrite V-shaped, with only vaguely defined mesoventral cavity; all abdominal ventrites free; paramere without latero-apical projection (Figs. 1 – 3; Kovalev et al. 2019).	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
038C133CD21ED4371A96B2C9FDA47F23.taxon	description	(Figs. 1 – 3)	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
038C133CD21ED4371A96B2C9FDA47F23.taxon	materials_examined	Type Locality. SW Iran, Lorestan province, 20 km N Pol-e Dokhtar, Baba Zeyd vill. Type Material. Holotype, male, “ SW Iran, Lorestan province, 20 km N Pol-e Dokhtar, Baba Zeyd vill., 2. V. 2007, A. A. Klimenko leg. ” (ZIN). The re-examination of type material was unnecessary because the original description, figures and drawings (Kovalev et al. 2019) allowed us to diagnose and identify this easily recognizable species. Material Examined. 1 male, “ Iran, Khuzestan [? Lorestan] Province, Pol-e / Dokhtar, 20. IV. 2018 / S. Azadbakhsh leg. ” (PCAA). Comparison. This is the only species classified in the genus Drilorhinus, and it clearly differs from Drilus species in the region by the yellowish to reddish brown head, the strongly projected frontoclypeal part of head, the subtrapezoidal, less transverse pronotum (1.30 – 1.35 times as wide as long), divergent elytra, and male genitalia 1.90 times as long as maximum phallobase width (head dark brown, frontoclypeal region short and widely concave, pronotum distinctly transverse, 1.60 – 1.80 times as wide as long, elytra adjacent to each other for almost their whole length, and male genitalia 1.65 – 1.70 times as long as maximum phallobase width in Drilus iljini and D. iranicus) (Figs. 1 – 5). = 2.0 mm. Variability. Body 6.8 – 7.7 mm long, frontal distance between eyes 1.85 – 1.95 times eye diameter, pronotum 1.30 – 1.35 times as wide as long, and elytra combined 1.9 – 2.0 times as long as wide.	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
038C133CD21ED4371A96B2C9FDA47F23.taxon	distribution	Distribution. Iran (Lorestan prov.).	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
038C133CD21FD4301977B440FEA37BE0.taxon	description	(Figs. 4 – 5) Drilus Olivier, 1790. Type species: Ptilinus flavescens Geoffroy, 1785.	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
038C133CD21FD4301977B440FEA37BE0.taxon	diagnosis	Diagnosis. Eyes relatively small to medium-sized, their minimal interocular distance 1.60 – 3.00 times eye diameter; antenna slightly serrate to weakly flabellate; pronotum 1.20 – 1.70 times as wide as long, with lateral sides sinuate or rounded, lateral carina almost complete; prosternal process strongly reduced; mesoventrite V-shaped, with only vaguely defined mesoventral cavity; all abdominal ventrites free; sternite IX and tergite X usually less than 1.80 times as long as wide; paramere without latero-apical projection (Figs. 4 – 5; Kundrata and Bocak 2019).	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
038C133CD218D43118ACB0FEFCD37CDB.taxon	description	(Fig. 4)	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
038C133CD218D43118ACB0FEFCD37CDB.taxon	materials_examined	Type Locality. Iran: Gīlān prov., Hassankiadeh. Type Material. Holotype, male, “ Persia septentrionalis [North Iran]: Hassan-Kiadeh (B. Iljin 27. IV. 1916 leg.) ” (coll. Barovskij, ZIN?) (Barovskij 1922). The type specimen has not been located at the ZIN collection (M. G. Volkovitsh and A. Kovalev, personal communication). Other Material Examined. 1 male, “ N-IRAN, Pr. Mazandaran / N-Elburz, Now Shahr, Om / 36 ° 36 ’ 17 ” N, 51 ° 38 ’ 23 ” E / 6. V. 2012. leg. D. Frenzel ” (NKME); 1 male, “ N-IRAN, Pr. Mazandaran / NElburz, Now Shahr, Om / 36 ° 36 ’ 17 ” N, 51 ° 38 ’ 23 ” E / 6. V. 2012. leg. D. Frenzel ” (NKME); 1 male, “ Iran, Mazandaran, 1400 m / 20 km südlich Nowshahr, / Weysser, 9. – 14. 6. 1977 / leg. Holzschuh & Ressl // D. / iljini / Barov. // Naturhistorisches / Museum Basel / Coll. W. Wittmer ” (NHMB); 1 male, “ ♂ // Persia / [unreadable data] // coll. Richard / Hicker, Wien // Naturhistorisches / Museum Basel / Coll. W. Wittmer ” (NHMB); 1 male, “ IRAN, Ardabil prov., 4. - 5. VI. 2006 / 5 km NW KOLUR / ” Shahrud ” river valley / 37 ° 26, 1 ’ N 48 ° 41,2 ’ E; 1670 m / Jiří Hájek & Pavel Chvojka leg. ” (NMPC). The following specimens were examined based only on photographs kindly provided by Dr. A. Kovalev (ZIN): 3 males, “ Астрабад [Astrabad; = Gorgan] / 13. IV. [19] 14. [other two specimens: “ 24. IV. [19] 14 ” and “ 2. V. [19] 14 ”] [the dates in Russia before 1918 are in the Julian calendar, with the difference in the 20 th century of 13 days compared to the Gregorian calendar, e. g., 13. IV. 1914 is 26. IV. 1914] / А. Кириченко [A. N. Kiritshenko] ” (ZIN). Comparison. This species differs from Drilorhinus klimenkoi in the dark brown to blackish head, pro- and mesothorax (yellowish to reddish brown in D. klimenkoi), the short, widely concave frontoclypeal region (strongly projected forwards in D. klimenkoi), the distinctly transverse pronotum, which is 1.6 – 1.8 times as wide as long (subtrapezoidal, 1.30 – 1.35 times as wide as long in D. klimenkoi), and elytra adjacent to each other for almost their whole length (adjacent only at basal third, then divergent in D. klimenkoi) (Figs. 1 – 2, 4). Drilus iljini differs from D. iranicus in the dark brown to blackish pronotum, hypomeron and scutellar shield (yellowish to reddish brown in D. iranicus), pronotum widest at about middle (at posterior angles in D. iranicus), and relatively slenderer parameres (more robust and relatively wider in D. iranicus) (Figs. 4 – 5). Diagnostic Redescription. Body (Fig. 4 A) 5.0 – 7.2 mm long, 2.8 – 3.3 times as long as width at humeri. Body dark brown to black, antenna brown, with antennomeres I and II slightly lighter, mouthparts and legs yellowish brown to brown (femur and tibia darker, tarsus distinctly lighter), and apical abdominal segment yellowish brown. Head (Figs. 4 B, C) including eyes 0.70 – 0.80 times as wide as pronotum, frontoclypeal region between antennal insertions widely concave; eyes medium-sized, their frontal distance 1.95 – 2.35 times eye diameter; mandible moderately long and curved; antenna (Fig. 4 A) pectinate, antennomere III subtriangular; antennomeres IV – X each with moderately long, slightly flattened branch, which is 1.00 – 1.25 times as long as its respective stem; antennomere XI approximately as twice as long as antennomere X. Pronotum (Figs. 4 B, C) transverse, widest around middle, 1.60 – 1.80 times as wide as long; sides more or less rounded, narrowed anteriorly; posterior angles subrectangular, apically rounded. Elytra (Fig. 4 A) elongate, covering almost whole abdomen, adjacent to each other for almost whole length, both combined 2.05 – 2.45 times as long as wide, 0.70 – 0.75 times as long as body, 3.90 – 4.40 times as long as pronotum. Abdominal sternite IX 1.90 – 1.95 times as long as wide. Male genitalia (Fig. 4 D) 1.65 – 1.70 times as long as maximum phallobase width; median lobe elongate, 1.3 times as long as phallobase; paramere robust, without latero-apical projection, apically rounded, partly membranous; phallobase U-shaped, 1.15 – 1.20 as wide as long. Female and immature stages unknown. Variability. There is variability in the coloration of labrum and body appendages, i. e., antennae, palpi, and legs. In darker specimens they are brown, with only tarsi always being lighter, while in lighter specimens they are yellowish brown to light brown, with tarsi again lighter (Figs. 4 A – C). Additionally, there is slight variability in the shape of the pronotum, especially the length / width ratio and the shape of the posterior angles (Figs. 4 A – C), and in the relative length of the elytra compared to the pronotum.	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
038C133CD218D43118ACB0FEFCD37CDB.taxon	distribution	Distribution. Iran (Ardabil, Gīlān and Mazandaran provinces).	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
038C133CD219D4311B4DB7F4FB9A7F67.taxon	description	(Fig. 5)	en	Packova, Gabriela, Hoffmannova, Johana, Kundrata, Robin (2021): Drilini (Coleoptera: Elateridae: Agrypninae) of Iran. The Coleopterists Bulletin 75 (3): 579-586, DOI: 10.1649/0010-065X-75.3.579, URL: http://dx.doi.org/10.1649/0010-065x-75.3.579
