identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038C5E69506EFF923DE37538990FFBD6.text	038C5E69506EFF923DE37538990FFBD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryaxis splendidus (Croissandeau 1891)	<div><p>Bryaxis splendidus species group</p> <p>Diffential diagnosis. The group is defined within the genus by the following combination of characters: (1) body unicoloured, reddish brown; (2) pronotum gibbous; (3) median occipital carina present; (4) eyes reduced, in females sometimes entirely absent; (5) maxillary palpomeres II and III irregularly granulate; (6) terminal maxillary palpomere large and flat with dense setation; (7) scape in males long, with well-projected secretory nodule at inner side; (8) pedicel longer than wide, oval, extended inwards by elevated edge with thickened and apically dentate margin; (9) inner side of pro- and metatibiae dentate close to apex; (10) aedeagus with parameres narrowed apically and forming proximally oriented lobes with variable shape, although the lobes are almost absent in B. monstrosetibialis (Stolz, 1923).</p> <p>Biology. All specimens have been found in caves or in deep, shady and cold cavities under wet leaves or large, deeply buried stones.</p> <p>Distribution. The species of the group are endemic to the Carpathians and the area of their distribution extends from northern Romania to central Slovakia (Fig. 8).</p></div> 	https://treatment.plazi.org/id/038C5E69506EFF923DE37538990FFBD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hlaváč, Peter	Hlaváč, Peter (2009): Taxonomic notes on the Bryaxis splendidus species group (Coleoptera: Staphylinidae: Pselaphinae), with the description of a new species from the Ukraine. Acta Entomologica Musei Nationalis Pragae 49 (2): 651-659, DOI: 10.5281/zenodo.5322469
038C5E69506EFF923D6873379E3DF9C5.text	038C5E69506EFF923D6873379E3DF9C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryaxis splendidus (Croissandeau 1891)	<div><p>Key to species of the Bryaxis splendidus species group</p> <p>Note. Because of the small number of specimens available for this study and that fact that the species are very similar in external features, the following key is only tentative. Reliable identification requires the study of the aedeagi.</p> <p>1 Scape robust, short, 1.4–1.6 times as long as wide, with triangular extension on inner side................................................................................................................................ 2</p> <p>– Scape slender and longer, 1.8–2.0 times as long as wide, almost parallel on inner side........................................................................................................................................ 3</p> <p>2 Eyes very small, composed of two facets in male (female unknown), last segment of maxillary palpus 0.45 mm long, Ukraine........................................... B. cepliki sp. nov.</p> <p>– Eyes large, composed of seven facets in males and 4–5 facets in females, last segment of maxillary palpus 0.38 mm long, Slovakia................... B. montrosetibialis (Stolz, 1923)</p> <p>3 Large species, body length 2.6 mm, Romania....................... B. goliath (Jeannel, 1922)</p> <p>– Smaller species, body length 2.0– 2.1 mm, Ukraine......................................................................................................................................... B. splendidus (Croissandeau, 1891)</p></div> 	https://treatment.plazi.org/id/038C5E69506EFF923D6873379E3DF9C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hlaváč, Peter	Hlaváč, Peter (2009): Taxonomic notes on the Bryaxis splendidus species group (Coleoptera: Staphylinidae: Pselaphinae), with the description of a new species from the Ukraine. Acta Entomologica Musei Nationalis Pragae 49 (2): 651-659, DOI: 10.5281/zenodo.5322469
038C5E69506DFF973DBD77B498CBFE52.text	038C5E69506DFF973DBD77B498CBFE52.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryaxis cepliki Hlaváč 2009	<div><p>Bryaxis cepliki sp. nov.</p> <p>(Figs. 1–4, 8)</p> <p>Type locality. Ukraine, Transcarpathian Ruthenia, Mala Ugolka, Zhemchuzhnaya cave</p> <p>Type material. HOLOTYPE: J, ‘ UKRAINE: Zakarpattya, Mala Ugolka, Žemčužnaja cave, 16.VII.2007, D. Čeplík lgt. / HOLOTYPE Bryaxis cepliki sp. nov., P. Hlaváč det., 2008’ (NMPC). PARATYPE: J, ‘ UKRAINE: Zakarpattya, Velika Ugolka, cave Moločnyj kameň, 15.VII.2007, D. Čeplík lgt.’ (CPH).</p> <p>the B. splendidus species group by the shape of the aedeagus (Figs. 3, 4).</p> <p>Etymology. Named after my friend David Čeplík, Košice, a very enthusiastic biospeleologist and collector of both specimens of the new species.</p> <p>Biology. The holotype was collected in wet scree near the wall at the end of the Zhemchuzhnaya cave, about 50 m from the entrance. The paratype was collected under large stones at the end of the Molochniy kamen cave, about 80 m from the entrance.</p> <p>Distribution. Ukraine (Transcarpathian Ruthenia).</p></div> 	https://treatment.plazi.org/id/038C5E69506DFF973DBD77B498CBFE52	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hlaváč, Peter	Hlaváč, Peter (2009): Taxonomic notes on the Bryaxis splendidus species group (Coleoptera: Staphylinidae: Pselaphinae), with the description of a new species from the Ukraine. Acta Entomologica Musei Nationalis Pragae 49 (2): 651-659, DOI: 10.5281/zenodo.5322469
038C5E69506BFF963DCD76B49B13FE6C.text	038C5E69506BFF963DCD76B49B13FE6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryaxis goliath (Jeannel 1922)	<div><p>Bryaxis goliath (Jeannel, 1922), stat. restit.</p> <p>(Figs. 5, 8)</p> <p>Megalobythus goliath Jeannel, 1922: 233. Type locality: Romania, cave Corobana Mandraţului, comm. de Scărişoara, jud. Turda-Arieş.</p> <p>Bryaxis splendidus goliath Jeannel: MACHULKA (1935: 131).</p> <p>Note. MACHULKA (1935) treated this species as a variety of B. splendidus. I had not the posibility to see the species but the shape of the apices of the parameres as illustrated in JEANNEL (1955: 40, Fig. 17b) and reproduced in Fig. 5 combined with the large body size indicate that B. goliath is a distinct species.</p> <p>Distribution. Known only from the type locality, Corobana Mandraţului cave in Munti Apuseni, Romania</p> <p>Bryaxis monstrosetibialis (Stolz, 1923)</p> <p>(Figs. 6, 8)</p> <p>Bythinus monstrosetibialis Stolz, 1923: 71–72. Type locality: Bad Rosenau in Ungarn (Komitat Gömör [Gemer region in southern Slovakia]).</p> <p>Bythinus Trojani Fleischer, 1925:17–18. Type: Slovakia, ‘okraj Demänovských jeskyň [edge of Demänovská cave]’, synonymised by MACHULKA (1929c: 120–121).</p> <p>Megalobythus Trojani Fleischer: MACHULKA (1929a: 22; 1929b: 23–24).</p> <p>Bythinus (Macrobythus) monstrosetibialis Stolz: ROUBAL (1930: 507–508).</p> <p>Bythinus monstrosetibialis Stolz var. ocularis Machulka, 1935: 131. Type locality: Slovakia, ‘okolie dobšinské jeskyně [surroundings of Dobšiná cave]’, synonymised by LÖBL &amp; BESUCHET (2004: 307).</p> <p>Balcanobythus (Megalobythus) monstrosetibialis Stolz: KARAMAN (1957: 187, 208, Fig. 6 (aedeagus)).</p> <p>Bryaxis monstrosetibialis Stolz: BESUCHET (1974: 349, Fig. F. Basis 18: 26 (first two antennal segments)); HLAVÁČ, (2000: 37, new records, biology, aedeagus and habitus drawings); FRANC &amp; MLEJNEK (2000: 31–34, new records,</p> <p>biology). New records (all data provided by Martin Švarc; all specimens deposited in his collection). SLOVAKIA: 13 JJ 17</p> <p>♀♀: Hrochoť, Hrochoťská dolina [valley], sifting of leaf-litter in cave, Švarc lgt.; 4 JJ 2 ♀♀: Liptovský Mikuláš, Demänovská jaskyňa [cave], under stones and sifting leaf-litter in cave, Švarc lgt.; 2 spec.: Muráňska planina, jaskyňa Maretkina [cave], Orszulík lgt.; 1 J: Slovenský raj, jaskyňa Duča [cave], trap, Košel lgt.; 3 JJ: Závadka nad Hronom, Veľká Stožka, Machnatá dolina [valley], sifting of litter near spring, Krátky lgt.; 1 ♀: Muráň, Javorníkova dolina [valley], sifting Fagetum, Mantič lgt.; 1 J: Kremnické vrchy, Diera pod oknom, 1190m, Mlejnek lgt.; 1 J: Nízke Tatry Mts., jaskyňa nad Patočinami [cave above Patočiny], 3,700 m a.s.l., Mlejnek lgt.; 1J: Malá Fatra, Veľký Rozsutec, Brestovanský lgt.; 2 ♀♀: Nízke Tatry Mts., Demänovská dolina [valley], Orszulík lgt.</p> <p>Records from the literature. ROUBAL (1930) and MACHULKA (1929a,b,c, 1935) summarized old data from the following localities: near Rožňava, near Košice, Gemer (exact locality not available), Krupina, Nová Baňa, Muráň, and Dobšiná. Additional recent record was published from Zádieľ in the eastern Slovakia by MAJZLAN &amp; RYCHLÍK (1994).A decade ago I summarized all old data and added some other localities: Slovenský kras (exact locality not available), Tisovec env. (Michňová jama), Handlová, Plešivecká planina (Lastovičia jama) (HLAVÁČ 2000). Finally, FRANC &amp; MLEJNEK (2000) studied the ecology and distribution of the species and mentioned the following localities: Malá Fatra (lúky nad Hornými dierami [meadows above Horné diery]); Sokolec; Fačkovské sedlo [saddle], Plešivecká planina (Lastovičia priepasť [abyss]), Revúcka vrchovina (jaskyňa [cave] Burda,Veľká drienčanská jaskyňa [cave]), Poľana (Hrochoťská jaskyňa [cave]), Horehronské podolie: Valaská (jaskyňa v kameňolome [cave in stone-pit] Potôčky), Horehronské podolie: Červená skala (Márnikova jaskyňa [cave]), Starohorské vrchy (jaskyňa pri Uľanke [cave near Uľanka]), Nízke Tatry (Demänovská dolina [valley], Tunelová jaskyňa [cave]).</p> <p>Distribution. Slovakia.</p></div> 	https://treatment.plazi.org/id/038C5E69506BFF963DCD76B49B13FE6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hlaváč, Peter	Hlaváč, Peter (2009): Taxonomic notes on the Bryaxis splendidus species group (Coleoptera: Staphylinidae: Pselaphinae), with the description of a new species from the Ukraine. Acta Entomologica Musei Nationalis Pragae 49 (2): 651-659, DOI: 10.5281/zenodo.5322469
038C5E69506AFF963D3476DE9980FD2C.text	038C5E69506AFF963D3476DE9980FD2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryaxis splendidus (Croissandeau 1891)	<div><p>Bryaxis splendidus (Croissandeau, 1891)</p> <p>(Figs. 7, 8)</p> <p>Bythinus splendidus Croissandeau, 1891: 134. Type locality: not provided.</p> <p>Bythinus splendidus Croissandeau: MACHULKA (1935: 131, new record).</p> <p>Material examined. UKRAINE: J: Kuzy. 1930. / Bythinus splendidus Croiss. Det. Ing. Machulka / 1 / Nár. Mus. Praha coll.Machulka / Bryaxis splendidus Croiss.Cl.Besuchet dét. I. 1977; ♀: Kuzy / Bythinus splendidus Croiss.Det. Ing.Machulka / Nár.Mus. Praha coll. Machulka;J [only aedeagus on a plastic slide, the whole body is missing]:Kuzy / 3 / Nár. Mus. Praha coll. Machulka.All specimens are deposited in NMPC and bear my identification label.</p> <p>Distribution. Known only from the locality of Kuzy in the Transcarpathian Ukraine.</p></div> 	https://treatment.plazi.org/id/038C5E69506AFF963D3476DE9980FD2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hlaváč, Peter	Hlaváč, Peter (2009): Taxonomic notes on the Bryaxis splendidus species group (Coleoptera: Staphylinidae: Pselaphinae), with the description of a new species from the Ukraine. Acta Entomologica Musei Nationalis Pragae 49 (2): 651-659, DOI: 10.5281/zenodo.5322469
