identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4F0D625B8AC2FFF2094B2AFC8EEC6B6E.text	4F0D625B8AC2FFF2094B2AFC8EEC6B6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheilosia barbafacies Vujic & Radenkovic	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Cheilosia
barbafacies 
Vujic
&amp; 
Radenkovic sp. n. Figs 1A2A, 2B4A, 4B5-78A, 8C9-1112A, 12B </p>
            <p> Cheilosia honesta of  Šimić , 1987 (in part). </p>
            <p> Cheilosia pascuorum of  Vujić , 1996 (in part). </p>
            <p>Type-locality.</p>
            <p> MONTENEGRO: Durmitor,  Škrčko-Sušički basen, 43˚11'7"N, 19˚3'28"E, broad-leaf forest, 25 June 1995, A.  Vujić leg. </p>
            <p>Type-specimen:</p>
            <p> Holotype ♂, in excellent condition. MONTENEGRO. Original label: "Durmitor YU / Skakala 25.06.'95. / leg.  Vujić.” 43°10'16"N; 18˚59'56"E (FSUNS 05768). </p>
            <p> Paratypes, in excellent condition. MONTENEGRO: ♂ Original label: "007. Durmitor / Skrcka jezera / 5.07.1983." 43°8'8"N; 19°0'56"E (published in  Šimić (1987) as  Cheilosia
honesta
 ) (FSUNS 05758); ♂ Original label: "199 G. Durmitor / Luke / 8.07.1991. YU." 43°7'37"N; 19°0'5"E (published in  Vujić (1996) as  Cheilosia pascuorum ) (FSUNS 05759); ♂ Original label: "199 H. Durmitor / Luke / 9.07.1991. YU." 43°7'37"N; 19°0'5"E (FSUNS 05763); 3♂ Original label: "199 H. Kanjon Susice / 9.07.1991. YU." 43°12'41"N; 18°59'44"E (FSUNS 05760, 05761, 05766); ♂ Original label: "Dur  mitor 8.07.92. / Skrcko Zdrijelo YU / leg.Vujic. " 43°7'7"N; 19°0'53"E (FSUNS 05765); ♂ Original label: "Durmitor 2.07.93. / ka Prutasu YU / leg. Radnovic S." 43°10'16"N; 18°59'56"E (FSUNS 05764); ♂ Original label: "Durmitor 1.6.1994 / KanjonSusice YU / leg. Vujic." 43°12'41"N; 18°59'44"E (FSUNS 05770); ♂ Original label: "Durmitor YU / Skakala 6.7.1994. / leg. Vujic." 43°11'16"N; 19°0'21"E (FSUNS 05767); ♂ Original label: "Durmitor 30.06.93. /  Skrčko jezero YU / leg. Radnovic S." 43°8'8"N; 19°0'56"E (FSUNS 05762); ♂ (FSUNS 05771) ♀ (FSUNS 05769) Original label: "Durmitor 25-26.5.96. / Susica-Skrke YU / leg. Vujic". 43°11'7"N; 19°0'28"E. BOSNIA-HERZEGOVINA: ♂ Original label: "1613 Bosna / Jahorina / 14.05.1989." 43°42'25"N; 18°34'13"E (published in  Vujić (1996) as  Cheilosia pascuorum ) (FSUNS 05757). </p>
            <p>Description.</p>
            <p>MALE (Figs 1A, 2A, 2B, 4A, 4B, 5, 6C, 8A, 8C, 9A, 11A, 12A, 12B).</p>
            <p> Head : Face with long, predominantly pale hairs, central prominence rounded (Fig. 5); orbital stripe with short, pale hairs. Frontal triangle small, undusted, covered with black hairs; eye contiguity longer than frontal triangle (Fig. 6C). Eyes completely covered with greyish hairs. Occiput narrow, white-grey dusted. Antennae dark, third antennal segment from dark-brown to reddish; arista bare and short (Fig. 11A). Clypeus dusted. </p>
            <p>Thorax: Scutum with dark-olive shine, laterally slightly dusted, covered with long, pale and black hairs (Fig. 9A); central disc shining, with fine puncturation. Scutellum covered with long hairs and numerous longer black hairs on posterior margin (Fig. 9A). Pleurae dusted, covered with predominantly pale hairs mixed with black hairs on anepisternum and anepimeron; katepisternum continuosly pilose. Wing brownish, with dark veins, completely covered with microtrichia; vein M1 meeting vein R4+5 at an obtuse angle (Fig. 7: x). Calypters yellowish-grey. Haltere yellowish-grey with dark capitulum. Legs dark, except pale apex of femora, basal 1/3-1/4 and apical 1/5-1/6 of tibiae and ventral surface of tarsi on fore and middle legs; hairs on legs predominately pale mixed with black.</p>
            <p>Abdomen: Tergites shining, except the whole tergite 2 and dull central area on tergites 2 and 3, which extends from anterior margin of tergite 2 to basal 6/7 of tergite 3, leaving the posterior margin of tergite 3 shining; tergites covered with erected, pale hairs, except few black hairs on posterior half of tergite 4 and on pregenital segments. Sternites grey dusted covered with pale hairs.</p>
            <p> Genitalia: Dorsal lobe of gonostylus broad basally (Fig. 8A), without distinct dorsolateral extension (Figs 1A, 2A, 2B), present in  Cheilosia pascuorum (Figs 1B, 1C, 2C, 2D); theca of hypandrium in ventral view with large quadrilateral excavation (Figs 4A, 4B). </p>
            <p> FEMALE (Figs 6A, 6B, 7, 9B, 10, 11B). Similar to the male, except for normal sexual dimorphism and the following characters: pile is general shorter and more extensively pale, and legs less dark, basal 1/4 of femora, basal 1/3 and apical 1/4 of tibiae pale; frons with two lateral channels, shiny, except dusted antero-lateral corners (Fig.  6 B), covered with pale hairs, except a few black hairs around ocellar triangle and above antennae; thorax pale haired, except for a few black hairs on post-alar calli and near wing base; tibiae of fore and middle legs pale, except dark central ring; basal tarsi of fore and middle legs pale; tergites covered with long and erect hairs, except adpressed hairs on central part of tergites 1-4. </p>
            <p>Size. Male, body length: 8.9-11.1 mm; wing length: 7.9-9.2 mm (14 specimens were measured). Female, body length: 10.5 mm; wing length: 8.7 mm (1 specimen was measured).</p>
            <p> Diagnosis . </p>
            <p> Species related to  Cheilosia pascuorum , but differs in the following characteristics: face covered with long hairs (Figs 5, 6), bare in  Cheilosia pascuorum ; clypeus dusted, shining in  Cheilosia pascuorum ; male genitalia: dorsal lobe of gonostylus without distinct dorsolateral extension (Figs 1A, 2A, 2B), present in  Cheilosia pascuorum (Figs 1B, 1C, 2C, 2D). </p>
            <p> Based on the identification keys for European hoverflies, this species can be confused with four other  Cheilosia species which have long facial hairs:  Cheilosia barbata (Fig. 3A),  Cheilosia lasiopa Kowarz, 1885 (Fig. 4D),  Cheilosia melanopa (Fig. 4E) and  Cheilosia vulpina (Fig. 4C). Identification of the new described species is possible based on a combination of the following characters: arista bare; central disc of scutum shining; vein M1 meeting vein R4+5 at an obtuse angle (Fig. 7: x); tergites 1-3 pale haired; sternites obviously grey dusted; male: dorsal lobe of gonostylus broader basally (Fig. 8A); female: arista about 3 times as long as third antennal segment (Fig. 11B); hairs on scutum long and erected (Fig. 9B); hind tibia on posterodorsal surface with few longer hairs (Fig. 10: y). </p>
            <p>Etymology.</p>
            <p> The specific epithet is derived from the Latin nouns (in apposition) of feminine gender in the nominative case:  “barba” (beard) and  “facies” (face). The name indicates the presence of long hairs on the face. </p>
            <p>Distribution</p>
            <p> (Fig. 13).  Cheilosia barbafacies sp. n. is found in two Dinaric mountains in the central part of the Balkan Peninsula, in Durmitor (Montenegro) and Jahorina (Bosnia-Herzegovina), while the related species,  Cheilosia pascuorum , has a wider range extending from the Alps, across the Balkan Peninsula, to Romania and the European part of Russia (Speight 2012). Both species appear sympatrically within one refuge area rich in endemics and relict species (Durmitor mountain, gorge of river  Sušica and glacial lakes  Škrčka jezera) (  Šimić 1987,  Vujić 1996). </p>
            <p>Discussion.</p>
            <p> Although  Cheilosia barbafacies has long facial hairs, it is closely related to  Cheilosia pascuorum with a non-hairy face. Species with long facial hairs were assigned to "group  B " of Sack (1928-1932), based on  Becker’s (1894) revision of the genus. Also recent authors (e.g. Bartsch et al. 2009) still use this old name ("group B") in their keys for practical reasons. In his recent classification of the genus, Barkalov (2002) included taxa with long facial hairs in seven out of his thirteen subgenera, i.e. subgenera  Cheiloisa s.str.,  Convocheila Barkalov, 2002,  Endoiasimyia Bigot, 1882,  Floccocheila Barkalov, 2002,  Hiatomyia Shannon, 1922,  Neocheilosia Barkalov, 1983 and  Taeniochilosia Oldenberg, 1916. Phylogenetic analysis of the relationship between the genus  Cheilosia and the  tribe Rhingiini , based on mtDNA COI gene sequence (  Ståhls et al. 2004), revealed that  Cheilosia species with this character state can be found in eight clades (see Fig. 1 in  Ståhls et al. 2004). It seems clear that the length of the facial pilosity is not a synapomorphic character of any species group, and might evolve several times within the genus  Cheilosia . </p>
            <p> In Bartsch et al. (2009), specimens of  Cheilosia barbafacies key out to  Cheilosia vulpina (group B), and in Van Veen (2004), males and females with dark-brown antennae can be identified as  Cheilosia vulpina , while specimens with reddish antennae are  Cheilosia barbata . For the separation of  Cheilosia barbafacies from  Cheilosia barbata , diagnostic characters of the proxima group can be used, and the distinction between  Cheilosia vulpina and  Cheilosia barbafacies sp. n. is presented in the following key. </p>
            <p> Some females of  Cheilosia barbafacies sp. n. are similar to females of  Cheilosia redi , and they can be separated by the following characters: </p>
            <p> Cheilosia barbafacies sp. n.: vein M1 meeting vein R4+5 with an obtuse angle (Fig. 7: x); hairs on scutum and tergites long and erected (Fig. 9B); hind tibia on posterodorsal surface with few longer hairs (Fig. 10: y). </p>
            <p> Cheilosia redi : vein M1 meeting vein R4+5 with an acute angle; hairs on scutum and tergites shorter and significantly adpressed; hind tibia on posterodorsal surface without long hairs. </p>
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	https://treatment.plazi.org/id/4F0D625B8AC2FFF2094B2AFC8EEC6B6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Vujic, Ante;Radenkovic, Snezana;Trifunov, Sonja;Nikolic, Tijana	Vujic, Ante, Radenkovic, Snezana, Trifunov, Sonja, Nikolic, Tijana (2013): Key for European species of the Cheilosia proxima group (Diptera, Syrphidae) with a description of a new species. ZooKeys 269: 33-50, DOI: http://dx.doi.org/10.3897/zookeys.269.4619, URL: http://dx.doi.org/10.3897/zookeys.269.4619
