taxonID	type	description	language	source
038D87AFE042A522A7EBFDB20960FCB6.taxon	materials_examined	Type species: Halozercon karacholana Wiśniewski et al., 1992, by original designation. Note. Diagnosis of Halozercon was given in a previous article by Marchenko (2018).	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE042A524A7EBFC450F89F82E.taxon	diagnosis	Diagnosis. Vertex with two pairs of setae j 1 and z 1, pore-like structures ip 1 absent. Podonotum ornamented by small tubercles and festoon reticulation over the entire surface. Dorsal shields neotrichous. Many dorsal setae lanceolate or leaf-shaped. Median region of opisthonotum with 17 – 22 setae of J series, with slightly linear reticulation. First sternal platelet entire with pair of lyrifissures iv 2. Metapodal shields large. Soft cuticle between genital and ventri-anal shields with two pairs of setae Jv 1, Zv 1 and two lightly sclerotised plates. Adults peritremes long, reaching level of spine top of coxae II anteriorly; in deutonymph peritremes shorter, reaching middle level of coxae II. Ventri-anal shield entire. Adult chelicera without sexual dimorphism.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE042A524A7EBFC450F89F82E.taxon	description	Description. Female (Figs 1 – 10, 18 – 27 and 30 – 41, n = 10) Dorsal idiosoma (Figs 1, 18, 19, 30 – 37). Idiosoma suboval, 490 – 525 long and 315 – 350 wide (Figs 1, 18, 30 – 31). Anterior margin of the podonotum curved ventrally to form a vertex; two setae inserted on the ventral side of idiosoma: seta j 1 (17 – 20) stout, slightly pilose and seta z 1 smooth (8 – 12), pore-like structures ip 1 absent. Podonotal shield strongly neotrichous (Figs 30 – 36), with about 69 – 72 setae on each side, including marginal r setae (Fig. 37). Podonotum ornamented with small tubercles and festoon reticulation over the entire surface of the shield (Figs 32 – 34). Setae in central region smooth, slightly lanceolate, 15 – 20 long (Figs 32 – 34); lateral setae leafshaped or lanceolate, pilose or smooth, 15 – 20 long (Figs 19, 36), the most posterior row of setae lanceoate and slightly pilose (Fig. 35). Four pairs of pore-like structures (glands) po 1 – po 4 located in podonotum, glands po 3 the smallest. Opisthonotum with 63 – 71 setae on each side, including marginal R setae. Median region neotrichous, with 17 – 22 setae of J series, arrangement asymmetrical, lanceolate, slightly serrated, 17 – 20 long (Figs 30 – 31, 33). Setae of Z, S series inserted closely in lateral regions, asymmetrical, neotrichous, lanceolate or leaf-shaped, smooth or serrated, 17 – 22 long (Figs 18, 19). Reticulation indistinct in median region between setae of J series and lateral setae, ornamented small tubercles in lateral regions. Four pairs of pore-like structures (glands) Po 1 – Po 4 inserted in opisthonotum. All dorsal setae surrounded by enlarged basal rings. Marginal dorsal setae of r – R series neotrichous, inserted on high tubercles, elongated, 17 – 25 long, slightly curved and usually smooth (Fig. 37). Ventral idiosoma. (Figs 2, 20 – 23 and 38 – 41). Base of tritosternum 25 – 30 long and 18 – 20 wide, paired pilose laciniae free from each other along entire length, 47 – 55 long (Fig. 2). Presternal (jugular) platelets irregular shape, weakly sclerotised, with pair of St 1 (23 – 25) setae (Figs 20 – 23). First sternal platelet entire, 33 – 38 long and 50 – 63 wide; with pair of St 2 setae (18 – 20) and pair of lyrifissures iv 2 (Figs 23, 40). Second pair of sternal platelets suboval, with two pairs of setae St 3, St 4 (15 – 17) or in some specimens St 4 seta from one side is inserted in soft cuticle; with two pairs of lyrifissures iv 3, iv 4 (Figs 22, 23). Genital shield elongated, 77 – 80 long, 35 – 42 wide, irregularly-shaped, with individual shape in each specimen; expanded anteriorly and posteriorly; anterior margin with fine folding; with genital setae St 5 (13 – 15) and lyrifissures iv 5 on soft cuticle laterad of shield (Figs 38 – 41). Subtriangular membrane (Figs 20 – 22) surrounds the genital shield. Genital shield with complex formation, including one median unpaired cap and two pairs of fine sclerites, a short anterior pair and elongate posterior pair. Endo- and exopodal shields absent. Metapodal shields large, suboval, located between posterior margins of periterematal and ventri-anal shields. Adgenital gland pores gv 2 are multiple, dispersed over the surface: two gland openings located in soft cuticle posterolaterad of genital shield, another three to four openings located in ventri-anal shield. Soft cuticle between genital and ventri-anal shields with two pairs of setae Jv 1, Zv 1 and two slightly sclerotised plates. Peritrematal shields fused anteriorly forming a vertex and fused with dorsal shield laterally; strongly sclerotised, postero-lateral ends are drawn back in an oblique angle; ornamented with festoon reticulation along entire length. Four pairs of pore-like structures inserted in peritrematal shield: gp 1, gp 2 and ip 2, ip 3. Peritremes straight, 150 – 163 long, reaching level of spine top of coxae II anteriorly; with internal cell structure. Ventri-anal shield entire, broad, 130 – 173 long and 290 – 320 wide, fused to opisthonotal shield around the outer edge; with festoon reticulation, with 7 – 8 pairs of smooth opisthogastric setae inserted asymmetrically (10 – 12), and about 8 – 12 pairs of opisthonotal leaf-shaped smooth setae inserted posteriorly (18 – 20). The most posterior marginal row with elongated smooth setae (20 – 25) on raised tubercles duplicates opisthonotal rows of setae as in Fig. 37. Anal area with smooth para-anal and post-anal setae (12 – 15); anal opening 30 – 33 long; with two lyrifissure on each valve; cribrum located posteriorly of post-anal seta. Pair of glands gv 3 located antero-laterad of para-anal setae. Gnathosoma. (Figs 3 – 5 and 27). Fixed digit of chelicera 42 – 45 long, with five teeth in addition to apical hook and leaf-shaped pilus dentilis (Fig. 3); movable digit the same length with three teeth in additional to apical hook. Chelicera with long dorsal seta (20), lateral (antiaxial) and dorsal (paraxial) lyrifissures; with arthrodial corona. Epistome subtriangular, with irregularly serrated edges and smooth pointed median projection (Figs 4, 27). Corniculi 27 – 30 long and 15 – 17 wide. Internal malae slightly longer than corniculi, with complex three-layer structure as in Fig. 5. Deutosternal groove with 7 – 8 transverse denticulate rows, usually with three paired smooth lateral transverse lines. The posterior fourth pair of lateral transverse lines arch-shaped, with large denticles, located at the level of pc setae. Setae h 1 – h 3 smooth: h 1 (33 – 35) longest, seta h 2 (11 – 13) shorter than h 3 (15 – 19); pc (15 – 17) serrated. Dorsal side of gnathosoma with the pair of protruding wedge-shaped structures between palpal coxa and trochanter, lateral to epistome. Palpal chaetotaxy 2 – 5 – 6 – 12 – 15, palp with five free segments; trochanter with seta al 1 long and pilose in the distal third; with seta al 2 short and smooth; palp genu with setae al 1 and al 2 pilose in distal third, palp apotele two-tined. Legs. (Figs 6 – 10). Lengths: I 337 – 350, II 270 – 280, III 265 – 275, IV 325 – 350 µm. Chaetotaxy of legs I – IV: coxae 2, 2, 2, 1; trochanters 6 (1 1 / 3 1), 5 (1 1 / 3 0), 5 (1 1 / 3 0), 5 (1 1 / 3 0), femora 13 (2 5 / 4 2), 11 (2 5 / 3 1), 6 (1 4 / 1 0), 6 (1 4 / 1 0); genua 13 (2 6 / 3 2), 12 (2 6 / 2 2), 10 (2 4 / 2 2), 10 (2 5 / 2 1); tibiae 14 (2 6 / 4 2), 10 (2 4 / 2 2), 9 (2 3 / 2 2), 10 (2 4 / 2 2); tarsi 49, 18 (3 7 / 5 3), 18 (3 7 / 5 3), 18 (3 7 / 5 3). Tarsi of leg I with 49 setae, including 6 antero-lateral, 5 posterolateral, 9 ventral and 29 dorsal setae (Fig. 10). All legs with pair of sclerotised claws and pulvillus with five lobes. Pretarsus of legs II – IV with ambulacral stalk, legs I with sessile claws. Coxae I – IV are grouped closely together vertically; coxae I split on dorsal side (Fig. 6), coxae III – IV with recesses on anterolateral side (Figs 8, 9). Coxae II – III with antero-dorsal spines: coxae II with large sharp spine, coxae III with small one (Figs 7, 8). Male. (Figs 11 – 12, 28 – 29 and 42 – 44, n = 10). Dorsal idiosoma. Dorsal shield suboval shape, 435 – 460 long and 265 – 285 wide. Ornamentation and chaetotaxy similar to that of female. Ventral idiosoma (Figs 11, 28 – 29 and 42 – 44). Base of tritosternum 25 – 27 long and 15 – 17 wide, laciniae pilose, 37 – 45 long. Pair weakly sclerotised presternal (jugular) platelets with pair of St 1 setae (Fig. 11). First sternal platelet entire, with pair of St 2 setae, with festoon ornamentation, 30 – 32 long and 52 – 62 wide. Second sternal platelet entire, subcordate shape, slightly ornamented by lines and folds; with two pairs of setae St 3, St 4 and two pairs of lyrifissures iv 3, iv 4 in antero-lateral margins; 25 – 27 long on median line and 47 – 50 wide at level of St 3 setae; surrounds the genital opening. Genital opening located at level of coxae III; with a pair of eugenital setae, covering by two platelets (Figs 11, 28 – 29, 43 – 44). Third sternal platelet entire, triangular, 15 – 30 long and 30 – 45 wide, with pair of St 5 setae and pair of iv 5 lyrifissures. Measurements of St 1 – St 5 setae: 19 – 20, 18 – 19, 15 – 17, 15 – 17, 11 – 13 µm respectively. Endo- and exopodal shields absent. Metapodal shields fused with ventri-anal shield and not distinct. Peritrematal shields similar to that female, with four pairs of pore-like structures: gp 1, gp 2, i 2, ip 3. Peritremes similar to those of female. Arch of vertex with two dorsal setae: pilose j 1 (17 – 19) and smooth thornshaped z 1 (10 – 12). Ventri-anal shield broad, 145 – 160 long and 255 – 265 wide, fused to opisthonotal shield around the outer edge; with festoon reticulation; 6 – 10 pairs of opisthogastric smooth setae inserted asymmetrically (12 – 15) with 7 – 9 dorsal leaf-shaped smooth setae (12 – 15) inserted posteriorly. Adgenital gland pores gv 2 are multiple, dispersed over the surface: one pair of glands located on soft cuticle posteriorly of coxae IV, other 3 – 4 openings located in ventri-anal shield. The most posterior marginal row with elongated smooth setae (17 – 20) on raised tubercles duplicates opisthonotal rows of setae as in female. Anal area with pre-anal and post-anal setae 10 – 12 long, anal opening 25 – 27 long; with two lyrifissure on each valve; cribrum present. Pair of glands gv 3 located antero-laterad of para-anal setae. Gnathosoma. Fixed digit of chelicera with five teeth in addition to apical hook and leaf-shaped pilus dentilis (Fig. 12). Movable digit of chelicera the same length as fixed digit (38 – 40), tridentate in addition to apical hook, with arthrodial corona at base of digit. Chelicera with long dorsal seta, lateral (antiaxial) and dorsal (paraxial) lyrifissures. Male chelicera lacking spermatodactyl, without sexual dimorphism. Epistome, corniculi, internal malae, hypostomal and palpal structures as in female. Legs. Lengths: I 300 – 330, II 250 – 275, III 250 – 275, IV 300 – 330 µm. Chaetotaxy and morphology of legs as in female. Leg II without sexual dimorphism (Fig. 28). Deutonymph (Fig. 13, n = 10). Dorsal idiosoma. Dorsal shield suboval shape, 415 – 430 long and 260 – 290 wide. Anterior margin of the podonotum curved ventrally to form a vertex, with setae j 1 and z 1. Podonotal and opisthonotal shields neotrichous as in adults. Ventral idiosoma (Fig. 13). Presternal (jugular) and first sternal platelets slightly distinct with pair of St 1 and St 2 setae accordingly; other setae St 3 – St 5 of sternal region inserted in soft cuticle. Peritrematal shields narrow, fused with dorsal shield laterally, with festoon reticulation along entire length. Four pairs of pore-like structures present, three of them inserted in soft cuticle: gp 1, gp 2 and ip 2; ip 3 in small platelet posterior to stigmata. Lyrifissures iv 5 located laterad of St 5 setae. Peritremes straight, 100 – 112 long, shorter than in adults, reaching level of middle of coxae II anteriorly; with internal cell structure. Three pairs of adgenital gland pores gv 2 located on soft cuticle posteriorly to coxae IV. Opisthogastric area with 6 – 7 pairs of simple setae inserted in soft cuticle asymmetrically. Anal shield with folded ornamentation, 75 – 120 long and 70 – 75 wide, with pair of Jv 3, para-anal and post-anal setae; with cribrum and pair of gv 3 glands located antero-laterad to para-anal setae. Dorsal shield is curved on the ventral side posteriorly, does not connect with anal shield, with 6 – 7 pairs of smooth leaf-shaped setae inserted asymetrically. Gnathosoma. As in adults. Legs. Lengths: I 265 – 290, II 210 – 240, III 210 – 240, IV 250 – 290. Chaetotaxy of legs as in adults. Protonymph (Figs 14 – 15, 45 – 46, n = 10). Dorsal idiosoma (Figs 14, 15). Dorsal idiosoma 335 – 345 long and 230 – 235 wide, with weakly sclerotised podonotal and opisthonotal shields. Anterior margin of podonotum curved ventrally to form a vertex, with setae j 1. Podonotum with about 31 – 32 setae on each side (including marginal) and four pairs of pores. Row j with six setae j 1 – j 6 and one additional asymmetrical seta in some specimens, z — row with recognisable z 5 and z 6 setae, homologies of other dorsal setae are obscure. Pygidial region with about 27 – 28 setae on each side and four pairs of pores; divided on two shields. Row J with four pairs of setae: J 1, J 3 – J 5. Other opisthonotal setae multiple. Ventral idiosoma (Figs 15, 46). Presternal and sternal platelets not distinct. Setae St 1 – St 3 and St 5 inserted in soft cuticle; St 5 minute seta, 4 – 5 long. Peritrematal shields and pore-like structuresof periteremaral regions not distinct. Peritremes short (23 – 25), with internal cell structure; with thin ducts leading from stigmata. One pair of adgenital gland pores gv 2 located in soft cuticle posteriorly to coxae IV. Opisthogastric area with 4 – 5 pairs of simple setae inserted in soft cuticle: Zv 1 as long as St 5 setae. Anal shield with folded ornamentation and cribrum, 57 – 63 long and 50 – 57 wide, with para-anal and post-anal setae; pair of glands gv 3 located laterad to para-anal setae. Dorsal shield is curved on the ventral side, does not connect with anal shield; with 2 – 4 pairs of asymmetrical setae, with posterior row of smooth marginal setae. Gnathosoma. As in adults. Legs. Lengths: I 265 – 280, II 200 – 210, III 210 – 215, IV 235 – 250. Chaetotaxy of legs I – IV: coxae 2, 2, 2, 1; trochanters 4, 4,4, 4; femora 10, 8, 5, 4; genua 8, 6, 6, 5; tibiae 8, 6, 7, 7; tarsi II – IV: 17, 17, 17. Structures of pretarsus and coxae I – IV as in adults and deutonymph, coxae III without distinct spine. Larva (Figs 16 – 17, 47, n = 1). Dorsal idiosoma (Figs 16, 47). Dorsal idiosoma 270 long and 260 wide with podonotal, pygidial shields and two pairs of mesonotal sclerites in pygidium region. Posterior surface of pygidium with two pairs of bubbles. Podonotal shield with nine pairs of setae, including five pairs of j row setae; setae j 1 the longest (20) located on anterior margin of shield; stout, pilose, inserted on tubercles. Pygidium with ten pairs of setae: simple and brushform. J - row with three pairs of setae. Ventral idiosoma (Fig. 17). Tritosternum as in adults. Sternal setae St 1 – St 3 inserted in soft cuticle. Periremes and stigmas are absent. Pore-like structures not visible. Opisthogastric area with five pairs of setae inserted in soft cuticle, of which seta Jv 2 the longest; most posterior pair of setae pilose. Anal shield 50 long and 40 wide with pair of para-anal and post-anal setae, simple form. Gnathosoma. Gnathosoma in forward position not curved in ventral side of idiosoma.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE042A524A7EBFC450F89F82E.taxon	materials_examined	Legs. Lengths: I 237, II 188, III 200 µm. Structures of pretarsus and coxae of leg I – III as in protonymphs. Material examined. Holotype female, Russia, Altaiskii krai, North-Western Altai Mountains, Tigeretskii Range, Tigerekskii Nature Reserve, 1300 m a. s. l., 51 ̊ 04 ʹ N, 83 ̊ 02 ʹ E, Betula tortuosa forest, in litter, 18 June 2017, leg. I. I. Marchenko. Paratypes: 29 females, 19 males, 5 deutonymphs, 6 protonymphs, 1 larva, same data as holotype. Other material: 42 females, 15 males, 4 deutonymphs, geographical and locality data as paratypes, 7 August 2013, leg. I. I. Marchenko; 68 females, 53 males, 9 deutonymphs, same geographical region, Picea obovata - Abies sibirica - Pinus sibirica forest, 1000 m a. s. l., in litter, 7 August 2013, leg. I. I. Marchenko; 12 females, 7 males, 11 protonymphs, 5 larvae, same geographical region, 1700 m a. s. l., highland woodland with Pinus sibirica - Picea obovata - Juniperus sabina, 5 June 2017, leg. T. M. Krugova.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE042A524A7EBFC450F89F82E.taxon	etymology	Etymology. The species is named after Tigerek village and Tigeretskii Range in the Altai Mountains, where this species inhabits.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE042A524A7EBFC450F89F82E.taxon	discussion	Remarks. Adults of H. tigerek most similar to H. barguzin Marchenko, 2018, but differ from those having podonotal shield ornamented with small tubercles and festoon reticulation over the entire surface; median region of opisthonotal shield neotrichous, with 17 – 22 setae of J series. Female of H. tigerek with first sternal shield entire in all specimens, ornamented by folds, with pair of iv 2 lyrifissures; soft cuticle between genital and ventri-anal shields with two slightly sclerotised plates. Adults of H. barguzin have podonotal shield ornamented only in anterior and lateral regions; median region of opisthonotum usually with 12 – 13 setae of J series. Female of H barguzin with first sternal shield entire or divided into two, ornamented by folds and lines of tubercles, without lyrifissures; soft cuticle between genital and ventri-anal shields without sclerites. In adult and deutonymph of H. barguzin, the peritremes are of equal length, while deutonymph of H. tigerek have peritremes shorter than adults. Deutonymph periteremes of H. tigerek reaching the level of middle of coxae II anteriorly; versus H. barguzin with periteremes reaching level of spine top of coxae II anteriorly.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE04BA530A7EBFA540F3CFE64.taxon	diagnosis	Diagnosis. Vertex with two pairs of setae: j 1, z 1 and pair of ip 1 glands, inserted very close to j 1 and z 1 setae anteriorly. Podonotum ornamented by small tubercles over the entire surface; median region with sclerotised rounded thickening between po 3 glands. All dorsal setae long, pilose, slightly curved. Median region of opisthonotal shield with 25 – 27 setae of J series, ornamented by tubercles network. First sternal shield divided in pair of platelets with pair of lyrifissures iv 2. Metapodal shields absent. Soft cuticle between genital and ventri-anal shields with pair of setae Jv 1. Adult peritremes shorter, than in deutonymph, reaching anterior level of coxae III. Deutotonymph peritremes reaching top of spine of coxae II. Ventri-anal shield with deep incisions from outer margins of shield to level of posterior edge of anal opening. Adult chelicera without sexual dimorphism.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE04BA530A7EBFA540F3CFE64.taxon	description	Description. Female (Figs 48 – 55, 66 – 73 and 75 – 80, n = 10) Dorsal idiosoma (Figs 48, 66, 75 – 76). Idiosoma suboval, 500 – 525 long and 345 – 380 wide. Anterior margin of the podonotum curved ventrally to form a vertex; with two pairs of setae at arisen bases: j 1 (20 – 22) stout, slightly pilose and z 1 smooth (10 – 12); with pair of ip 1 glands, inserted very close to j 1 and z 1 setae. Podonotal shield strongly neotrichous, with about 83 – 86 setae on each side, including marginal r setae; with four pairs of pore-like structures (glands) po 1 – po 4. Podonotum ornamented with small tubercles over the entire surface of the shield; median region with sclerotised rounded thickening between po- 3 glands. All podonotal setae pilose, slightly curved, 22 – 25 long. Opisthonotal shield ornamented by network of tubercles over the entire surface, with 58 – 60 setae on each side, including marginal R setae, all setae long, pilose, slightly curved; with four pairs of pore-like structures (glands) Po 1 – Po 4 (Fig. 76). Median region neotrichous, with 25 – 27 setae of J series, arrangement asymmetrically, 30 – 35 long. Setae of Z, S series inserted closely in lateral regions, asymmetrically, neotrichous, 30 – 37 long. All dorsal setae surrounded by enlarged basal rings. Marginal dorsal setae of r – R series neotrichous, inserted on high tubercles, elongated, 35 – 37 long. Ventral idiosoma. (Figs 49 – 50, 67 – 72 and 77 – 80). Base of tritosternum 25 – 30 long and 15 – 17 wide, pilose laciniae free from each other along entire length, 55 – 60 long (Fig. 49). Presternal (jugular) platelets irregular shape, weakly sclerotised, with pair of St 1 (20 – 23) setae. First sternal shield divided in two suboval platelets, 30 – 33 long and 18 – 23 wide, with pair of St 2 setae (18 – 20) and pair of lyrifissures iv 2; with a folded surface (Figs 49, 68 – 70, 77 – 80). Median area around first sternal platelets ornamented by rounded tubercles. Second sternal shield divided in pair of suboval platelets, 25 – 30 long, 15 – 17 wide, with two pairs of setae St 3, St 4 (15 – 17) or in some specimens St 4 seta inserted in soft cuticle; with two pairs of lyrifissures iv 3, iv 4. Genital shield 62 – 70 long, 30 – 48 wide, irregularly-shaped, with individual shape in each specimen; expanded anteriorly and posteriorly; anterior margin with fine folding; with genital setae St 5 (13 – 15) and lyrifissures iv 5 on genital shield or in soft cuticle, laterad of shield (Fig. 50). Subtriangular membrane surrounds the genital shield. Genital shield with complex formation, including one median unpaired cap and two pairs of fine sclerites. Endo-, exopodal and metapodal shields absent. Adgenital gland pores gv 2 are multiple, dispersed over the surface: 1 – 2 glands openings located in soft cuticle postero-laterad of genital shield, another three to five openings located in ventri-anal shield. Soft cuticle between genital and ventri-anal shields with pair of setae Jv 1. Peritrematal shields fused anteriorly forming a vertex and fused with dorsal shield laterally; strongly sclerotised, postero-lateral ends are drawn back; ornamented with festoon reticulation along entire length. Five pairs of pore-like structures inserted in peritrematal shield: gp 1, gp 2 and ip 1 – ip 3. Peritremes straight or slightly curved, short, 90 – 102 long, reaching anterior level of coxae III; with internal cell structure. Ventri-anal shield broad, 138 – 148 long and 275 – 285 wide, fused to opisthonotal shield or with deep incisions from outer margins of shield to level of posterior edge of anal opening; with festoon reticulation, with 8 – 9 pairs of smooth opisthogastric setae (13 – 15) inserted anteriorly of incisions. Opisthonotal shield curved on ventral side, with about 15 pilose setae (27 – 30), inserted posteriorly of incisions; with pair of Po 4 glands and the most posterior row of smooth setae. Anal area with pilose para-anal and smooth post-anal setae (15 – 17); anal opening 30 – 33 long; with two lyrifissure on each valve; cribrum located posteriorly of post-anal seta. Pair of glands gv 3 located antero-laterad of para-anal setae. Gnathosoma. (Figs 51 – 55, 73). Fixed digit of chelicera 5 0 – 55 long, with five teeth in addition to apical hook and leaf-shaped pilus dentilis (Fig. 51); movable digit the same length with three teeth in additional to apical hook. Chelicera with long dorsal seta (18 – 20), lateral (antiaxial) and dorsal (paraxial) lyrifissures; with arthrodial corona. Epistome subtriangular, with irregularly serrated edges and smooth pointed median projection (Fig. 52). Corniculi 29 – 30 long and 13 – 15 wide. Internal malae slightly longer than corniculi, with complex three-layer structure as in Fig. 53. Deutosternal groove with 7 – 8 transverse denticulate rows, usually with three paired smooth lateral transverse lines (Fig. 54). The posterior fourth pair of lateral transverse lines arch-shaped, with large denticles, located at the level of pc setae. Setae h 1 – h 3 smooth: h 1 (35 – 37) longest, seta h 2 (13 – 15) shorter than h 3 (25 – 30); pc (18 – 20) serrated. Dorsal side of gnathosoma with the pair of protruding wedge-shaped structures between palpal coxa and trochanter, lateral to epistome (Fig. 55). Palpal trochanter with seta al 1 long and pilose in the distal third; with seta al 2 short and smooth; palp genu with setae al 1 and al 2 pilose in distal third, palp apotele two-tined. Legs. Lengths: I 325 – 340, II 275 – 285, III 27 5 – 285, IV 325 – 340 µm. Chaetotaxy of legs I – IV: coxae 2, 2, 2, 1; trochanters 6 (1 1 / 3 1), 5 (1 1 / 3 0), 5 (1 1 / 3 0), 5 (1 1 / 3 0), femora 13 (2 5 / 4 2), 11 (2 5 / 3 1), 6 (1 4 / 1 0), 6 (1 4 / 1 0); genua 13 (2 6 / 3 2), 12 (2 6 / 2 2), 10 (2 4 / 2 2), 10 (2 5 / 2 1); tibiae 14 (2 6 / 4 2), 10 (2 4 / 2 2), 9 (2 3 / 2 2), 10 (2 4 / 2 2); tarsi 49, 18 (3 7 / 5 3), 18 (3 7 / 5 3), 18 (3 7 / 5 3). All legs with pair of sclerotised claws and pulvillus with five lobes. Pretarsus of legs II – IV with ambulacral stalk, legs I with sessile claws. Coxae I – IV grouped closely together vertically; coxae I split on dorsal side, coxae III – IV with recesses on anterolateral side. Coxae II – III with anterodorsal spines: coxae II with large sharp spine, coxae III with small one. Male. (Figs 56 – 59, 74 and 81, n = 10). Dorsal idiosoma. Dorsal shield suboval shape, 470 – 480 long and 315 – 335 wide. Ornamentation and chaetotaxy similar to that of female. Ventral idiosoma. Tritosternum as in female. Pair weakly sclerotised presternal (jugular) platelets with pair of St 1 setae (Figs 56, 74, 81). First sternal platelet entire, with pair of St 2 setae, with festoon and folds ornamentation, 28 – 30 long and 45 – 50 wide. Second sternal platelet entire, subcordate shape, ornamented by festoon lines and folds; with two pairs of setae St 3, St 4 and one pair of lyrifissures iv 3 in antero-lateral margins; 25 – 35 long on median line and 30 – 45 wide at level of St 3 setae; surrounds the genital opening. Genital opening located at level of coxae III; with a pair of eugenital setae, covering by two platelets (Fig. 57), with pair of genital sclerites as in Fig. 58. Third sternal platelet entire, triangular, 25 – 35 long and 25 – 45 wide, with St 5 setae and pair of iv 5 lyrifissures; in some specimens one opening of iv 5 asymmetrically inserted in soft cuticle. Endo-, exopodal and metapodal shields absent. Peritrematal shields similar to that female, with five pairs of pore-like structures: gp 1, gp 2, i 1 — ip 3. Peritremes short, similar to those of female. Arch of vertex with two dorsal setae: pilose j 1 and smooth thorn-shaped z 1. Ventri-anal shield broad, 145 – 150 long and 235 – 250 wide, entire or with deep lateral incisions as in female; with festoon reticulation; with 7 – 10 pairs of opisthogastric smooth setae and lateral rows of opisthonotal pilose setae, inserted posteriorly of incisions, each with 8 – 9 setae. The most posterior marginal row with elongated smooth setae on raised tubercles. Adgenital gland pores gv 2 are multiple, dispersed over the surface: one pair of glands located in soft cuticle posteriorly of coxae IV, other 4 – 6 openings located in ventri-anal shield. Anal area as in female. Pair of glands gv 3 located antero-laterad of para-anal setae. Opisthonotal shield curved on ventral side with pair of Po 4 glands. Gnathosoma. Fixed digit of chelicera 48 – 50 long, with five teeth in addition to apical hook and leaf-shaped pilus dentilis (Fig. 59). Movable digit of chelicera the same length as fixed digit, tridentate in addition to apical hook, with arthrodial corona at base of digit. Chelicera with long dorsal seta, lateral (antiaxial) and dorsal (paraxial) lyrifissures. Male chelicera lacking spermatodactyl, without sexual dimorphism. Epistome, corniculi, internal malae, hypostomal and palpal structures as in female. Legs. Lengths: I 325 – 340, II 270 – 285, III 270 – 285, IV 315 – 340 µm. Chaetotaxy and morphology of legs as in female. Leg II without sexual dimorphism. Deutonymph (Fig. 60, n = 10). Dorsal idiosoma. Dorsal shield suboval shape, 415 – 450 long and 280 – 315 wide. Anterior margin of the podonotum curved ventrally to form a vertex, with pair of setae j 1, z 1 and lyrifissures ip 1 (Fig. 60). Podonotal and opisthonotal shields neotrichous, pore-like structures as in adults. Ventral idiosoma. Presternal (jugular), first and second sternal platelets slightly distinct with pairs of St 1 – St 3 setae accordingly; other setae St 4, St 5 inserted in soft cuticle. Peritrematal shields narrow, fused with dorsal shield laterally, with festoon reticulation along entire length; separated from peritremes. Five pairs of pore-like structures present, three of them inserted in soft cuticle: gp 1, gp 2 and ip 2; ip 3 in small platelet posterior to stigmata. Peritremes straight or slightly curved, longer than in adults (150 – 157), reaching anterior level of coxae II; with internal cell structure. One pair of adgenital gland pores gv 2 located in soft cuticle posteriorly to coxae IV. Opisthogastric area with 7 – 8 simple and pilose setae inserted in soft cuticle from each side asymmetrically. Anal shield elongated, 75 – 90 long and 50 – 55 wide, with pair of Jv 3, para-anal and post-anal setae; with cribrum and pair of gv 3 glands located antero-laterad to para-anal setae. Opisthonotal shield curved on the ventral side posteriorly, does not connect with anal shield, with 9 – 11 pilose setae inserted asymmetrically and the most posterior row with 14 – 16 smooth, curved setae from each side. Gnathosoma. As in adults. Legs. Lengths: I 280 – 305, II 245 – 250, III 245 – 250, IV 270 – 280. Chaetotaxy of legs as in adults. Protonymph (Figs 61 – 62, n = 10). Dorsal idiosoma (Fig. 61). Dorsal idiosoma 315 – 335 long and 215 – 225 wide, with weakly sclerotised podonotal and pygidial shields. Anterior margin of podonotum curved ventrally to form a vertex, with setae j 1 and z 1. Podonotum with about 37 – 38 setae on each side (including marginal) and four pairs of pores; row j with six pairs of setae j 1 – j 6; z row with recognisable z 5 setae, homologies of other dorsal setae are obscure. Pygidial shield with about 30 – 32 setae on each side from dorsal view and three pairs of pores; pair of Po 4 inserted in curved margins of pygidial shield on ventral side. Row J with three pairs of setae: J 1, J 3 – J 4 and some additional asymmetrical. Other pygidial setae multiple. Ventral idiosoma (Fig. 62). Presternal and sternal platelets not distinct. Setae St 1 – St 3 and St 5 inserted in soft cuticle; St 5 minute seta, 3 – 4 long. Peritrematal shields very narrow, pore-like structures of periteremaral regions not distinct. Peritremes short (23 – 25), with internal cell structure; with thin ducts leading from stigmata. Opisthogastric area with 5 – 6 simple and pilose setae inserted in soft cuticle from each side. Anal shield with cribrum, as long as wide (45 – 50), with pilose para-anal and smooth post-anal setae; pair of glands gv 3 located laterad to para-anal setae. Dorsal shield is curved on the ventral side, does not connect with anal shield; with 4 – 5 pairs of pilose setae, pairs of pores Po 4, and posterior row of pilose curved marginal setae. Gnathosoma. As in adults. Legs. Lengths: I 220 – 250, II 195 – 210, III 195 – 210, IV 220 – 250. Chaetotaxy of legs I – IV: coxae 2, 2, 2, 1; trochanters 4, 4,4, 4; femora 10, 8, 5, 4; genua 8, 6, 6, 5; tibiae 8, 6, 7, 7; tarsi II – IV: 17, 17, 17. Structures of pretarsus and coxae I – IV as in adults and deutonymph, coxae III without distinct spine. Larva (Figs 63 – 65, n = 2). Dorsal idiosoma 225 – 230 long and 165 – 170 wide with podonotal, pygidial shields and three pairs of mesonotal sclerites (Fig. 63). Podonotal shield with nine pairs of setae, including five pairs of j row setae; setae j 1 the longest (18 – 20) located on anterior margin of shield; stout and pilose. Pygidium with eight pairs of setae: four short, brush-form and four marginal on large bases — long, curved, pilose. J - row with two pairs of setae. Ventral idiosoma (Fig. 65). Tritosternum as in adults. Sternal setae St 1 – St 3 and St 5 inserted in soft cuticle. Periremes and stigmas are absent. Pore-like structures not visible. Opisthogastric area with three pairs of setae inserted in soft cuticle, of which seta Jv 2 the longest and smooth; three most posterior pairs of setae pilose, inserted on tubercles. Anal shield 20 long and 40 wide with pair of pilose para-anal and simple post-anal setae. Gnathosoma. Gnathosoma in forward position not curved in ventral side of idiosoma (Figs 64 – 65) with two simple hypostomal setae. Legs. Structures of pretarsus and coxae of leg I – III as in protonymph.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE04BA530A7EBFA540F3CFE64.taxon	materials_examined	Material examined. Holotype female, Russia, Altaiskii krai, North-Western Altai Mountains, Tigeretskii Range, Tigerekskii Nature Reserve, 1100 m a. s. l., 51 ̊ 04 ʹ N, 83 ̊ 02 ʹ E, Picea obovata-Abies sibirica-Pinus sibirica forest, in litter, 24 June 2017, leg. I. I. Marchenko. Paratypes: 7 females, same data as holotype; 7 females, 8 males, 700 m a. s. l., same geographical and locality data, in litter, 20 June 2017, leg. I. I. Marchenko; 1 male, same geographical and locality data, in litter, 7 August 2013, leg. I. I. Marchenko; 1 male, same geographical data, 1500 m a. s. l., Betula tortuosa forest, in litter, 7 August 2013, leg. I. I. Marchenko; 1 female, 3 males, 1 protonymph, same geographical data, 500 m a. s. l., Betula sp. - Abies sibirica forest, in litter, 5 August 2013, leg. I. I. Marchenko; 13 females, 3 males, same geographical data, 900 m a. s. l., in litter, 7 August 2013, leg. I. I. Marchenko; 3 females, 4 males, same geographical data, 1800 m a. s. l., subalpine woodlands with Pinus sibirica-Picea obovata-Juniperus sabina, 5 June 2017, leg. T. M. Krugova. Other material: 32 females, 2 males, 9 deutonymphs, 5 protonymphs, 1 larva, same geographical data as paratypes, 700, 1100 and 1800 m a. s. l., Picea obovata-Abies sibirica-Pinus sibirica forest, in litter, 7 August 2013, 20 and 24 June 2017, leg. I. I. Marchenko; 4 females, 2 males, same geographical data as paratypes, 600 m a. s. l., Caragana sp. - Spirea sp. bush, in litter, 22 June 2017, leg. I. I. Marchenko; 11 females, 1 deutonymph, 1 protonymph, same geographical data, 500 m a. s. l., Populus tremula- Abies sibirica forest, in litter, 7 August 2013, leg. I. I. Marchenko; 1 female, 1 male, 2 deutonymphs, same geographical data, environs of Tigerek village, Kozyr Mt., 500 m a. s. l., Betula sp. - Populus tremula forest, in litter, 24 June 2017, leg. I. I. Marchenko; 10 females, 7 males, 3 deutonymphs, 1 protonymph, Altaiskii krai, North- Western Altai Mountains, Baschelakskii Range, 51 ̊ 06 ʹ N, 84 ̊ 24 ʹ E, 1250 m a. s. l., Larix sibirica forest, in litter, 25 June 2012, leg. R. Yu. Dudko; 2 females, Central Altai Mountains, Ust’ - Koksa Distr., environs of lower Multinskoe lake, Katunskii Range, 50 ̊ 00 ʹ N, 85 ̊ 49 ʹ E, 1500 m a. s. l., Abies sibirica-Pinus sibirica forest, in litter, 12 July 2013, leg. I. I. Marchenko.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE04BA530A7EBFA540F3CFE64.taxon	etymology	Etymology. The species is named after russian folk dance “ kazachok ”, as the Cossacks lived in the Tigirek fort of Altai Mountains since the 18 th century.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE04BA530A7EBFA540F3CFE64.taxon	discussion	Remarks. Adults of Halozercon kazachok most similar to H. karacholana Wiśniewski et al. 1992, but has a number of differences. Vertex of H. kazachok with pairs of j 1, z 1 setae and pore-like structures ip 1 grouped closely together in anterior margin of shield; H. karacholana with z 1 setae inserted far behind j 1 setae. Dorsal setae of H. kazachok long and curved: podonotal setae 22 – 25 µm, opisthonotal setae 30 – 35 µm; H. karacholana with podonotal setae shorter and straight: 10 – 20 µm and opisthonotal 22 – 25 µm. Ventri-anal shield of H. kazachok with incisions extending to posterior edge of anal opening; female of H. kazachok with opisthogastral setae Jv 1 inserted in soft cuticle. Ventri-anal shield of H. karacholana with incisions extending to anterior edge of anal opening; female opisthogastral setae Jv 1 inserted in ventri-anal shield. Adult chelicera of H. kazachok sp. n. without sexual dimorphism, whereas fixed digit of male chelicera of H. karacholana with two basal and 5 – 6 small distal teeth and female fixed digit with five teeth.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE055A53DA7EBFB3E0E61FA6B.taxon	diagnosis	Diagnosis. Vertex with two pairs of setae: j 1 and z 1, with pair of glands ip 1 inserted in middle between j 1 and z 1 setae. Surface of podonotal shield ornamented by small tubercles in anterior and lateral regions, median region with indistinct reticulation. All podonotal setae smooth. Opisthonotal shield with linear reticulation in median region, with small tubercles in lateral regions. Median region with 14 – 17 slightly serrated setae of J series. First sternal platelets divided, with pair of lyrifissures iv 2. Metapodal shields absent. Setae Jv 1 located in anterior margin of ventri-anal shields or in some specimens in soft cuticle between genital and ventri-anal shields. Peritremes reaching level of posterior 1 / 3 part of coxae II. Ventri-anal shield entire. Adult chelicera without sexual dimorphism.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE055A53DA7EBFB3E0E61FA6B.taxon	description	Description. Female (Figs 82 – 86, 88 – 98, n = 3) Dorsal idiosoma. Idiosoma suboval, 580 – 595 long and 377 – 390 wide. Anterior margin of the podonotum curved ventrally to form a vertex with two pairs of setae at arisen bases: j 1 (12 – 15) stout, smooth and z 1 setae (10 – 12) smooth, thorn-like; with pair of glands ip 1 inserted in middle between j 1 and z 1 setae. Podonotal shield strongly neotrichous with about 63 – 66 setae on each side, including marginal r setae; ornamented by small tubercles in anterior and lateral regions, median region with indistinct reticulation (Figs 82, 88 – 89). All podonotal setae smooth, 12 – 15 long. Four pairs of pore-like structures (glands) po 1 – po 4 located in podonotum, glands po 1 visible in ventral side. Opisthonotal shield linear reticulated in median region, with small tubercles in lateral regions; with four pairs of pore-like structures (glands) Po 1 – Po 4 and with 48 – 53 neotrichous setae on each side, including marginal R setae. Median region with 14 – 17 slightly serrated setae of J series (Fig. 90), lateral region with setae of Z, S series inserted closely, smooth or pilose (Fig. 91), all setae arrangement asymmetrically, 15 – 20 long. All dorsal setae surrounded by enlarged basal rings. Marginal setae of r – R series neotrichous, elongated, pilose, 25 – 30 long, inserted on high tubercles. Ventral idiosoma. (Figs 83, 92 – 98). Base of tritosternum 30 – 35 long and 20 – 22 wide, pilose laciniae free from each other along entire length, 60 – 65 long. Presternal (jugular) platelets irregular shape, weakly sclerotised, with pair of St 1 setae (23 – 25). First pair of sternal platelets divided, suboval, 25 – 35 long and 13 – 18 wide, with pair of St 2 setae (20 – 22) and pair of lyrifissures iv 2 (Figs 83, 92 – 95). Area between jugular and first sternal platelets ornamented by small rounded tubercles. Second pair of sternal platelets divided, suboval, 30 – 33 long and 12 – 15 wide, with two pairs of setae St 3, St 4 (15 – 17) and two pairs of lyrifissures iv 3, iv 4 (Figs 92 – 95). Genital shield stocky form, 58 – 67 long, 48 – 52 wide, irregularly-shaped, with individual shape in each specimen; expanded anteriorly and posteriorly; anterior margin with fine folding; with genital setae St 5 (15 – 16) and lyrifissures iv 5 on genital shield (Fig. 95). Subtriangular membrane surrounds the genital shield. Endo-, exopodal and metapodal shields absent. Adgenital gland pores gv 2 are multiple, dispersed over the surface: one gland opening located in soft cuticle postero-laterad of genital shield, another 4 – 6 openings located in ventri-anal shield. Setae Jv 1 located in anterior margin of ventri-anal shields or in some specimens in soft cuticle between genital and ventri-anal shields. Peritrematal shields fused anteriorly forming a vertex and fused with dorsal shield laterally; strongly sclerotised, postero-lateral ends are drawn back; ornamented with festoon reticulation along entire length. Five pairs of pore-like structures inserted in peritrematal shield: gp 1, gp 2 and ip 1 – ip 3. Peritremes straight or slightly curved, 123 – 127 long, reaching level of posterior 1 / 3 part of coxae II; with internal cell structure (Fig. 96). Ventrianal shield entire, broad (Figs 93, 97 – 98), 190 – 200 long and 325 – 335 wide, fused to opisthonotal shield; with festoon reticulation, with 8 – 9 pairs of smooth opisthogastric setae (15 – 20). Opisthonotal shield curved on ventral side, with 5 – 6 pilose setae (23 – 25). Anal area with simple pre-anal (13 – 15) and post-anal setae (17); anal opening 37 long; with two lyrifissure on each valve; cribrum located posteriorly of post-anal seta. Pair of glands gv 3 located antero-laterad of para-anal setae. Gnathosoma. (Figs 84 – 86). Fixed digit of chelicera 60 – 61 long, with five teeth in addition to apical hook and leaf-shaped pilus dentilis (Fig. 84); movable digit the same length with three teeth in additional to apical hook. Chelicera with long dorsal seta, lateral (antiaxial) and dorsal (paraxial) lyrifissures; with arthrodial corona. Epistome subtriangular, with irregularly serrated edges and smooth pointed median projection (Fig. 85). Corniculi 30 – 31 long and 14 – 15 wide. Internal malae slightly longer than corniculi, with complex three-layer structure (Fig. 86). Deutosternal groove with 7 – 8 transverse denticulate rows. The posterior pair of lateral transverse lines archshaped, with large denticles, located at the level of pc setae. Setae h 1 – h 3 smooth: h 1 (45 – 47) longest, seta h 2 (15 – 17) shorter than h 3 (25 – 30); pc (20 – 22) serrated. Palpal trochanter with seta al 1 long and pilose in the distal third; with seta al 2 short and smooth; palp genu with setae al 1 and al 2 pilose in distal third, palp apotele two-tined. Legs. Lengths: I 380 – 385, II 315 – 325, III 31 5 – 325, IV 375 – 385 µm. Chaetotaxy of legs I – IV: coxae 2, 2, 2, 1; trochanters 6 (1 1 / 3 1), 5 (1 1 / 3 0), 5 (1 1 / 3 0), 5 (1 1 / 3 0), femora 13 (2 5 / 4 2), 11 (2 5 / 3 1), 6 (1 4 / 1 0), 6 (1 4 / 1 0); genua 13 (2 6 / 3 2), 12 (2 6 / 2 2), 10 (2 4 / 2 2), 10 (2 5 / 2 1); tibiae 14 (2 6 / 4 2), 10 (2 4 / 2 2), 9 (2 3 / 2 2), 10 (2 4 / 2 2); tarsi 49, 18 (3 7 / 5 3), 18 (3 7 / 5 3), 18 (3 7 / 5 3). All legs with pair of sclerotised claws and pulvillus with five lobes. Pretarsus of legs II – IV with ambulacral stalk, legs I with sessile claws. Coxae I – IV grouped closely together vertically; coxae I split on dorsal side, coxae III – IV with recesses on anterolateral side. Coxae II with antero-dorsal large sharp spine. Male. (87, 99 – 101, n = 2). Dorsal idiosoma. Dorsal shield suboval shape, 515 – 575 long and 310 – 360 wide. Ornamentation and chaetotaxy similar to that of female. Ventral idiosoma. Tritosternum as in female. Pair weakly sclerotised presternal (jugular) platelets with pair of St 1 setae (Figs 87, 99 – 101). First sternal platelet entire, with pair of St 2 setae, with line ornamentation, 30 – 32 long and 50 – 55 wide. Second sternal platelet entire, subcordate shape, with indistinct reticulation; with two pairs of setae St 3, St 4 and one pair of lyrifissures iv 3 in antero-lateral margins; 40 – 42 long on median line and 48 – 55 wide at level of St 3 setae; surrounds the genital opening. Genital opening located at level of coxae III; with a pair of eugenital setae, covering by two platelets (Figs 100 – 101), with pair of genital sclerites. Third sternal platelet triangular, divided in three fragments, with St 5 setae and pair of iv 5 lyrifissures inserted in soft cuticle. Endo-, exopodal and metapodal shields absent. Peritrematal shields similar to that female, with five pairs of pore-like structures: gp 1, gp 2, i 1 — ip 3. Peritremes similar to those of female, 115 – 125 long. Arch of vertex with two dorsal setae: smooth, stout j 1 and smooth, thorn-like z 1. Ventri-anal shield broad, entire, with festoon reticulation (Figs 87, 99), 165 – 180 long and 275 – 310 wide; with 7 – 10 pairs of opisthogastric smooth setae and 4 – 6 opisthonotal slightly pilose setae. Adgenital gland pores gv 2 are multiple, dispersed over the surface: one pair of glands located in soft cuticle posteriorly of coxae IV, other 3 – 5 openings located in ventri-anal shield. Anal area as in female. Pair of glands gv 3 located antero-laterad of para-anal setae. Gnathosoma. Fixed digit of chelicera 55 long, with five teeth in addition to apical hook and leaf-shaped pilus dentilis. Movable digit of chelicera the same length as fixed digit, tridentate in addition to apical hook. Male chelicera lacking spermatodactyl, without sexual dimorphism. Epistome, corniculi, internal malae, hypostomal and palpal structures as in female. Legs. Lengths: I 325 – 340, II 270 – 285, III 270 – 285, IV 315 – 340 µm. Chaetotaxy and morphology of legs as in female. Leg II without sexual dimorphism.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE055A53DA7EBFB3E0E61FA6B.taxon	materials_examined	Material examined. Holotype female, Russia, Altaiskii krai, North-Western Altai Mountains, Tigeretskii Range, Tigerekskii Nature Reserve, 1300 m a. s. l., 51 ̊ 04 ʹ N, 83 ̊ 02 ʹ E, Betula tortuosa forest, in litter, 18 June 2017, leg. I. I. Marchenko. Paratypes: 2 females, same data as holotype; 1 male, same data as holotype; 1 male, Tigerekskii Nature Reserve, environs of Tigerek village, Kozyr Mt., 500 m a. s. l., 51 ̊ 04 ʹ N, 83 ̊ 02 ʹ E, in litter, 24 June 2017, leg. I. I. Marchenko.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE055A53DA7EBFB3E0E61FA6B.taxon	etymology	Etymology. The species is named capitaneus, as captain of team, due to its large size compared with other species of Halozercon found in Tigerekskii Nature Reserve.	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
038D87AFE055A53DA7EBFB3E0E61FA6B.taxon	discussion	Remarks. Adults of Halozercon capitaneus are most similar to H. karacholana Wiśniewski et al. 1992, but differ from those having larger size of idiosoma; ornamentation of dorsal shields — median regions of podonotal and opisthonotal shields with smooth reticulation; pore-like structures Po 4 (glands) located in posterior part of opisthonotum at dorsal side; all podonotal setae smooth except marginal; opisthonotal setae of J series slightly serrated (which is visible only at 1000 magnification). Adults of Halozercon karacholana having smaller size of idiosoma; median regions of podonotal and opisthonotal shields with tubercles and festoon network, with tubercle or festoon patterns around J 5 setae and po 3 glands (Figs 102 – 103); Po 4 glands located in posterior part of opisthonotum at ventral side; all dorsal setae pilose (Fig. 104).	en	Marchenko, Irina I. (2019): Three new species of Halozercon (Acari: Mesostigmata: Zerconidae) from Altai Mountains in South Siberia (Russia). Zootaxa 4568 (3): 401-434, DOI: 10.11646/zootaxa.4568.3.1
