taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
038BAE5EFFD3A229F968FB05FD20F7A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037238/files/figure.png	https://doi.org/10.5281/zenodo.5037238	Fig. 15. Elytral sculpture and male genitalia of Eumetopus Balfour-Browne, 1949. A–H – elytral sculpture, same specimens as in Fig. 14: A – E. acutimontis Ji & Jäch, 1998; B – E. asperatus (Champion, 1919); C – E. bullatus (Sharp, 1875); D – E. flavidulus (Sharp, 1890); E – E. maindroni (Régimbart, 1903); F – E. schuelkei Jäch, 2002; G – E. tibialis Ji & Jäch, 1998; H – E. weigeli Skale & Jäch, 2003. I–L – male genitalia of examined specimens, large basal portion of the phallobase omitted (dorsal and lateral view): I – E. acutimontis from Vietnam; J – E. flavidulus from India: Andhra Pradesh; K – E. maindroni from India: Gujarat; L – E. weigeli from India: Uttarakhand.	Fig. 15. Elytral sculpture and male genitalia of Eumetopus Balfour-Browne, 1949. A–H – elytral sculpture, same specimens as in Fig. 14: A – E. acutimontis Ji & Jäch, 1998; B – E. asperatus (Champion, 1919); C – E. bullatus (Sharp, 1875); D – E. flavidulus (Sharp, 1890); E – E. maindroni (Régimbart, 1903); F – E. schuelkei Jäch, 2002; G – E. tibialis Ji & Jäch, 1998; H – E. weigeli Skale & Jäch, 2003. I–L – male genitalia of examined specimens, large basal portion of the phallobase omitted (dorsal and lateral view): I – E. acutimontis from Vietnam; J – E. flavidulus from India: Andhra Pradesh; K – E. maindroni from India: Gujarat; L – E. weigeli from India: Uttarakhand.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A229F968FB05FD20F7A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037236/files/figure.png	https://doi.org/10.5281/zenodo.5037236	Fig. 14. Habitus photographs of the known species of Eumetopus Balfour-Browne, 1949. A – E. acutimontis Ji & Jäch, 1998, paratype from China: Hainan (NHMW); B – E. asperatus (Champion, 1919), specimen from Nepal: Koshi Valley (NMPC); C – E. bullatus (Sharp, 1875), specimen from India: Maharashtra, Sawantwadi (NMPC); D – E. flavidulus (Sharp, 1890), specimen from India: Meghalaya, Bagmara (NHMW); E – E. maindroni (Régimbart, 1903), specimen from India: Maharashtra, Sawantwadi (NHMW); F – E. schuelkei Jäch, 2002, paratype from central Laos; G – E. tibialis Ji & Jäch, 1998 from Thailand, Ban Na env. (NMPC); H – E. weigeli Skale & Jäch, 2003, holotype from Nepal (NHMW). Scale bars: 1 mm.	Fig. 14. Habitus photographs of the known species of Eumetopus Balfour-Browne, 1949. A – E. acutimontis Ji & Jäch, 1998, paratype from China: Hainan (NHMW); B – E. asperatus (Champion, 1919), specimen from Nepal: Koshi Valley (NMPC); C – E. bullatus (Sharp, 1875), specimen from India: Maharashtra, Sawantwadi (NMPC); D – E. flavidulus (Sharp, 1890), specimen from India: Meghalaya, Bagmara (NHMW); E – E. maindroni (Régimbart, 1903), specimen from India: Maharashtra, Sawantwadi (NHMW); F – E. schuelkei Jäch, 2002, paratype from central Laos; G – E. tibialis Ji & Jäch, 1998 from Thailand, Ban Na env. (NMPC); H – E. weigeli Skale & Jäch, 2003, holotype from Nepal (NHMW). Scale bars: 1 mm.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A229F968FB05FD20F7A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037218/files/figure.png	https://doi.org/10.5281/zenodo.5037218	Fig. 6. Male genitalia and associated structures of Epimetopidae. A–H – Eupotemus Ji & Jäch, 1998 (A–F – E. smithi sp. nov.; G – E. cameroonensis sp. nov.; H – E. carinaticollis (Basilewsky, 1956)). I–R – Epimetopus Lacordaire, 1854 (I–N – E. mendeli Fikáček et al., 2011; O – E. cf. burruyacu Oliva, 1986; P – E. multiporus Perkins, 2012; Q – E. clandestinus Perkins, 2012; R – E. thermarum Schwarz & Barber, 1917). S–Z – Eumetopus Balfour-Browne, 1949 (S–Y – E. acutimontis Ji & Jäch, 1998; Z – E. bullatus (Sharp, 1875)).A, I, S – sternite VIII; B, J, T – tergite VIII; C, K, U – sternite IX; D, L, V – aedeagus dorsally; E, M, W – aedeagus laterally; F, N, X – aedeagus ventrally (basal part omitted in X); Y – sperm pump. Color coding: green – ventral projections of the median lobe; red – median lobe; blue – paramere (pale blue – dorsal lobe; dark blue – ventral lobe). Not to scale.	Fig. 6. Male genitalia and associated structures of Epimetopidae. A–H – Eupotemus Ji & Jäch, 1998 (A–F – E. smithi sp. nov.; G – E. cameroonensis sp. nov.; H – E. carinaticollis (Basilewsky, 1956)). I–R – Epimetopus Lacordaire, 1854 (I–N – E. mendeli Fikáček et al., 2011; O – E. cf. burruyacu Oliva, 1986; P – E. multiporus Perkins, 2012; Q – E. clandestinus Perkins, 2012; R – E. thermarum Schwarz & Barber, 1917). S–Z – Eumetopus Balfour-Browne, 1949 (S–Y – E. acutimontis Ji & Jäch, 1998; Z – E. bullatus (Sharp, 1875)).A, I, S – sternite VIII; B, J, T – tergite VIII; C, K, U – sternite IX; D, L, V – aedeagus dorsally; E, M, W – aedeagus laterally; F, N, X – aedeagus ventrally (basal part omitted in X); Y – sperm pump. Color coding: green – ventral projections of the median lobe; red – median lobe; blue – paramere (pale blue – dorsal lobe; dark blue – ventral lobe). Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A229F968FB05FD20F7A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037224/files/figure.png	https://doi.org/10.5281/zenodo.5037224	Fig. 9. Habitus photographs of the species of the Eupotemus limicola species group, holotypes: A–C – E. bilobatus sp. nov.; D–F – E. cameroonensis sp. nov.; G–I – E. limicola (Delève, 1967).	Fig. 9. Habitus photographs of the species of the Eupotemus limicola species group, holotypes: A–C – E. bilobatus sp. nov.; D–F – E. cameroonensis sp. nov.; G–I – E. limicola (Delève, 1967).	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A229F968FB05FD20F7A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037226/files/figure.png	https://doi.org/10.5281/zenodo.5037226	Fig. 10. Habitus photographs of the species of the Eupotemus limicola species group, holotypes (A–F) and differences between species groups in pronotal morphology (G–H). A–C – E. smithi sp. nov.; D–F – E. ophioglossus sp. nov.; G–I – pronotal sculpture: G – E. carinaticollis species group; H – E. limicola species group.	Fig. 10. Habitus photographs of the species of the Eupotemus limicola species group, holotypes (A–F) and differences between species groups in pronotal morphology (G–H). A–C – E. smithi sp. nov.; D–F – E. ophioglossus sp. nov.; G–I – pronotal sculpture: G – E. carinaticollis species group; H – E. limicola species group.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A229F968FB05FD20F7A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037230/files/figure.png	https://doi.org/10.5281/zenodo.5037230	Fig. 12. Habitus photographs of the species of the Eupotemus carinaticollis species group, holotypes: A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. uluguru sp. nov.; G–I – E. taianus sp. nov.	Fig. 12. Habitus photographs of the species of the Eupotemus carinaticollis species group, holotypes: A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. uluguru sp. nov.; G–I – E. taianus sp. nov.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A229F968FB05FD20F7A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037240/files/figure.png	https://doi.org/10.5281/zenodo.5037240	Fig. 16. Photographs of Epimetopus species, including two specimens of questionable provenance (E–G).A – E. flavicaptus Fikáček, Barclay & Perkins, 2011; B – E. deceptus Perkins, 2012; C – E. tridens Perkins, 2012; D – E. hintoni Balfour-Browne, 1949; E – Epimetopus sp. (ʻZambiaʼ); F–G – E. cf. burruyacu Oliva, 1986 (ʻSaudi Arabiaʼ) (F – dorsal view; G – aedeagus). Scale bars for A–F = 0.5 mm. A–D adopted from Pൾ*©κංඇඌ (2012).	Fig. 16. Photographs of Epimetopus species, including two specimens of questionable provenance (E–G).A – E. flavicaptus Fikáček, Barclay & Perkins, 2011; B – E. deceptus Perkins, 2012; C – E. tridens Perkins, 2012; D – E. hintoni Balfour-Browne, 1949; E – Epimetopus sp. (ʻZambiaʼ); F–G – E. cf. burruyacu Oliva, 1986 (ʻSaudi Arabiaʼ) (F – dorsal view; G – aedeagus). Scale bars for A–F = 0.5 mm. A–D adopted from Pൾ*©κංඇඌ (2012).	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A229F968FB05FD20F7A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037212/files/figure.png	https://doi.org/10.5281/zenodo.5037212	Fig. 3. Thoracic morphology of the genera of Epimetopidae. A–B, J, Q – Eupotemus smithi sp. nov.; C–D, L–O – Eumetopus schuelkei Jäch, 2002; E–F – Epimetopus mendeli Fikáček, Barclay & Perkins, 2012; G–I, P – Epimetopus costaricensis Perkins, 1979. A–I – prothorax in ventral view (B, D, F, H – detail of ventral sculpture of the pronotal hood; I – detail of closed procoxal cavity. J–L – meso- and metathorax in ventral view. M–N – elytron in ventral view (M – general view; N – detail of the ventral ridge, notice the spiny surface on inner face of the ridge (in) and on the elytral plectrum posteriorly of it (pl)). O – mesotrochanter. P–Q – details of pretarsus with the leaf-like empodial seta. Not to scale.	Fig. 3. Thoracic morphology of the genera of Epimetopidae. A–B, J, Q – Eupotemus smithi sp. nov.; C–D, L–O – Eumetopus schuelkei Jäch, 2002; E–F – Epimetopus mendeli Fikáček, Barclay & Perkins, 2012; G–I, P – Epimetopus costaricensis Perkins, 1979. A–I – prothorax in ventral view (B, D, F, H – detail of ventral sculpture of the pronotal hood; I – detail of closed procoxal cavity. J–L – meso- and metathorax in ventral view. M–N – elytron in ventral view (M – general view; N – detail of the ventral ridge, notice the spiny surface on inner face of the ridge (in) and on the elytral plectrum posteriorly of it (pl)). O – mesotrochanter. P–Q – details of pretarsus with the leaf-like empodial seta. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A237FB43F8A2FF02FE17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037222/files/figure.png	https://doi.org/10.5281/zenodo.5037222	Fig. 8. Male genitalia of the species of the Eupotemus limicola species group, holotypes.A–C, R–S – E. bilobatus sp. nov.; D–F, T–U – E. cameroonensis sp. nov.; G–I, V–W – E. ophioglossus sp. nov.; J–L, X–Y – E. smithi sp. nov.; M–Q – E. limicola (Delève, 1967). A, D, G, J, M – dorsal view; B, E, H, K, N – lateral view; C, F, I, L – ventral view; O – apex of the median lobe, dorsal view; P, R, T, V, X – fork of the median lobe; Q, S, U, W, Y – apex of parameres in lateral view.	Fig. 8. Male genitalia of the species of the Eupotemus limicola species group, holotypes.A–C, R–S – E. bilobatus sp. nov.; D–F, T–U – E. cameroonensis sp. nov.; G–I, V–W – E. ophioglossus sp. nov.; J–L, X–Y – E. smithi sp. nov.; M–Q – E. limicola (Delève, 1967). A, D, G, J, M – dorsal view; B, E, H, K, N – lateral view; C, F, I, L – ventral view; O – apex of the median lobe, dorsal view; P, R, T, V, X – fork of the median lobe; Q, S, U, W, Y – apex of parameres in lateral view.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A237FB43F8A2FF02FE17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037226/files/figure.png	https://doi.org/10.5281/zenodo.5037226	Fig. 10. Habitus photographs of the species of the Eupotemus limicola species group, holotypes (A–F) and differences between species groups in pronotal morphology (G–H). A–C – E. smithi sp. nov.; D–F – E. ophioglossus sp. nov.; G–I – pronotal sculpture: G – E. carinaticollis species group; H – E. limicola species group.	Fig. 10. Habitus photographs of the species of the Eupotemus limicola species group, holotypes (A–F) and differences between species groups in pronotal morphology (G–H). A–C – E. smithi sp. nov.; D–F – E. ophioglossus sp. nov.; G–I – pronotal sculpture: G – E. carinaticollis species group; H – E. limicola species group.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A237FB43F8A2FF02FE17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037228/files/figure.png	https://doi.org/10.5281/zenodo.5037228	Fig. 11. Male genitalia of the Eupotemus carinaticollis species group, holotypes.A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. taianus sp. nov.; G–I – E. uluguru sp. nov. A, D, G – dorsal view; B, E, H – lateral view; C, F, I – ventral view.	Fig. 11. Male genitalia of the Eupotemus carinaticollis species group, holotypes.A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. taianus sp. nov.; G–I – E. uluguru sp. nov. A, D, G – dorsal view; B, E, H – lateral view; C, F, I – ventral view.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A237FB43F8A2FF02FE17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037234/files/figure.png	https://doi.org/10.5281/zenodo.5037234	Fig. 13. Known distribution of Eupotemus species.A – species of E. limicola species group; B – species of E. carinaticollis species group.	Fig. 13. Known distribution of Eupotemus species.A – species of E. limicola species group; B – species of E. carinaticollis species group.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A237FB43F8A2FF02FE17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037218/files/figure.png	https://doi.org/10.5281/zenodo.5037218	Fig. 6. Male genitalia and associated structures of Epimetopidae. A–H – Eupotemus Ji & Jäch, 1998 (A–F – E. smithi sp. nov.; G – E. cameroonensis sp. nov.; H – E. carinaticollis (Basilewsky, 1956)). I–R – Epimetopus Lacordaire, 1854 (I–N – E. mendeli Fikáček et al., 2011; O – E. cf. burruyacu Oliva, 1986; P – E. multiporus Perkins, 2012; Q – E. clandestinus Perkins, 2012; R – E. thermarum Schwarz & Barber, 1917). S–Z – Eumetopus Balfour-Browne, 1949 (S–Y – E. acutimontis Ji & Jäch, 1998; Z – E. bullatus (Sharp, 1875)).A, I, S – sternite VIII; B, J, T – tergite VIII; C, K, U – sternite IX; D, L, V – aedeagus dorsally; E, M, W – aedeagus laterally; F, N, X – aedeagus ventrally (basal part omitted in X); Y – sperm pump. Color coding: green – ventral projections of the median lobe; red – median lobe; blue – paramere (pale blue – dorsal lobe; dark blue – ventral lobe). Not to scale.	Fig. 6. Male genitalia and associated structures of Epimetopidae. A–H – Eupotemus Ji & Jäch, 1998 (A–F – E. smithi sp. nov.; G – E. cameroonensis sp. nov.; H – E. carinaticollis (Basilewsky, 1956)). I–R – Epimetopus Lacordaire, 1854 (I–N – E. mendeli Fikáček et al., 2011; O – E. cf. burruyacu Oliva, 1986; P – E. multiporus Perkins, 2012; Q – E. clandestinus Perkins, 2012; R – E. thermarum Schwarz & Barber, 1917). S–Z – Eumetopus Balfour-Browne, 1949 (S–Y – E. acutimontis Ji & Jäch, 1998; Z – E. bullatus (Sharp, 1875)).A, I, S – sternite VIII; B, J, T – tergite VIII; C, K, U – sternite IX; D, L, V – aedeagus dorsally; E, M, W – aedeagus laterally; F, N, X – aedeagus ventrally (basal part omitted in X); Y – sperm pump. Color coding: green – ventral projections of the median lobe; red – median lobe; blue – paramere (pale blue – dorsal lobe; dark blue – ventral lobe). Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A237FB43F8A2FF02FE17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037224/files/figure.png	https://doi.org/10.5281/zenodo.5037224	Fig. 9. Habitus photographs of the species of the Eupotemus limicola species group, holotypes: A–C – E. bilobatus sp. nov.; D–F – E. cameroonensis sp. nov.; G–I – E. limicola (Delève, 1967).	Fig. 9. Habitus photographs of the species of the Eupotemus limicola species group, holotypes: A–C – E. bilobatus sp. nov.; D–F – E. cameroonensis sp. nov.; G–I – E. limicola (Delève, 1967).	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFD3A237FB43F8A2FF02FE17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037216/files/figure.png	https://doi.org/10.5281/zenodo.5037216	Fig. 5. Morphology of Epimetopidae. A–B – ventral morphology: A – Eupotemus smithi sp. nov.; B – Epimetopus mendeli Fikáček, Perkins & Barclay, 2011. C–F – hind wings: C – Eumetopus schulkei Jäch, 2002; D – Eupotemus smithi sp. nov.; E – Epimetopus mendeli, adopted from Fංκගඹൾκ et al. (2011); F – Epimetopus costaricensis Perkins, 1979. G – dorsal part of the metathorax. H – scutellum. I–K – abdominal ventrites (I – Eumetopus schuelkei; J – Eupotemus smithi sp. nov.; K – Epimetopus mendeli). L–M – ovipositor (L – Eumetopus schuelkei; M – Eupotemus smithi). O – elytral punctation, slide-mounted elytra of Epimetopus costaricensis. P – female of Eumetopus acutimontis in ventral view, with the egg cases carried under the abdomen. Not to scale.	Fig. 5. Morphology of Epimetopidae. A–B – ventral morphology: A – Eupotemus smithi sp. nov.; B – Epimetopus mendeli Fikáček, Perkins & Barclay, 2011. C–F – hind wings: C – Eumetopus schulkei Jäch, 2002; D – Eupotemus smithi sp. nov.; E – Epimetopus mendeli, adopted from Fංκගඹൾκ et al. (2011); F – Epimetopus costaricensis Perkins, 1979. G – dorsal part of the metathorax. H – scutellum. I–K – abdominal ventrites (I – Eumetopus schuelkei; J – Eupotemus smithi sp. nov.; K – Epimetopus mendeli). L–M – ovipositor (L – Eumetopus schuelkei; M – Eupotemus smithi). O – elytral punctation, slide-mounted elytra of Epimetopus costaricensis. P – female of Eumetopus acutimontis in ventral view, with the egg cases carried under the abdomen. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037222/files/figure.png	https://doi.org/10.5281/zenodo.5037222	Fig. 8. Male genitalia of the species of the Eupotemus limicola species group, holotypes.A–C, R–S – E. bilobatus sp. nov.; D–F, T–U – E. cameroonensis sp. nov.; G–I, V–W – E. ophioglossus sp. nov.; J–L, X–Y – E. smithi sp. nov.; M–Q – E. limicola (Delève, 1967). A, D, G, J, M – dorsal view; B, E, H, K, N – lateral view; C, F, I, L – ventral view; O – apex of the median lobe, dorsal view; P, R, T, V, X – fork of the median lobe; Q, S, U, W, Y – apex of parameres in lateral view.	Fig. 8. Male genitalia of the species of the Eupotemus limicola species group, holotypes.A–C, R–S – E. bilobatus sp. nov.; D–F, T–U – E. cameroonensis sp. nov.; G–I, V–W – E. ophioglossus sp. nov.; J–L, X–Y – E. smithi sp. nov.; M–Q – E. limicola (Delève, 1967). A, D, G, J, M – dorsal view; B, E, H, K, N – lateral view; C, F, I, L – ventral view; O – apex of the median lobe, dorsal view; P, R, T, V, X – fork of the median lobe; Q, S, U, W, Y – apex of parameres in lateral view.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037224/files/figure.png	https://doi.org/10.5281/zenodo.5037224	Fig. 9. Habitus photographs of the species of the Eupotemus limicola species group, holotypes: A–C – E. bilobatus sp. nov.; D–F – E. cameroonensis sp. nov.; G–I – E. limicola (Delève, 1967).	Fig. 9. Habitus photographs of the species of the Eupotemus limicola species group, holotypes: A–C – E. bilobatus sp. nov.; D–F – E. cameroonensis sp. nov.; G–I – E. limicola (Delève, 1967).	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037226/files/figure.png	https://doi.org/10.5281/zenodo.5037226	Fig. 10. Habitus photographs of the species of the Eupotemus limicola species group, holotypes (A–F) and differences between species groups in pronotal morphology (G–H). A–C – E. smithi sp. nov.; D–F – E. ophioglossus sp. nov.; G–I – pronotal sculpture: G – E. carinaticollis species group; H – E. limicola species group.	Fig. 10. Habitus photographs of the species of the Eupotemus limicola species group, holotypes (A–F) and differences between species groups in pronotal morphology (G–H). A–C – E. smithi sp. nov.; D–F – E. ophioglossus sp. nov.; G–I – pronotal sculpture: G – E. carinaticollis species group; H – E. limicola species group.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037234/files/figure.png	https://doi.org/10.5281/zenodo.5037234	Fig. 13. Known distribution of Eupotemus species.A – species of E. limicola species group; B – species of E. carinaticollis species group.	Fig. 13. Known distribution of Eupotemus species.A – species of E. limicola species group; B – species of E. carinaticollis species group.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037210/files/figure.png	https://doi.org/10.5281/zenodo.5037210	Fig. 2. Morphology of mouthparts and head appendages of the genera of Epimetopidae.A–I – Eupotemus Ji & Jäch, 1998: A, C, E, F, H – E. smithi sp. nov., B, D, G – E. limicola (Delève, 1967), I – E. cameroonensis sp. nov. J–Q – Eumetopus Balfour-Browne, 1949: J, L, M, N, P – E. schuelkei Jäch, 2002, Q – E. bullatus (Sharp, 1875), K, L*, O – E. sp. from Sri Lanka. R–Z, a–l – Epimetopus Lacordaire, 1854: R–W – E. mendeli Fikáček et al., 2011, X–Z, a–c – E. costaricensis Perkins, 1979, g, j – E. thermarum Schwarz & Barber, 1917, e, k – E. trogoides (Sharp, 1874), h – E. costatus group, f, i, l – E. punctipennis Perkins, 1979. Body parts:A–B, J–K, R, X, g–i – labrum; C, L, S, Y – mandibles; D, L*, e–f – mandibular apex; E, M, T, Z – maxilla; U – detail of basal part of ultimate palpomere; F–G, N–O, V, a–b, j–l – mentum and prementum; H–I, P–Q, W, c – antenna. Not to scale.	Fig. 2. Morphology of mouthparts and head appendages of the genera of Epimetopidae.A–I – Eupotemus Ji & Jäch, 1998: A, C, E, F, H – E. smithi sp. nov., B, D, G – E. limicola (Delève, 1967), I – E. cameroonensis sp. nov. J–Q – Eumetopus Balfour-Browne, 1949: J, L, M, N, P – E. schuelkei Jäch, 2002, Q – E. bullatus (Sharp, 1875), K, L*, O – E. sp. from Sri Lanka. R–Z, a–l – Epimetopus Lacordaire, 1854: R–W – E. mendeli Fikáček et al., 2011, X–Z, a–c – E. costaricensis Perkins, 1979, g, j – E. thermarum Schwarz & Barber, 1917, e, k – E. trogoides (Sharp, 1874), h – E. costatus group, f, i, l – E. punctipennis Perkins, 1979. Body parts:A–B, J–K, R, X, g–i – labrum; C, L, S, Y – mandibles; D, L*, e–f – mandibular apex; E, M, T, Z – maxilla; U – detail of basal part of ultimate palpomere; F–G, N–O, V, a–b, j–l – mentum and prementum; H–I, P–Q, W, c – antenna. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037212/files/figure.png	https://doi.org/10.5281/zenodo.5037212	Fig. 3. Thoracic morphology of the genera of Epimetopidae. A–B, J, Q – Eupotemus smithi sp. nov.; C–D, L–O – Eumetopus schuelkei Jäch, 2002; E–F – Epimetopus mendeli Fikáček, Barclay & Perkins, 2012; G–I, P – Epimetopus costaricensis Perkins, 1979. A–I – prothorax in ventral view (B, D, F, H – detail of ventral sculpture of the pronotal hood; I – detail of closed procoxal cavity. J–L – meso- and metathorax in ventral view. M–N – elytron in ventral view (M – general view; N – detail of the ventral ridge, notice the spiny surface on inner face of the ridge (in) and on the elytral plectrum posteriorly of it (pl)). O – mesotrochanter. P–Q – details of pretarsus with the leaf-like empodial seta. Not to scale.	Fig. 3. Thoracic morphology of the genera of Epimetopidae. A–B, J, Q – Eupotemus smithi sp. nov.; C–D, L–O – Eumetopus schuelkei Jäch, 2002; E–F – Epimetopus mendeli Fikáček, Barclay & Perkins, 2012; G–I, P – Epimetopus costaricensis Perkins, 1979. A–I – prothorax in ventral view (B, D, F, H – detail of ventral sculpture of the pronotal hood; I – detail of closed procoxal cavity. J–L – meso- and metathorax in ventral view. M–N – elytron in ventral view (M – general view; N – detail of the ventral ridge, notice the spiny surface on inner face of the ridge (in) and on the elytral plectrum posteriorly of it (pl)). O – mesotrochanter. P–Q – details of pretarsus with the leaf-like empodial seta. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037214/files/figure.png	https://doi.org/10.5281/zenodo.5037214	Fig. 4. Head and leg morphology of the genera of Epimetopidae. A–B, H, L – Eumetopus acutimontis Ji & Jäch, 1998; D, M – Eumetopus schuelkei Jäch, 2002; C, K–L, Q – Eupotemus smithi sp. nov.; F–G – Epimetopus punctipennis Perkins, 1979 (adopted from Pൾ*©κංඇඌ 2012); E, I–J, R – Epimetopus mendeli Fikáček, Barclay & Perkins, 2011; N–O – Epimetopus costaricensis Perkins, 1979. A – head in dorsolateral view (can – clypeal canthus causing the eye emargination); B–C – head in dorsal view (arrow – the ridge dividing anterior declined part of clypeus); D – mentum and maxilla; E – head in ventral view; F–G – occipital part of the head with median raised tubercle, dorsolateral view (arrows: parallel impressions corresponding to ventral ridges of pronotal hood); H–I – antenna, scapus largely omitted; J – labial palp; K – apical tibial armature, ventral view, mesothoracic leg; L – apical tibial armature and tarsus, ventral view, metathoracic leg; M – mesotibia; N–O – tarsus (N – posterior; O – anterior); P–R – metatibia, dorsal view. Not to scale.	Fig. 4. Head and leg morphology of the genera of Epimetopidae. A–B, H, L – Eumetopus acutimontis Ji & Jäch, 1998; D, M – Eumetopus schuelkei Jäch, 2002; C, K–L, Q – Eupotemus smithi sp. nov.; F–G – Epimetopus punctipennis Perkins, 1979 (adopted from Pൾ*©κංඇඌ 2012); E, I–J, R – Epimetopus mendeli Fikáček, Barclay & Perkins, 2011; N–O – Epimetopus costaricensis Perkins, 1979. A – head in dorsolateral view (can – clypeal canthus causing the eye emargination); B–C – head in dorsal view (arrow – the ridge dividing anterior declined part of clypeus); D – mentum and maxilla; E – head in ventral view; F–G – occipital part of the head with median raised tubercle, dorsolateral view (arrows: parallel impressions corresponding to ventral ridges of pronotal hood); H–I – antenna, scapus largely omitted; J – labial palp; K – apical tibial armature, ventral view, mesothoracic leg; L – apical tibial armature and tarsus, ventral view, metathoracic leg; M – mesotibia; N–O – tarsus (N – posterior; O – anterior); P–R – metatibia, dorsal view. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037216/files/figure.png	https://doi.org/10.5281/zenodo.5037216	Fig. 5. Morphology of Epimetopidae. A–B – ventral morphology: A – Eupotemus smithi sp. nov.; B – Epimetopus mendeli Fikáček, Perkins & Barclay, 2011. C–F – hind wings: C – Eumetopus schulkei Jäch, 2002; D – Eupotemus smithi sp. nov.; E – Epimetopus mendeli, adopted from Fංκගඹൾκ et al. (2011); F – Epimetopus costaricensis Perkins, 1979. G – dorsal part of the metathorax. H – scutellum. I–K – abdominal ventrites (I – Eumetopus schuelkei; J – Eupotemus smithi sp. nov.; K – Epimetopus mendeli). L–M – ovipositor (L – Eumetopus schuelkei; M – Eupotemus smithi). O – elytral punctation, slide-mounted elytra of Epimetopus costaricensis. P – female of Eumetopus acutimontis in ventral view, with the egg cases carried under the abdomen. Not to scale.	Fig. 5. Morphology of Epimetopidae. A–B – ventral morphology: A – Eupotemus smithi sp. nov.; B – Epimetopus mendeli Fikáček, Perkins & Barclay, 2011. C–F – hind wings: C – Eumetopus schulkei Jäch, 2002; D – Eupotemus smithi sp. nov.; E – Epimetopus mendeli, adopted from Fංκගඹൾκ et al. (2011); F – Epimetopus costaricensis Perkins, 1979. G – dorsal part of the metathorax. H – scutellum. I–K – abdominal ventrites (I – Eumetopus schuelkei; J – Eupotemus smithi sp. nov.; K – Epimetopus mendeli). L–M – ovipositor (L – Eumetopus schuelkei; M – Eupotemus smithi). O – elytral punctation, slide-mounted elytra of Epimetopus costaricensis. P – female of Eumetopus acutimontis in ventral view, with the egg cases carried under the abdomen. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037218/files/figure.png	https://doi.org/10.5281/zenodo.5037218	Fig. 6. Male genitalia and associated structures of Epimetopidae. A–H – Eupotemus Ji & Jäch, 1998 (A–F – E. smithi sp. nov.; G – E. cameroonensis sp. nov.; H – E. carinaticollis (Basilewsky, 1956)). I–R – Epimetopus Lacordaire, 1854 (I–N – E. mendeli Fikáček et al., 2011; O – E. cf. burruyacu Oliva, 1986; P – E. multiporus Perkins, 2012; Q – E. clandestinus Perkins, 2012; R – E. thermarum Schwarz & Barber, 1917). S–Z – Eumetopus Balfour-Browne, 1949 (S–Y – E. acutimontis Ji & Jäch, 1998; Z – E. bullatus (Sharp, 1875)).A, I, S – sternite VIII; B, J, T – tergite VIII; C, K, U – sternite IX; D, L, V – aedeagus dorsally; E, M, W – aedeagus laterally; F, N, X – aedeagus ventrally (basal part omitted in X); Y – sperm pump. Color coding: green – ventral projections of the median lobe; red – median lobe; blue – paramere (pale blue – dorsal lobe; dark blue – ventral lobe). Not to scale.	Fig. 6. Male genitalia and associated structures of Epimetopidae. A–H – Eupotemus Ji & Jäch, 1998 (A–F – E. smithi sp. nov.; G – E. cameroonensis sp. nov.; H – E. carinaticollis (Basilewsky, 1956)). I–R – Epimetopus Lacordaire, 1854 (I–N – E. mendeli Fikáček et al., 2011; O – E. cf. burruyacu Oliva, 1986; P – E. multiporus Perkins, 2012; Q – E. clandestinus Perkins, 2012; R – E. thermarum Schwarz & Barber, 1917). S–Z – Eumetopus Balfour-Browne, 1949 (S–Y – E. acutimontis Ji & Jäch, 1998; Z – E. bullatus (Sharp, 1875)).A, I, S – sternite VIII; B, J, T – tergite VIII; C, K, U – sternite IX; D, L, V – aedeagus dorsally; E, M, W – aedeagus laterally; F, N, X – aedeagus ventrally (basal part omitted in X); Y – sperm pump. Color coding: green – ventral projections of the median lobe; red – median lobe; blue – paramere (pale blue – dorsal lobe; dark blue – ventral lobe). Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037228/files/figure.png	https://doi.org/10.5281/zenodo.5037228	Fig. 11. Male genitalia of the Eupotemus carinaticollis species group, holotypes.A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. taianus sp. nov.; G–I – E. uluguru sp. nov. A, D, G – dorsal view; B, E, H – lateral view; C, F, I – ventral view.	Fig. 11. Male genitalia of the Eupotemus carinaticollis species group, holotypes.A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. taianus sp. nov.; G–I – E. uluguru sp. nov. A, D, G – dorsal view; B, E, H – lateral view; C, F, I – ventral view.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCDA233FBD4FF1AFC78F97C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037230/files/figure.png	https://doi.org/10.5281/zenodo.5037230	Fig. 12. Habitus photographs of the species of the Eupotemus carinaticollis species group, holotypes: A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. uluguru sp. nov.; G–I – E. taianus sp. nov.	Fig. 12. Habitus photographs of the species of the Eupotemus carinaticollis species group, holotypes: A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. uluguru sp. nov.; G–I – E. taianus sp. nov.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC9A231FBACF8DDFDFAF8A9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037228/files/figure.png	https://doi.org/10.5281/zenodo.5037228	Fig. 11. Male genitalia of the Eupotemus carinaticollis species group, holotypes.A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. taianus sp. nov.; G–I – E. uluguru sp. nov. A, D, G – dorsal view; B, E, H – lateral view; C, F, I – ventral view.	Fig. 11. Male genitalia of the Eupotemus carinaticollis species group, holotypes.A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. taianus sp. nov.; G–I – E. uluguru sp. nov. A, D, G – dorsal view; B, E, H – lateral view; C, F, I – ventral view.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC9A231FBACF8DDFDFAF8A9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037230/files/figure.png	https://doi.org/10.5281/zenodo.5037230	Fig. 12. Habitus photographs of the species of the Eupotemus carinaticollis species group, holotypes: A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. uluguru sp. nov.; G–I – E. taianus sp. nov.	Fig. 12. Habitus photographs of the species of the Eupotemus carinaticollis species group, holotypes: A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. uluguru sp. nov.; G–I – E. taianus sp. nov.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC9A231FBACF8DDFDFAF8A9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037234/files/figure.png	https://doi.org/10.5281/zenodo.5037234	Fig. 13. Known distribution of Eupotemus species.A – species of E. limicola species group; B – species of E. carinaticollis species group.	Fig. 13. Known distribution of Eupotemus species.A – species of E. limicola species group; B – species of E. carinaticollis species group.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA231F951F898FAD9F8C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037228/files/figure.png	https://doi.org/10.5281/zenodo.5037228	Fig. 11. Male genitalia of the Eupotemus carinaticollis species group, holotypes.A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. taianus sp. nov.; G–I – E. uluguru sp. nov. A, D, G – dorsal view; B, E, H – lateral view; C, F, I – ventral view.	Fig. 11. Male genitalia of the Eupotemus carinaticollis species group, holotypes.A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. taianus sp. nov.; G–I – E. uluguru sp. nov. A, D, G – dorsal view; B, E, H – lateral view; C, F, I – ventral view.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA231F951F898FAD9F8C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037230/files/figure.png	https://doi.org/10.5281/zenodo.5037230	Fig. 12. Habitus photographs of the species of the Eupotemus carinaticollis species group, holotypes: A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. uluguru sp. nov.; G–I – E. taianus sp. nov.	Fig. 12. Habitus photographs of the species of the Eupotemus carinaticollis species group, holotypes: A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. uluguru sp. nov.; G–I – E. taianus sp. nov.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA231F951F898FAD9F8C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037234/files/figure.png	https://doi.org/10.5281/zenodo.5037234	Fig. 13. Known distribution of Eupotemus species.A – species of E. limicola species group; B – species of E. carinaticollis species group.	Fig. 13. Known distribution of Eupotemus species.A – species of E. limicola species group; B – species of E. carinaticollis species group.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA23EFBCEF84AFB28F786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037210/files/figure.png	https://doi.org/10.5281/zenodo.5037210	Fig. 2. Morphology of mouthparts and head appendages of the genera of Epimetopidae.A–I – Eupotemus Ji & Jäch, 1998: A, C, E, F, H – E. smithi sp. nov., B, D, G – E. limicola (Delève, 1967), I – E. cameroonensis sp. nov. J–Q – Eumetopus Balfour-Browne, 1949: J, L, M, N, P – E. schuelkei Jäch, 2002, Q – E. bullatus (Sharp, 1875), K, L*, O – E. sp. from Sri Lanka. R–Z, a–l – Epimetopus Lacordaire, 1854: R–W – E. mendeli Fikáček et al., 2011, X–Z, a–c – E. costaricensis Perkins, 1979, g, j – E. thermarum Schwarz & Barber, 1917, e, k – E. trogoides (Sharp, 1874), h – E. costatus group, f, i, l – E. punctipennis Perkins, 1979. Body parts:A–B, J–K, R, X, g–i – labrum; C, L, S, Y – mandibles; D, L*, e–f – mandibular apex; E, M, T, Z – maxilla; U – detail of basal part of ultimate palpomere; F–G, N–O, V, a–b, j–l – mentum and prementum; H–I, P–Q, W, c – antenna. Not to scale.	Fig. 2. Morphology of mouthparts and head appendages of the genera of Epimetopidae.A–I – Eupotemus Ji & Jäch, 1998: A, C, E, F, H – E. smithi sp. nov., B, D, G – E. limicola (Delève, 1967), I – E. cameroonensis sp. nov. J–Q – Eumetopus Balfour-Browne, 1949: J, L, M, N, P – E. schuelkei Jäch, 2002, Q – E. bullatus (Sharp, 1875), K, L*, O – E. sp. from Sri Lanka. R–Z, a–l – Epimetopus Lacordaire, 1854: R–W – E. mendeli Fikáček et al., 2011, X–Z, a–c – E. costaricensis Perkins, 1979, g, j – E. thermarum Schwarz & Barber, 1917, e, k – E. trogoides (Sharp, 1874), h – E. costatus group, f, i, l – E. punctipennis Perkins, 1979. Body parts:A–B, J–K, R, X, g–i – labrum; C, L, S, Y – mandibles; D, L*, e–f – mandibular apex; E, M, T, Z – maxilla; U – detail of basal part of ultimate palpomere; F–G, N–O, V, a–b, j–l – mentum and prementum; H–I, P–Q, W, c – antenna. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA23EFBCEF84AFB28F786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037212/files/figure.png	https://doi.org/10.5281/zenodo.5037212	Fig. 3. Thoracic morphology of the genera of Epimetopidae. A–B, J, Q – Eupotemus smithi sp. nov.; C–D, L–O – Eumetopus schuelkei Jäch, 2002; E–F – Epimetopus mendeli Fikáček, Barclay & Perkins, 2012; G–I, P – Epimetopus costaricensis Perkins, 1979. A–I – prothorax in ventral view (B, D, F, H – detail of ventral sculpture of the pronotal hood; I – detail of closed procoxal cavity. J–L – meso- and metathorax in ventral view. M–N – elytron in ventral view (M – general view; N – detail of the ventral ridge, notice the spiny surface on inner face of the ridge (in) and on the elytral plectrum posteriorly of it (pl)). O – mesotrochanter. P–Q – details of pretarsus with the leaf-like empodial seta. Not to scale.	Fig. 3. Thoracic morphology of the genera of Epimetopidae. A–B, J, Q – Eupotemus smithi sp. nov.; C–D, L–O – Eumetopus schuelkei Jäch, 2002; E–F – Epimetopus mendeli Fikáček, Barclay & Perkins, 2012; G–I, P – Epimetopus costaricensis Perkins, 1979. A–I – prothorax in ventral view (B, D, F, H – detail of ventral sculpture of the pronotal hood; I – detail of closed procoxal cavity. J–L – meso- and metathorax in ventral view. M–N – elytron in ventral view (M – general view; N – detail of the ventral ridge, notice the spiny surface on inner face of the ridge (in) and on the elytral plectrum posteriorly of it (pl)). O – mesotrochanter. P–Q – details of pretarsus with the leaf-like empodial seta. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA23EFBCEF84AFB28F786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037214/files/figure.png	https://doi.org/10.5281/zenodo.5037214	Fig. 4. Head and leg morphology of the genera of Epimetopidae. A–B, H, L – Eumetopus acutimontis Ji & Jäch, 1998; D, M – Eumetopus schuelkei Jäch, 2002; C, K–L, Q – Eupotemus smithi sp. nov.; F–G – Epimetopus punctipennis Perkins, 1979 (adopted from Pൾ*©κංඇඌ 2012); E, I–J, R – Epimetopus mendeli Fikáček, Barclay & Perkins, 2011; N–O – Epimetopus costaricensis Perkins, 1979. A – head in dorsolateral view (can – clypeal canthus causing the eye emargination); B–C – head in dorsal view (arrow – the ridge dividing anterior declined part of clypeus); D – mentum and maxilla; E – head in ventral view; F–G – occipital part of the head with median raised tubercle, dorsolateral view (arrows: parallel impressions corresponding to ventral ridges of pronotal hood); H–I – antenna, scapus largely omitted; J – labial palp; K – apical tibial armature, ventral view, mesothoracic leg; L – apical tibial armature and tarsus, ventral view, metathoracic leg; M – mesotibia; N–O – tarsus (N – posterior; O – anterior); P–R – metatibia, dorsal view. Not to scale.	Fig. 4. Head and leg morphology of the genera of Epimetopidae. A–B, H, L – Eumetopus acutimontis Ji & Jäch, 1998; D, M – Eumetopus schuelkei Jäch, 2002; C, K–L, Q – Eupotemus smithi sp. nov.; F–G – Epimetopus punctipennis Perkins, 1979 (adopted from Pൾ*©κංඇඌ 2012); E, I–J, R – Epimetopus mendeli Fikáček, Barclay & Perkins, 2011; N–O – Epimetopus costaricensis Perkins, 1979. A – head in dorsolateral view (can – clypeal canthus causing the eye emargination); B–C – head in dorsal view (arrow – the ridge dividing anterior declined part of clypeus); D – mentum and maxilla; E – head in ventral view; F–G – occipital part of the head with median raised tubercle, dorsolateral view (arrows: parallel impressions corresponding to ventral ridges of pronotal hood); H–I – antenna, scapus largely omitted; J – labial palp; K – apical tibial armature, ventral view, mesothoracic leg; L – apical tibial armature and tarsus, ventral view, metathoracic leg; M – mesotibia; N–O – tarsus (N – posterior; O – anterior); P–R – metatibia, dorsal view. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA23EFBCEF84AFB28F786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037216/files/figure.png	https://doi.org/10.5281/zenodo.5037216	Fig. 5. Morphology of Epimetopidae. A–B – ventral morphology: A – Eupotemus smithi sp. nov.; B – Epimetopus mendeli Fikáček, Perkins & Barclay, 2011. C–F – hind wings: C – Eumetopus schulkei Jäch, 2002; D – Eupotemus smithi sp. nov.; E – Epimetopus mendeli, adopted from Fංκගඹൾκ et al. (2011); F – Epimetopus costaricensis Perkins, 1979. G – dorsal part of the metathorax. H – scutellum. I–K – abdominal ventrites (I – Eumetopus schuelkei; J – Eupotemus smithi sp. nov.; K – Epimetopus mendeli). L–M – ovipositor (L – Eumetopus schuelkei; M – Eupotemus smithi). O – elytral punctation, slide-mounted elytra of Epimetopus costaricensis. P – female of Eumetopus acutimontis in ventral view, with the egg cases carried under the abdomen. Not to scale.	Fig. 5. Morphology of Epimetopidae. A–B – ventral morphology: A – Eupotemus smithi sp. nov.; B – Epimetopus mendeli Fikáček, Perkins & Barclay, 2011. C–F – hind wings: C – Eumetopus schulkei Jäch, 2002; D – Eupotemus smithi sp. nov.; E – Epimetopus mendeli, adopted from Fංκගඹൾκ et al. (2011); F – Epimetopus costaricensis Perkins, 1979. G – dorsal part of the metathorax. H – scutellum. I–K – abdominal ventrites (I – Eumetopus schuelkei; J – Eupotemus smithi sp. nov.; K – Epimetopus mendeli). L–M – ovipositor (L – Eumetopus schuelkei; M – Eupotemus smithi). O – elytral punctation, slide-mounted elytra of Epimetopus costaricensis. P – female of Eumetopus acutimontis in ventral view, with the egg cases carried under the abdomen. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA23EFBCEF84AFB28F786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037218/files/figure.png	https://doi.org/10.5281/zenodo.5037218	Fig. 6. Male genitalia and associated structures of Epimetopidae. A–H – Eupotemus Ji & Jäch, 1998 (A–F – E. smithi sp. nov.; G – E. cameroonensis sp. nov.; H – E. carinaticollis (Basilewsky, 1956)). I–R – Epimetopus Lacordaire, 1854 (I–N – E. mendeli Fikáček et al., 2011; O – E. cf. burruyacu Oliva, 1986; P – E. multiporus Perkins, 2012; Q – E. clandestinus Perkins, 2012; R – E. thermarum Schwarz & Barber, 1917). S–Z – Eumetopus Balfour-Browne, 1949 (S–Y – E. acutimontis Ji & Jäch, 1998; Z – E. bullatus (Sharp, 1875)).A, I, S – sternite VIII; B, J, T – tergite VIII; C, K, U – sternite IX; D, L, V – aedeagus dorsally; E, M, W – aedeagus laterally; F, N, X – aedeagus ventrally (basal part omitted in X); Y – sperm pump. Color coding: green – ventral projections of the median lobe; red – median lobe; blue – paramere (pale blue – dorsal lobe; dark blue – ventral lobe). Not to scale.	Fig. 6. Male genitalia and associated structures of Epimetopidae. A–H – Eupotemus Ji & Jäch, 1998 (A–F – E. smithi sp. nov.; G – E. cameroonensis sp. nov.; H – E. carinaticollis (Basilewsky, 1956)). I–R – Epimetopus Lacordaire, 1854 (I–N – E. mendeli Fikáček et al., 2011; O – E. cf. burruyacu Oliva, 1986; P – E. multiporus Perkins, 2012; Q – E. clandestinus Perkins, 2012; R – E. thermarum Schwarz & Barber, 1917). S–Z – Eumetopus Balfour-Browne, 1949 (S–Y – E. acutimontis Ji & Jäch, 1998; Z – E. bullatus (Sharp, 1875)).A, I, S – sternite VIII; B, J, T – tergite VIII; C, K, U – sternite IX; D, L, V – aedeagus dorsally; E, M, W – aedeagus laterally; F, N, X – aedeagus ventrally (basal part omitted in X); Y – sperm pump. Color coding: green – ventral projections of the median lobe; red – median lobe; blue – paramere (pale blue – dorsal lobe; dark blue – ventral lobe). Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA23EFBCEF84AFB28F786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037220/files/figure.png	https://doi.org/10.5281/zenodo.5037220	Fig. 7. Biology of Epimetopidae. A–D – Eumetopus acutimontis Ji & Jäch, 1998 in Vietnam, Khánh Hòa Prov., Ba Ho Waterfalls National Park, 12°23.131′N 109°08.052′E:A–B – alive specimens on wet sandy shores; C – specimen buried in wet sand; D – general view of the locality. E–F – locality of Epimetopus surinamensis Perkins, 2012 in Suriname: Sipaliwini, Wehepai, 2°21.776′N 56°41.861′W (SR10-0904-01A) (E – general view; F – detail of sandy banks from which specimens were collected). G–H – locality of Epimetopus simplex Perkins, 1979 in Venezuela, Guarico, Rio San Antonio, 9°46.32′N 67°21.177′W (VZ09-0108-02A) (G – detail of gravelly shore show which specimens were collected; H – general view). I–J – marshy roadside ditch with Epimetopus venezuelensis Perkins, 2012 in Venezuela, Apure, ca. 1 km N Rio Claro, 7°10.162′N 67°38.69′W (AS-06-009). (I – general view; J – detail of the microhabitat). Photos by A. Sazhnev (A–C), A. Prokin (D), A. Short (E–J).	Fig. 7. Biology of Epimetopidae. A–D – Eumetopus acutimontis Ji & Jäch, 1998 in Vietnam, Khánh Hòa Prov., Ba Ho Waterfalls National Park, 12°23.131′N 109°08.052′E:A–B – alive specimens on wet sandy shores; C – specimen buried in wet sand; D – general view of the locality. E–F – locality of Epimetopus surinamensis Perkins, 2012 in Suriname: Sipaliwini, Wehepai, 2°21.776′N 56°41.861′W (SR10-0904-01A) (E – general view; F – detail of sandy banks from which specimens were collected). G–H – locality of Epimetopus simplex Perkins, 1979 in Venezuela, Guarico, Rio San Antonio, 9°46.32′N 67°21.177′W (VZ09-0108-02A) (G – detail of gravelly shore show which specimens were collected; H – general view). I–J – marshy roadside ditch with Epimetopus venezuelensis Perkins, 2012 in Venezuela, Apure, ca. 1 km N Rio Claro, 7°10.162′N 67°38.69′W (AS-06-009). (I – general view; J – detail of the microhabitat). Photos by A. Sazhnev (A–C), A. Prokin (D), A. Short (E–J).	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA23EFBCEF84AFB28F786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037236/files/figure.png	https://doi.org/10.5281/zenodo.5037236	Fig. 14. Habitus photographs of the known species of Eumetopus Balfour-Browne, 1949. A – E. acutimontis Ji & Jäch, 1998, paratype from China: Hainan (NHMW); B – E. asperatus (Champion, 1919), specimen from Nepal: Koshi Valley (NMPC); C – E. bullatus (Sharp, 1875), specimen from India: Maharashtra, Sawantwadi (NMPC); D – E. flavidulus (Sharp, 1890), specimen from India: Meghalaya, Bagmara (NHMW); E – E. maindroni (Régimbart, 1903), specimen from India: Maharashtra, Sawantwadi (NHMW); F – E. schuelkei Jäch, 2002, paratype from central Laos; G – E. tibialis Ji & Jäch, 1998 from Thailand, Ban Na env. (NMPC); H – E. weigeli Skale & Jäch, 2003, holotype from Nepal (NHMW). Scale bars: 1 mm.	Fig. 14. Habitus photographs of the known species of Eumetopus Balfour-Browne, 1949. A – E. acutimontis Ji & Jäch, 1998, paratype from China: Hainan (NHMW); B – E. asperatus (Champion, 1919), specimen from Nepal: Koshi Valley (NMPC); C – E. bullatus (Sharp, 1875), specimen from India: Maharashtra, Sawantwadi (NMPC); D – E. flavidulus (Sharp, 1890), specimen from India: Meghalaya, Bagmara (NHMW); E – E. maindroni (Régimbart, 1903), specimen from India: Maharashtra, Sawantwadi (NHMW); F – E. schuelkei Jäch, 2002, paratype from central Laos; G – E. tibialis Ji & Jäch, 1998 from Thailand, Ban Na env. (NMPC); H – E. weigeli Skale & Jäch, 2003, holotype from Nepal (NHMW). Scale bars: 1 mm.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFCBA23EFBCEF84AFB28F786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037238/files/figure.png	https://doi.org/10.5281/zenodo.5037238	Fig. 15. Elytral sculpture and male genitalia of Eumetopus Balfour-Browne, 1949. A–H – elytral sculpture, same specimens as in Fig. 14: A – E. acutimontis Ji & Jäch, 1998; B – E. asperatus (Champion, 1919); C – E. bullatus (Sharp, 1875); D – E. flavidulus (Sharp, 1890); E – E. maindroni (Régimbart, 1903); F – E. schuelkei Jäch, 2002; G – E. tibialis Ji & Jäch, 1998; H – E. weigeli Skale & Jäch, 2003. I–L – male genitalia of examined specimens, large basal portion of the phallobase omitted (dorsal and lateral view): I – E. acutimontis from Vietnam; J – E. flavidulus from India: Andhra Pradesh; K – E. maindroni from India: Gujarat; L – E. weigeli from India: Uttarakhand.	Fig. 15. Elytral sculpture and male genitalia of Eumetopus Balfour-Browne, 1949. A–H – elytral sculpture, same specimens as in Fig. 14: A – E. acutimontis Ji & Jäch, 1998; B – E. asperatus (Champion, 1919); C – E. bullatus (Sharp, 1875); D – E. flavidulus (Sharp, 1890); E – E. maindroni (Régimbart, 1903); F – E. schuelkei Jäch, 2002; G – E. tibialis Ji & Jäch, 1998; H – E. weigeli Skale & Jäch, 2003. I–L – male genitalia of examined specimens, large basal portion of the phallobase omitted (dorsal and lateral view): I – E. acutimontis from Vietnam; J – E. flavidulus from India: Andhra Pradesh; K – E. maindroni from India: Gujarat; L – E. weigeli from India: Uttarakhand.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC5A23FF97AFF65FB54FE9B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037236/files/figure.png	https://doi.org/10.5281/zenodo.5037236	Fig. 14. Habitus photographs of the known species of Eumetopus Balfour-Browne, 1949. A – E. acutimontis Ji & Jäch, 1998, paratype from China: Hainan (NHMW); B – E. asperatus (Champion, 1919), specimen from Nepal: Koshi Valley (NMPC); C – E. bullatus (Sharp, 1875), specimen from India: Maharashtra, Sawantwadi (NMPC); D – E. flavidulus (Sharp, 1890), specimen from India: Meghalaya, Bagmara (NHMW); E – E. maindroni (Régimbart, 1903), specimen from India: Maharashtra, Sawantwadi (NHMW); F – E. schuelkei Jäch, 2002, paratype from central Laos; G – E. tibialis Ji & Jäch, 1998 from Thailand, Ban Na env. (NMPC); H – E. weigeli Skale & Jäch, 2003, holotype from Nepal (NHMW). Scale bars: 1 mm.	Fig. 14. Habitus photographs of the known species of Eumetopus Balfour-Browne, 1949. A – E. acutimontis Ji & Jäch, 1998, paratype from China: Hainan (NHMW); B – E. asperatus (Champion, 1919), specimen from Nepal: Koshi Valley (NMPC); C – E. bullatus (Sharp, 1875), specimen from India: Maharashtra, Sawantwadi (NMPC); D – E. flavidulus (Sharp, 1890), specimen from India: Meghalaya, Bagmara (NHMW); E – E. maindroni (Régimbart, 1903), specimen from India: Maharashtra, Sawantwadi (NHMW); F – E. schuelkei Jäch, 2002, paratype from central Laos; G – E. tibialis Ji & Jäch, 1998 from Thailand, Ban Na env. (NMPC); H – E. weigeli Skale & Jäch, 2003, holotype from Nepal (NHMW). Scale bars: 1 mm.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC5A23FF97AFF65FB54FE9B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037238/files/figure.png	https://doi.org/10.5281/zenodo.5037238	Fig. 15. Elytral sculpture and male genitalia of Eumetopus Balfour-Browne, 1949. A–H – elytral sculpture, same specimens as in Fig. 14: A – E. acutimontis Ji & Jäch, 1998; B – E. asperatus (Champion, 1919); C – E. bullatus (Sharp, 1875); D – E. flavidulus (Sharp, 1890); E – E. maindroni (Régimbart, 1903); F – E. schuelkei Jäch, 2002; G – E. tibialis Ji & Jäch, 1998; H – E. weigeli Skale & Jäch, 2003. I–L – male genitalia of examined specimens, large basal portion of the phallobase omitted (dorsal and lateral view): I – E. acutimontis from Vietnam; J – E. flavidulus from India: Andhra Pradesh; K – E. maindroni from India: Gujarat; L – E. weigeli from India: Uttarakhand.	Fig. 15. Elytral sculpture and male genitalia of Eumetopus Balfour-Browne, 1949. A–H – elytral sculpture, same specimens as in Fig. 14: A – E. acutimontis Ji & Jäch, 1998; B – E. asperatus (Champion, 1919); C – E. bullatus (Sharp, 1875); D – E. flavidulus (Sharp, 1890); E – E. maindroni (Régimbart, 1903); F – E. schuelkei Jäch, 2002; G – E. tibialis Ji & Jäch, 1998; H – E. weigeli Skale & Jäch, 2003. I–L – male genitalia of examined specimens, large basal portion of the phallobase omitted (dorsal and lateral view): I – E. acutimontis from Vietnam; J – E. flavidulus from India: Andhra Pradesh; K – E. maindroni from India: Gujarat; L – E. weigeli from India: Uttarakhand.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC6A23CF890FF66FA54FE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037236/files/figure.png	https://doi.org/10.5281/zenodo.5037236	Fig. 14. Habitus photographs of the known species of Eumetopus Balfour-Browne, 1949. A – E. acutimontis Ji & Jäch, 1998, paratype from China: Hainan (NHMW); B – E. asperatus (Champion, 1919), specimen from Nepal: Koshi Valley (NMPC); C – E. bullatus (Sharp, 1875), specimen from India: Maharashtra, Sawantwadi (NMPC); D – E. flavidulus (Sharp, 1890), specimen from India: Meghalaya, Bagmara (NHMW); E – E. maindroni (Régimbart, 1903), specimen from India: Maharashtra, Sawantwadi (NHMW); F – E. schuelkei Jäch, 2002, paratype from central Laos; G – E. tibialis Ji & Jäch, 1998 from Thailand, Ban Na env. (NMPC); H – E. weigeli Skale & Jäch, 2003, holotype from Nepal (NHMW). Scale bars: 1 mm.	Fig. 14. Habitus photographs of the known species of Eumetopus Balfour-Browne, 1949. A – E. acutimontis Ji & Jäch, 1998, paratype from China: Hainan (NHMW); B – E. asperatus (Champion, 1919), specimen from Nepal: Koshi Valley (NMPC); C – E. bullatus (Sharp, 1875), specimen from India: Maharashtra, Sawantwadi (NMPC); D – E. flavidulus (Sharp, 1890), specimen from India: Meghalaya, Bagmara (NHMW); E – E. maindroni (Régimbart, 1903), specimen from India: Maharashtra, Sawantwadi (NHMW); F – E. schuelkei Jäch, 2002, paratype from central Laos; G – E. tibialis Ji & Jäch, 1998 from Thailand, Ban Na env. (NMPC); H – E. weigeli Skale & Jäch, 2003, holotype from Nepal (NHMW). Scale bars: 1 mm.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC6A23CF890FF66FA54FE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037238/files/figure.png	https://doi.org/10.5281/zenodo.5037238	Fig. 15. Elytral sculpture and male genitalia of Eumetopus Balfour-Browne, 1949. A–H – elytral sculpture, same specimens as in Fig. 14: A – E. acutimontis Ji & Jäch, 1998; B – E. asperatus (Champion, 1919); C – E. bullatus (Sharp, 1875); D – E. flavidulus (Sharp, 1890); E – E. maindroni (Régimbart, 1903); F – E. schuelkei Jäch, 2002; G – E. tibialis Ji & Jäch, 1998; H – E. weigeli Skale & Jäch, 2003. I–L – male genitalia of examined specimens, large basal portion of the phallobase omitted (dorsal and lateral view): I – E. acutimontis from Vietnam; J – E. flavidulus from India: Andhra Pradesh; K – E. maindroni from India: Gujarat; L – E. weigeli from India: Uttarakhand.	Fig. 15. Elytral sculpture and male genitalia of Eumetopus Balfour-Browne, 1949. A–H – elytral sculpture, same specimens as in Fig. 14: A – E. acutimontis Ji & Jäch, 1998; B – E. asperatus (Champion, 1919); C – E. bullatus (Sharp, 1875); D – E. flavidulus (Sharp, 1890); E – E. maindroni (Régimbart, 1903); F – E. schuelkei Jäch, 2002; G – E. tibialis Ji & Jäch, 1998; H – E. weigeli Skale & Jäch, 2003. I–L – male genitalia of examined specimens, large basal portion of the phallobase omitted (dorsal and lateral view): I – E. acutimontis from Vietnam; J – E. flavidulus from India: Andhra Pradesh; K – E. maindroni from India: Gujarat; L – E. weigeli from India: Uttarakhand.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC7A238F95DFF66FE6EFB57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037210/files/figure.png	https://doi.org/10.5281/zenodo.5037210	Fig. 2. Morphology of mouthparts and head appendages of the genera of Epimetopidae.A–I – Eupotemus Ji & Jäch, 1998: A, C, E, F, H – E. smithi sp. nov., B, D, G – E. limicola (Delève, 1967), I – E. cameroonensis sp. nov. J–Q – Eumetopus Balfour-Browne, 1949: J, L, M, N, P – E. schuelkei Jäch, 2002, Q – E. bullatus (Sharp, 1875), K, L*, O – E. sp. from Sri Lanka. R–Z, a–l – Epimetopus Lacordaire, 1854: R–W – E. mendeli Fikáček et al., 2011, X–Z, a–c – E. costaricensis Perkins, 1979, g, j – E. thermarum Schwarz & Barber, 1917, e, k – E. trogoides (Sharp, 1874), h – E. costatus group, f, i, l – E. punctipennis Perkins, 1979. Body parts:A–B, J–K, R, X, g–i – labrum; C, L, S, Y – mandibles; D, L*, e–f – mandibular apex; E, M, T, Z – maxilla; U – detail of basal part of ultimate palpomere; F–G, N–O, V, a–b, j–l – mentum and prementum; H–I, P–Q, W, c – antenna. Not to scale.	Fig. 2. Morphology of mouthparts and head appendages of the genera of Epimetopidae.A–I – Eupotemus Ji & Jäch, 1998: A, C, E, F, H – E. smithi sp. nov., B, D, G – E. limicola (Delève, 1967), I – E. cameroonensis sp. nov. J–Q – Eumetopus Balfour-Browne, 1949: J, L, M, N, P – E. schuelkei Jäch, 2002, Q – E. bullatus (Sharp, 1875), K, L*, O – E. sp. from Sri Lanka. R–Z, a–l – Epimetopus Lacordaire, 1854: R–W – E. mendeli Fikáček et al., 2011, X–Z, a–c – E. costaricensis Perkins, 1979, g, j – E. thermarum Schwarz & Barber, 1917, e, k – E. trogoides (Sharp, 1874), h – E. costatus group, f, i, l – E. punctipennis Perkins, 1979. Body parts:A–B, J–K, R, X, g–i – labrum; C, L, S, Y – mandibles; D, L*, e–f – mandibular apex; E, M, T, Z – maxilla; U – detail of basal part of ultimate palpomere; F–G, N–O, V, a–b, j–l – mentum and prementum; H–I, P–Q, W, c – antenna. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC7A238F95DFF66FE6EFB57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037212/files/figure.png	https://doi.org/10.5281/zenodo.5037212	Fig. 3. Thoracic morphology of the genera of Epimetopidae. A–B, J, Q – Eupotemus smithi sp. nov.; C–D, L–O – Eumetopus schuelkei Jäch, 2002; E–F – Epimetopus mendeli Fikáček, Barclay & Perkins, 2012; G–I, P – Epimetopus costaricensis Perkins, 1979. A–I – prothorax in ventral view (B, D, F, H – detail of ventral sculpture of the pronotal hood; I – detail of closed procoxal cavity. J–L – meso- and metathorax in ventral view. M–N – elytron in ventral view (M – general view; N – detail of the ventral ridge, notice the spiny surface on inner face of the ridge (in) and on the elytral plectrum posteriorly of it (pl)). O – mesotrochanter. P–Q – details of pretarsus with the leaf-like empodial seta. Not to scale.	Fig. 3. Thoracic morphology of the genera of Epimetopidae. A–B, J, Q – Eupotemus smithi sp. nov.; C–D, L–O – Eumetopus schuelkei Jäch, 2002; E–F – Epimetopus mendeli Fikáček, Barclay & Perkins, 2012; G–I, P – Epimetopus costaricensis Perkins, 1979. A–I – prothorax in ventral view (B, D, F, H – detail of ventral sculpture of the pronotal hood; I – detail of closed procoxal cavity. J–L – meso- and metathorax in ventral view. M–N – elytron in ventral view (M – general view; N – detail of the ventral ridge, notice the spiny surface on inner face of the ridge (in) and on the elytral plectrum posteriorly of it (pl)). O – mesotrochanter. P–Q – details of pretarsus with the leaf-like empodial seta. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC7A238F95DFF66FE6EFB57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037214/files/figure.png	https://doi.org/10.5281/zenodo.5037214	Fig. 4. Head and leg morphology of the genera of Epimetopidae. A–B, H, L – Eumetopus acutimontis Ji & Jäch, 1998; D, M – Eumetopus schuelkei Jäch, 2002; C, K–L, Q – Eupotemus smithi sp. nov.; F–G – Epimetopus punctipennis Perkins, 1979 (adopted from Pൾ*©κංඇඌ 2012); E, I–J, R – Epimetopus mendeli Fikáček, Barclay & Perkins, 2011; N–O – Epimetopus costaricensis Perkins, 1979. A – head in dorsolateral view (can – clypeal canthus causing the eye emargination); B–C – head in dorsal view (arrow – the ridge dividing anterior declined part of clypeus); D – mentum and maxilla; E – head in ventral view; F–G – occipital part of the head with median raised tubercle, dorsolateral view (arrows: parallel impressions corresponding to ventral ridges of pronotal hood); H–I – antenna, scapus largely omitted; J – labial palp; K – apical tibial armature, ventral view, mesothoracic leg; L – apical tibial armature and tarsus, ventral view, metathoracic leg; M – mesotibia; N–O – tarsus (N – posterior; O – anterior); P–R – metatibia, dorsal view. Not to scale.	Fig. 4. Head and leg morphology of the genera of Epimetopidae. A–B, H, L – Eumetopus acutimontis Ji & Jäch, 1998; D, M – Eumetopus schuelkei Jäch, 2002; C, K–L, Q – Eupotemus smithi sp. nov.; F–G – Epimetopus punctipennis Perkins, 1979 (adopted from Pൾ*©κංඇඌ 2012); E, I–J, R – Epimetopus mendeli Fikáček, Barclay & Perkins, 2011; N–O – Epimetopus costaricensis Perkins, 1979. A – head in dorsolateral view (can – clypeal canthus causing the eye emargination); B–C – head in dorsal view (arrow – the ridge dividing anterior declined part of clypeus); D – mentum and maxilla; E – head in ventral view; F–G – occipital part of the head with median raised tubercle, dorsolateral view (arrows: parallel impressions corresponding to ventral ridges of pronotal hood); H–I – antenna, scapus largely omitted; J – labial palp; K – apical tibial armature, ventral view, mesothoracic leg; L – apical tibial armature and tarsus, ventral view, metathoracic leg; M – mesotibia; N–O – tarsus (N – posterior; O – anterior); P–R – metatibia, dorsal view. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC7A238F95DFF66FE6EFB57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037216/files/figure.png	https://doi.org/10.5281/zenodo.5037216	Fig. 5. Morphology of Epimetopidae. A–B – ventral morphology: A – Eupotemus smithi sp. nov.; B – Epimetopus mendeli Fikáček, Perkins & Barclay, 2011. C–F – hind wings: C – Eumetopus schulkei Jäch, 2002; D – Eupotemus smithi sp. nov.; E – Epimetopus mendeli, adopted from Fංκගඹൾκ et al. (2011); F – Epimetopus costaricensis Perkins, 1979. G – dorsal part of the metathorax. H – scutellum. I–K – abdominal ventrites (I – Eumetopus schuelkei; J – Eupotemus smithi sp. nov.; K – Epimetopus mendeli). L–M – ovipositor (L – Eumetopus schuelkei; M – Eupotemus smithi). O – elytral punctation, slide-mounted elytra of Epimetopus costaricensis. P – female of Eumetopus acutimontis in ventral view, with the egg cases carried under the abdomen. Not to scale.	Fig. 5. Morphology of Epimetopidae. A–B – ventral morphology: A – Eupotemus smithi sp. nov.; B – Epimetopus mendeli Fikáček, Perkins & Barclay, 2011. C–F – hind wings: C – Eumetopus schulkei Jäch, 2002; D – Eupotemus smithi sp. nov.; E – Epimetopus mendeli, adopted from Fංκගඹൾκ et al. (2011); F – Epimetopus costaricensis Perkins, 1979. G – dorsal part of the metathorax. H – scutellum. I–K – abdominal ventrites (I – Eumetopus schuelkei; J – Eupotemus smithi sp. nov.; K – Epimetopus mendeli). L–M – ovipositor (L – Eumetopus schuelkei; M – Eupotemus smithi). O – elytral punctation, slide-mounted elytra of Epimetopus costaricensis. P – female of Eumetopus acutimontis in ventral view, with the egg cases carried under the abdomen. Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC7A238F95DFF66FE6EFB57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037218/files/figure.png	https://doi.org/10.5281/zenodo.5037218	Fig. 6. Male genitalia and associated structures of Epimetopidae. A–H – Eupotemus Ji & Jäch, 1998 (A–F – E. smithi sp. nov.; G – E. cameroonensis sp. nov.; H – E. carinaticollis (Basilewsky, 1956)). I–R – Epimetopus Lacordaire, 1854 (I–N – E. mendeli Fikáček et al., 2011; O – E. cf. burruyacu Oliva, 1986; P – E. multiporus Perkins, 2012; Q – E. clandestinus Perkins, 2012; R – E. thermarum Schwarz & Barber, 1917). S–Z – Eumetopus Balfour-Browne, 1949 (S–Y – E. acutimontis Ji & Jäch, 1998; Z – E. bullatus (Sharp, 1875)).A, I, S – sternite VIII; B, J, T – tergite VIII; C, K, U – sternite IX; D, L, V – aedeagus dorsally; E, M, W – aedeagus laterally; F, N, X – aedeagus ventrally (basal part omitted in X); Y – sperm pump. Color coding: green – ventral projections of the median lobe; red – median lobe; blue – paramere (pale blue – dorsal lobe; dark blue – ventral lobe). Not to scale.	Fig. 6. Male genitalia and associated structures of Epimetopidae. A–H – Eupotemus Ji & Jäch, 1998 (A–F – E. smithi sp. nov.; G – E. cameroonensis sp. nov.; H – E. carinaticollis (Basilewsky, 1956)). I–R – Epimetopus Lacordaire, 1854 (I–N – E. mendeli Fikáček et al., 2011; O – E. cf. burruyacu Oliva, 1986; P – E. multiporus Perkins, 2012; Q – E. clandestinus Perkins, 2012; R – E. thermarum Schwarz & Barber, 1917). S–Z – Eumetopus Balfour-Browne, 1949 (S–Y – E. acutimontis Ji & Jäch, 1998; Z – E. bullatus (Sharp, 1875)).A, I, S – sternite VIII; B, J, T – tergite VIII; C, K, U – sternite IX; D, L, V – aedeagus dorsally; E, M, W – aedeagus laterally; F, N, X – aedeagus ventrally (basal part omitted in X); Y – sperm pump. Color coding: green – ventral projections of the median lobe; red – median lobe; blue – paramere (pale blue – dorsal lobe; dark blue – ventral lobe). Not to scale.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC7A238F95DFF66FE6EFB57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037240/files/figure.png	https://doi.org/10.5281/zenodo.5037240	Fig. 16. Photographs of Epimetopus species, including two specimens of questionable provenance (E–G).A – E. flavicaptus Fikáček, Barclay & Perkins, 2011; B – E. deceptus Perkins, 2012; C – E. tridens Perkins, 2012; D – E. hintoni Balfour-Browne, 1949; E – Epimetopus sp. (ʻZambiaʼ); F–G – E. cf. burruyacu Oliva, 1986 (ʻSaudi Arabiaʼ) (F – dorsal view; G – aedeagus). Scale bars for A–F = 0.5 mm. A–D adopted from Pൾ*©κංඇඌ (2012).	Fig. 16. Photographs of Epimetopus species, including two specimens of questionable provenance (E–G).A – E. flavicaptus Fikáček, Barclay & Perkins, 2011; B – E. deceptus Perkins, 2012; C – E. tridens Perkins, 2012; D – E. hintoni Balfour-Browne, 1949; E – Epimetopus sp. (ʻZambiaʼ); F–G – E. cf. burruyacu Oliva, 1986 (ʻSaudi Arabiaʼ) (F – dorsal view; G – aedeagus). Scale bars for A–F = 0.5 mm. A–D adopted from Pൾ*©κංඇඌ (2012).	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC7A238F95DFF66FE6EFB57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037208/files/figure.png	https://doi.org/10.5281/zenodo.5037208	Fig. 1. Distribution and phylogenetic relationships of Epimetopidae genera. A – summary of all known records of Epimetopidae, mapped by genera. Records for American Epimetopus adopted from Pൾ*©κංඇඌ (2012). B–E – phylogenetic relationship among genera based on morphological characters (B–C) and DNA data (D–E): B–C – strict consensus tree of the morphology-based maximum parsimony analysis (B – complete tree with bootstrap supports; C – Georissidae + Epimetopidae subtree with characters mapped). D – Bayesian maximum credibility tree based on cox1, 16S, 18S and 28S sequences, with posterior probability of particular clades. E – morphological characters mapped on the molecular topology of Epimetopidae constrained as sister to all other hydrophilid families following the topology by Lඳ et al. (2020).	Fig. 1. Distribution and phylogenetic relationships of Epimetopidae genera. A – summary of all known records of Epimetopidae, mapped by genera. Records for American Epimetopus adopted from Pൾ*©κංඇඌ (2012). B–E – phylogenetic relationship among genera based on morphological characters (B–C) and DNA data (D–E): B–C – strict consensus tree of the morphology-based maximum parsimony analysis (B – complete tree with bootstrap supports; C – Georissidae + Epimetopidae subtree with characters mapped). D – Bayesian maximum credibility tree based on cox1, 16S, 18S and 28S sequences, with posterior probability of particular clades. E – morphological characters mapped on the molecular topology of Epimetopidae constrained as sister to all other hydrophilid families following the topology by Lඳ et al. (2020).	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC7A238F95DFF66FE6EFB57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037228/files/figure.png	https://doi.org/10.5281/zenodo.5037228	Fig. 11. Male genitalia of the Eupotemus carinaticollis species group, holotypes.A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. taianus sp. nov.; G–I – E. uluguru sp. nov. A, D, G – dorsal view; B, E, H – lateral view; C, F, I – ventral view.	Fig. 11. Male genitalia of the Eupotemus carinaticollis species group, holotypes.A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. taianus sp. nov.; G–I – E. uluguru sp. nov. A, D, G – dorsal view; B, E, H – lateral view; C, F, I – ventral view.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
038BAE5EFFC7A238F95DFF66FE6EFB57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5037230/files/figure.png	https://doi.org/10.5281/zenodo.5037230	Fig. 12. Habitus photographs of the species of the Eupotemus carinaticollis species group, holotypes: A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. uluguru sp. nov.; G–I – E. taianus sp. nov.	Fig. 12. Habitus photographs of the species of the Eupotemus carinaticollis species group, holotypes: A–C – E. carinaticollis (Basilewsky, 1956); D–F – E. uluguru sp. nov.; G–I – E. taianus sp. nov.	2021-02-03	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.		Zenodo	biologists	Fikáček, Martin;Matsumoto, Keita;Perkins, Philip;Prokin, Alexander;Sazhnev, Alexey;Litovkin, Stanislav;Jäch, Manfred A.			
