taxonID	type	description	language	source
038887B4FFD5151DFE9FFCC9FEB5F863.taxon	description	(Figs 1 – 7, 19)	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD5151DFE9FFCC9FEB5F863.taxon	materials_examined	Type material. HOLOTYPE: ♂ (NMPC): RUSSIA: ALTAI REPUBLIC: Belyashi (Dzhazator) env. (25 km NW), confluence of Argut and Karagem rivers, 49 ° 51 ′ 56 ″ N, 87 ° 10 ′ 22 ″ E, rocky steppe, 1400 m a. s. l., 27. – 28. vii. 2017 (Barcode NMPC-LEP- 0381), J. Šumpich leg. PARATYPES: 51 ♂♂ 1 ♀ (49 ♂♂ 1 ♀ NMPC, 2 ♂♂ ZMKU): the same collecting data as holotype, gen. prep. Šumpich 18239, 21353, 21357.	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD5151DFE9FFCC9FEB5F863.taxon	description	Description. Adult. Male (Figs 1 – 3). Wingspan 17 – 21 mm (on average 20 mm). Segment 2 of labial palpus comparatively long, at upper border of outer surface white dusted with brown scales, lower part densely dusted with brown scales, exceptionally almost brown (Fig. 3), inner surface similarly coloured. Segment 3 white, mottled with brown scales, especially near top. Antennal scape with several long pecten scales. Flagellum brown, distinctly ringed with creamy ochre to whitish. Head, thorax and tegula creamy whitish, weakly mottled with brown scales. Forewing grey, slightly darker near costa, two small blackish dots at 1 / 3 and 2 / 3 of wing, fringes light grey. Hindwing dark grey with grey fringes. Female (Fig. 4). Wingspan 19.5 mm. Similar to male but generally lighter, hindwings whitish. Head of the only available female is lost. Variation. The degree of variability is very low. Fresh specimens appear darker and more contrasting. Male genitalia (Figs 5 – 6). Uncus broad, almost square, slightly rounded in corners of apical part. Gnathos stout, broad, slightly longer than uncus, middle part is straight, at end curved and sharply pointed. Tegumen broad, in shape of isosceles triangle, pedunculi broadly oval. Valva broad, regularly equal, extending to near top of uncus, apex distinctly rounded laterally and inwardly ended sharply, covered with setae, more densely in apex area. Separated sacculus not developed. Posterior margin of vinculum with comparatively deeper medial emargination. Saccus sub-triangular with weakly pointed apex, ratio maximum width to length approximately 0.7, lateral sclerites approximately 0.7 times maximum width of saccus. Phallus straight, bulbous coecum, in middle of phallus small sclerotised microplate with minute spines, ductus ejaculatorius twisted several times, apically with short interior sclerotisation. Female genitalia (Fig. 7). Papilla analis large, elongated, apically rounded. Anterior apophysis very short, posterior apophysis 3.5 times longer than anterior apophysis. Segment VIII distinct, slightly sclerotized distally. Subgenital plate with triangular subostial sclerotisation, distally pointed, anteriorly delimiting oblong ostium bursae, anterior margin smooth, slightly concave, connecting with anterior apophysis. Colliculum comparatively short, tube-shaped, sclerotised, slightly expanding anteriorly. Ductus bursae gradually widening to weakly delimited, comparatively broad corpus bursae, entire length of ductus and corpus bursae only slightly longer than entire length of papilla analis and posterior apophysis. Signum small, in shape of rounded spiny plate.	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD5151DFE9FFCC9FEB5F863.taxon	diagnosis	Differential diagnosis. Megacraspedus bidzilyai sp. nov. is characterised by grey coloration and two small black dots on the forewings in combination with a fairly large wingspan. It is most similar to M. majorella Caradja, 1920, but M. bidzilyai sp. nov. is overall darker, a particularly distinct difference is in the hindwing, which is very light in M. majorella. Moreover, in M. bidzilyai sp. nov., the forewing is densely dusted with darker scales and appears finely speckled, while in M. majorella the overall coloring gives a solid plain impression. Male genitalia of M. bidzilyai sp. nov. are unique within the genus due to broadly sub-rectangular uncus, stout gnathos and a small sclerotised spined patch on the phallus. Similarity is evident only with species from the M. majorella group, but M. majorella has distinctly thinner valvae and broader saccus while M. latiuncus Huemer & Karsholt, 2018 has a less stout gnathos, posteriorly tapering valves and differently shaped spined patch on the phallus. Molecular data. BIN: BOLD: ADR 7289. The average intraspecific distance of the barcode region is unknown (n = 1). The minimum distance to the nearest neighbour, Megacraspedus latiuncus (BIN: BOLD: AEK 4287), is 5.93 % (p-dist) (Fig. 18).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD5151DFE9FFCC9FEB5F863.taxon	etymology	Etymology. The species name is dedicated to the excellent Ukrainian lepidopterologist Oleksiy Bidzilya (Institute for Evolutionary Ecology of the National Academy of Sciences of Ukraine, Kyiv, Ukraine), who has made an exceptionally significant contribution to the knowledge of the gelechiids of Siberia including the Russian Altai Mountains.	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD5151DFE9FFCC9FEB5F863.taxon	biology_ecology	Biology. Host plant and early stages are unknown. The adults were collected at an altitude of 1400 m at the end of July. The type locality is extremely low-lying in the geomorphological conditions of the high parts of the Russian Altai, surrounded by several high mountain ridges with glacier-covered peaks, difficult to reach and substantially isolated from places with comparable natural conditions (Figs 19 – 20).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD5151DFE9FFCC9FEB5F863.taxon	distribution	Distribution. Russia: Altai Republic.	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD5151DFE9FFCC9FEB5F863.taxon	discussion	Remarks. Based on male genitalia characters, Megacraspedus bidzilyai sp. nov. belongs to the Megacraspedus majorella species group, which was established by HUEMER & KARSHOLT (2018). This group comprised only two species so far, and no female is known from either of them. HUEMER & KARSHOLT (2018) point to the uncertain systematic position of this species group due to the absence of supportive molecular data. However, since molecular data have been obtained for two species of this group, an updated annotated checklist of the M. majorella group is attached. Specific structures in the male genitalia of all three species and the close genetic relationship between M. latiuncus and M. bidzilyai sp. nov. confirm the correctness of establishing a separate species group. The description of the female of M. bidzilyai sp. nov. can be considered as the initial basis for the characteristics of the females of M. majorella group.	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD7151DFC3AFC41FC2EFBAA.taxon	type_taxon	Type locality. Russia (Altai Mountains).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD7151DFC3AFC41FC2EFBAA.taxon	distribution	Distribution. Russia, probably endemic to the Altai Mountains (this paper).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD7151DFCC5FE4CFBE6FC33.taxon	materials_examined	Type locality. Kazahkstan (Zailiskiy Alatau). Material examined. KYRGYZSTAN: North Tian-Shan Mts, Issyk-Kul Region, Kungei Ala-Too range, 13 km NW of Balykchy, 42 ° 32.5 ′ N, 76 ° 4.0 ′ E, 3050 m, 24. vii. 2019, 1 ♂ (gen. prep. 21351 Šumpich), (Barcode NMPC-LEP- 0664), I. Dvořák leg. (NMPC). Molecular data. BIN BOLD: AEK 4287. The average intraspecific distance of the barcode region is unknown (n = 1). The distance to the nearest neighbour, M. bidzilyai sp. nov. (BIN: BOLD: ADR 7289), is 5.93 % (p-dist) (Fig. 18).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD7151DFCC5FE4CFBE6FC33.taxon	biology_ecology	Biology. In Kyrgyzstan, it was collected in a grassy steppe slope at an altitude of 3050 m (Fig. 21).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD7151DFCC5FE4CFBE6FC33.taxon	distribution	Distribution. Northern Tian-Shan Mts in south-eastern Kazahkstan (HUEMER & KARSHOLT 2018) and northern Kyrgyzstan (this paper).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD7151DFC07FF2EFA45FEC5.taxon	materials_examined	Type locality. Kyrgyzstan (Alai Mountains).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD7151DFC07FF2EFA45FEC5.taxon	distribution	Distribution. Kyrgyzstan, probably endemic to the Alai Mountains (CARADJA 1920, HUEMER & KARSHOLT 2018).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD7151BFC03FBA4FB76FEA3.taxon	description	(Figs 8 – 9)	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD7151BFC03FBA4FB76FEA3.taxon	materials_examined	Material examined. RUSSIA: ALTAI REPUBLIC: Aktash vill., 50 ° 19 ′ 12 ″ N, 87 ° 36 ′ 00 ″ E, grassy steppe, rocks, 1400 m, 11. vii. 2014, 1 ♂ (Barcode NMPC-LEP- 0383), J. Šumpich leg. (NMPC); the same locality but 21. vi. 2015, 1 ♂, J. Šumpich leg. (NMPC); the same locality but 24. vi. 2019, 8 ♂♂, J. Šumpich leg. (NMPC); Kosh-Agach Distr., Kurai env. (15 km SW), Dzhangyskol lake, 50 ° 10 ′ 49 ″ N; 87 ° 44 ′ 19 ″ E, coniferous forest / steppe, 1830 m, 24. – 25. vi. 2015, 1 ♂ (Barcode NMPC-LEP- 0384), J. Šumpich leg. (NMPC); the same locality but 29. – 30. vi. 2019, 6 ♂♂, J. Šumpich leg. (NMPC); Shebalino Distr., Cherga vill. (8 km W), 51 ° 34 ′ 04 ″ N; 85 ° 28 ′ 33 ″ E, rocky slopes, steppe, 580 m, 4 ♂♂ (Barcode NMPC-LEP- 0615), J. Šumpich leg. (NMPC). Molecular data. A genetically extremely variable species resulted in 23 BINs (HUEMER & KARSHOLT 2018), which is a unique phenomenon within the Gelechiidae (HUEMER et al. 2020). The mean intraspecific divergence of the barcode region is 7.5 % and the maximum is 13.8 % (HUEMER & KARSHOLT 2018). Altaic specimens are clustered in two branches with different BINs: BOLD: ACB 3319 (n = 2; 2 public, both from Altai) [as “ BIN dolo 20 ” in HUEMER & KARSHOLT 2018], where the average intraspecific divergence of the barcode region is 0.96 % (p-dist), and BOLD: ACZ 3530 (n = 8, 8 public, all from Altai) [as “ BIN dolo 21 ” in HUEMER & KARSHOLT 2018], where the average intraspecific divergence of the barcode region is 0.03 % (pdist) (maximum distance 0.16 % (p-dist )). Distance between these clusters is 1.38 % (p-dist).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD7151BFC03FBA4FB76FEA3.taxon	distribution	Distribution. From Europe (not recorded in northern parts of the continent) to Central Asia (HUEMER & KARSHOLT 2018, GAST ṒN & VIVES MORENO 2021). Based on the current knowledge, the Altai Mountains seem to be the easternmost area of its occurrence.	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD11519FC10FEE6FD21FF42.taxon	description	(Figs 10 – 12) Nothris leuca Filipjev, 1929. Type locality: Russia (Sajan Mts).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD11519FC10FEE6FD21FF42.taxon	materials_examined	Material examined. RUSSIA: ALTAI REPUBLIC: Aktash vill., 50 ° 19 ′ 12 ″ N; 87 ° 36 ′ 00 ″ E, grassy steppe, rocks, 1400 m, 11. vii. 2014, 1 ♂, J. Šumpich leg. (NMPC); Kosh-Agach Distr., Kurai env. (6.5 km SW), 50 ° 10 ′ 35 ″ N; 87 ° 53 ′ 55 ″ E, grassy steppe, 1550 m, 9. – 10. vii. 2014, 19 ♂♂, J. Šumpich leg. (NMPC); the same locality but 30. vii. 2017, 9 ♂♂ (Barcode NMPC-LEP- 0385), J. Šumpich leg.; Belyashi (Dzhazator) env. (25 km NW), confluence of Argut and Karagem rivers, 49 ° 51 ′ 56 ″ N, 87 ° 10 ′ 22 ″ E, rocky steppe, 1400 m, 27. – 28. vii. 2017, 101 ♂♂ (Barcode NMPC-LEP- 0386), J. Šumpich leg.; Kosh-Agach Distr., Chagan-Uzun env., Krasnaya Gorka hill, 50 ° 05 ′ 00 ″ N; 88 ° 25 ′ 15 ″ E, rocky steppe, 1870 m, 29. vi. 2015, 1 ♂, J. Šumpich leg.; the same locality but 1. – 3. vii. 2019, 5 ♂♂, J. Šumpich leg.; Belyashi env. (56 km SE), 49 ° 38 ′ N; 88 ° 12 ′ E Dzhazator valley, mountain meadows near Tara river, 2300 m, 25. – 26. vii. 2017, 1 ♂, J. Šumpich leg.; Kosh-Agach Distr., Zhana-Aul env., 49 ° 48 ′ 28 ″ N, 88 ° 56 ′ 17 ″ E, steppe, rocky steppe, 2000 m, 24. vii. 2017, 17 ♂♂, J. Šumpich leg. (all NMPC). Molecular data. BIN BOLD: ACB 3260 (n = 12; 10 public, 10 from Altai). The average intraspecific divergence of the barcode region is 0.05 % (maximum 0.32 %). The distance to the nearest neighbours, M. attritellus Staudinger, 1871 (BIN: BOLD: ACE 2700) and M. skulei Huemer & Karsholt, 2018 (BIN: BOLD: ACM 0982), is 6.6 % (p-dist). Distribution. Russia (Altai Republic, Buryatia, Chitin Region, Tuva) (PONOMARENKO 2019), Mongolia (POVOLN Ỷ 1982).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD11519FC10FEE6FD21FF42.taxon	discussion	Remark. The species is widely distributed in the Altai Mountains and is the most common gelechiid species there.	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD31516FF7BFEC6FE1DFDA3.taxon	description	(Figs 13 – 14)	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD31516FF7BFEC6FE1DFDA3.taxon	materials_examined	Material examined. RUSSIA: ALTAI REPUBLIC: Aktash vill., 50 ° 19 ′ 12 ″ N; 87 ° 36 ′ 00 ″ E, grassy steppe, rocks, 1400 m, 11. vii. 2014, 1 ♂ (Barcode TLMF Lep 19983) (gen. prep. GU 16 / 1459 P. Huemer), J. Šumpich leg. (NMPC); 45 km N of Ulagan vill., Chulyshman valley, 51 ° 01 ′ 03 ″ N; 88 ° 00 ′ 39 ″ E, grassy steppe, rocks, 600 m, 26. – 27. vi. 2019, 2 ♂♂, J. Šumpich leg. (NMPC); the same locality but 4. – 5. vii. 2019, 3 ♂♂ (gen. prep. Šumpich 21379), J. Šumpich leg. (NMPC). Molecular data. BIN BOLD: ADB 8683 (n = 5; 1 public, 1 from Altai). The average intraspecific divergence of the barcode region is 0.33 % (maximum 0.88 %). The distance to the nearest neighbour, a specimen from Croatia, in BOLD currently identified as M. balneariellus (Chrétien, 1907) (BIN: AEC 7522, n = 1), is 2.24 % (p-dist). The distance to the second nearest neighbour, M. balneariellus (BIN: ADR 2637, n = 2), is 6.3 % (p-dist).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD31516FF7BFEC6FE1DFDA3.taxon	distribution	Distribution. Austria, Hungary, Romania, Russia (S Ural, Altai Mts), Ukraine (HUEMER & KARSHOLT 2018).	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
038887B4FFD31516FF7BFEC6FE1DFDA3.taxon	discussion	Remark. The first report from the Altai Mountains (HUEMER & KARSHOLT 2018) was based on the above presented record from Aktash in 2014.	en	Šumpich, Jan (2022): Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group. Acta Entomologica Musei Nationalis Pragae 62 (2): 401-409, DOI: 10.37520/aemnp.2022.022, URL: http://dx.doi.org/10.37520/aemnp.2022.022
