taxonID	type	description	language	source
038887C6FFC7FFCBFE4386171BEB0FF3.taxon	type_taxon	Typus: Abrahamia ditimena (H. Perrier) Randrian. & Lowry (≡ Protorhus ditimena H. Perrier).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFC7FFCBFE4386171BEB0FF3.taxon	diagnosis	Diagnosis Dioecious trees or shrubs. Leaves simple, persistent, opposite, subopposite, verticillate or alternate, chartaceous, subcoriaceous or coriaceous, lateral veins parallel, arcuate, forming an obtuse to acute angle with the midrib toward the base of the blade. Inflorescence erect, a thyrse or panicle. Flowers (4 -) 5 (- 6) - merous; pedicel short, articulated; sepals much smaller than petals; stamens alternipetalous, filament broadened at the base (sterile in female flowers, forming staminodes), anthers dorsifixed, introrse, glabrous; ovary pubescent or glabrous (rudimentary or absent in male flowers), with 1 locule containing a single subpendulous or pendulous ovule. Fruits drupaceous, ellipsoidal or ovoid, symmetrical or slightly asymmetrical, resiniferous; seeds ruminate; cotyledons usually inseparable (except in Abrahamia ibityensis (H. Perrier) Randrian. & Lowry, in which the resiniferous canals are only peripheral).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFC7FFCBFE4386171BEB0FF3.taxon	description	Description Dioecious trees or shrubs; bark with white, translucent or red resinous latex. Leaves alternate, opposite or subopposite (verticillate in one species), simple, coriaceous, subcoriaceous or chartaceous, persistent (deciduous in A. humbertii). Venation pinnate, craspedodromous or semicraspedromous, midvein usually flat, rarely very slightly raised above, prominently raised below (rarely on both surfaces), secondary veins parallel or nearly so, straight or arcuate, prominent below or on both surfaces, forming an obtuse to acute angle with the midrib toward the base of the blade. Inflorescence terminal and / or axillary, an erect thyrse or panicle, or intermediate and forming a paniculiform thyrse. Flowers usually 5 - merous (sometimes 4 - or 6 - merous); sepals much smaller than petals; stamens 5 or 4 (rarely 6), alternipetalous, filaments straight, flattened, broadened at the base, inserted basally on the outer surface of the disk, anthers dorsifixed, introrse, dehiscent with longitudinal slits (stamens reduced in size and sterile in female flowers, forming staminodes); disk usually cupuliform and crenulate in male flowers, annular and cupuliform in female flowers; ovary with 1 locule containing a single pendulous or subpendulous or ovule on a short funiculus, placentation subapical (ovary rudimentary or absent in male flowers); style one, short, apically trifid or unbranched with 3 capitate stigmatic lobes. Fruits drupaceous, ellipsoidal or ovoid, symmetrical or slightly asymmetrical, exocarp usually with longitudinal striations on surface, glabrous or velutinous, mesocarp resinous, endocarp thin, papyraceous; seed solitary, with resiniferous canals throughout (only peripheral in A. ibityensis), ruminate; cotyledons usually inseparable (separable in A. ibityensis).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFC7FFCBFE4386171BEB0FF3.taxon	etymology	Etymology The genus is named in honor of the late Jean Prosper Abraham (1936 - 1996), a wellknown Malagasy forester with a remarkable knowledge of the woody plants of Madagascar and who contributed much to our knowledge of the island’s rich and diverse flora.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFC7FFCBFE4386171BEB0FF3.taxon	discussion	Notes Abrahamia includes a total of 34 species, all endemic to Madagascar, including 15 species formerly placed in Protorhus (e. g., by PERRIER DE LA BâTHIE, 1944, 1946), to which 19 additional species are newly described here. With the recognition of Abrahamia as a separate genus, Protorhus is now restricted to a single species in Africa, P. longifolia. The two genera differ in several flower and fruit characters, as indicated by PELL (2004) and PELL et al. (2011), and summarized in Table 1. Abrahamia is represented in all of Madagascar’s major vegetation types (MOAT & SMITH, 2007) and broad bio-climatic zones (CORNET, 1974; SCHATZ, 2000). Species can be found in humid forest in the eastern and Sambirano regions (A. antongilensis, A. sambiranensis), wooded grassland and bushland in the center of the island (A. ibityensis, A. buxifolia), western dry forest (A. deflexa, A. humbertii), and southwestern subarid spiny forest-thicket (A. phillipsonii Randrian. & Lowry). Members of the genus occur on many types of substrate and at a wide range of elevations, from littoral forest on sand at the sea level (A. nitida (Engl.) Randrian. & Lowry, A. suarezensis Randrian. & Lowry) to humid forest on lateritic soil at altitudes up to 2000 m (A. buxifolia, A. itremoensis). Species of Abrahamia were described by PERRIER DE LA BâTHIE (1946) as polygamo-dioecious, indicating that he interpreted them as having both unisexual and hermaphroditic flowers on the same individual. However, careful observation has shown that all members of the genus are functionally dioecious. Female flowers have rudimentary, sterile stamens that are reduced in size, whereas male flowers either have an ovary that is rudimentary and non-functional or lack an ovary altogether.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFD5FFF4FE4383651B860F22.taxon	materials_examined	Typus: MADAGASCAR. Prov. Toamasina: Rég. Analanjirofo, Anjanaharibe forest, 15 ° 10 ’ 48 ’’ S 49 ° 37 ’ 30 ’’ E, 860 m, 4. II. 2003, fr., Antilahimena 1862 (holo-: MO- 6671431!; iso-: G [G 00341711]!, K!, NY!, P [P 00580355]!, TAN [TAN 002003]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFD5FFF4FE4383651B860F22.taxon	diagnosis	Diagnosis Abrahamia antongilensis Randrian. & Lowry can be distinguished from the other members of the genus by the combination of its leaves with semicraspedodromous venation and a distinctly acuminate apex (10 - 15 mm long), and inflorescences that are very short (c. 1.8 cm) and terminal.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFD5FFF4FE4383651B860F22.taxon	description	Description Trees 8 - 10 m tall; young twigs covered with dense, whitish sericeous indument. Leaves alternate, clustered at the end of branches; blade oblanceolate to obovate, 7 - 17 3 3 - 6 cm, chartaceous, apex abruptly acuminate, the acumen 10 - 15 mm long, margin undulate and very slightly revolute, base cuneate, adaxial surface glabrous or with scattered whitish sericeous indument, abaxial surface covered with dense whitish, appressed sericeous indument when younger, much less dense on older leaves, venation semicraspedodromous, midvein prominent abaxially, secondary veins 25 - 35 pairs, 2 - 7 mm apart, forming a wide angle with the midvein (almost perpendicular from the base to the leaf mid-section), raised on both surfaces, tertiary veins visible on both surfaces; petiole 10 - 18 mm long, c. 1.5 mm in diam., canaliculate on adaxial surface, covered with whitish sericeous indument. Inflorescence short, terminal, a panicle, c. 1.8 cm long, axes covered with dense, appressed indument. Male flowers unknown. Female flowers 5 - merous, small, c. 3 mm long, subtended by a very small bract with dense indument; pedicel c. 3 mm long, densely sericeous; calyx lobes small, widely ovate, 1 3 1 mm, valvate, adaxial surface glabrous, abaxial surface pubescent; corolla lobes ovate, 2.5 3 1.5 mm, thick, imbricate, adaxial surface glabrous, abaxial surface pubescent; staminodes 5; disk cupuliform, c. 1.5 mm in diam., glabrous; ovary slightly ovoid to spherical, c. 1 3 1 mm, glabrous; style short, thick, c. 1 mm long, glabrous, stigmas capitate. Fruits 2.5 - 3.5 3 1.7 - 2 cm, ellipsoid, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFD5FFF4FE4383651B860F22.taxon	distribution	Distribution, ecology and phenology This species is known only from the area around the northern end of the Baie d’Antongil in northeastern Madagascar (Map 1). Flowering material has been collected in November and fruiting specimens in February.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFD5FFF4FE4383651B860F22.taxon	conservation	Conservation status With a minimum AOO of 8 km 2 and two known localities, one within the newly established Makira natural park and the other based on a historical collection that can be assumed to be have come from within the buffer zone of Masoala national park, comprising 2 locations, the latter of which is subjected to projected decline in habitat quality, A. antongilensis is assigned a preliminary conservation status of “ Endangered ” [EN B 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFD5FFF4FE4383651B860F22.taxon	discussion	Notes Abrahamia antongilensis most closely resembles A. lenticellata and A. viguieri but differs by having the abaxial leaf surface covered with whitish appressed indument (vs glabrous in A. lenticellata and with reddish lanate indument in A. viguieri). Moreover, A. antongilensis can be distinguished from A. lenticellata by having glabrous fruits (vs velutinous) and from A. viguieri by its leaves with semicraspedodromous (vs craspedodromous) venation. Abrahamia antongilensis is only known from two collections, both from the area around the Baie d’Antongil in northeastern Madagascar. The type collection (Antilahimena 1862), from northwestern of Maroantsetra, has older leaves with less indument on the abaxial surface than on the much younger leaves of the second collection (Service Forestier 8770) from the northern part of the Masoala Peninsula, approximately half way between Antalaha and Maroantsetra.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFD5FFF4FE4383651B860F22.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Toamasina: Rég. SAVA, S du Col d’Ambatondradama, [15 ° 16 ’ S 50 ° 04 ’ E], c. 750 m, 17. XI. 1953, fl., Service Forestier 8770 (MO, P [2 sheets], TEF).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE8FFF6FE4383651B5208C6.taxon	materials_examined	Typus: MADAGASCAR. Prov. Toamasina: Rég. Atsinanana, Betampona RN, [17 ° 55 ’ S 49 ° 13 ’ E], 24. X. 1955, bud, Réserves Naturelles 7701 (holo-: P [P 06774885]!; iso-: TEF [TEF 000880]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE8FFF6FE4383651B5208C6.taxon	diagnosis	Diagnosis Abrahamia betamponensis Randrian. & Lowry can easily be distinguished from other members of the genus by its large leaves (9 - 24 3 4.3 - 8.5 cm) with a remarkably prominent midvein on the abaxial surface, a long, thick petiole (15 - 18 mm in length), numerous secondary veins (50 - 60 pairs), and fruit surface covered with dense, short indument.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE8FFF6FE4383651B5208C6.taxon	description	Description Trees c. 7 m tall; bark with lenticels; young twigs whitish-gray to light tan, pubescent at the very tips. Leaves opposite or subopposite; blade oblanceolate, 9 - 24 3 4.3 - 8.5 cm, subcoriaceous to coriaceous, apex shortly acuminate, retuse or emarginate, margin slightly revolute, base cuneate, adaxial surface glabrous, abaxial surface sparsely strigose along midvein, venation craspedodromous, midvein strikingly prominent abaxially, secondary veins 50 - 60 pairs, 2 - 5 mm apart, raised on both surfaces, tertiary veins visible on both surfaces, petiole 15 - 18 mm long, canaliculate on adaxial surface, c. 2.5 - 3 mm in diam., glabrous. Inflorescence terminal and axillary, a panicle, 2 - 7 cm long, axes thick, covered with dense, short, appressed indument; bracts triangular, c. 4 3 2 mm, adaxial surface glabrous, abaxially surface with short, appressed indument. Male flowers 5 - merous, small, c. 3 mm long, subtended by a small, deltoid bract; pedicel short, c. 1 mm long, strigose; calyx lobes triangular, 2.5 3 1 mm, adaxial surface glabrous, abaxial surface pubescent; corolla lobes very widely ovate, 2 3 2 mm, adaxial surface glabrous, abaxial surface pubescent; stamens 5, filaments 1.2 - 1.3 mm long, anthers c. 1 mm long, ovoid; disk thin, dish-shaped and glabrous. Female flowers unknown. Fruits 2.4 - 3.2 3 1.4 - 1.8 cm, ellipsoid, covered with dense, short indument, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE8FFF6FE4383651B5208C6.taxon	vernacular_names	Vernacular names “ Hazombarorana ”, “ Hazombarorana à grandes feuilles ”, “ Vatodinga’ ’, “ Voatodinga ”.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE8FFF6FE4383651B5208C6.taxon	distribution	Distribution, ecology and phenology Abrahamia betamponensis is known only from Betampona reserve and its vicinity, with a single collection from Zahamena national park (Map 1). It occurs in mid-elevation humid forest. Material in bud has been collected in October and fruiting specimens have been made in January.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE8FFF6FE4383651B5208C6.taxon	conservation	Conservation status Abrahamia betamponensis has a minimum EOO of 8 km 2, but its two known localities are both situated within protected areas (Betampona reserve and Zahamena national park) so in the absence of any data on population size, no future decline is expected in terms of AOO, EOO, or quality of habitat. It has therefore assigned a preliminary status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE8FFF6FE4383651B5208C6.taxon	discussion	Notes Abrahamia betamponensis belongs to a group of species with large leaves (up to 37 cm long) that have numerous (40 - 70) secondary veins. Within this group, A. betamponensis most closely resembles A. sambiranensis, but differs by the dense, short indument on the surface of its mature fruit (vs glabrescent or glabrous).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE8FFF6FE4383651B5208C6.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Toamasina: Rég. Antisnana, Fotsimavo, [Betampona RN], [17 ° 55 ’ S 49 ° 13 ’ E], 300 - 400 m, 20. I. 1950, fr., Réserves Naturelles 2366 (P, TAN); ibid. loco, Réserves Naturelles 2489 (P, TAN); Anjiro, Ambodiriana, [Betampona RN], [17 ° 55 ’ S 49 ° 13 ’ E], 26. X. 1957, bud, Réserves Naturelles 9148 (P, TEF); Mangabe, [Zahamena PN], [17 ° 42 ’ S 48 ° 54 ’ E], 20. I. 1963, old fr., Réserves Naturelles 12446 (MO, P, TEF); Betampona RN, 17 ° 54 ’ 29 ” S 49 ° 12 ’ 16 ” E, 530 m, 13. I. 2017, ster., Schatz et al. 4472 (MO, P, TAN).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE2FFFFFE4383651D490CBC.taxon	materials_examined	Lectotypus (designated here): MADAGASCAR. Prov. Toliara: Mt. Belambana, bassin du Mananara (Sud), [22 ° 08 ’ S 46 ° 21 ’ E], 1000 m, VII. 1910, Perrier de la Bâthie 5852 (P [P 06775177]!; isolecto-: P [P 00722637]!). Syntypi: MADAGASCAR. Prov. Fianarantsoa: massif de l’Andringitra, [22 ° 13 ’ S 47 ° 01 ’ E], c. 1800 m, IV. 1921, fr., Perrier de la Bâthie 13696 (P [P 00722638]!). Prov. Toliara: Mont Vohipolaka, N Betroka, [22 ° 08 ’ S 46 ° 21 ’ E], 1100 - 1300 m, XI. 1933, ster., Humbert 11637 bis (P [P 06775327]!); ibid. loco, 1100 - 1200 m, Humbert 11667 bis (P [P 00722639]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE2FFFFFE4383651D490CBC.taxon	description	Description Trees 2 - 7 m tall, bark with white milky latex; young twigs pubescent or glabrescent, lenticellate or waxy. Leaves subopposite or opposite; blades obovate to elliptic, 1.5 - 6.8 3 1 - 2.7 cm, subcoriaceous, apex retuse or rounded, margin entire, base cuneate, adaxial surface glabrous, glaucous, often chocolate brown when dry, abaxial surface glabrous (except for sericeous indument on the midvein, and sometimes scattered indument elsewhere on the blade), venation craspedodromous, midvein prominent abaxially, secondary veins 15 - 25 pairs (rarely to 36 pairs), 1 - 3 mm apart, prominent on both surfaces, tertiary veins raised on both surfaces; petiole 5 - 7 mm long, c. 1 - 1.1 mm in diam., shallowly canaliculate on adaxial surface, glabrescent or pubescent. Inflorescences axillary and terminal, a thyrse, 1.5 - 4.5 cm long, pubescent. Male and female flowers 5 - merous; pedicel c. 0.2 - 0.3 mm long, pubescent; calyx lobes deltate to widely depressed ovate, 1 - 1.2 3 1 - 1.5 mm, valvate, persistent on fruit, adaxial surface glabrous, abaxial surface pubescent; corolla lobes ovate, 2 - 3 3 1.5 - 2 mm, pale yellow, imbricate, adaxial surface glabrous, abaxial surface pubescent. Male flowers with 5 or 6 stamens, filaments c. 1 - 1.5 mm long, glabrous or with sparse indument, white, anthers c. 0.8 mm long, obloid to slightly ovoid, yellow, glabrous; disk c. 2 mm in diam., glabrous; ovary rudimentary. Female flowers with 5 or 6 staminodes, filaments c. 1 mm long, glabrous, white, anthers c. 0.5 mm long, ovoid, sterile, yellowish, glabrous; disk c. 1.5 mm in diam., glabrous; ovary widely ovoid, 1 - 2 3 1 - 1.5 mm, pubescent; style shortly trifid, inserted apically, c. 1 mm long, pubescent, stigmas capitate. Fruits c. 1.5 3 1.1 cm, ellipsoid to slightly obovoid, surface smooth, velutinous when young, glabrescent.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE2FFFFFE4383651D490CBC.taxon	vernacular_names	Vernacular names “ Sohihy’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE2FFFFFE4383651D490CBC.taxon	distribution	Distribution, ecology and phenology Abrahamia buxifolia occurs in the southern and southwestern parts of central Madagascar, with populations extending toward the western slopes and the eastern escarpment (Map 1). It grows at elevation between 900 - 1800 m in areas where ecological conditions are harsh, including low rainfall and poor soils. It flowers in September and October, and sets fruits between November and February.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE2FFFFFE4383651D490CBC.taxon	conservation	Conservation status With an EOO of 37,259 km ², a minimum AOO of 60 km ², and 18 recorded localities, most of which are subjected to frequent burning and only 3 of which are within protected areas (Andringitra, Ibity, and Itremo), representing 11 locations, A. buxifolia is assigned a preliminary conservation status of “ Near Threatened’ ’ [NT] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE2FFFFFE4383651D490CBC.taxon	discussion	Notes Abrahamia buxifolia can easily be distinguished from its most similar congener, A. itremoensis, by its glaucous and subcoriaceous leaves (vs non-glaucous and coriaceous) that are somewhat larger in size (1.5 - 6.8 3 1 - 2.7 vs 2 - 6 3 0.6 - 2 cm), and by its flowers with a glabrous disk (vs with evident indument). Of the four collections cited in the protologue that we examined, we have selected Perrier de la Bâthie 5852 as the lectotype because it is more complete than the other syntypes and is represented by two specimens, whereas each of the three other syntypes comprises only a single sheet, two of which (Humbert 11637 bis and Humbert 11667 bis) are sterile and one (Perrier de la Bâthie 13696) has just a few fruits. Two other collections were also mentioned in the protologue (Decary 13041 and Humbert 14048) but we have not been able to find these specimens (although a sheet of the latter is reported to be at US) and we have not listed them below in the material examined. The label data for Service Forestier 23500 and 23501 indicate a trunk diameter of 0.5 m, a value that far exceeds any observation we have made in the field. We therefore regard this as a label error and have not included this information in the description.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE2FFFFFE4383651D490CBC.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Antananarivo: Antsirabe, Betafo, Antakoboka, 20 ° 03 ’ 46 ’’ S 46 ° 49 ’ 30 ’’ E, 1807 m, 17. IX. 2004, y. fl., Andriamihajarivo 371 (G, MO, P, TAN, TEF); forêt d’Analandraisoa (SW Tsiroanimandidy), [19 ° 09 ’ S 45 ° 49 ’ E], 1200 - 1300 m, 17 - 20. XI. 1952, fr., Leandri 1852 (G, MO, P [2 sheets], TAN); Mont Ibity, E slope of Kibory, 20 ° 06 ’ S 46 ° 59 ’ E, 1600 m, 2. X. 1996, ster., A. Randrianasolo 488 (MO, P, TAN); Tsiroanomandidy, forêt d’Analandraisoa, [18 ° 59 ’ 30 ’’ S 45 ° 41 ’ 30 ’’ E], c. 1100 m, XI. 1952, fr., Service Forestier 6726 (G, MO, P, TEF). Prov. Fianarantsoa: Ambatofinandrahana, Itremo, forêt d’Antsirakambiaty, 20 ° 35 ’ 42 ’’ S 46 ° 33 ’ 55 ’’ E, 1605 m, 11. XII. 2003, fr., Andriamihajarivo 258 (G, MO, P, TAN); Lovokaloha, 20 ° 29 ’ 58 ’’ S 46 ° 51 ’ 37 ’’ E, 1411 m, 2. XI. 2004, fr., Andriamihajarivo 408 (G, MO, P, TAN); Ambatofinandrahana, 10 km N de RN 35, Antanimena, 20 ° 31 ’ 05 ’’ S 46 ° 34 ’ 02 ’’ E, 1541 m, 24. VIII. 2010, fr., Andrianaivoravelona et al. 363 (K, MO, TAN); entre Ambatofinandrahana et Itremo, [20 ° 33 ’ S 46 ° 45 ’ E], IX. 1956, bud, Bosser 10037 (MO, P [3 sheets], TAN); Itremo, [20 ° 32 ’ S 46 ° 33 ’ E], 1600 m, IX. 1956, bud, Bosser 10075 (P, TAN); Inselbergs, PK 475, W of Ambalavao, 950 - 1050 m, 1. II. 1975, fr., Croat 30326 (MO); N of Sendrisoa, c. 1200 m, 23. II. 1975, fr., Croat 32165 (MO, P, TAN); Itremo massif, along Ambalo-ranogolana creek, 20 ° 34 ’ 31 ’’ S 46 ° 34 ’ 50 ’’ E, 1500 m, 9. XI. 2002, y. fr., Lowry et al. 5861 (G, MO, P, TAN); Itremo massif, montane forest on quartz, 20 ° 34 ’ 40 ’’ S 46 ° 34 ’ 53 ’’ E, 1550 m, 29. X. 1997, fl., Malcomber et al. 2849 (G, MO, P, TAN); Anja Parc, Inselberg Ambohilady, 21 ° 51 ’ 11 ’’ S 46 ° 51 ’ 14 ’’ E, 1224 m, 10. XII. 2010, fr., Ramahefaharivelo 272 (G, MO, P, TAN); Itremo chain, W of Itremo village, 20 ° 35 ’ 00 ’’ S 46 ° 36 ’ 10 ’’ E, 1500 m, 4. X. 1996, fl., A. Randrianasolo 489 (G, MO, P, TAN); ibid. loco, 1450 m, 6. X. 1996, bud, A. Randrianasolo 493 (G, MO, P, TAN); Ambalavao, Sendrisoa, [22 ° 07 ’ S 46 ° 49 ’ E], 15. XI. 1953, fr., Réserves Naturelles 5858 (P, TEF); Ivohibe, [Ivohibe RN], Antambohobe, 12. IX. 1962, bud, Réserves Naturelles 12154 (P); forêt Ambondrozana, Ihosy, [21 ° 57 ’ S 46 ° 36 ’ E], 17. II. 1955, bud, Service Forestier 13771 (P, TEF); massif granitique d’Ifandana, entre Ankaramena et Ihosy, [22 ° 01 ’ 00 ’’ S 46 ° 22 ’ 30 ’’ E], 1300 - 1400 m, 7. X. 1964, fl., Service Forestier 23500 (G, MO, P [2 sheets], TEF); ibid. loco, Service Forestier 23501 (G, MO, NY, P, TEF); massif d’Iandrambaky, SW d’Iarintsena (Ambalavao), [21 ° 51 ’ S 46 ° 51 ’ E], 4. XI. 1967, y. fr., Service Forestier 27830 (G, MO, P, TEF); rocher de l’Ifandana, N de Zazafotsy, [22 ° 01 ’ 00 ’’ S 46 ° 22 ’ 30 ’’ E], 2. XII. 1968, fr., Service Forestier 28473 (G, K, MO, NY, P [2 sheets], TEF); Ambatofinandrahanana, Andranomivoaka, 18. III. 1952, ster., Service Forestier 45 - R- 207 (P); Ambalavao, forêt d’Amboandrozana, 9. VI. 1954, ster., Service Forestier 63 - R- 141 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE1FFF9FE4383651B5C0E06.taxon	materials_examined	Typus: MADAGASCAR. Prov. Antsiranana: Rég. SAVA, Dist. Sambava, S de l’Anjehabe, [14 ° 18 ’ 30 ’’ S 49 ° 44 ’ 40 ’’ E], c. 600 m, 7. XI. 1950, fl., Service Forestier 767 (holo-: P [P 06774827]!; iso-: G [G 00341710]!, MO- 6684132!, NY!, P [P 06774828, P 06774829]!, TEF [TEF 000881]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE1FFF9FE4383651B5C0E06.taxon	diagnosis	Diagnosis Abrahamia capuronii Randrian. & Lowry can be distinguished from all other species of Abrahamia by a combination of its elliptic leaves, narrow corolla lobes (3 3 1.2 mm), and slender inflorescence axes.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE1FFF9FE4383651B5C0E06.taxon	description	Description Trees 10 - 15 m tall, 20 cm DBH, bark with white or translucid latex; young twigs only pubescent at the tips. Leaves alternate or sub-opposite; blade obovate to elliptic, 4 - 7.8 3 2 - 3.6 cm, chartaceous to subcoriaceous, apex rounded or retuse, margin undulate, base cuneate, adaxial surface glabrous, abaxial surface with short appressed indument, mainly along the midvein and at the base, venation craspedodromous, midvein prominent abaxially, secondary veins 18 - 33 pairs, 1 - 4 mm apart, prominent abaxially, slightly raised adaxially, tertiary veins slightly visible on adaxial surface; petiole 10 - 25 mm long, c. 0.5 - 1 mm in diam., canaliculate on adaxial surface, glabrescent. Inflorescence axillary or terminal, a panicle, 2 - 8 cm long, axes slender, strigose. Male flowers 5 - merous, small, c. 2 - 2.5 mm long, subtended by a very small, pubescent bract; pedicel short, c. 0.3 - 0.5 mm long, pubescent; calyx lobes small, widely ovate, 1 3 1 mm, valvate, connate at the base, adaxial surface glabrous, abaxial surface sparsely covered with short indument, glabrescent; corolla lobes 3 3 1.2 mm, lanceolate, imbricate, adaxial surface glabrous, abaxial surface pubescent along its longitudinal axis or glabrescent to glabrous, white (according to field notes); stamens 5, filament c. 1.5 mm, glabrous, anthers c. 1 mm long, obloid, glabrous; disk glabrous, c. 1.2 - 1.5 mm in diam.; ovary rudimentary. Mature and opened female flowers unknown, developing buds with calyx lobes triangular to deltate, c. 1 3 0.5 - 0.8 mm, adaxial surface glabrous, abaxial surface with some indument; corolla lobes ovate, c. 1.5 3 1 mm, mostly glabrous but abaxial surface with a few appressed short trichomes; stamens vestigial; disk glabrous; ovary depressed, ovoid, 0.3 - 0.5 3 0.5 mm, glabrous; style c. 1 mm, glabrous, stigmas capitate. Young fruits ellipsoid, very slightly asymmetrical, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE1FFF9FE4383651B5C0E06.taxon	etymology	Etymology The species epithet honors René Capuron, whose extensive field work contributed so much to our knowledge of Madagascar’s woody flora, and who personally collected material of 24 of the 34 species of Abrahamia.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE1FFF9FE4383651B5C0E06.taxon	distribution	Distribution, ecology and phenology Abrahamia capuronii is restricted to far northeastern Madagascar (Map 1), where it occurs in mid-elevation humid and transitional forests on rocky substrate up to an elevation of 700 m. Material with buds and flowers has been collected in October and early November, and specimens with young fruits have been gathered in late November.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE1FFF9FE4383651B5C0E06.taxon	conservation	Conservation status With an EOO of 2,585 km ² and a minimum AOO of 24 km ², A. capuronii would qualify as “ Endangered ” under Criteria B 1 and B 2 of the IUCN Red List (IUCN, 2012) if other qualifying criteria were met. However, it is known from a total of 6 locations, one of which is situated within a protected area (Loky Manambato) while the others are at sites potentially subjected to clearing and habitat degradation, and should thus be assigned a preliminary status of “ Vulnerable’ ’ [VU B 1 ab (iii) + 2 ab (iii)].	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE1FFF9FE4383651B5C0E06.taxon	discussion	Notes Abrahamia capuronii resembles both A. grandidieri and A. sericea by its relatively closely spaced, parallel and craspedodromous secondary veins, but can be distinguished from these two species by its smaller number of secondary veins (10 - 33 in A. capuronii vs 35 - 40 (- 47) in A. grandidieri and 40 - 70 in A. sericea).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE1FFF9FE4383651B5C0E06.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Antsiranana: Mont Anjenabe, [14 ° 17 ’ S 49 ° 46 ’ E], 650 m, 3 - 7. XI. 1950, fl., Humbert 24055 (G, K, MO, P [2 sheets], TAN); Vohémar, forêt de Binara, 13 ° 15 ’ S 49 ° 37 ’ E, 700 m, 8. XI. 2001, fl., Gautier et al. 4125 (G, MO, TAN); Vohémar, Analamazava forest, 13 ° 15 ’ 43 ’’ S 49 ° 35 ’ 22 ’’ E, 25. XI. 2004, y. fr., Ranarivelo 446 (MO); Daraina, Ambilobe, 13 ° 44 ’ 10 ’’ S 49 ° 23 ’ 07 ’’ E, 1299 m, 27. X. 2007, bud, Randriambololomamonjy et al. 166 (MO, P, TAN); Vohémar, Daraina, forêt d’Analamazava, [13 ° 15 ’ 58 ’’ S 49 ° 36 ’ 14 ’’ E], 700 m, 29. X. 2005, fl., Randrianaivo 1259 (CNARP, MO, P, TAN); Vohémar, Daraina, forêt d’Antsahabe, 13 ° 12 ’ 37 ’’ S 49 ° 33 ’ 29 ’’ E, 545 m, 31. X. 2005, fl., Razafitsalama 741 (CNARP, MO, P, TAN); Vohémar, Andrafainkona, Ampisarahana, 13 ° 38 ’ 28 ’’ S 49 ° 31 ’ 51 ’’ E, 1009 m, 9. XI. 2007, bud, Razakamalala et al. 3716 (G, MO, P, TAN).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE5FFFAFE4383651B5A0D86.taxon	materials_examined	Typus: MADAGASCAR. Prov. Antsiranana: Rég. SAVA, Daraina, forêt d’Antsahabe, 13 ° 12 ’ 53 ’’ S 49 ° 31 ’ 53 ’’ E, 460 m, 25. I. 2006, fr., Ranirison & Nusbaumer 1118 (holo-: G [G 00090546]!; iso-: MO- 6671432!, P [P 00580353]!, TEF [TEF 000882]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE5FFFAFE4383651B5A0D86.taxon	diagnosis	Diagnosis Abrahamia darainensis Randrian. & Lowry can be distinguished from other members of the genus by its small, chartaceous, glabrous, elliptic to obovate leaves with a short petiole (5 - 7 mm long), and its glabrous, ellipsoid fruits that measure 2.6 3 1.5 cm.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE5FFFAFE4383651B5A0D86.taxon	description	Description Trees 6 m tall, 9 cm DBH. Leaves alternate; blade elliptic to obovate, 4 - 7.5 3 1.2 - 2.2 cm, chartaceous, apex shortly acuminate, margin undulate, base cuneate, both surfaces glabrous, venation craspedodromous, midvein visible but not prominent on both surfaces, secondary veins 14 - 18 pairs, 3 - 5 mm apart, almost impressed adaxially, slightly raised baxially, tertiary veins few and only weakly visible adaxially; petiole 5 - 7 mm long, c. 0.5 - 1 mm in diam., canaliculate on adaxial surface, glabrous. Inflorescence and flowers unknown. Fruits 2.6 3 1.5 cm, ellipsoid, light red, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE5FFFAFE4383651B5A0D86.taxon	distribution	Distribution, ecology and phenology Abrahamia darainensis is known only from humid forest on the eastern slope of Antsahabe in the Daraina region (Map 2). It was collected in fruit in January.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE5FFFAFE4383651B5A0D86.taxon	conservation	Conservation status While Abrahamia darainensis is only known from a single locality, it is situated within a large block of forest in the Loky Manambato protected area (Daraina), and in the absence of data on population size, no future decline is expected in terms of AOO, EOO, or quality of habitat, so we therefore assign a preliminary conservation status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFE5FFFAFE4383651B5A0D86.taxon	discussion	Notes Abrahamia darainensis is similar to A. lenticellata in having chartaceous leaves with a short acuminate apex and undulate margins, and fruits of similar size (2.6 3 1.5 cm in A. darainensis and 2 - 2.6 3 1.3 - 1.5 cm in A. lenticellata). It differs, however, by having secondary veins that are fewer in number (14 - 18 vs 20 - 25) and are craspedodromous (vs semicraspedodromous), a shorter petiole (5 - 7 mm vs 7 - 13 mm), and glabrous (vs velutinous) fruits. Abrahamia darainensis is only known from the type gathering.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFFAFFE4FE4383631A280D11.taxon	materials_examined	Lectotypus (designated here): MADAGASCAR. Prov. Mahajanga: Ambongo, XI. 1903, fl., Perrier de la Bâthie 1627 (P [P 00580302]!; isolecto-: MO- 6684133!, P [P 00580303]!). Syntypi: MADAGASCAR. Prov. Mahajanga: Ankatsepy, près Majujnga, s. d., fr., Perrier de la Bâthie 1627 bis (P [P 00580304]!). Prov. Toliara: Mt. Vohipolaka, N de Betroka, s. d., ster., Humbert 11694 ter (P, not seen); Mt. Apika, au-dessus de Mahamavo, bassin de réception de la Mananara, affluent du Mandrare, s. d., fr., Humbert 13850 (P, not seen).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFFAFFE4FE4383631A280D11.taxon	description	Description Trees 8 - 15 m tall, 15 - 20 cm DBH, bark with red latex, young twigs with lenticels, waxy, the very tips scabrous. Leaves alternate, subopposite or opposite; blades ovate, oblong, or elliptic, 2.5 - 6.5 3 1 - 3.5 cm, apex retuse, rounded or shortly acuminate, sometimes emarginate, margine entire, base cuneate, adaxial surface glabrous, abaxial surface ferruginous to scabrous when young, later glabrous or glabrescent along the midvein, venation craspedodromous, midvein prominent abaxially, secondary veins 15 - 20 pairs, more or less parallel, 1 - 4 mm apart, prominent abaxially, slightly prominent adaxially, tertiary veins very widely spaced, slightly raised on both surfaces; petiole 5 - 10 mm long, canaliculate on adaxial surface, covered with very short, appressed indument when young, glabrescent. Inflorescence terminal, a thyrse, 2 - 5 cm long, ferruginous strigose. Male flowers 5 - merous; pedicel c. 0.3 - 0.4 mm long, strigose; calyx lobes depressed ovate, 0.8 - 1 3 0.5 - 0.8 mm, imbricate, adaxial surface glabrous, abaxial surface glabrous or pubescent, persistent on the fruit; corolla lobes ovate, c. 2 3 1.5 mm, white or cream, adaxial surface glabrous, abaxial surface glabrous or with appressed short indument, imbricate; stamens 5, filaments 1 mm long, glabrous, anthers c. 0.5 mm long, ovoid to subglobose, yellow, glabrous; disk dish-shaped to cupuliform, c. 1 mm in diam., glabrous; ovary rudimentary. Female flowers unknown. Fruits 1.5 - 3 3 1 - 1.5 cm, ellipsoid to ovoid, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFFAFFE4FE4383631A280D11.taxon	vernacular_names	Vernacular names “ Hazombora’ ’, “ Sohihy’ ’, “ Sohy’ ’, “ Tarata’ ’, “ Motsalahy’ ’ (Sakalava).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFFAFFE4FE4383631A280D11.taxon	distribution	Distribution, ecology and phenology Abrahamia deflexa occurs in dry forests of western and northwestern Madagascar (Map 1), usually on sandy or calcareous soils. It flowers between September and November, and has been collected in fruit in November and December.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFFAFFE4FE4383631A280D11.taxon	conservation	Conservation status With an EOO of 51,339 km 2, a minimum AOO of 40 km 2, and an estimated maximum AOO of less than 2,000 km 2, as well as 10 known locations, only 2 of which are situated within protected areas (Ankarafantsika and Bemaraha) with the remainder projected to be subjected to continuing decline in habitat quality due to a combination of fire and land clearing, A. deflexa is assigned a preliminary conservation status of “ Vulnerable’ ’ [VU B 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFFAFFE4FE4383631A280D11.taxon	discussion	Notes Abrahamia deflexa can easily be distinguished from all its congeners by the combination the small, glabrous leaves with a long, thin petiole and glabrous fruits. Of the five collections cited in the protologue, we have selected Perrier de la Bâthie 1627 as the lectotype because it is represented by three specimens, whereas one of the other syntypes (Perrier de la Bâthie 1627 bis) comprises only a single sheet, another (Humbert 5063 = Humbert & Swingle 5063 = Service Forestier 50) is a collection of A. phillipsonii, and the remaining two (Humbert 11694 ter and 13850) could not be located.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFFAFFE4FE4383631A280D11.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Mahajanga: Katsepy, station d’Antrema, [15 ° 42 ’ 37 ’’ S 46 ° 10 ’ 30 ’’ E], 10. XI. 2008, fr., Hladick 7200 (P); Bemaraha, S of the Manambolo River, 19 ° 09 ’ S 44 ° 49 ’ E, 50 m, 11. XII. 1996, fr., Jongkind 3505 (MO, P, TAN, WAG); Antsalova, Bemaraha, 18 ° 38 ’ S 44 ° 42 ’ E, 100 - 200 m, 5. XII. 1992, fr., Labat et al. 2308 (MO, P, TAN); Mitsinjo, Katsepy, forêt classée d’Antrema, 15 ° 42 ’ 37 ’’ S 46 ° 10 ’ 25 ’’ E, 15 m, 12. XI. 2005, fr., Labat 3560 (G, MO, P, TAN); 33 km Ankirika-Analalava, 14 ° 49 ’ S 47 ° 41 ’ E, 100 m, 16. XI. 1996, fl., Leeuwenberg et al. 14728 (MO, P, TAN, WAG); NW Madagascar, 1841, Pervillé 93 (P [5 sheets]); Ambongo, 1841, Pervillé 590 (P [3 sheets]); Katsepy, station d’Antrema, [15 ° 42 ’ S 46 ° 10 ’ E], 11. XI. 2009, fl., Ramanamisata 157 (P); Port-Berger, Tsingia, Marosely, 15 ° 38 ’ 58 ’’ S 47 ° 35 ’ 03 ’’ E, 217 m, 16. XI. 2004, fl., Ramananjanahary et al. 49 (MO, P, TAN); Marovoay, W of Ampijoroa reserve, garden A, [16 ° 18 ’ S 46 ° 48 ’ E], 11. XI. 1996, ster., A. Randrianasolo 528 (G, MO, P, TAN); forêt sèche de Marosely, 17 ° 38 ’ 58 ’’ S 47 ° 35 ’ 03 ’’ E, 217 m, 17. XI. 2004, y. fr., Razakamalala et al. 1711 (G, K, MO, P, TEF); Port-Bergé, forêt d’Ambohimanga, 6. XII. 2004, 15 ° 57 ’ 05 ’’ S 47 ° 26 ’ 09 ’’ E, 232 m, fr., Razakamalala et al. 1831 (G, K, MO, NY, P, TEF); forêt d’Ankarafantsika, aux env. d’Ampijoroa, [16 ° 18 ’ S 46 ° 48 ’ E], 19 - 21. XI. 1957, fl., Service Forestier 18470 (MO, P [2 sheets], TEF); Analalava, env. d’Antsangabe, [15 ° 01 ’ S 47 ° 16 ’ E], 4. XI. 1958, fr., Service Forestier 18872 (G, MO, P [2 sheets], TEF); Ambongo, près de Benetsy, [15 ° 04 ’ S 47 ° 14 ’ E], 21. XI. 1965, Service Forestier 24266 (P, TEF); Ampijoroa station forestière, fonkontany Ampombolava, [16 ° 18 ’ S 46 ° 49 ’ E], 15. XII. 1991, fr., Service Forestier 34333 (MO, TEF); ibid. loco, 200 m, 23. I. 1992, fr., Service Forestier 34358 (MO, TEF).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF9FFE1FE4383651ABA0E73.taxon	materials_examined	Typus: MADAGASCAR. Prov. Toliara: Rég. Anosy, Vinanibe, W de Fort-Dauphin, [25 ° 03 ’ S 46 ° 56 ’ E], 7. XII. 1968, fr., Service Forestier 28606 (holo-: P [P 06774807]!; iso-: G [G 00341712]!, K!, MO- 5056712!, NY!, P [P 00580305, P 00580306]!, TEF [TEF 000883]!)	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF9FFE1FE4383651ABA0E73.taxon	diagnosis	Diagnosis Abrahamia delphinensis Randrian. & Lowry is easily distinguished from its congeners by its semicraspedodromous secondary veins, its thin petiole, and its fruits densely covered with short indument.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF9FFE1FE4383651ABA0E73.taxon	description	Description Trees 5 - 12 m tall, 20 - 21 cm DBH, bark with white latex; tips of young twigs with short, appressed indument. Leaves opposite or sub-opposite; blade obovate, 2.8 - 5.8 - (6) 3 1 - 3 cm, chartaceous, apex retuse, margins entire, base broadly cuneate, adaxial surface glabrous, abaxial surface sparsely strigose on midvein, venation semicraspedodromous, midvein prominent abaxially, secondary veins 18 - 25 pairs, 2 - 3 mm apart, raised and prominent abaxially, slightly raised adaxially, tertiary veins visible only on abaxial surface; petiole 7 - 15 mm, c. 0.5 - 1 mm in diam., usually glabrous but sometimes with very sparse short, appressed indument. Inflorescence axillary or terminal, a panicle, 1.5 - 5.5 cm long, pubescent. Male and female flowers small, 4 - merous, c. 2 mm long; calyx lobes 0.5 3 1 mm, triangular, adaxial surface glabrous, abaxial surface pubescent; corolla lobes c. 2 3 1 - 1.2 mm, ovate, adaxial surface glabrous, abaxial surface pubescent. Male flowers with filaments 1 - 1.2 mm long, pubescent or sometimes glabrous, anthers c. 0.5 - 0.8 mm, subglobose, glabrous; disk slightly cupuliform, c. 1.5 mm in diam., glabrous; ovary completely absent. Female flowers with vestigial stamens, filaments c. 1 mm long, pubescent, anthers very small, c. 0.2 mm, sterile; disk glabrous, cupuliform, c. 1.5 mm in diam.; ovary c. 0.6 - 0.8 3 1 mm, depressed ovoid, completely covered with indument; style very short, inserted apically, partly covered with indument, stigmas capitate. Fruits 2.3 - 2.5 3 1.2 - 1.4 cm, ellipsoid, densely covered with short, appressed and slightly tangled indument, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF9FFE1FE4383651ABA0E73.taxon	distribution	Distribution, ecology and phenology Abrahamia delphinensis occurs in low elevation humid forest, including littoral forest on sand, and is restricted to the area around Fort Dauphin (Map 2). It flowers between September and November, and fruits between October and December.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF9FFE1FE4383651ABA0E73.taxon	conservation	Conservation status With an EOO of 1442 km 2, a minimum AOO of 48 km 2, and 12 known localities representing a total of 7 locations, 3 of which are situated within protected areas (Andohahela, Mandena, and Sainte Luce) while the others are variously subjected mining as well as fire and land clearing leading to a projected decline in all 5 sub-criteria, A. delphinensis is assigned a preliminary conservation status of “ Vulnerable ” [VU B 1 ab (i, ii, iii, iv, v) + 2 ab (i, ii, iii, iv, v)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF9FFE1FE4383651ABA0E73.taxon	discussion	Notes Abrahamia delphinensis most closely resembles A. lenticellata but can be distinguished by its retuse (vs short acuminate to acuminate) leaf apex and the complete absence (vs presence) of a rudimentary ovary in male flowers.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF9FFE1FE4383651ABA0E73.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Toliara: Andohahela PN, NW of Tolagnaro, 24 ° 45 ’ S 46 ° 51 ’ E, 250 - 500 m, 17 - 20. X. 1992, fl., Malcomber et al. 1678 (BM, CAS, G, K, MO [2 sheets], NY, P, TAN, US, WAG); N of Fort-Dauphin, near Ste. Luce, 24 ° 47 ’ S 47 ° 10 ’ E, 10 m, 21. X. 1989, y. fr., McPherson 14196 (MO, P, TAN, TEF); Mandena, botanical garden, 24 ° 57 ’ S 47 ° 00 ’ E, 25 m, 25. X. 1989, fl., McPherson et al. 14244 (MO, P, TAN, TEF); Ste. Luce, N de Mahatalaky, 0 - 10 m, 28. IV. 1989, ster., Rabevohitra 1959 (MO, P, TAN); Andohahela PN, Isaka-Ivondro, 24 ° 47 ’ S 46 ° 50 ’ E, 0 - 500 m, 10 - 27. IX. 1993, fl., Randriamampionona 631 (A, BM, BR, BRIT, CAS, F, G, K, MO [2 sheets], NY, P [2 sheets], PRE, S, TAN, US, W, WAG, Z); Andohahela PN, Andranolava, 24 ° 41 ’ S 46 ° 45 ’ E, 28. VIII. 1996, ster., Randriamampionona 1338 (MO, P, TAN); Mandena forestry station, 24 ° 57 ’ S 47 ° 00 ’ E, 0 - 10 m, 18. XI. 1996, ster., A. Randrianasolo 546 (MO, P, TAN); N of Tolagnaro, Ste. Luce area, [24 ° 47 ’ S 49 ° 09 ’ E], 0 - 10 m, 19. XI. 1996, ster., A. Randrianasolo 569 (MO, P, TAN); Bevoay, Androranga, [24 ° 29 ’ 39 ’’ S 47 ° 05 ’ 39 ’’ E], 171 m, 18. XI. 2009, fr., Razakamalala 4680 (G, MO, P, TAN); Ifarantsa, Ivorona, 24 ° 29 ’ 09 ’’ S 47 ° 04 ’ 43 ’’ E, 35 m, 10. XII. 2010, fr., Razakamalala 5033 (G, K, MO, P, TAN); Mandena, [24 ° 57 ’ S 47 ° 00 ’ E], 16. XI. 1948, fl., Réserves Naturelles 1348 (G, MO, P); ibid. loco, 18. IX. 1950, Service Forestier 1545 (P, TEF); Marofanenitra, 25. IX. 1951, Service Forestier 4043 (P, TEF); Mandena, [17 ° 30 ’ S 48 ° 38 ’ E], 29. X. 1953, fl., Service Forestier 7826 (P, TEF); ibid. loco, 3. IX. 1953, fl., Service Forestier 8170 (MO, P, TEF); Col du Maningotry, 300 - 400 m, 27. IX. 1953, fl., Service Forestier 8510 (G, MO, P, TEF); Mandena, 15. XII. 1954, fr., Service Forestier 15622 (MO, P, TEF).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF6FFE8FE4383651DD00A86.taxon	materials_examined	Typus: MADAGASCAR. Prov. Toliara: Rég. Anosy, Vinanibe, à l’W de Fort-Dauphin, [25 ° 03 ’ S 46 ° 56 ’ E], 7. XII. 1968, fr., Service Forestier 28611 (P [P 00722641]!; iso-: G [G 00341714]!, K!, MO- 6684134!, P [P 00722642]!, TEF [TEF 000884]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF6FFE8FE4383651DD00A86.taxon	diagnosis	Diagnosis Abrahamia ellipticarpa Randrian. & Lowry can be distinguished from other member of the genus by its large, glabrous, ellipsoid fruits measuring 3 - 3.5 3 1.5 - 1.8 cm.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF6FFE8FE4383651DD00A86.taxon	description	Description Trees c. 10 - 12 tall, bark with white latex; young twigs with appressed indument. Leaves alternate, subopposite or opposite; blade oblanceolate to narrowly elliptic, 3 - 9.5 3 1.3 - 3 cm, subcoriaceous, apex acute or sometimes retuse or rarely emarginate, margin entire, base cuneate, adaxial surface glabrous, abaxial surface with appressed indument, more so along the midvein and margin, venation craspedodromous, midvein prominent abaxially, secondary veins 18 - 30 pairs, more or less parallel, 2 - 3 mm apart, slightly raised adaxially and prominent abaxially, tertiary veins very few, impressed and barely visible; petiole 5 - 10 mm long, c. 1 mm in diam., canaliculate on adaxial surface, pubescent. Inflorescences and flowers unknown. Fruits 3 - 3.5 3 1.5 - 1.8 cm, ellipsoid, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF6FFE8FE4383651DD00A86.taxon	distribution	Distribution, ecology and phenology Abrahamia ellipticarpa has been recorded from sublittoral forest on sand at several sites in the Fort-Dauphin area, including Mandena, Vinanibe, and Petriky (Map 1). It has been collected in flower in September and in fruit in December.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF6FFE8FE4383651DD00A86.taxon	conservation	Conservation status With an EOO of 28 km 2, an AOO of 12 km 2, and three known locations, 2 of which are situated within protected areas (Mandena and Petriky) whereas the third one is highly degraded and subjected to continued degradation from uncontrolled harvesting and possibly also fire leading to a projected decline in all 5 sub-criteria, A. ellipticarpa is assigned a preliminary conservation status of “ Endangered’ ’ [EN B 1 ab (i, ii, iii, iv, v) + 2 ab (i, ii, iii, iv, v)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF6FFE8FE4383651DD00A86.taxon	discussion	Notes Abrahamia ellipticarpa is vegetatively similar to A. oblongifolia but can be distinguished by its fewer secondary veins (18 - 30 vs 35 - 40), impressed and barely visible (vs slightly raised and very visible) tertiary venation, and large fruits (3 - 3.5 3 1.5 - 1.8 vs 2 3 1 cm) with evident (vs obscure) longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF6FFE8FE4383651DD00A86.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Toliara: Petriky, N shore of Lake Andranany, 25 ° 03 ’ S 46 ° 53 ’ E, 0 - 10 m, 14. IV. 1989, ster., Gereau 3376 (MO, P, TAN); env. de Fort-Dauphin, entre le Pic St. Louis et la mer, 1 - 25 m, 20. IX. - 6. X. 1928, fl., Humbert 5984 (G, P [2 sheets]); Man-dena forestry station, 24 ° 57 ’ S 47 ° 00 ’ E, 0 - 10 m, 20. XI. 1996, ster., A. Randrianasolo 578 (MO, P, TAN); ibid. loco, 3. IX. 1953, bud, Service Forestier 8171 (MO, P, TEF); ibid. loco, 12. XII. 1952, ster., Service Forestier 207 - R- 16 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF4FF95FE4383651A760EE3.taxon	materials_examined	Typus: MADAGASCAR. Prov. Toamasina: Rég. Analanjirofo, Maroantsetra, Nosy Mangabe, 15 ° 30 ’ S 49 ° 46 ’ E, 0 - 330 m, 28. IX- 5. X. 1989, fl., Schatz 2770 (holo-: MO- 3759750!; iso-: K, P [P 06169108]!, TAN [TAN 001988]!, WAG [WAG 1435428] image seen).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF4FF95FE4383651A760EE3.taxon	diagnosis	Diagnosis Abrahamia elongata Randrian. & Lowry can be distinguished from its congeners by its oblanceolate or narrowly elliptic leaves that are 8.5 - 26.5 3 2.5 - 6.5 cm, combined with very pronounced secondary and tertiary veins, long petiole (1.5 - 3 cm), and a relatively large and glabrous fruits (2 - 4 3 1 - 1.5 cm).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF4FF95FE4383651A760EE3.taxon	description	Description Trees 4 - 25 m tall, 10 - 15 cm DBH, bark with white milky latex; young twigs waxy or lanate to pubescent at the tips. Leaves alternate; blades oblanceolate to narrowly elliptic, 8.5 - 26.5 3 2.5 - 6.5 cm, coriaceous, apex emarginate, rounded, shortly acuminate or sometimes acuminate, margin entire, base acute to cuneate, adaxial surface glabrous or with short appressed indument, abaxial surface sericeous to glabrescent, venation craspe-dodromous, midvein prominent abaxially, secondary veins 25 - 50 pairs, more or less par-allel, 1.5 - 10 mm apart, very prominent abaxially, tertiary veins raised on both surfaces; petiole 15 - 30 mm long, canaliculate on adaxial surface, with short, appressed indument when young, later glabrescent or glabrous. Inflorescence axillary, a panicle, 1.5 - 6 cm long, the branches regular, pubescent or tomentose. Male and female flowers 5 - merous; pedicel 0.1 - 0.3 mm long, pubescent; calyx lobes shallowly deltate to shallowly triangular, c. 1 3 1.2 - 1.5 mm, adaxial surface glabrous, abaxial surface pubescent, slightly imbricate in male flowers, valvate in female flowers, persistent in fruit; corolla lobes ovate, 2.5 - 3 3 1.5 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent, white or pale yellow; stamens 5. Male flowers with filaments 1.5 - 2 mm long, glabrous, white, anthers 1 mm long, obloid to slightly ovoid, yellow, glabrous; disk c. 1 mm in diam., glabrous; ovary rudimentary. Female flowers with filaments c. 1 mm long, glabrous, white, anthers 0.3 - 0.5 mm long, ovoid, sterile, yellow, glabrous; disk annular, cupuliform, c. 1.5 - 2 mm in diam., glabrous; ovary very widely ovoid or obovoid, slightly asymmetrical, 1.5 - 2 3 1.5 - 1.8 mm, glabrous, placentation subapical; style unbranched, latero-subapically inserted, c. 1 mm long, stigmas capitate, glabrous. Fruits 2 - 4 3 1 - 1.5 cm, ellipsoid or slightly asymmetrical, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF4FF95FE4383651A760EE3.taxon	vernacular_names	Vernacular names “ Ditimena’ ’, “ Hazombarorana’ ’, “ Tarantana’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF4FF95FE4383651A760EE3.taxon	distribution	Distribution, ecology and phenology Abrahamia elongata occurs from eastern coastal littoral forest on sand to premontane forests on laterite up to elevations of 900 m, and extends from Marojejy and near Ambi-lobe in the north to Manombo in the south (Map 2). It flowers between September and November, and fruits in November and December.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF4FF95FE4383651A760EE3.taxon	conservation	Conservation status With an EOO of 147,171 km 2, an AOO of 120 km 2, and 32 recorded localities, many of which occur within protected areas (Ambalabe, Anjanaharibe-Sud, Betampona, Manombo, Marojejy, Masoala, Nosy Mangabe, and Zahamena), A. elongata is assigned a preliminary conservation assessment of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF4FF95FE4383651A760EE3.taxon	discussion	Notes Abrahamia elongata most closely resembles A. latifolia, with which it shares large leaves with many secondary veins and exclusively axillary inflorescences. However, A. elon-gata differs by its coriaceous (vs subcoriaceous) leaves with craspedodromous (vs semi-craspedodromous) venation, and its longer petiole (15 - 30 vs 10 mm). It also resembles A. sambiranensis vegetatively, but differs by having leaves with evident, raised tertiary veins on both surfaces (vs lacking in A. sambiranensis) and whose apex is flat in pressed material (vs folded), and by its fruits that are glabrous (vs pubescent when young and glabrescent at maturity in A. sambiranensis).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFF4FF95FE4383651A760EE3.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Antsiranana: Sambava, forêt de Tsihomanaomby, 14 ° 06 ’ 08 ’’ S 50 ° 02 ’ 50 ’’ E, 103 m, 15. II. 2006, ster., Andrianjafy 1582 (MO, P, TAN); Marojejy PN, campement 1, 14 ° 26 ’ 13 ’’ S 49 ° 46 ’ 32 ’’ E, 460 m, 12. X. 1996, y. fr., Rakotomalaza et al. 752 (MO, P, TAN); Andapa, Anjanaharibe-Sud RS, [14 ° 36 ’ S 49 ° 27 ’ E], 3 - 7. XII. 1996, fr., Ravelonarivo et al. 1052 (MO, P, TAN); Antalaha, forêt Ampohafana, [14 ° 48 ’ 30 ’’ S 49 ° 51 ’ 30 ’’ E], 9. IX. 1955, fl., Service Forestier 15064 (P [2 sheets], TEF); Andapa, Andilandrano, [14 ° 45 ’ S 49 ° 34 ’ E], 12. XII. 1964, fr., Service Forestier 25019 (G, MO, NY, P, TEF); S de Sambava, [14 ° 18 ’ S 50 ° 11 ’ E], 30. XI- 1. XII. 1966, fr., Service Forestier 27110 (P, TEF); Vohémar, Fanambana, 22. XII. 1954, ster., Service Forestier 54 - R- 151 (P). Prov. Fianarantsoa: forêt de Manombo, S de Farafangana, [23 ° 02 ’ 10 ’’ S 47 ° 44 ’ 50 ’’ E], 17. X. 1964, fl., Service Forestier 23654 (P, TEF); forêt entre Manakara et Vohipeno (PK 30), [22 ° 22 ’ S 47 ° 51 ’ E], 21. X. 1964, y. fr., Service Forestier 23696 (P, TEF). Prov. Toamasina: Manompana, forêt d’AntanambaoAmbodimanga, 16 ° 45 ’ 38 ’’ S 49 ° 42 ’ 22 ’’ E, 7 m, 5. X. 2003, fr., Andrianarivelo 30 (G, MO, P, TEF); Maroantsetra, Ambatoledama forest, 15 ° 17 ’ S 50 ° 00 ’ E, VI. 2003, y. fr., Antilahimena 1996 (G, K, MO, NY, P, TAN); Rantabe, Behovana, 15 ° 47 ’ 48 ’’ S 49 ° 32 ’ 10 ’’ E, 815 m, 31. X. 2003, fr., Antilahimena 2251 (G, K, MO, P, TAN); Antsirabe, Sahatany, Anjiakely, 15 ° 24 ’ 19 ’’ S 49 ° 30 ’ 28 ’’ E, 360 m, 27. VI. 2004, fr., Antilahimena 2579 (G, K, MO, P, TAN); Nosy Mangabe, 15 ° 29 ’ S 49 ° 45 ’ E, 10 m, 18 - 26. IV. 1990, ster., Carlson 126 (MO); ibid. loco, Carlson 150 (MO); ibid. loco, Carlson 188 (MO); Sahalampy (Onibe), [17 ° 50 ’ S 48 ° 25 ’ E], 700 m, XI. 1938, fl., Cours 1160 (MO, P); Mananara, 1. IX. 1920, fl., Decary 59 (P, TAN); Soanierana-Ivongo, forêt d’Antanambao-Ambodimanga, 16 ° 45 ’ 40 ’’ S 49 ° 42 ’ 35 ’’ E, s. d., fr., Rabehivohitra 4354 (MO, P, TAN, TEF); Sainte Marie, forêt d’Ambohidena, 16 ° 51 ’ 11 ’’ S 49 ° 57 ’ 10 ’’ E, 10 m, 14. V. 2003, ster., Rabenantoandro 1480 J (MO); Fénérive-Est, Tampolo station forestière, 17 ° 17 ’ S 49 ° 23 ’ E, 10 m, 11. X. 1999, ster., A. Randrianasolo 596 (MO); Vatomandry, Ambalabe, Ambinanindrano II, 19 ° 09 ’ 38 ’’ S 48 ° 35 ’ 02 ’’ E, 600 m, 9. X. 2005, fl., A. Randrianasolo et al. 1045 (MO, NY, P, TAN); ibid. loco, 19 ° 09 ’ 01 ’’ S 48 ° 34 ’ 55 ’’ E, 604 m, 6. X. 2009, fl., A. Randrianasolo 1182 (G, K, MO, NY, P, TAN); ibid. loco, 19 ° 09 ’ 39 ’’ S 48 ° 36 ’’ E, 568 m, 7. X. 2005, ster., Razanatsima et al. 38 (MO, P, TAN); Farankaraina, s. d., fl., Réserves Naturelles 65 (P); Zahamena NP, Imerimandroso, [17 ° 30 ’ S 48 ° 38 ’ E], 16. IX. 1951, fl., Réserves Naturelles 3171 (P, TAN); Marojejy PN, [14 ° 18 ’ S 49 ° 33 ’ E], 15. IX. 1956, fl., Réserves Naturelles 8265 (G, MO, P); Nosy Mangabe, 15 ° 30 ’ S 49 ° 46 ’ E, 0 - 330 m, 28. IX- 5. X. 1989, fl., Schatz 2757 (MO, P, TAN); Soanierana-Ivongo, [16 ° 54 ’ 30 ’’ S 49 ° 35 ’ 00 ’’ E], 27. XII. 1949, y. fr., Service Forestier 2361 (P, TAN, TEF); Farankaraina station forestière, [15 ° 25 ’ S 49 ° 52 ’ E], 26. IX. 1952, fl., Service Forestier 6154 (G, MO, P [3 sheets], TEF); Betampona RNI, près d’Ambodiriana, [17 ° 55 ’ S 49 ° 13 ’ E], 6. XI. 1953, fl., Service Forestier 8600 (MO, P, TEF); massif de l’Ambohitsitondroina de Mahalevona (NW de la presqu’île Masoala), [15 ° 25 ’ S 49 ° 58 ’ E], c. 500 m, 30. XI. 1953, fr., Service Forestier 8678 (G, MO, NY, P, TEF); Brickaville, Sahamamy, Anivorano, [18 ° 33 ’ S 48 ° 58 ’ E], s. d., ster., Service Forestier 10884 (P, TEF); Farankaraina station forestière, [15 ° 25 ’ S 49 ° 52 ’ E], 11. X. 1955, fl. & y. fr., Service Forestier 15361 (MO, P [3 sheets], TEF); PK 154, quelques km S de Soanierana-Ivongo, [17 ° 00 ’ S 49 ° 32 ’ E], 2. IX. 1957, fl., Service Forestier 18182 (P [2 sheets], TEF); Baie d’Antongil, NW de l’embouchure de la Rantabe, [15 ° 40 ’ 30 ’’ S 49 ° 38 ’ 30 ’’ E], 50 - 150 m, 12. IX. 1957, fl., Service Forestier 18262 (MO, TEF); Farankaraina, [15 ° 25 ’ S 49 ° 52 ’ E], 28. IX. 1963, fl., Service Forestier 21392 (G, MO, P [2 sheets], TEF); Tampolo, [17 ° 17 ’ 00 ’’ S 49 ° 23 ’ 30 ’’ E], 26. XI. 1962, fr., Service Forestier 22121 (G, K, MO, NY, P [3 sheets], TEF, WAG); Tamatave, Tanambao-Tamboro, 17. XII. 1954, ster., Service Forestier 9 - R- 195 (P); Lohan’i Sahatenany, N de Beanana, 2. II. 1952, ster., Service Forestier 29 - R- 139 (P); Maroantsetra, Farankaraina, [15 ° 25 ’ S 49 ° 52 ’ E], 13. VIII. 1952, ster., Service Forestier 44 - R- 139 (P); ibid. loco, 27. XII. 1952, ster., Service Forestier 127 - R- 199 (P); Tampolo, Fénérive Est, 13. I. 1951, ster., Service Forestier 16 - R- 7 (P, TEF); ibid. loco, Service Forestier 323 - R- 107 (P, TEF); Masoala Peninsula, S of Ambanizana, 15 ° 40 ’ 24 ’’ S 49 ° 57 ’ 51 ’’ E, 110 - 260 m, 25. X. 1994, ster., Vasey et al. 55 (MO, P, TAN).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF89FF93FE4383651DF20CB9.taxon	materials_examined	Typus: MADAGASCAR. Prov. Toliara: Morondava, s. d., fl., Grevé s. n. (holo-: P [P 00722643]!; iso-: P [P 00722644]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF89FF93FE4383651DF20CB9.taxon	description	Description Trees 4 - 20 m tall, 20 - 45 cm DBH, bark with milky white latex; young twigs pubescent, with lenticels. Leaves alternate, rarely subopposite; blades oblanceolate or narrowly oblong, 3 - 16 3 1.5 - 4 cm, subcoriaceous, apex emarginate, retuse or rounded, margin entire, base acute or cuneate, adaxial surface glaucous, glabrous or sometimes with sericeous indument, especially on and alongside the midvein, abaxial surface with appressed sericeous indument, more so on and alongside the midvein, venation craspedodromous, midvein prominent abaxially, secondary veins 35 - 40 (- 47) pairs, parallel, 0.5 - 5 mm apart, tertiary veins barely visible on both surfaces; petiole 5 - 15 mm long, canaliculate on adaxial surface, pubescent or glabrescent. Inflorescence axillary or terminal, a panicle, 1.5 - 16 cm long, pubescent or villous. Male and female flowers 5 - merous; pedicel 0 - 0.5 mm long, pubescent; calyx lobes deltate, 1 - 1.2 3 c. 1 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent, persistent on fruit; corolla lobes ovate, 2 - 2.5 3 1 - 1.5 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent, white to light yellow or yellow green; stamens 5. Male flowers with filaments 1 - 1.2 mm long, glabrous, white, anthers 0.5 mm long, ovoid to subglobose, yellow, glabrous; disk c. 1 mm in diam., glabrous; ovary rudimentary. Female flowers with filaments 1 - 1.3 mm long, glabrous, white, anthers 0.5 mm long, subglobose or ovoid, sterile, yellow, glabrous; disk 1 - 1.5 mm in diam., glabrous; ovary depressed ovoid, slightly asymmetrical, c. 1 3 1 - 1.2 mm, glabrous, placentation subapical; style shortly trifid, inserted apically, c. 1 mm long, glabrous. Fruits 2 - 2.8 3 1.1 - 1.7 cm, ellipsoid, slightly asymmetrical, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF89FF93FE4383651DF20CB9.taxon	vernacular_names	Vernacular names “ Ditimena’ ’, “ Lakalaka’ ’, “ Manavodrevo’ ’, “ Sandramiramy’ ’, “ Soahihy’ ’, “ Sohihy’ ’, “ Sohy’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF89FF93FE4383651DF20CB9.taxon	distribution	Distribution, ecology and phenology Abrahamia grandidieri occurs in western and southern Madagascar (Map 4), at elevations ranging from sea level to 1,500 m, and is very often found near rivers or streams. It flowers from August to November and fruits from November through January.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF89FF93FE4383651DF20CB9.taxon	conservation	Conservation status With an EOO of 412,300 km 2, a minimum AOO of 272 km 2, and 70 known localities, including subpopulations in 6 protected areas (Analavelona, Andohahela, Andringitra, Bemarivo, Namoroka, and Pic d’Ivohibe), A. grandidieri is assigned a preliminary conservation status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF89FF93FE4383651DF20CB9.taxon	discussion	Notes Abrahamia grandidieri most closely resembles A. sericea, with which it shares leaves with parallel and very closely spaced secondary veins, but can easily be distinguished by its narrower leaves that are glaucous (vs broader and not glaucous), its flowers with a glabrous (vs pubescent) disk and ovary, and its glabrous (vs velutinous) fruits. While the protologue of Protorhus heckelii indicates that it is based on “ Perrier de la Bâthie, no. 3, in Herb. Mus ”, it is clear that the authors prepared their description using the specimen of Perrier de la Bâthie 859 that bears the original label written in Perrier’s hand [P 06775232], along with another label added later that indicates “ No. 3 = 859 Perrier de la Bâthie ”. Curiously, this specimen also bears a note added some time later by Perrier that reads “ Ces spécimens proviennent d’une autre localité que le no. 859 (les bords de l’Iubohago près de l’Ankirihitra (Boina) ”. A second specimen in the Paris herbarium [P 06775233], comprising a single leaf and a small part of an inflorescence in a fragment packet on which someone (clearly not Perrier) has indicated “ Protorhus (au sp. nov.). Madagascar. M. Perrier de la Bâthie, envoyé par Mr. Heckel au Lab. Colonial, fin 1906 ”, which appears to have been the basis for the choice of the epithet used by Dubard and Dop. This second specimen does not have an original label, but it does bear a note in Perrier’s hand, dated October 1947, that reads “ Perrier no. 859! Type (?) de Protorhus Heckelei Dub. & Dop, récolté sur les bords de la Boina Kely, petit affluent de droite de la Betsiboka, à l’est de Mevatenana (Boina) ”. It is not clear whether Dubard and Dop examined this specimen, and because according to Perrier de la Bâthie it is not part of the same gathering as the holotype, we have refrained from regarding it as either an isotype or a syntype.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF89FF93FE4383651DF20CB9.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Antsiranana: Antsiranana, bord de la rivière Betsieka, I. 1938, ster., Humbert 19093 ter (P); Ambanja, Beampongy, [13 ° 40 ’ 30 ’’ S 48 ° 27 ’ 00 ’’ E], 4. X. 1953, Service Forestier 7715 (P, TEF); Befandriana-Nord, Marosakana, le long de la rivière Tsinjomolava, [14 ° 56 ’ S 47 ° 40 ’ E], 11. IX. 1956, fl., Service Forestier 16094 (P, TEF). Prov. Fianarantsoa: Ihosy, entre Ankaramena et Zazafotsy, [22 ° 12 ’ S 46 ° 22 ’ E], 9. X. 1966, fl., Boiteau 339 (G, MO, P); ibid. loco, Boiteau 341 A-C (P); Ambalavao, entre Vohitromby et Vohitrosy, [21 ° 50 ’ S 46 ° 56 ’ E], 8. XII. 1974, fr., Cremers 3547 (P [2 sheets], TAN); entre Ihosy et Fianarantsoa, 9. VIII. 1938, ster., Grandidier s. n. (P); env. d’Ihosy, [22 ° 23 ’ 00 ’’ S 46 ° 07 ’ 30 ’’ E], 800 - 900 m, III. 1934, ster., Humbert 14452 ter (P); Ambohimahasoa, Ikalamavony, [21 ° 10 ’ 00 ’’ S 46 ° 35 ’ 30 ’’ E], 1955, ster., Humbert 30255 bis (P); RN Andringitra, Vohitsaoka, 17. IX. 1951, fl. & bud, Réserves Naturelles 3112 (P [2 sheets], TAN); ibid. loco, 8. XI. 1952, bud, Réserves Naturelles 4802 (P [2 sheets], TAN); ibid. loco, 27. IX. 1957, fr., Réserves Naturelles 9300 (P, TEF); Ivohibe, [22 ° 31 ’ 30 ’’ S 46 ° 59 ’ 00 ’’ E], 8. XI. 1950, Service Forestier 1449 (P, TAN, TEF); forêt de Mangidy, [21 ° 15 ’ 30 ’’ S 46 ° 47 ’ 30 ’’ E], 15. X. 1951, fl., Service Forestier 4726 (P, TAN, TEF); Fianarantsoa, forêt Isahafa, [21 ° 27 ’ 00 ’’ S 47 ° 04 ’ 30 ’’ E], 27. IX. 1951, fl., Service Forestier 4762 (P, TEF); Maintinandrina, [21 ° 15 ’ 30 ’’ S 46 ° 47 ’ 30 ’’ E], 9. IX. 1952, fl., Service Forestier 5958 (P, TEF); Ihosy, [22 ° 23 ’ 00 ’’ S 46 ° 07 ’ 30 ’’ E], 30. X. 1953, bud, Service Forestier 7615 (P, TEF). Prov. Mahajanga: bord de ruisseau, env. de Bekodoka, [16 ° 58 ’ E 45 ° 06 ’ E], 10. IX. 1930, fl., Decary 8303 (G, P, US); Bekodoka, [16 ° 58 ’ S 45 ° 06 ’ E], 19. IX. 1930, fl., Decary 8096 (P); Ambanja, Forêt d’Andranomatavy, forêt dense humide, bord de rivière, 13 ° 40 ’ 29 ’’ S 47 ° 59 ’ 12 ’’ E, 123 m, 28. IX. 2013, fl., Tahinarivony 746 (G, MO, P, TAN); Befandriana Nord, 14. IX. 1942, fl., Herb. Jard. Bot. Tananarive 5253 (P); Ambongo et Boeny, IX. 1904, Perrier de la Bâthie 859 (P); Antajiandava, Soalala, [16 ° 33 ’ S 45 ° 49 ’ E], 2. VIII. 1951, bud, Service Forestier 3990 (P, TEF); Soalala, bassin moyen du Bemarivo, [17 ° 34 ’ S 45 ° 43 ’ E], IX. 1907, fl., Perrier de la Bâthie 4519 (P [2 sheets], TAN); Bassin du Bemarivo, [16 ° 56 ’ S 44 ° 24 ’ E], c. 80 m, IX. 1907, fl., Perrier de la Bâthie 5863 (P [3 sheets]); ibid. loco, Perrier de la Bâthie 5865 (P); plateau d’Ankara, [17 ° 13 ’ S 46 ° 16 ’ E], VIII. 1922, bud, Perrier de la Bâthie 14825 (P [2 sheets]); env. de Mandritsara, [15 ° 50 ’ 00 ’’ S 48 ° 49 ’ 30 ’’ E], 600 m, IX. 1922, fr., Perrier de la Bâthie 15047 (P); Tsaratanana, forêt de Koakala, 16 ° 48 ’ 09 ’’ S 47 ° 28 ’ 58 ’’ E, 385 m, 7. XII. 2003, fr., A. Randrianaivo et al. 994 (CNARP, MO, P, TAN); Namoroka RN, Andranomavo, [16 ° 36 ’ S 45 ° 34 ’ E], 10. IV. 1952, ster., Réserves Naturelles 3888 (P [2 sheets]); ibid. loco, 8. IX. 1956, fl., Réserves Naturelles 8139 (= Service Forestier 17172) (P, TEF); ibid. loco, 17. XI. 1960, fr., Réserves Naturelles 11437 (P); Maevatanana, Ankaboka, Ambalajanakomby, [17 ° 30 ’ 50 ’’ S 46 ° 04 ’ 30 ’’ E], 8. I. 1955, fr., Service Forestier 13034 (P, TEF); Morafenobe, Beravina, [18 ° 10 ’ 30 ’’ S 45 ° 13 ’ 30 ’’ E], 19. VIII. 1956, fl., Service Forestier 16351 (P, TEF); Befandriana Nord, Canton Ambararata, [16 ° 36 ’ S 46 ° 25 ’ E], 16. I. 1959, fr., Service Forestier 19403 (P, TEF). Prov. Toliara: PN Andohahela, 24 ° 56 ’ 23 ’’ S 46 ° 38 ’ 40 ’’ E, 120 m, 9 - 12. V. 1997, fr., Birkinshaw et al. 421 (MO, P, TAN); Rivière de l’Onilahy, E de Maromala, s. d., y. fr., Geay 6053 (P); Betroka, Ampandrandava, 8. IV. 1942, y. fr., Herb. Jard. Bot. Tananarive 4971 (P); Bekily, Ampandrandava, IX-X. 1943, bud & fl., Herb. Jard. Bot. Tananarive 5793 (P); Andohahela PN, Talakimangery, 24 ° 51 ’ 47 ’’ S 46 ° 42 ’ 34 ’’ E, 250 - 500 m, 10. X. 2000, fl., Hoffmann et al. 172 (G, K, P); Betioky, vallée de l’Onilahy, env. de Tongobory, [23 ° 32 ’ S 44 ° 20 ’ E], 100 - 200 m, 1 - 8. X. 1924, Humbert 2699 (G, P); Amboasary, vallée de la Menarahaka, [22 ° 33 ’ S 46 ° 30 ’ E], 700 - 800 m, 31. X. 1924, fl. & fr., Humbert 3046 (P); près de Bevilany, [25 ° 00 ’ 30 ’’ S 46 ° 36 ’ E], 200 - 300 m, 14. IX. 1928, fl., Humbert & Swingle 5712 (P [2 sheets], TAN); haute vallée du Mandrare, [24 ° 39 ’ S 46 ° 26 ’ E], 8 - 15. XI. 1928, ster., Humbert 6487 bis (P); Tsivory à Anadabolava, [24 ° 12 ’ 40 ’’ S 46 ° 19 ’ 00 ’’ E], 300 - 400 m, XII. 1933, fr., Humbert 12287 (P [2 sheets]); ibid. loco, 200 m, Humbert 12367 (P [2 sheets]); vallée de la Manambolo, env. d’Esomony, [24 ° 32 ’ S 46 ° 41 ’ E], 400 - 900 m, XII. 1933, fr., Humbert 12882 (G, P); vallée de l’Onilahy, en aval de Tongobory, [23 ° 31 ’ 30 ’’ S 44 ° 19 ’ 30 ’’ E], 0 - 300 m, 8. XI. 1960, ster., Leandri 3759 (P); Beroroha, Makay massif, 21 ° 13 ’ 02 ’’ S 45 ° 19 ’ 08 ’’ E, 480 m, 22. XI. 2010, fr., Phillipson et al. 6188 (G, K, MO, NY, P, TAN); PN Andohahela, 22. IX. 1994, fl., Rakotomalaza 273 (G, K, MO, NY, P, TAN); PN Andohahela, 25 ° 01 ’ S 46 ° 38 ’ E, 15. V. 2001, fr., Randriamampionona 1013 (G, K, MO, NY, P, TAN, WAG); ibid. loco, 24 ° 50 ’ 20 ’’ S 46 ° 32 ’ 17 ’’ E, 6. IV. 1996, ster., Randriamampionona 1241 (MO, TAN); PN Andohahela, Eso-mony, coté W de la réserve, 24 ° 31 ’ S 46 ° 37 E, 20. IX. 1991, fl., Randriamanantena & Durbin 58 (MO, P, TAN); Ambatoabo, forêt d’Ankoba, 24 ° 47 ’ 28 ’’ S 46 ° 42 ’ 01 ’’ E, 205 m, 24. XI. 2008, fr., Randrianaivo 1716 (G, K, MO, P, TAN); Sakaraha, Mahaboboka, Marotsiraka Betsileo, 22 ° 44 ’ 20 ’’ S 44 ° 13 ’ 24 ’’ E, 594 m, 21. III. 2011, fr., Randrianarivony et al. 294 (G, MO, P, TAN); Ranobe, Behalomboro, 24 ° 18 ’ 28 ’’ S 46 ° 11 ’ 18 ’’ E, 210 m, 1. XI. 2007, fr., Randriatsivery 295 (MO, P, TAN, TEF); Ampasinampoana, Enato, 24 ° 53 ’ 05 ’’ S 46 ° 58 ’ 52 ’’ E, 425 m, 16. XII. 2009, fr., Razakamalala et al. 5102 (G, MO, P, TAN); Betorabato, forêt d’Androtsy, 21 ° 05 ’ 17 ’’ S 45 ° 05 ’ 39 ’’ E, 235 m, 11. I. 2011, fr., Razakamalala 5920 (MO, P, TAN); PN Andohahela, Behara, [24 ° 48 ’ S 46 ° 32 ’ E], 4. X. 1951, fl., Réserves Naturelles 3407 (MO, P [2 sheets], TAN); Andohahela PN, Canton Enanihila, [24 ° 31 ’ S 46 ° 37 ’ E], 2. VIII. 1952, y. fr., Réserves Naturelles 3958 (P [2 sheets], TAN); PN Andohahela, Behara, [24 ° 53 ’ S 46 ° 35 ’ E], 25. XI. 1953, fr., Réserves Naturelles 5956 (G, P [2 sheets], TEF); Imanombo, Behara, [24 ° 26 ’ 00 ’’ S 45 ° 49 ’ 30 ’’ E], 16. IX. 1950, fl. & y. fr., Service Forestier 1518 (P, TAN, TEF); Anapaby, [23 ° 00 ’ S 44 ° 36 ’ E], 1. X. 1951, fl., Service Forestier 3862 (P, TEF); Morondava, Dabara, [20 ° 24 ’ S 44 ° 47 ’ E], 12. VIII. 1951, bud, Service Forestier 3915 (P); Ranopiso, [25 ° 03 ’ 00 ’’ S 46 ° 41 ’ 30 ’’ E], 10. IX. 1951, y. fr., Service Forestier 3967 (P, TEF); Betroka, Merarano, [20 ° 30 ’ 30 ’’ S 45 ° 00 ’ 30 ’’ E], 16. IX. 1952, bud, Service Forestier 6337 (P [2 sheets], TEF); Belo sur Tsiribihina, Tsaramandroso, [19 ° 54 ’ 30 ’’ S 45 ° 08 ’ 00 ’’ E], 1. VII. 1953, fr., Service Forestier 7797 (MO, P, TEF); Bekily, [24 ° 13 ’ 20 ’’ S 45 ° 19 ’ E], 14. I. 1954, fr., Service Forestier 8392 (P, TEF); vallée de l’Onilahy, en aval de Tongobory, [23 ° 31 ’ 30 ’’ S 44 ° 19 ’ 30 ’’ E], 17. IX. 1953, bud & fl., Service Forestier 8462 (P, TEF); Betroka, Benonoka-I, [23 ° 33 ’ 30 ’’ S 46 ° 16 ’ 30 ’’ E], 10. IV. 1954, y. fr., Service Forestier 10199 (P, TEF); Sakaraha, Sohara, [24 ° 08 ’ S 45 ° 28 ’ E], 26. V. 1954, y. fr., Service Forestier 10293 (MO, P, TEF); forêt de Befitroa, [20 ° 00 ’ 30 ’’ S 45 ° 32 ’ E], 20. IX. 1954, fl., Service Forestier 10876 (P [2 sheets], TEF); km 455, route Ft. Dauphin, [25 ° 02 ’ 00 ’’ S 46 ° 39 ’ 30 ’’ E], 9. IX. 1954, y. fr., Service Forestier 10907 (MO, P, TEF); Sakaraha, Bekily, [24 ° 08 ’ S 45 ° 28 ’ E], 25. XI. 1954, fr., Service Forestier 11158 (P, TEF); Miandrivazo, Ankilimadinika, [19 ° 31 ’ S 45 ° 28 ’ E], 23. X. 1954, ster., Service Forestier 11185 (P, TEF); Sakatà, Betroka, [22 ° 53 ’ 30 ’’ S 45 ° 57 ’ 30 ’’ E], 17. IX. 1954, fl., Service Forestier 12124 (P, TEF); Manambolo, [24 ° 34 ’ S 46 ° 36 ’ E], 5. XI. 1954, y. fr., Service Forestier 12199 (P, TEF); Sakamarakely, [22 ° 11 ’ S 45 ° 10 ’ E], 14. XI. 1954, fr., Service Forestier 12363 (P); ibid. loco, Service Forestier 12364 (P, TEF); Vineta, [23 ° 03 ’ 30 ’’ S 44 ° 15 ’ 30 ’’ E], 20. XII. 1955, fr., Service Forestier 15594 (P, TEF); Morondava, Canton Befasy, Tananadava, [20 ° 28 ’ S 44 ° 08 ’ E], 4. X. 1956, fl., Service Forestier 16556 (P, TEF); Behara, Amboasary, [24 ° 53 ’ S 46 ° 35 ’ E], 3. X. 1956, bud, Service Forestier 17038 [= Réserves Naturelles 7920] (MO, P, TEF); Mahabo, Canton Ankilizato, Migodo, [20 ° 24 ’ S 45 ° 03 ’ E], 4. XII. 1957, fr., Service Forestier 17869 (P, TEF); Bekily, Ankaranabo-Sud, [24 ° 13 ’ S 45 ° 19 ’ E], 9. IX. 1963, bud, Service Forestier 21370 (P, TEF); Bords des ruisseaux, versant W du massif de l’Analavelona, W de Mitia, [22 ° 37 ’ S 44 ° 07 ’ E], 15. XII. 1962, fr., Service Forestier 22203 (G, MO, P [2 sheets], TEF); Tsihombe, Manindro, bord rivière Manambovo, [25 ° 19 ’ S 45 ° 29 ’ E], 27. IV. 1967, y. fr., Service Forestier 26502 (MO, P, TEF); Ampanihy, Bekitro, Tranoroa, [24 ° 42 ’ S 45 ° 04 ’ E], 13. IX. 1968, bud & fl., Service Forestier 26745 (P, TEF); Tanambao Befasy, [20 ° 40 ’ 31 ’’ S 44 ° 25 ’ E], 1. IX. 1954, ster., Service Forestier 64 - R- 19 (P, TEF); Ampandrandava, [24 ° 05 ’ S 45 ° 42 ’ E], 700 - 800 m, IX. 1942, fl., Seyrig 1 (P [2 sheets]). Sine loco: s. d., Baron 9796 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF8DFF9DFE4383651CBB0EF5.taxon	materials_examined	Lectotypus (designated here): MADAGASCAR. Prov. Antsiranana: Rég. Diana, collines et plateaux calcaires de l’Ankarana, forêt tropophile, [12 ° 54 ’ S 49 ° 08 ’ E], XII. 1937 - I. 1938, fl., Humbert 18793 (P [P 00722645]!; isolecto-: P [P 00722646]!). Syntypi: MADAGAS-CAR. Prov. Antsiranana: Ankarana, [12 ° 54 ’ S 49 ° 08 ’ E], 300 - 350 m, XII. 1937 - I. 1938, fr., Humbert 18963 (P [P 00722648]!); ibid. loco, Humbert 18967 (P [P 00722647]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF8DFF9DFE4383651CBB0EF5.taxon	description	Description Trees 3 - 20 m tall, 5 - 13 cm DBH, bark with translucid latex; young twigs glabrous to pubescent towards the tips, with lenticels, waxy. Leaves opposite, deciduous; blades widely elliptic to elliptic or circular to obovate, 3 - 9 3 1.7 - 6 cm, subcoriaceous, apex emarginate, retuse, margin entire, base obtuse and oblique, adaxial surface glabrous or waxy, abaxial surface glabrous or sometimes with very scarce short indument, venation craspedodromous, midvein prominent abaxially, secondary veins 10 - 18 pairs, 2 - 6 mm apart, prominent abaxially, tertiary veins visible on both surfaces; petiole 7 - 15 mm long, canaliculate on adaxial surface, pubescent and waxy. Inflorescence axillary and terminal, a thyrse, 1.5 - 3 cm long, branches flattened, pubescent. Male and female flowers 5 - merous; pedicel c. 0.2 - 0.3 mm long, glabrous; calyx lobes widely depressed ovate, 0.5 - 0.7 3 1 mm, imbricate, glabrous on both surfaces, persistent on fruit; corolla lobes ovate, 2.5 3 1.5 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent; stamens 5 or 6. Male flowers with filaments 1 - 1.5 mm long, glabrous, anthers 0.5 - 1 mm long, obloid to slightly ovoid, yellow, glabrous; disk cupuliform and crenulate, c. 1 mm in diam., glabrous; ovary rudimentary. Female flowers with filaments 1.5 mm long, glabrous, anthers 0.7 - 1 mm long, ovoid, sterile, yellow, glabrous; disk c. 1 - 1.5 mm in diam., glabrous; ovary widely to very widely ovoid, 1.5 - 2.5 3 1 - 2 mm, glabrous, placentation parietal or subapical; style trifid, apically inserted, c. 0.5 mm long, glabrous, stigmas capitate. Fruits 2.5 - 3 3 1.5 - 2 cm, ellipsoid, symmetrical or very slightly asymmetrical, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF8DFF9DFE4383651CBB0EF5.taxon	vernacular_names	Vernacular names “ Taimbarika’ ’, “ Mampisaraka’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF8DFF9DFE4383651CBB0EF5.taxon	distribution	Distribution, ecology and phenology Abrahamia humbertii occurs in deciduous and dry forests in northwestern and western Madagascar (Map 2), particularly in several areas with karstic limestone (locally known as ‘ tsingy’), viz. Ankarana, Bemaraha, and Kasinga (Tamponketsa). Flowers have been collected in November and December, and fruits are known from November through March.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF8DFF9DFE4383651CBB0EF5.taxon	conservation	Conservation status With an EOO of 22,183 km 2, a minimum AOO of 32 km 2, and 7 known locations, only 1 of which occurs within a protected area (Ankarana) whereas the others are very likely subjected to ongoing pressure from fire and forest degradation and clearing that is pro-jected to lead to a reduction in quality of habitat, A. humbertii is assigned a preliminary conservation status of “ Vulnerable ” [VU B 2 (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF8DFF9DFE4383651CBB0EF5.taxon	discussion	Notes The most distinctive character of A. humbertii that sets it apart from the other members of the genus is its widely elliptic to circular leaf shape. Among the syntypes of Protorhus humbertii, we have selected Humbert 18793 as the lectotype because it, along with the isolectotype, comprise more complete specimens (albeit in flower) than the other collections cited in the protologue (Humbert 18963 and 18967), both of which are in fruit.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF8DFF9DFE4383651CBB0EF5.taxon	materials_examined	Additional material examined Madagascar. Prov. Antsiranana: Ankarana, env. du Campement des Anglais, 12 ° 55 ’ 57 ’’ S 49 ° 04 ’ 20 ’’ E, 180 - 225 m, 19. XI. 1996, fr., Andrianantoanina et al. 1018 (G, MO, P, TAN); Ankarana, zone du lac Vert, 2. XI. 1990, fl., Bardot-Vaucoulon 197 (P); Ankarana, camp des Anglais, 10. XI. 1990, fl., Bardot-Vaucoulon 270 (P); ibid. loco, 25. XI. 1990, fr., Bardot-Vaucoulon 338 (P); ibid. loco, 12 ° 54 ’ 42 ’’ S 49 ° 6 ’ 42 ’’ E, 180 m, 15. X. 1997, fr., Bardot-Vaucoulon 762 (P); ibid. loco, Bardot-Vaucoulon 763 (MO, P, TAN); Ankarana, XII. 1991, fl., Foury s. n. (P); Sakaramy, Montagne des Français, [12 ° 26 ’ S 49 ° 16 ’ E], 8. XI. 1944, y. fr., Homolle 307 (P); Ankarana, near campement des Anglais, 12 ° 54 ’ S 49 ° 08 ’ E, 150 m, 29. I. 1994, fr., Leeuwenberg 14345 (MO, P, TAN, WAG); Andranovondronina, Anjiabe, forêt de Belamoty, 12 ° 06 ’ 09 ’’ S 49 ° 19 ’ 34 ’’ E, 85 m, fr., Ratovoson 1154 (CNARP, MO, P, TAN); Ankara (= Ankarana), Jardin Botanique n ° 8, [12 ° 49 ’ S 49 ° 01 ’ E], 28. XI. 1952, fr., Service Forestier 6185 (P [2 sheets], TEF); S de Marotaolana (Anivorano-Nord), [12 ° 49 ’ 30 ’’ S 49 ° 15 ’ 30 ’’ E], 8. III. 1964, fr., Service Forestier 23366 (P). Prov. Mahajanga: Antsalova, forêt de Tsiampihy et forêts cotières près de Besaraha, de Bemiha et de Soahanina, 0 - 20 m, 23. XII. 1952, fl., Leandri 2240 (MO, P); Antsalova, env. de Besara, [18 ° 40 ’ S 44 ° 15 ’ E], 23. XII. 1952, fl., Service Forestier 6863 (G, MO, P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF80FF9EFE4383651DA90DE9.taxon	materials_examined	Lectotypus (designated here): Madagascar. Prov. Antananarivo: Mont Ibity, quartzites, [20 ° 07 ’ S 47 ° 01 ’ E], c. 2000 m, IX. 1921, fl., Perrier de la Bâthie 13976 (P [P 00722649]!; isolecto-: P [P 00722650, P 00722651]!). Syntypi: MADAGASCAR. Prov. Antananarivo: Ibity, [20 ° 07 ’ S 47 ° 01 ’ E], 1900 - 2000 m, s. d., ster., Perrier de la Bâthie 5840 (P [P 06169079]!); ibid. loco, c. 2000 m, III. 1921, fr., Perrier de la Bâthie 13571 (P [P 00722653, P 00722654]!); ibid. loco, Perrier de la Bâthie 13571 bis (P [P 00722652]!). Sine loco: s. d., Decary 13085 “ A ” (P [P 06169081]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF80FF9EFE4383651DA90DE9.taxon	description	Description Shrubs 0.25 - 2 m tall, bark with white or translucid latex; young twigs glabrescent, with lenticels, waxy. Leaves alternate or subopposite; blades oblong to elliptic, 3 - 7 3 1.2 - 3 cm, subcoriaceous, apex retuse or emarginate, margin entire and slightly revolute, base cuneate to obtuse, adaxial surface glabrous, sometimes glossy, abaxial surface with scattered, appressed indument, more so along the midvein, venation craspedodromous, midvein prominent abaxially, secondary veins 15 - 30 pairs, visible, parallel, 0.5 - 2 mm apart, very slightly raised on both surface, tertiary veins impressed and very rarely visible on both surfaces; petiole 5 - 7 mm long, shallowly canaliculate on adaxial surface, pubescent when young, glabrescent. Inflorescence axillary or terminal, a thyrse, 1.5 - 3 cm long, pubescent. Male and female flowers 5 - merous; pedicel c. 0.3 - 0.5 mm long, pubescent; calyx lobes ovate to deltate, 1.5 - 1.8 3 1.5 - 2 mm, imbricate, glabrous on both surfaces, margin sparesly ciliate; corolla lobes ovate, 2.5 - 3 3 1.5 - 2 mm, imbricate, glabrous, white; stamens 5. Male flowers with filaments 1 - 1.3 mm long, glabrous on both surfaces, white, anthers c. 1 mm long, ovoid or obloid, sterile, yellow, glabrous; disk dish-shaped, c. 2 mm in diam., glabrous; ovary rudimentary. Female flowers with filaments 1 - 1.5 mm long, glabrous, white, anthers 0.5 mm long, ovoid, yellow, glabrous; disk c. 1.5 - 2 mm in diam., glabrous; ovary very widely ovoid, 1 3 1 mm, glabrous, placentation parietal; style apical, shortly trifid, 0.5 mm long, glabrous, stigmas capitate. Fruits 1.5 - 1.8 3 1.5 cm, widely depressed-ovate, glabrous, with longitudinal striations, pyrenes with a few peripheral resiniferous canals, cotyledons separable.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF80FF9EFE4383651DA90DE9.taxon	distribution	Distribution, ecology and phenology Abrahamia ibityensis is only known from Mt. Ibity and its surroundings, to the SW of the town of Antsirabe (Map 2), at elevations between 1500 and 2000 m, where it grows mostly on rocky soil in areas that are frequently subjected to fire. Flowers have been recorded in September and October, and fruiting material has been collected in February and March.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF80FF9EFE4383651DA90DE9.taxon	conservation	Conservation status With an EOO of 34 km 2, a minimum AOO of 28 km 2, and 9 known localities representing 7 locations, most of which are situated within a newly established protected area (Ibity) but are still subjected to fire and possible degradation leading to a projected decline in all quality of habitat, A. ibityensis is assigned a provisional conservation status of “ Vulnerable’ ’ [VU B 1 ab (iii) + 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF80FF9EFE4383651DA90DE9.taxon	discussion	Notes Abrahamia ibityensis can be distinguished from all of its congeners by its oblong to elliptic leaves with inconspicuous tertiary venation and the presence of a few peripheral resiniferous canals in the seeds, which results in the two cotyledons being distinct and easily separable. Among the syntypes of Protorhus ibityensis, we have selected Perrier de la Bâthie 13976 as the lectotype because it, along with the two isolectotypes, comprise more complete specimens than the other collections cited in the protologue (Decary 13085 “ A ”, Perrier de la Bâthie 5840, 13571, and 13571 bis [incorrectly cited in the protologue as 1357 bis]). We have added the letter “ A ” to the Decary specimen because this same number has been applied to a specimen of Beilschmiedia microphylla (Kosterm.) Kosterm. (Lauraceae) that bears an original label, whereas the label accompanying the syntype of Protorhus ibityensis was clearly prepared by someone else.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF80FF9EFE4383651DA90DE9.taxon	materials_examined	Additional material examined Madagascar. Prov. Antananarivo: Antsirabe, Ibity (S), W of Ambohimanjaka, [20 ° 15 ’ S 47 ° 03 ’ E], 2000 m, 22. XII. 1985, y. fr., Dorr & Rakotozafy 4528 (MO, NY, P, TAN); Ibity, 13. X. 1970, fl., Keraudren-Aymonin & Aymonin 24565 (P); Ibity, Ambero, S end of Amboropotsy, 20 ° 10 ’ 17 ’’ S 47 ° 02 ’ 07 ’’ E, 1700 - 1800 m, 23. II. 2003, fr. & fl., Koopman 170 (MO, P, TAN); Ibity, 27 km S of Antsirabe, 20 ° 05 ’ S 47 ° 10 ’ E, 1500 - 2240 m, 13. X. 1993, fl., Lewis et al. 741 (G, K, MO, NY, P, TAN); Ibity, 23 km E de la cimenterie, 20 ° 04 ’ 40 ’’ S 47 ° 00 ’ 10 ’’ E, 1830 m, 3. X. 1996, fl., Randrianaivo et al. 6 (MO, P, TAN); ibid. loco, Randrianaivo et al. 9 (MO, P, TAN); Ibity, E of Alatsinainy Ibity village and the cement factory, 20 ° 06 ’ S 46 ° 59 ’ E, 1600 m, 2. X. 1996, fl., A. Randrianasolo 485 (G, K, MO, P, TAN); ibid. loco, A. Randrianasolo 486 (MO, P, TAN); Ibity, Falirano, 20 ° 04 ’ 24 ’’ S 47 ° 00 ’ 10 ’’ E, 1470 m, 20. I. 2003, fr., Rogers et al. 82 (G, MO, P, TAN); abords du sommet de l’Analabe, dans la chaine d’Ampandrianombilapa, S de l’Ibity, [20 ° 05 ’ 43 ’’ S 47 ° 00 ’ 14 ’’ E], 1900 m, 19. XI. 1967, fr., Service Forestier 27394 (P, TEF). Prov. Fianarantsoa: Ambatofinanadrahana, Itremo, E. du Mt. Ambatoandrano, 20 ° 05 ’ 25 ’’ S 47 ° 00 ’ 07 ’’ E, 1693 m, 1. XI. 2001, fl., Andriamihajarivo 49 (MO, P, TAN).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF86FF98FE4383651A510F53.taxon	materials_examined	Typus: MADAGASCAR. Prov. Fianarantsoa: Itremo, W of Itremo village, slope NE of the waterfall, 20 ° 35 ’ S 46 ° 36 ’ E, 1450 m, 6. X. 1996, fl., A. Randrianasolo 491 (holo-: MO- 4895266!; iso-: P [P 06169070]!, TAN [TAN 001990]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF86FF98FE4383651A510F53.taxon	diagnosis	Diagnosis Abrahamia itremoensis Randrian. & Lowry differs from the species it most closely resembles, A. buxifolia, by its leaves that are pubescent (vs glabrous) and the presence of indument on the disk in male flowers (vs glabrous).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF86FF98FE4383651A510F53.taxon	description	Description Shrubs or small trees 0.3 - 4 m tall. Leaves opposite or subopposite; blades oblanceolate, 2 - 6 3 0.6 - 2 cm, coriaceous, apex rounded, sometimes retuse or slightly emarginate, margin entire, base cuneate to slightly obtuse, adaxial surface puberulous, abaxial surface pubescent, venation craspedodromous, midvein prominent abaxially, secondary veins 18 - 26 pairs, parallel, 0.5 - 1 mm apart, prominent on both surfaces, tertiary veins weakly visible on upper surface, completely invisible on lower surface, petiole short, 1 - 3 mm long, canaliculate on adaxial surface, pubescent. Inflorescence axillary or terminal, a panicle, short, c. 1 - 2.5 cm, pubescent; bracts triangular to deltate, c. 1 - 2 3 0.5 - 0.8 mm. Male flowers small, 5 - merous, 2 - 4 mm long; pedicel very short, c. 0.1 - 0.2 mm, pubescent; calyx lobes triangular to deltate, c. 1 - 1.5 3 1 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent; corolla lobes ovate to oblong, c. 2 - 4 3 1 - 2 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent; stamens 5, filaments c. 1 mm long, pubescent, anthers c. 1 3 0.3 - 0.4 mm, oblong to slightly ovoid, glabrous; disk c. 1 mm in diam., cupuliform, pubescent; ovary rudimentary. Female flowers unknown. Fruits 1.5 - 2.3 3 1.2 - 1.5 cm, obovoid to ellipsoid, velutinous.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF86FF98FE4383651A510F53.taxon	distribution	Distribution, ecology and phenology Abrahamia itremoensis occurs mainly at Itremo and in the surrounding area (Map 5), at elevations between 1000 and 1800 m. It grows on rocky soil in areas that are regularly subjected to fire. Flowering material has been collected in October and specimens with fruit have been gathered between January and April.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF86FF98FE4383651A510F53.taxon	conservation	Conservation status With an EOO of 154 km 2, a minimum AOO of 32 km 2, and 6 known locations, most of which are situated within a newly established protected area (Itremo) but are still subjected to fire and possible degradation leading to a projected decline in quality of habitat, A. itremoensis is assigned a preliminary conservation status of “ Vulnerable’ ’ [VU B 1 ab (iii) + 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF86FF98FE4383651A510F53.taxon	discussion	Notes Abrahamia itremoensis can be confused with A. minutifolia due to its small, coriaceous leaves with craspedodromous venation, and it velutinous fruits, but it clearly differs by its oblanceolate (vs obovate) leaf blades, its paniculate (vs thyrsiform) inflorescence, and its larger fruits (1.5 - 2.3 3 1.2 - 1.5 vs 1 - 1.5 3 0.5 - 0.8 cm). One of the collections assigned to this species, Decary 13041, was cited in the protologue of Protorhus buxifolia H. Perrier and therefore represents a syntype of that name.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF86FF98FE4383651A510F53.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Fianarantsoa: Ambatofinandrahana, Itremo, SW d’Antsirakambiaty, 20 ° 36 ’ 34 ’’ S 46 ° 33 ’ 30 ’’ E, 1688 m, 10. XII. 2003, fr., Andriamihajarivo 251 (G, MO, P, TAN); Itremo, Andaobatofotsivava, 20 ° 37 ’ 54 ’’ S 46 ° 35 ’ 53 ’’ E, 1387 m, 21. XII. 2003, fr., Antilahimena et al. 2465 (MO, P, TAN); col d’Itremo, 2 km from Vodiharana, 3. VIII. 1984, ster., Baum 47 (MO, TAN); Itremo, [20 ° 32 ’ S 46 ° 33 ’ E], 1700 m, IX. 1956, bud, Bosser 10074 (TAN [2 sheets]); ibid. loco, X. 1963, bud, Bosser 18107 (MO, P, TAN); env. d’Ambatofinandrahana, [20 ° 33 ’ S 46 ° 48 ’ E], 1600 - 1800 m, 17. II. 1938, fr., Decary 13041 (P); W d’Itremo, [20 ° 36 ’ S 46 ° 39 ’ E], 1500 - 1700 m, 17 - 22. I. & 18 - 22. IV. 1955, fr., Humbert 28151 (MO, P); Itremo, 20 ° 34 ’ 40 ’’ S 46 ° 34 ’ 53 ’’ E, 1550 m, 29. X. 1997, fl., Malcomber et al. 2841 (G, MO, P, TAN); forêt d’Ianasana, 7 km W d’Itremo, 20 ° 36 ’ S 46 ° 34 ’ E, 1630 m, 5. II. 1999, fr., Messmer et al. 794 (G, MO, P, TAN); Itremo, [20 ° 32 ’ S 46 ° 33 ’ E], X. 1964, fl., Morat 905 (P, TAN); Ambatofinandrahana, [20 ° 33 ’ S 46 ° 48 ’ E], 15. X [no year given], bud, Rauh 813 (TAN); Mt. Ambatomenaloha, W d’Itremo, [20 ° 32 ’ S 46 ° 33 ’ E], 20. I. 1955, fr., Service Forestier 11576 (P, TEF); env. du col d’Itremo (W d’Ambatofinandrahana), [20 ° 32 ’ S 46 ° 33 ’ E], 1600 - 1700 m, 27. XI. 1969, y. fr., Service Forestier 28897 (P, TEF); Ambatofinandrahana, massif d’Itremo, 20 ° 37 ’ S 46 ° 35 ’ E, 1500 m, 27. X. 1994, fl., van der Werff 13563 (G, MO, P, TAN).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF84FF9AFE4383651B6A0F99.taxon	materials_examined	Lectotypus (designated here): MADAGASCAR: sine loco, s. d., fl., Thouars s. n. (P [P 00722655]!; isolecto-: P [P 00722656, P 00722657]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF84FF9AFE4383651B6A0F99.taxon	description	Description Trees 6 - 16 m tall, 30 cm DBH, bark with translucid, white or reddish milky latex; young twigs usually with white corky exoderm, glabrous, with lenticels. Leaves alternate; blades oblanceolate, 11 - 37 3 3.5 - 9 cm, subcoriaceous, apex shortly acuminate, retuse or rounded, margin entire, base acute or cuneate, adaxial surface glabrous, abaxial surface glabrous, venation craspedodromous, midvein prominent abaxially, secondary veins 40 - 60 pairs, parallel, 1 - 4 mm apart, prominent abaxially, tertiary veins visible on abaxial surface, slightly less so on adaxial surface; petiole 10 mm long, shallowly canaliculate on adaxial surface, glabrous or glabrescent. Inflorescence axillary, a panicle, 2.5 - 8 cm long, branches flattened or slightly triangular, pubescent or glabrescent. Male and female flowers 5 - merous; pedicel c. 0.5 mm long, pubescent; calyx lobes deltate, c. 0.8 3 1 mm, valvate, adaxial surface glabrous, abaxial surface pubescent, persistent on fruit; corolla lobes ovate, 3 3 1.5 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent, pale yellow; stamens 5. Male flowers with filaments 1.5 mm long, glabrous, white, anthers 0.5 - 0.7 mm long, subglobose, yellow, glabrous; disk, c. 1 - 1.5 mm in diam., cupuliform, glabrous; ovary rudimentary. Female flowers with filaments 0.7 - 0.8 mm long, anthers 0.5 mm long, ovoid, sterile, yellow, glabrous; disk c. 1.5 mm in diam., glabrous; ovary depressed ovoid, c. 1 3 1 - 1.5 mm, glabrous, placentation subapical; style shortly trifid, apicaly inserted, c. 1 mm long, glabrous, stigmas capitate. Fruits 2.5 3 1.7 cm, ellipsoid, surface smooth, glabrous.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF84FF9AFE4383651B6A0F99.taxon	vernacular_names	Vernacular names “ Hazombarora’ ’, “ Hazombarora beravina’ ’, “ Hazombarorana’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF84FF9AFE4383651B6A0F99.taxon	distribution	Distribution, ecology and phenology Abrahamia latifolia occurs at low- to mid-altitude sites (at elevations from sea level to 780 m) on sandy or lateritic soils in eastern coastal and sublittoral humid forests (Map 5). It has been recorded in flower in October and November and in fruit in December.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF84FF9AFE4383651B6A0F99.taxon	conservation	Conservation status With an EOO of 15,645 km 2, a minimum AOO of 44 km 2, but a maximum AOO that is estimated to exceed 2,000 km 2, and 11 known localities representing 9 locations, only 2 of which are in protected areas (Makira and Mananara) while the remainder are subjected to land clearing leading to a projected decline in quality of habitat, A. latifolia is assigned a preliminary conservation status of “ Vulnerable’ ’ [VU B 1 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF84FF9AFE4383651B6A0F99.taxon	discussion	Notes Abrahamia latifolia can be distinguished from all other members of the genus by its particularly large leaves (11 - 37 3 3.5 - 9 cm), which are glabrous on both surfaces. Among the three syntypes collected by du Petit-Thouars, we have selected as the lectotype the sheet that is accompanied by a hand-written description and a line drawing of a flower.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF84FF9AFE4383651B6A0F99.taxon	materials_examined	Additional material examined Madagascar. Prov. Mahajanga: Mandritsara, village le plus proche Lambafotsy, [16 ° 10 ’ S 48 ° 50 ’ E], 26. X. 1955, fl., Service Forestier 15129 (MO, P [3 sheets], TEF). Prov. Toamasina: Antsirabe, Anjiahely, Ankahaminivola forest, 15 ° 23 ’ 57 ’’ S 49 ° 26 ’ 08 ’’ E, 780 m, 15. XII. 2002, fr., Antilahimena 1493 (G, K, MO, P, TAN); Foulpointe, [17 ° 41 ’ S 49 ° 31 ’ E], s. d., Humblot s. n. (P [4 sheets]); Saharamy, [16 ° 24 ’ 30 ’’ S 49 ° 41 ’ 00 ’’ E], 14. X. 1990, fl., Raharimalala 2320 (P); Tampolo forest station, 0 - 3 m, 31. VII. 1996, ster., A. Randrianasolo 470 (G, MO, P, TAN); Mahatsara forest station, 17 ° 38 ’ S 49 ° 29 ’ E, 0 - 5 m, 26. X. 1996, fl., A. Randrianasolo 511 (MO, P, TAN); Sainte Marie, Lonkintsy, forêt d’Ambohidena, 16 ° 46 ’ 23 ’’ S 49 ° 42 ’ 47 ’’ E, 15. V. 2003, ster., Razakamalala et al. 451 B (MO, P, TAN); Maroantsetra, massif de l’Androrona, [15 ° 51 ’ S 49 ° 31 ’ E], 700 m, II. 1954, fr., Service Forestier 8999 (G, MO, P [2 sheets]); Tampolo, Jardin Botanique n ° 21, [17 ° 17 ’ S 49 ° 23 ’ E], 29. XI. 1956, fl., Service Forestier 16469 (MO [2 sheets], P [3 sheets], TEF); ibid. loco, [17 ° 17 ’ 00 ’’ S 49 ° 23 ’ 30 ’’ E], 21. XI. 1957, bud & fl., Service Forestier 17807 (G, MO, P [2 sheets], TEF); ibid. loco, [17 ° 17 ’ S 49 ° 23 ’ E], 16. X. 1958, fl., Service Forestier 19229 (MO, P [3 sheets], TEF); Vavatenina, forêt de Sahavolo, [17 ° 40 ’ 50 ’’ S 49 ° 06 ’ 30 ’’ E], 25. X. 1964, fl., Service Forestier 21919 (P, TEF); forêt de Mangalimaso, W de Foulpointe, [17 ° 41 ’ 30 ’’ S 49 ° 29 ’ 30 ’’ E], 23. XI. 1962, fl., Service Forestier 22103 (MO, P, TEF); Nosy Sainte Marie, forêt de Kalalao, 5. VIII. 1964, ster., Service Forestier 8 - R- 64 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9EFF80FE4383651AE30C4B.taxon	materials_examined	Lectotypus (designated here): MADAGASCAR. Prov. Toamasina: Rég. Atsinanana, Vatomandry, bois littoraux en arrière des lagunes, fl., IX. 1921, Perrier de la Bâthie 14301 (P [P 00722658]!; isolecto-: MO- 6684135!, P [P 00722659]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9EFF80FE4383651AE30C4B.taxon	description	Description Trees 5 - 15 m tall, 5 - 20 cm DBH, bark with white latex; young twigs glabrous, waxy, with lenticels, white or pale brown, spongy, sometimes the very tip with sericeous red brown indument. Leaves alternate, closely spaced, more so toward the branch ends; blades, oblanceolate, rarely widely elliptic, 5.3 - 17.5 3 1.5 - 5 cm, subcoriaceous, apex shortly acuminate, acuminate or rarely rounded, margin entire, base acute or cuneate to obtuse, adaxial surface glabrous, abaxial surface sericeous on midvein, sometimes sparsely so, venation semicraspedodromous, midvein prominent abaxially, secondary veins 20 - 30 pairs, more or less parallel, 2 - 8 mm apart, rather prominent abaxially, tertiary veins often obscure on adaxial surface, fairly visible on abaxial surface; petiole 5 - 8 - (10) mm long, shallowly canaliculate on adaxial surface, red-brown pubescent when young, glabrescent. Inflorescence axillary, a panicle, 1 - 5 cm long, axes regular, red-brown pubescent; bracts triangular, 1 - 2 3 0.5 - 1 mm, abaxial surface red-brown pubescent outside. Male and female flowers 5 - (or 6) - merous; pedicel c. 0.3 - 0.5 mm long, red-brown pubescent; calyx lobes ovate to deltate, 1 - 1.2 3 0.8 - 1.5 mm, imbricate (valvate in female flowers), adaxial surface glabrous, abaxial surface glabrescent or with red brown indument; corolla lobes ovate or oblong, 2 - 3 3 1 - 1.5 mm, imbricate, adaxial surface glabrous, abaxial surface glabrous or with sparse reddish indument, yellow green; stamens 5. Male flowers with filaments 0.5 - 0.7 mm long, glabrous, anthers 0.5 - 0.6 mm long, obloid to slightly ovoid, white, glabrous; disk cupuliform, c. 1 mm in diam., glabrous; ovary completely absent. Female flowers with filaments c. 0.8 mm long, glabrous, anthers c. 0.5 mm long, ovoid to obloid, sterile, yellow, glabrous and pubescent; disk c. 1 - 1.5 mm in diam., glabrous; ovary very widely ovoid, c. 1 3 1 mm, glabrous, placentation subapical (nearly apical); style simple, subapically to apically inserted, 1 mm long, glabrous, stigmas capitate. Fruits c. 1.7 3 1 cm, ellipsoid, symmetrical, glabrous, with weakly visible longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9EFF80FE4383651AE30C4B.taxon	vernacular_names	Vernacular names “ Ditimena’ ’, “ Hazombarorana’ ’, “ Tavolo malamatra’ ’, “ Tsiramiramy mena’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9EFF80FE4383651AE30C4B.taxon	distribution	Distribution, ecology and phenology Abrahamia lecomtei occurs in eastern rainforest from coastal areas to elevations as high as 950 m (Map 4). It has been recorded in flower between September and December, and fruits have been collected in December and January.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9EFF80FE4383651AE30C4B.taxon	conservation	Conservation status With an EOO of 7,413 km 2, a minimum AOO of 32 km 2 but an estimated AOO> 500 km 2, and 7 recorded localities, only some of which are in protected areas (Ambalabe, Betampona, and Zahamena), A. lecomtei is assigned a preliminary conservation status of “ Vulnerable ” [VU B 1 ab (iii) + 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9EFF80FE4383651AE30C4B.taxon	discussion	Notes Abrahamia lecomtei can be distinguished from all other members of the genus by the presence of spongy, pale or white bark on the twigs and small branches, its leaves with semicraspedodromous venation and a short petiole (5 - 8 mm long), and its male flowers with short filaments (not exceeding 0.8 mm) and a disk that lacks a rudimentary ovary. Perrier de la Bâthie 5283 (P) was cited in the protologue of Protorhus lecomtei and thus represents a syntype, but it is clearly referable to Abrahamia thouvenotii.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9EFF80FE4383651AE30C4B.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Toamasina: Zahamena PN, W d’Ampasimaventy, 17 ° 44 ’ 42 ’’ S 48 ° 58 ’ 26 ’’ E, 300 - 350 m, 9. VII. 2003, fl., Andrianjafy 396 (CNARP, MO, P); Ambatondrazaka, Sahalampy, Onibe, [17 ° 45 ’ S 48 ° 51 ’ E], 700 m, XI. 1938, bud, Cours 1171 (P [2 sheets]); RNI Betampona, 17 ° 51 ’ 55 ’’ S 49 ° 12 ’ 15 ’’ E, 275 - 650 m, 19 - 25. VIII. 1990, ster., Carlson 364 (MO, P, TAN); Ambodimangavolo, 3 km W d’Ambatoharanana, 17 ° 34 ’ 17 ’’ S 48 ° 53 ’ 49 ’’ E, 950 m, 30. IV. 2003, fl., Rakotondrajaona 266 (CNARP, G, K, MO, P, TEF); Vatomandry, Ambalabe, forêt de Vohibe, 19 ° 09 ’ 07 ’’ S 48 ° 34 ’ 33 ’’ E, 704 m, 20. I. 2012, fr., Rakotoarivelo 608 (G, K, MO, P, TAN); ibid. loco, 19 ° 09 ’ 42 ’’ S 48 ° 34 ’ 41 ’’ E, 667 m, 20. XI. 2013, y. fr., A. Randrianasolo 1456 (G, K, MO, P, TAN); Zahamena PN, Ihofika, 17 ° 39 ’ 13 ’’ S 48 ° 57 ’ 50 ’’ E, 550 m, 22. X. 2002, fl., S. Randrianasolo 335 (CNARP, MO, P); Ambalabe, forêt de Vohibe, 19 ° 09 ’ 32 ’’ S 48 ° 34 ’ 39 ’’ E, 617 m, 16. X. 2008, fl., Razanatsima 629 (G, K, MO, P, TAN); ibid. loco, 624 m, 1. XII. 2008, y. fr. & fl., Razanatsima & Celestin 635 (MO, P, TAN); Betampona RNI, [17 ° 55 ’ S 49 ° 13 ’ E], 22. XII. 1950, fl., Réserves Naturelles 2656 (MO, P [2 sheets]); Betampona RNI, Betavoana, [17 ° 56 ’ S 49 ° 17 ’ E], 12. XI. 1957, fl., Réserves Naturelles 9156 (P, TEF). Sine loco: 19. XII. 1938, y. fr., Lam & Meeuse 6022 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9CFF8CFE4383641D760F39.taxon	materials_examined	Typus: MADAGASCAR. Prov. Antsiranana: Rég. Analanjorofo, N de Nosiarina (basse Bemarivo), [14 ° 14 ’ 30 ’’ S 50 ° 03 ’ 30 ’’ E], 50 m, 7. XII. 1966, fr., Service Forestier 27167 (holo-: P [P 06775145]; iso-: G [G 00341713]!, MO- 6684136!, P [P 00580350]!, TEF [TEF 000885]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9CFF8CFE4383641D760F39.taxon	diagnosis	Diagnosis Abrahamia lenticellata Randrian. & Lowry most closely resembles A. ditimena but differs by having chartaceous (vs coriaceous) leaves with an acuminate or shortly acuminate apex (vs emarginate or retuse) and semicraspedodromous (vs craspedodromous) secondary veins.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9CFF8CFE4383641D760F39.taxon	description	Description Trees 5 - 20 m tall, 10 - 35 cm DBH, bark with white milky latex; young twigs glabrous or with ferruginous pubescence, with lenticels, waxy. Leaves alternate or subopposite; blades obovate or oblanceolate, 2.5 - 8 (- 14.5) 3 1.2 - 4 cm, chartaceous, apex shortly acuminate or acuminate, rarely rounded, margin sometimes undulate, base cuneate, adaxial surface glabrous, abaxial surface with sparse short, appressed indument, sometimes glabrous, venation semicraspedodromous, midvein prominent abaxially, secondary veins 20 - 25 pairs, 1 - 5 mm apart, slightly prominent abaxially, tertiary veins raised on abaxial surface; petiole 7 - 13 mm long, 0.5 - 1 mm in diam., canaliculate on adaxial surface, glabrous. Inflorescence axillary or terminal, a panicle or sometimes a thyrse, 1 - 4 cm long, axes terete, pubescent. Male and female flowers 4 - merous (rarely 5 - merous), small, c. 2 mm long; pedicel c. 0.5 - 0.8 mm, scabrous; calyx lobes deltate to triangular, c. 1 3 1 - 1.5 mm, valvate, adaxial surface glabrous, abaxial surface pubescent, persistent on fruit; corolla lobes ovate, 2 - 2.5 3 1 - 2 mm, imbricate, adaxial surface glabrous, light yellow or cream, abaxial surface pubescent; stamens 4. Male flowers with filaments 1 - 1.2 mm long, white, glabrous, sometimes with sparse trichomes toward the base, anthers 0.5 mm long, ovoid to subglobose, yellow, glabrous; disk 1 - 1.5 mm in diam., glabrous; ovary rudimentary. Female flowers with filaments c. 0.5 mm long, pubescent, anthers 0.2 - 0.3 mm long, sub-spherical, sterile, brown, glabrous; disk c. 2 mm in diam., slightly cupuliform, glabrous; ovary depressed ovoid, 1 3 1.1 mm, pubescent, slightly asymmetrical, style short, thick, pubescent, c. 0.3 - 0.4 mm long, stigmas capitate. Fruits 2 - 2.6 3 1.3 - 1.5 cm, ellipsoid to ovoid, slightly asymmetrical, surface velutinous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9CFF8CFE4383641D760F39.taxon	vernacular_names	Vernacular names “ Hazombarorana’ ’, “ Hazombarorana à petite feuille’ ’, “ Sandramy mena’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9CFF8CFE4383641D760F39.taxon	distribution	Distribution, ecology and phenology Abrahamia lenticellata occurs in eastern rainforest (Map 6) at elevations from sea level to 1200 m. Material in flower has been collected between August and October, and fruits have been gathered in November and December.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9CFF8CFE4383641D760F39.taxon	conservation	Conservation status With an EOO of 69,556 km 2, a minimum AOO of 104 km 2, and 28 recorded localities, with subpopulations in at least 11 protected areas (Zahamena, Betampona, Ambalabe, Pic d’Ivohibe, Ranomafana, Analamazoatra, Makira, Nosy Mangabe, Masoala, Anjanaharibe-Sud, and Marojejy) that comprise about 22 locations, A. lenticellata is assigned a preliminary conservation status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9CFF8CFE4383641D760F39.taxon	discussion	Notes Material attributable to this species was assigned by PERRIER DE LA BâTHIE (1944, 1946) to an invalidly published variety of Protorhus ditimena (≡ Abrahamia ditimena). Despite an unquestionable similarity to that species, A. lenticellata clearly warrants recognition at the species level, differing from A. ditimena in having leaves with an apex that is almost always acuminate, sometimes shortly so (vs emarginate or retuse) and whose blades are chartaceous (vs coriaceous) in texture.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF9CFF8CFE4383641D760F39.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Antsiranana: Sambava, Marojejy PN, N of Mandena, 14 ° 27 ’ S 49 ° 47 ’ E, 230 - 550 m, 21. X. 1989, fl., Miller 4327 (MO, P, TAN); Sambava, Marojejy PN, Mandena, près du campement 1, WP- 2 - 4, 14 ° 26 ’ 13 ’’ S 49 ° 46 ’ 32 ’’ E, 460 m, 10. X. 1996, fl., Rakotomalaza et al. 728 (G, K, MO, NY, P, TAN); Andapa, Anjanaharibe Sud RS, 14 ° 43 ’ 10 ’’ S 49 ° 27 ’ 12 ’’ E, 1700 m, 14. II. 1995, fr., Ravelonarivo et al. 669 (MO, P, TAN); Doany, Andrinomilolo, 13 km W d’Andranopositra, 14 ° 20 ’ 04 ’’ S 49 ° 17 ’ 50 ’’ E, 856 m, 13. XI. 2006, fr., Ravelonarivo et al. 2088 (MO, P, TAN); Sambava, [Marojejy PN], 24. IX. 1956, fl., Réserves Naturelles 8266 (P, TAN). Prov. Fianarantsoa: Pic d’Ivohibe RS, forêt de Marovitsika, 22 ° 28 ’ 49 ’’ S 46 ° 56 ’ 49 ’’ E, 950 m, fl., s. d., Hoffmann et al. 241 (G, K, P); Ranomafana Park, Parcelle n ° 3, Talatakely, 21 ° 15 ’ S 47 ° 27 ’ E, 800 - 1000 m, 3 - 8. IX. 1993, fl., Kotozafy et al. 230 (MO, P, TAN); Ranomafana PN, c. 15 km W de Tsarata-nana, 21 ° 09 ’ S 47 ° 32 ’ E, 750 m, 8. X. 1994, ster., Kotozafy 703, 724 (MO, P, TAN); Ivohibe, 7.5 km ENE d’Ivohibe, 22 ° 28 ’ 12 ’’ S 46 ° 57 ’ 36 ’’ E, 1000 m, 11. X. 1997, fl., Rakotovao et al. 830 (G, P, MO, TAN, G); Ranomafana PN, piste T 200, 21 ° 16 ’ S 47 ° 25 ’ E, 800 - 900 m, 8. X. 1996, ster., A. Randrianasolo 501 (G, K, MO, P, TAN); Nosy Varika, Besakoana-Ampasinambo, 7. VIII. 1952, ster., Service Forestier 68 - R- 116 (P). Prov. Toamasina: Maroantsetra, Ambinanihely, 15 ° 26 ’ S 49 ° 29 ’ E, 358 m, 20. XI. 2003, y. fr. & fl., Antilahimena 2328 (G, MO, P, TAN); Vinanibe, Andogona, Makira forest, 15 ° 27 ’ 40 ’’ S 49 ° 17 ’ 46 ’’ E, 1153 m, 28. XI. 2003, fr., Antilahimena 2427 (G, K, MO, NY, P, TAN); Toamasina, Betampona RN, 17 ° 51 ’ 55 ’’ S 49 ° 12 ’ 15 ’’ E, 275 - 650 m, 25. VIII. 1990, ster., Carlson 383 (MO, P, TAN); Ambatondrazaka, Manambato à Amboditafonana, [17 ° 27 ’ S 48 ° 46 ’ E], 1200 m, 11. X. 1945, fl., Cours 2826 (MO, P, TAN); Ambatondrazaka, Sahalampy, Onibe, [17 ° 45 ’ S 48 ° 15 ’ E], 700 m, XI. 1938, fl., Cours 1042 (P); Moramanga, Analamazaotra, [18 ° 56 ’ S 48 ° 26 ’ E], 800 - 900 m, VIII. 1925, bud & fl., Louvel 48 (P); near Andasibe, forest of Mantadia, 18 ° 55 ’ S 48 ° 25 ’ E, 900 m, 5. XI. 1994, fl., McPherson et al. 16505 (G, K, MO, P, TAN); Masoala PN, Antalavia, 15 ° 47 ’ S 50 ° 02 ’ E, 22. XI. 1994, ster., Rahajasoa et al. 965 (MO, P, TAN); Amparafaravola, SE d’Ambodisaina, 17 ° 17 ’ 25 ’’ S 48 ° 40 ’ 30 ’’ E, 850 m, 10. XII. 2001, fr., Rakotondrafara et al. 98 (CNARP, MO, P, TAN); Vatomandry, Commune Ambalabe, Tobin’i Foara, 19 ° 09 ’ 28 ’’ S 48 ° 34 ’ 30 ’’ E, 560 m, 4. X. 2005, fl., Ranaivojaona et al. 1197 (G, K, MO, P, TAN); Commune Ambalabe, N of Tsitondroina waterfall, 19 ° 10 ’ 15 ’’ S 48 ° ’ 33 ’ 02 ’’ E, 500 m, s. d., fr., A. Randrianasolo et al. 927 (G, K, MO, NY, P, TAN, WAG); Commune Ambalabe, fokontany Sahanionaka, 19 ° 11 ’ 24 ’’ S 48 ° 34 ’ 30 ’’ E, 288 m, 13. XI. 2010, fr., A. Randrianasolo 1313 (G, K, MO, NY, P, TAN); Commune Ambalabe, Vohibe forest, 19 ° 09 ’ 15 ’’ S 48 ° 34 ’ 44 ’’ E, 629 m, 21. XI. 2013, fr., A. Randrianasolo 1458 (G, K, MO, P, TAN); Périnet, [18 ° 56 ’ S 48 ° 26 ’ E], 800 - 900 m, 23. XII. 1947, y. fr., Réserves Naturelles 1261 (P); Nosy Mangabe, 15 ° 30 ’ S 49 ° 46 ’ E, 0 - 330 m, 28. IX- 5. X. 1989, fl., Schatz et al. 2777 (MO, P, TAN); ibid. loco, 17. XII. 1989, fr., Schatz & Carlson 2848 (MO, NY, P, TAN); Soanieran’Ivongo, [16 ° 54 ’ 30 ’’ S 49 ° 35 ’ 00 ’’ E], 0 - 50 m, 27. XII. 1949, fr., Service Forestier 2370 (P, TAN, TEF); Farankaraina, [15 ° 25 ’ S 49 ° 52 ’ E], 20 - 25 m, 9. X. 1952, bud, Service Forestier 6248 (G, P); massif de l’Ambatotsitondroina de Mahalevona, NW de la presqu’île Masoala, [15 ° 25 ’ S 49 ° 58 ’ E], 600 m, 21. XII. 1953, y. fr., Service Forestier 8697 (MO, P, TEF); forêt orientale, entre Ankovana et le Col d’Ambatondradama, 500 m, 13. XII. 1953, fr., Service Forestier 8750 (MO, P, TEF); Farankaraina, Andranofotsy, [15 ° 25 ’ S 49 ° 52 ’ E], 20 - 25 m, 18. X. 1963, fl., Service Forestier 21483 (G, MO, NY, P, TEF); Vavatenina, sommet Mahatsinjo, [17 ° 27 ’ S 49 ° 00 ’ E], 20. IX. 1966, bud, Service Forestier 26139 (P); Mahatsinjo, Sahatavy, 27. IX. 1966, fl., Service Forestier 26246 (P [2 sheets]); Brickaville, Lohariandava, Ankarangina, 14. VI. 1954, ster., Service Forestier 70 - R- 77 (P); Maroantsetra, Jardin Botanique 17, [Farankaraina], 12. XII. 1952, ster., Service Forestier 99 - R- 199 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF90FF89FE4083651D9E0CF1.taxon	materials_examined	Typus: MADAGASCAR. Prov. Toamasina: Rég. Analanjirofo, Fénérive-Est, Tampolo forestry station, along the “ Grand Layon ”, just 20 m before the first bridge going toward the Indian Ocean, 0 - 5 m, 25. X. 1996, fl., A. Randrianasolo 509 (holo-: MO- 04895278!; iso-: G!, K!, P [P 06775163]!, TAN [TAN 001996]!, WAG!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF90FF89FE4083651D9E0CF1.taxon	diagnosis	Diagnosis Abrahamia littoralis Randrian. & Lowry is easily distinguished from its congeners by its medium sized (4 - 17 3 1.3 - 5.2 cm), oblanceolate leaves bearing scattered, very often obscure tertiary venation, its long petiole (10 - 30 mm), and its axillary inflorescences.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF90FF89FE4083651D9E0CF1.taxon	description	Description Trees 4 - 15 m tall, 12 - 18 cm DBH, bark with white milky or translucid latex; young twigs pubescent or ferruginously scabrous. Leaves alternate; blades oblanceolate, 4 - 17 3 1.3 - 5.2 cm, coriaceous to subcoriaceous, apex very often shortly acuminate, rounded, retuse or emarginate, margin entire, base acute, adaxial surface sericeous when young, later glabrescent, abaxial surface sericeous when young, glabrescent, venation craspedodromous, slightly prominent adaxially, midvein prominent abaxially, secondary veins c. 25 pairs, more or less parallel, 2 - 10 mm apart, prominent abaxially, forming an acute angle with the midrib, tertiary veins widely spaced, often obscure on adaxial surface; petiole 10 - 30 mm long, canaliculate on adaxial surface, pubescent when young, later glabrescent. Inflorescence axillary, a panicle, 2 - 5 cm long, axes, strigose. Male and female flowers 5 - merous; pedicel 0.3 - 0.5 mm long, with short, appressed indument; calyx lobes triangular, 1.5 3 1 mm, imbricate, adaxial surface glabrous, abaxial surface with short appressed indument, persistent on fruit; corolla lobes narrowly ovate, 2 - 3 3 c. 1 mm, imbricate, adaxial surface glabrous, abaxial surface strigose, pale yellow, whiter inside; stamens 5. Male flowers with filaments 1.2 - 1.5 mm long, glabrous, white, anthers 0.8 - 1 mm long, obloid to slightly ovoid, white, glabrous; disk cupuliform, c. 1.5 mm in diam., glabrous; ovary rudimentary. Female flowers with filaments c. 1 mm long, glabrous, white, anthers c. 0.2 - 0.3 mm long, subglobose, sterile, white, glabrous; disk c. 1.5 mm in diam., glabrous; ovary widely ovoid to obovoid, c. 1.5 3 c. 1 mm, glabrous, placentation subapical; style very shortly trifid, apical, c. 1 - 1.1 mm long, glabrous, stigmas capitate. Fruits 2 - 2.5 3 1 - 1.5 cm, ellipsoid, slightly asymmetrical, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF90FF89FE4083651D9E0CF1.taxon	vernacular_names	Vernacular names “ Hazombarorana’ ’, “ Tarantana’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF90FF89FE4083651D9E0CF1.taxon	distribution	Distribution, ecology and phenology Abrahamia littoralis is found in eastern littoral or sublittoral forests near sea level on sandy substrates (Map 7). It flowers in October and very young fruits have been collected in November.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF90FF89FE4083651D9E0CF1.taxon	conservation	Conservation status With an EOO of 18,722 km 2, a minimum AOO of 56 km 2, and 10 recorded localities, with a subpopulation in a single protected area (Agnalazaha), comprising 8 locations subjected to pressure from habitat degradation due to forest exploitation and land clearing, A. littoralis is assigned a preliminary conservation assessment of “ Vulnerable ” [VU B 1 ab (iii) + 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF90FF89FE4083651D9E0CF1.taxon	discussion	Notes With its oblanceolate leaf blade, craspedodromous venation, axillary inflorescence, presence of a rudimentary ovary in male flowers, and glabrous fruit, A. littoralis most closely resembles A. elongata but differs by its much smaller leaves (4 - 17 3 1.3 - 5.2 vs 8.5 - 26.5 3 2.5 - 6.5 cm), fewer secondary veins (25 vs 25 - 50), and obscure (vs prominent) tertiary venation on the abaxial surface.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF90FF89FE4083651D9E0CF1.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Fianarantsoa: Farafangana, Mahabo, forêt littorale d’Apoakafo, 23 ° 11 ’ 02 ’’ S 47 ° 41 ’ 38 ’’ E, 34 m, 19. IX. 2003, fl., Ludovic et al. 435 (MO, P, TAN); ibid. loco, 23 ° 11 ’ 10 ’’ S 47 ° 43 ’ 02 ’’ E, 30 m, 5. VIII. 2004, fl., Ludovic et al. 880 (G, MO, P, TAN); ibid. loco, 23 ° 11 ’ S 47 ° 41 ’ E, 21 m, 29. X. 2004, fr., Ludovic et al. 928 (MO, TAN); Mahabo, Baboaka, forêt d’Andranomainty, 23 ° 10 ’ S 47 ° 44 ’ E, 21 m, s. d., fr., Ludovic & Emilson 1006 (G, K, MO, P, TAN); near Mahabo village, 23 ° 10 ’ 51 ’’ S 47 ° 42 ’ 29 ’’ E, 30 m, 5. XI. 2001, fr., McPherson & Rabenantoandro 18338 (G, K, MO, NY, P, TEF, WAG); Fokontany Mahabo, Firaisana Mahabo-Mananivo, 23 ° 11 ’ 01 ’’ S 47 ° 42 ’ 44 ’’ E, 17 m, 6. XI. 2001, fr., Rabenantoandro et al. 701 (MO, P, TAN, TEF); ibid. loco, 23 ° 11 ’ 01 ’’ S 47 ° 42 ’ 44 ’’ E, 17 m, 6. XI. 2001, y. fr., Rabenantoandro et al. 706 (MO, P, TAN, TEF); ibid. loco, 23 ° 10 ’ 39 ’’ S 47 ° 43 ’ 06 ’’ E, 10 m, 25. IX. 2002, fr., Rabenantoandro et al. 1011 (G, MO, P, TEF); ibid. loco, 23 ° 11 ’ 26 ’’ S 47 ° 43 ’ 17 ’’ E, 11 m, 1. IX. 2007, bud, Rakotoarivelo 198 (MO, P, TAN); ibid. loco, 23 ° 11 ’ 13 ’’ S 47 ° 42 ’ 27 ’’ E, 22 m, 26. VIII. 2003, fl., Razakamalala et al. 708 (G, K, MO, P, TEF); Farafangana, Nosiala, [23 ° 12 ’ 00 ’’ S 47 ° 42 ’ 30 ’’ E], 19. XI. 1955, fr., Service Forestier 16211 (P, TEF). Prov. Toamasina: Fénérive-Est, Station forestière de Tampolo, 17 ° 16 ’ 52 ’’ S 49 ° 24 ’ 44 ’’ E, 0 - 15 m, 24. I. 1995, ster., Raholivelo et al. 150 (MO); ibid. loco, 0 - 5 m, 1. VIII. 1996, ster., A. Randrianasolo 467, 467 a (MO, P, TAN); ibid. loco, 25. X. 1996, y. fr., A. Randrianasolo 510 (MO, P, TAN); Brickaville, Vohibola forest, 18 ° 33 ’ 39 ’’ S 49 ° 14 ’ 33 ’’ E, 27 m, 28. XI. 2002, fl., A. Randrianasolo et al. 737 (G, K, MO, P); Ambila Lemaitso, [18 ° 49 ’ S 49 ° 08 ’ E], 26. X. 1950, y. fr., Service Forestier 1583 (G, MO, P, TAN, TEF); Tampolo, [17 ° 17 ’ 00 ’’ S 49 ° 23 ’ 30 ’’ E], 15. X. 1954, fl., Service Forestier 11071 (MO, P [2 sheets]); ibid. loco, [17 ° 17 ’ 00 ’’ S 49 ° 23 ’ 30 ’’ E], 19. X. 1955, fl., Service Forestier 15187 (MO, P, TEF); ibid. loco, [17 ° 17 ’ S 49 ° 23 ’ E], 28. IX. 1957, fl., Service Forestier 17797 (G, MO, NY, P, TEF); forêt sublittorale de Vohibola, W de Tampina, [18 ° 31 ’ S 49 ° 17 ’ E], 12 - 14. VIII. 1957, y. fr., Service Forestier 18086 (G, MO, P [2 sheets]); forêt sublittorale, entre Foulpointe et l’embouchure de l’Onibe, [17 ° 40 ’ 30 ’’ S 49 ° 29 ’ 30 ’’ E], 29. IX. 1963, fl., Service Forestier 22764 (P). Prov. Toliara: Tolagnaro (Fort-Dauphin), Mandena Forest, along road SW of QIT Fer et Titane work camp, 24 ° 57 ’ S 47 ° 00 ’ E, 0 - 10 m, 19. IV. 1989, ster., Gereau et al. 3393 (MO, P, TAN); Tolagnaro, Ambinanibe, port d’Ehoala, 25 ° 03 ’ S 46 ° 56 ’ E, 2 m, 5. III. 2007, ster., Rabenantoandro et al. 1777 (MO). Sine loco: s. d., fl., Humblot 358 (P); s. d., fl., Louvel 147 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF97FF8BFE4383651D7F0936.taxon	materials_examined	Typus: MADAGASCAR. Prov. Antsiranana: Rég. Diana, RNI Lokobe, 13 ° 24 ’ 40 ’’ S 48 ° 18 ’ 50 ’’ E, 40 - 80 m, 24. XI. 1994, fr., Antilahimena 198 (holo-: MO- 6671433!; iso-: G!, K!, NY!, P [P 06775168]!, TAN [TAN 001991]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF97FF8BFE4383651D7F0936.taxon	diagnosis	Diagnosis Abrahamia lokobensis Randrian. & Lowry can be distinguished from its congeners by its large (14.6 - 23.9 3 4.8 - 7.2 cm), chartaceous to subcoriaceous, oblanceolate leaves and its obovoid fruit borne in a terminal infructescence.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF97FF8BFE4383651D7F0936.taxon	description	Description Trees c. 8 - 25 m tall, 10 - 80 cm DBH, bark with white latex; young twigs lenticellate, glabrous. Leaves opposite to subopposite or alternate; blade oblanceolate, 14.6 - 23.9 3 4.8 - 7.2 cm, chartaceous to subcoriaceous, apex short acuminate, retuse or sometimes emarginate, margin entire, base cuneate, sometimes asymmetric, adaxial surface glabrous, abaxial surface nearly glabrous except sparsely scabrous indument on midvein or around it, venation craspedodromous, midvein prominent abaxially, secondary veins c. 30 - 45 pairs, 2 - 10 mm apart, forming wide angle with the midvein, raised on both surfaces, tertiary veins visible on both surfaces; petiole 13 - 30.5 mm long, 1.5 - 3 mm in diam., canaliculate on adaxial surface, very often pubescent, glabrescent or sometimes glabrous. Inflorescence terminal or pseudoterminal, a panicle, 2.5 3 7 cm, with buds at very early stage of development, dark brown pubescent. Flowers unknown. Fruits 2 - 3.1 3 1 - 1.5 cm, obovoid, pale green, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF97FF8BFE4383651D7F0936.taxon	distribution	Distribution, ecology and phenology Abrahamia lokobensis is only known from northwestern Madagascar; it occurs at elevations between 10 and 100 m at the Lokobe reserve on the island of Nosy-be, and up to 1200 - 1400 m in Tsaratanana national park (Map 5). Flowers have been collected in August and fruits in November.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF97FF8BFE4383651D7F0936.taxon	conservation	Conservation status Abrahamia lokobensis has an EOO of 3,124 km 2 and a minimum AOO of 16 km 2, with two subpopulations occurring within a protected area (Lokobe and Tsaratanana) whereas the other subpopulation was collected in the area of Maromandia where future decline is expected in terms of quality of habitat, and thus exists at three locations with respect to the threat of habitat degradation. It has therefore been assigned a preliminary status of “ Endangered ” [EN B 1 ab (iii) + 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF97FF8BFE4383651D7F0936.taxon	discussion	Notes Abrahamia lokobensis most closely resembles A. sambiranensis but can be distinguished by its smaller number of secondary veins (35 - 45 vs 50 - 60), its tertiary venation that is visible on both leaf surfaces (vs not visible), its shorter petiole (13 - 19 vs (20 -) 25 - 65 mm), and exclusively terminal (vs axillary) infructescence.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF97FF8BFE4383651D7F0936.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Antsiranana: Rég. Diana, Lokobe RNI, 13 ° 25 ’ S 48 ° 18 ’ E, 10 - 100 m, 14. XI. 1996, fr., Antilahimena et al. 315 (G, K, MO, P, TAN); ibid. loco, [13 ° 25 ’ S 48 ° 19 ’ E], 13 - 15. III. 1964, fr., Service Forestier 23459 (MO, P); Maromandia, Ambanja, [14 ° 12 ’ S 48 ° 05 ’ E], 15. VIII. 1952, ster., Service Forestier 64 - R- 152 (P [2 sheets]); Tsaratanana RNI, Ambinany Beangona, 14 ° 01 ’ S 48 ° 47 ’ E, 1200 - 1400 m, 1. VIII. 2000, y. bud, Antilahimena et al. 533 (G, K, MO, NY, P, TAN).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF95FFB5FE4383651B840CAF.taxon	materials_examined	Typus: MADAGASCAR. Prov. Antsiranana: Rég. Diana, commune Mahavonona, Montagne des Français, campement Ampitiantsambo, 12 ° 23 ’ 13 ’’ S 49 ° 22 ’ 53 ’’ E, 239 m, 11. IX. 2004, bud & fl., Randriambololomamonjy 7 (holo-: MO- 6278643!; iso-: P [P 04800624]!, TAN [TAN 001993]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF95FFB5FE4383651B840CAF.taxon	diagnosis	Diagnosis Abrahamia longipetiolata Randrian. & Lowry can be distinguished from other members of the genus by the combination of its strikingly long petiole (15 - 30 mm) and small leaves (6 - 11 3 1.5 - 3 cm) with impressed secondary veins on the adaxial surface.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF95FFB5FE4383651B840CAF.taxon	description	Description Trees c. 7 - 25 m tall, 19 - 60 cm DBH, bark with white latex. Leaves alternate, sometimes sub-opposite; blade narrowly elliptic to narrowly oblong, 6 - 11 3 1.5 - 3 cm, chartaceous to subcoriaceous, discolorous (glaucous above), apex acute and retuse, margin slightly revolute, base broadly cuneate, sometimes slightly asymmetric, adaxial surface glabrous, abaxial surface with appressed sericeous indument along the midvein, especially towards the base, venation craspedodromous, midvein prominent below, secondary veins 25 - 35 pairs, more or less parallel, 2 - 4 mm apart, impressed above, barely raised below, forming a wide angle with the midvein, tertiary veins very few, barely visible; petiole 15 - 30 mm long, 1 - 2 mm in diam., canaliculate on adaxial surface, strigose. Inflorescence terminal, a panicle, branched, 9 cm long, axes pubescent; bracts triangular, 1 - 2.5 3 0.5 - 1 mm, adaxial surface glabrous, abaxial surface pubescent. Male and female flowers 5 - merous, small, 3 - 4 mm long; pedicel c. 1 mm long, pubescent; calyx lobes 1 - 1.1 mm, ovate, valvate, adaxial surface glabrous, abaxial surface pubescent; corolla lobes ovate, 3 - 4 3 1.5 - 2 mm, imbricate, adaxial surface glabrous, abaxial surface sparsely pubescent (much less so than calyx); stamens 5 (rarely 6). Male flowers with filaments c. 1 - 1.2 mm long, glabrous, anthers 1.1 - 1.2 3 1 mm, ovoid, yellow, glabrous; disk c. 1.5 - 2 mm in diam., cupuliform, glabrous; ovary rudimentary. Female flowers with 5 staminodes, sometimes 6, filaments c. 1 mm long, glabrous, anthers 1 3 0.5 mm, ovoid, sterile, yellow, glabrous; disk c. 2 - 2.5 mm in diam., glabrous; ovary widely ovoid, 1 - 1.1 3 1.1. 5 mm, glabrous; style short, c. 0.5 mm, stigmas capitate. Fruits 2.4 - 2.6 3 1.5 - 1.8 cm, asymmetrical, ellipsoid, glabrous, with weak longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF95FFB5FE4383651B840CAF.taxon	distribution	Distribution, ecology and phenology Abrahamia longipetiolata occurs in forests in far northeastern Madagascar (Map 7). Flowering material has been collected in August and September, and fruits have been recorded from October to December.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF95FFB5FE4383651B840CAF.taxon	conservation	Conservation status With an EOO of 799 km 2, a minimum AOO of 20 km 2, and 6 localities, with subpopulations in only 2 protected areas (Loky Manambato and Montagne des Français), representing 5 locations, 3 of which are subjected to pressure from forest exploitation and land clearing, A. longipetiolata is assigned a provisional conservation status of “ Endangered’ ’ [EN B 1 ab (iii) + 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF95FFB5FE4383651B840CAF.taxon	discussion	Notes Abrahamia longipetiolata resembles A. ibityensis by the impressed venation of its leaves, but differs by its larger leaves (6 - 11 3 1.5 - 3 vs 3 - 7 3 1.2 - 3 cm), longer petiole (15 - 30 vs 5 - 7 mm), terminal paniculate (vs terminal or axillary thyrsiform) inflorescences, and fruit with united (vs separate) cotyledons.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF95FFB5FE4383651B840CAF.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Antsiranana: Vohémar, forêt d’Ambilondava, N du village de Daraina, 13 ° 10 ’ S 49 ° 39 ’ E, 400 m, 25. X. 2004, fr., Callmander & Anselme Be 268 (G, MO, P, TAN); Sadjoavato, forêt de Sahafary, 12 ° 35 ’ 17 ’’ S 49 ° 26 ’ 45 ’’ E, 207 m, 15. X. 2007, fl., Rakotoarivelo 172 (CNARP, G, MO, P, TAN); Mahavonona, Montagne des Français, campement Ampitiantsambo, 12 ° 23 ’ 13 ’’ S 49 ° 22 ’ 53 ’’ E, 239 m, 11. IX. 2004, y. fr., Randriambololomamonjy 2 (G, MO, P [2 sheets], TAN); Andrafiabe, Sahafary, 12 ° 35 ’ 43 ’’ E 49 ° 26 ’ 54 ’’ E, 233 m, 11. IX. 2004, fl. & y. fr., Razafitsalama et al. 638 (CNARP, G, MO, P, TAN, TEF); ibid. loco, [12 ° 34 ’ S 49 ° 26 ’ E], 10. XII. 1952, fr., Service Forestier 6287 (MO, P, TEF); ibid. loco, 9. IV. 1953, ster., Service Forestier 216 - R- 6 (P); N de Vohémar, versant du Massif de l’Ankerana, [13 ° 17 ’ S 49 ° 51 ’ E], 50 - 450 m, 17. XII. 1966, fr., Service Forestier 27359 (G, MO, P [2 sheets], TEF).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA9FFB1FE4383651CD109A6.taxon	materials_examined	Lectotypus (designated here): MADAGASCAR. Prov. Toamasina: Rég. Alaotra-Mangoro, Analamazaotra, c. 1000 m, fl., 1919, Thouvenot 36 (P [P 00722660]!; isolecto-: P [P 00722661]!, TAN [TAN 000075]!, US- 2494483 image seen).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA9FFB1FE4383651CD109A6.taxon	description	Description Trees 14 - 15 m tall, bark with white-yellow milky latex; young twigs glabrous but with scabrous indument at the tips, with a few lenticels. Leaves subopposite to opposite; blades obovate to oblanceolate, 2 - 4 3 1 - 2 cm, coriaceous to subcoriaceous, apex emarginate, retuse or rounded, margin entire, base cuneate, adaxial surface glabrous, abaxial surface with very sparse, appressed, short indument, venation craspedodromous, midvein prominent abaxially, secondary veins 15 - 28 pairs, 1 - 2 mm apart, parallel or nearly so, prominent abaxially, tertiary veins obscure on both surfaces; petiole 3 - 7 mm long, canaliculate on adaxial surface, glabrous. Inflorescence axillary and terminal, a panicle, 2.5 - 4.5 cm long, axes flattened or somewhat triangular, pubescent to strigose. Male flowers 5 - merous; pedicel 0.2 - 0.3 mm long, very sparsely strigose; calyx lobes deltate, 1 3 1 mm, valvate, glabrous; corolla lobes ovate, 2.5 3 1.5 mm, imbricate, glabrous; stamens 5, filaments 0.5 - 0.7 mm long, glabrous, anthers 0.5 - 0.6 mm long, obloid to slightly ovoid, glabrous; disk dish-shaped or slightly cupiliform, 1.5 mm in diam., glabrous; ovary rudimentary. Female flowers unknown. Fruits 1.5 - 1.7 3 1 - 1.2 cm, green when fresh, black when dry, ellipsoid to slightly obovoid, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA9FFB1FE4383651CD109A6.taxon	vernacular_names	Vernacular names “ Ditimena’ ’, “ Soretry’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA9FFB1FE4383651CD109A6.taxon	distribution	Distribution, ecology and phenology Abrahamia louvelii is known only from the Analamazaotra and Vohibe (Ambalabe) forests (Map 5) at elevations between 600 and 1000 m, in areas with eastern humid forest. Material has been collected in flower in November and in fruit in January.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA9FFB1FE4383651CD109A6.taxon	conservation	Conservation status With a minimum AOO of 8 km 2 and 3 localities, all of which are situated within protected areas (Ambalabe and Analamazaotra), A. louvelii is assigned a preliminary conservation status of “ Least Concern ” [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA9FFB1FE4383651CD109A6.taxon	discussion	Notes Abrahamia louvelii most closely resembles A. minutifolia but differs by having flowers borne in panicles (vs thyrses), a glabrous (vs abaxially pubescent) corolla, and glabrous (vs velutinous) fruits. 100 Boissiera 71 We have selected one of the two sheets of Thouvenot 36 in the Paris herbarium as the lectotype of Protorhus louvelii because it comprises more complete material.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA9FFB1FE4383651CD109A6.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Toamasina: Réserve spéciale au S d’Andasibe Mahaveriky, [18 ° 56 ’ S 48 ° 26 ’ E], 4. XI. 1967, ster., Jacquemin 687 (P); Vatomandry, Ambalabe, forêt de Vohibe, 19 ° 09 ’ 39 ’’ S 48 ° 34 ’ 43 ’’ E, 657 m, 22. I. 2012, fr., Rakotoarivelo et al. 643 (G, K, MO, P, TAN); ibid. loco, 19 ° 09 ’ 31 ’’ S 48 ° 34 ’ 40 ’’ E, 536 m, 19. II. 2009, ster., Razanatsima 667 (MO, P, TAN); Moramanga, Analamazaotra, Périnet, [18 ° 56 ’ S 48 ° 26 ’ E], II. 1966, ster., Service Forestier 25719 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFAFFFBDFE4383651DF50D24.taxon	materials_examined	Typus: MADAGASCAR. Prov. Toamasina: Rég. Alaotra-Mangoro, Moramanga, Analamazaotra RS, “ carrière de sable ”, [18 ° 56 ’ S 48 ° 26 ’ E], 900 - 950 m, 29. X. 1996, fl., A. Randrianasolo 516 (holo-: MO- 04895273!; iso-: G!, K!, P [P 06169142]!, TAN [TAN 002000]!)	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFAFFFBDFE4383651DF50D24.taxon	diagnosis	Diagnosis Abrahamia minutifolia Randrian. & Lowry can be distinguished by having leaves that are smaller (1 - 3 3 0.9 - 1.8 cm) than any other member of the genus and fruits that are among the smallest (1 - 1.5 3 0.5 - 0.8 cm).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFAFFFBDFE4383651DF50D24.taxon	description	Description Trees 3.5 - 12 m tall, 11.9 - 40 cm DBH, bark with translucid latex; young twigs waxy, lenticellate, with short, red, appressed indument at the tip. Leaves subopposite; blades, obovate, 1.8 - 4 3 0.9 - 1.8 cm, subcoriaceous to coriaceous, apex emarginate, retuse, or rounded, margin entire, base acute, adaxial surface glabrous, abaxial surface with very sparse appressed reddish indument when young, glabrescent, venation craspedodromous, midvein prominent below, secondary veins 15 - 20 pairs, 0.5 - 2 mm apart, prominent on both surfaces, tertiary veins raised on both surfaces; petiole 3 - 8 mm long, canaliculate on adaxial surface, reddish pubescent when young, later glabrescent. Inflorescence axillary or terminal, a thyrse, 1 - 3 cm long, axes somehow flattened, with ferruginous indument. Male and female flowers 4 - or 5 - merous; pedicel 0.2 - 0.3 mm long, pubescent; calyx lobes deltate, 1 3 1 mm, imbricate, glabrous, persistent on fruit; corolla lobes ovate, 2 - 2.2 3 1 - 1.5 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent; stamens 4 or 5. Male flowers with filaments 1 - 1.5 mm long, glabrous, white, anthers c. 0.8 mm long, obloid to slightly ovoid, yellow, glabrous; disk cupuliform, c. 1 mm in diam., glabrous; ovary rudimentary. Female flowers with filaments 0.3 - 0.5 mm long, glabrous to slightly pubescent, white, anthers c. 0.5 mm long, subglobose to ovoid, sterile, yellow, glabrous; disk c. 1.5 mm in diam., glabrous; ovary widely to very widely ovoid, c. 1.5 3 1 mm, pubescent, placentation subapical; style shortly trifid, subapically to apically inserted, 0.5 mm long, partly pubescent, stigmas capitate. Fruits 1 - 1.5 3 0.5 - 0.8 cm, ellipsoid, slightly asymmetrical, surface velutinous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFAFFFBDFE4383651DF50D24.taxon	distribution	Distribution, ecology and phenology Abrahamia minutifolia has been collected primarily in premontane forest along Madagas-car’s eastern escarpment at an elevation of c. 900 m and in central high elevation forest at c. 1500 m, but it also occurs in humid forests in both the eastern and northwestern parts of the island (Map 7). It flowers between September and November, and fruits have been collected from November to April.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFAFFFBDFE4383651DF50D24.taxon	conservation	Conservation status With an EOO of 117,262 km 2, a minimum AOO of 84 km 2, and 21 known localities, with subpopulations in at least 5 protected areas (Ambohitantely, Kalabenono, Maso-ala, Manongarivo, and Analamazaotra-Perinet), A. minutifolia is assigned a preliminary conservation status of “ Least Concern ” [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFAFFFBDFE4383651DF50D24.taxon	discussion	Notes Abrahamia minutifolia is part of a group of species, along with A. delphinensis, A. diti-mena, A. itremoensis, A. lenticellata and A. sericea, that has velutinous fruits, but is distinguished by its thyrsiform (vs paniculate) inflorescence and smaller fruits (1 - 1.5 3 0.5 - 0.8 vs 2 - 3 - 2.5 3 1.2 - 1.4 cm in A. delphinensis, 2 - 3 3 1.4 - 1.6 cm in A. ditimena, 1.5 - 2.3 3 1.2 - 1.5 cm in A. itremoensis, 2 - 2.6 3 1.3 - 1.5 cm in A. lenticellata and 2 - 3 3 0.8 - 1.3 cm in A. sericea). The name Protorhus ditimena var. minutifolia, published by PERRIER DE LA BâTHIE (1944) on the basis of a fragmentary specimen (Louvel 21), is invalid because it lacked a Latin diagnosis, as required for all names published between 1 January 1935 and 31 December 2011 under Article 39.1 of the International Code of Nomenclature (MCNEILL et al., 2012). We have chosen to retain Perrier de la Bâthie’s epithet, which aptly characterizes this taxon, by formally describing it as a new species.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFAFFFBDFE4383651DF50D24.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Antananarivo: Ankazobe, Ambohitantely RS, 18 ° 11 ’ S 47 ° 17 ’ E, 1620 m, 14. I. 1997, fl., Randrianaivo et al. 53 (G, K, MO, NY, P, TAN, WAG); forest E of Anjozorobe, E Antsahabe village, [18 ° 22 ’ S 47 ° 47 ’ E], 1400 m, 22. X. 1996, fl., A. Randrianasolo 508 (MO, P, TAN); Ambohitantely RS, 18 ° 10 ’ 42 ’’ S 47 ° 17 ’ 00 ’’ E, 1530 m, 23 - 24. XI. 1993, fl., Schatz et al. 3565 (G, K, MO, P, TAN); ibid. loco, [18 ° 09 ’ S 47 ° 14 ’ E], V. 1953, fr., Service Forestier 7361 (P, TEF); ibid. loco, [18 ° 08 ’ S 47 ° 16 ’ E], 1600 m, IV. 1953, fr., Service Forestier 8417 (P, TEF); ibid. loco, 26. VII. 1954, ster., Service Forestier 11 - R- 282 (P, TEF); Poste forestier Manankazo, Ankazobe, [18 ° 09 ’ S 47 ° 14 ’ E], 22. VI. 1955, fr., Service Forestier 16814 (P, TEF). Prov. Antsiranana: Ambanja, Bekolosy, 14 ° 02 ’ S 48 ° 18 ’ E, 1200 m, 28. III. 1996, fr., Gautier & Be 2909 (G, MO, P, TAN, TEF); ibid. loco, 14 ° 02 ’ S 48 ° 18 ’ E, 1070 m, 23. IX. 1996, fl., Gautier et al. 3123 (G, MO, P, TAN, TEF); Ambilobe, forêt de Kalabenono, 13 ° 38 ’ 45 ’’ S 48 ° 40 ’ 45 ’’ E, 934 m, 27. XI. 2006, fr., Callmander et al. 631 (G, MO); Andapa, Doany, forêt d’Andramanalana, 14 ° 23 ’ 02 ’’ S 49 ° 21 ’ 36 ’’ E, 1280 m, IV. 2006, fr., Rakotovao et al. 3074 (G, MO, P, TAN); Andramilolo, SE Anjanaha-ribe, 14 ° 19 ’ 43 ’’ S 49 ° 18 ’ 08 ’’ E, 1412 m, 12. XI. 2006, fl., Razakamalala et al. 2939 (G, MO, P, TAN). Prov. Fianarantsoa: Tolongoina, Andrambavato, 2 km S Tatamaly, 21 ° 30 ’ S 47 ° 24 ’ E, 1075 m, 13. X. 2000, fl., Randriatafika 142 (G, K, MO, P, TAN). Prov. Toamasina: Alaotra-Mangoro, Ambatovy, Analamay, 18 ° 48 ’ 28 ’’ S 48 ° 20 ’ 10 ’’ E, 1083 m, 2. XII. 2005, fl., Antilahimena et al. 4316 (G, MO, P, TAN); ibid. loco, fl., Anti-lahimena et al. 4318 (G, MO, P, TAN); Moramanga, Ambatovy / Andranovery forest, 18 ° 52 ’ 09 ’’ S 48 ° 18 ’ 15 ’’ E, 1013 m, 26. X. 2007, fl., Antilahimena 5937 (G, MO, P, TAN); Analamazaotra, IX. 1925, bud, Louvel 21 (P); Maroantsetra, Masoala PN, E slope of Ambohitsitondroinan’Mahalevona summit, 15 ° 25 ’ 34 ’’ S 49 ° 57 ’ 22 ’’ E, 1160 m, 25. II. 2003, fr., Lowry et al. 6190 (G, K, MO, P, TAN, WAG); along road to Lakato, 18 km S of RN 2, 19 ° 04 ’ 06 ’’ S 48 ° 22 ’ 01 ’’ E, 1050 m, 17. X. 2004, fl., Lowry et al. 6547 (G, MO, P, TAN); Ambatovy, Analamay, 18 ° 49 ’ 26 ’’ S 48 ° 20 ’ 16 ’’ E, 1000 m, 26. I. 1997, fr., Rakotomalaza et al. 1005 (MO, P, TAN); Analamazaotra, 950 m, 29. X. 1996, fl., A. Randrianasolo 515 (G, MO, P, TAN); Mantadia PN, c. 11 km N of village of Andasibe, 18 ° 49 ’ 47 ’’ S 48 ° 25 ’ 56 ’’ E, 939 m, 10. I. 2003, ster., A. Randrianasolo 781 (MO, P, TEF); Ambatovy, 18 ° 49 ’ 28 ’’ S 48 ° 19 ’ 23 ’’ E, 1134 m, 26. XI. 2005, bud, Razafindraibe 135 (MO, P, TAN); Analamazaotra-Périnet, [18 ° 56 ’ S 48 ° 26 ’ E], 6. XI. 1957, bud, Service Forestier 18400 (G, K, MO, NY, P [2 sheets]); Ankazomanitra, PK 45, route Anosi-be-Moramanga, [19 ° 58 ’ S 48 ° 08 ’ E], 6. XI. 1968, bud, Service Forestier 26800 (G, MO, NY, P, TEF); W d’Antanandava (PK 45 Moramanga-Anosibe), [19 ° 08 ’ S 48 ° 15 ’ E], XI. 1968, fl., Service Forestier 28453 (G, MO, P, TEF); ibid. loco, XI. 1968, bud & fl., Service Forestier 28454 (G, MO, NY, P, TEF).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA3FFBFFE4383651AF80C09.taxon	materials_examined	Typus: MADAGASCAR: sine loco, s. d., fl., Thouars s. n. (holo-: P [P 00722662]!; iso-: P [P 00722664]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA3FFBFFE4383651AF80C09.taxon	description	Description Trees 4 - 25 m m tall, 10 - 22 cm DBH, bark with red or white milky latex; young twigs glabrous or glabrescent, with lenticels, waxy. Leaves usually whorled, sometimes opposite; blades oblanceolate, 3.5 - 10.5 3 1.5 - 3.8 cm, coriaceous, apex emarginate or retuse, margin revolute, base cuneate, glabrous on both surfaces, venation craspedodromous, midvein prominent abaxially, secondary veins 20 - 30 pairs, parallel, 3 mm apart, prominent abaxially, tertiary veins very slightly raised on abaxial surface; petiole 5 - 15 mm long, canaliculate on adaxial surface, glabrous. Inflorescence axillary or terminal, a panicle, 2 - 4 cm long, axes flattened, glabrous or glabrescent. Male and female flowers 5 - merous; pedicel 0.5 mm long, pubescent; calyx lobes deltoid to triangular, 0.5 - 0.7 3 1 - 1.2 mm, valavate, glabrous, persistent on fruit; corolla lobes ovate, 2 - 2.2 3 1 - 1.5 mm, imbricate, adaxial surface glabrous, abaxial surface with short reddish indument; stamens 5. Male flowers with filaments c. 1 mm long, glabrous, anthers 0.5 mm long, ovoid to subglobose, glabrous; disk c. 1 mm in diam., glabrous; ovary rudimentary. Female flowers with filaments c. 1 mm long, anthers 0.5 3 0.5 mm, obloid to ovoid, sterile, glabrous; disk c. 1 mm in diam., cupuliform, glabrous; ovary c. 0.5 - 0.6 3 1 mm, broadly ovoid, glabrous, placenation subapical, style very short, c. 0.2 - 0.3 mm long, apical, stigmas capitate. Fruits 2 - 2.5 3 1 - 1.5 cm, ellipsoid, glabrous, surface smooth.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA3FFBFFE4383651AF80C09.taxon	vernacular_names	Vernacular names “ Ditimenafotsy’ ’, “ Hazombarorana’ ’, “ Tahaka’ ’, “ Tsiramiramy’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA3FFBFFE4383651AF80C09.taxon	distribution	Distribution, ecology and phenology Abrahamia nitida occurs in humid forest in coastal eastern Madagascar (Map 6). It has been recorded in flower in December and in fruit in late January.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA3FFBFFE4383651AF80C09.taxon	conservation	Conservation status With an EOO of 45,042 km 2, a minimum AOO of 76 km 2, and 14 known locations, with subpopulations in 4 protected areas (Agnalazaha, Makira, Nosy Mangabe, and Pointe à Lareée) and the remainder subjected to pressure from forest exploitation and land clearing, A. nitida is assigned a preliminary conservation status of “ Near Threatened ” [NT] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA3FFBFFE4383651AF80C09.taxon	discussion	Notes The leaves of A. nitida are usually whorled, a feature found in only a few other members of the genus, the most similar being A. sericea, from which A. nitida can be distinguished by its leaves with widely (vs very closely) spaced secondary veins, glabrous (vs pubescent) disk, and glabrous (vs velutinous) fruit. We have followed René Capuron (note in herb.) in considering the second collection made by Thouars listed above as an isotype despite the fact that we cannot be certain it represents part of a single gathering.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA3FFBFFE4383651AF80C09.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Antsiranana: Antalaha, PN Masoala, 15 ° 35 ’ S 50 ° 25 ’ E, 22. IX. 1994, ster., Rahajasoa et al. 510 (MO, TAN); ibid. loco, 22. IX. 1994, ster., Rahajasoa et al. 523 (MO, P, TAN). Prov. Fianarantsoa: Farafangana, Mahabo, forêt de Marovahatry, 23 ° 10 ’ 41 ’’ 47 ° 43 ’ 49 ’’ E, 23 m, 21. IV. 2004, y. fr., Ludovic 738 (MO, P, TAN); Mahabo Mananivo, Nosiala, bloc de Betaindambo, 23 ° 08 ’ 42 ’’ S 47 ° 42 ’ 18 ’’ E, 31 m, 22. IV. 2004, fr., Ludovic 766 (MO, P, TAN); ibid. loco, 23 ° 11 ’ S 47 ° 41 ’ E, 21 m, 29. X. 2004, fl., Ludovic et al. 927 (G, K, MO, P, TAN); Mahabo Mananivo, Baboaka, [23 ° 10 ’ S 47 ° 44 ’ E], 21 m, 20. XI. 2004, fl., Ludovic et al. 998 (G, K, MO, P, TAN); near village of Mahabo, 23 ° 10 ’ 51 ’’ S 47 ° 42 ’ 29 ’’ E, 30 m, 5. XI. 2001, fl., McPherson et al. 18334 (MO, TAN, TEF); ibid. loco, McPherson et al. 18347 (MO, P, TAN, TEF); ibid. loco, 23 ° 10 ’ 42 ’’ S 47 ° 42 ’ 24 ’’ E, 20 m, 7. XI. 2001, fl., McPherson et al. 18373 (MO, P, TAN, TEF); ibid. loco, 23 ° 08 ’ 53 ’’ S 47 ° 43 ’ 04 ’’ E, 22. IX. 2002, fr., Rabevohitra 4077 (MO, NY, P, TAN, TEF); ibid. loco, 10 m, 19. VI. 2004, ster., A. Randrianasolo et al. 867 (MO); ibid. loco, 23 ° 12 ’ 05 ’’ S 47 ° 42 ’ 48 ’’ E, 24 m, 26. VIII. 2003, fr., Razakamalala et al. 744 (G, MO, P, TEF); Mananjary, forêt d’Alimamba, 21 ° 28 ’ 04 ’’ S 48 ° 17 ’ 43 ’’ E, 13. X. 2004, fl., Razakamalala et al. 1655 (MO, NY, P, TAN, TEF); Amporoforo, [22 ° 29 ’ 00 ’’ S 47 ° 47 ’ 30 ’’ E], 26. VII. 1955, ster., Service Forestier 13912 (MO, P, TEF); Ambalavotaka, [21 ° 19 ’ 30 ’’ S 48 ° 21 ’ 30 ’’ E], 7. I. 1955, fr., Service Forestier 15459 (P, TEF). Prov. Toamasina: Maroantsetra, commune d’Ambinanitelo, Ankirindro, 15 ° 28 ’ 29 ’’ S 49 ° 33 ’ 10 ’’ E, 200 m, 21. XI. 2002, fl., Antilahimena et al. 1430 (MO, P, TAN); Fénérive-Est, Mahatsara forest station, [17 ° 38 ’ S 49 ° 29 ’ E], 23. I. 1986, fr., Dorr 4617 (MO, P, TAN); Soanieran’Ivongo, forêt d’Antanambao-Ambodimanga, 16 ° 46 ’ 24 ’’ S 49 ° 42 ’ 51 ’’ E, 30. I. 2003, fr., Rabevohitra et al. 4305 (G, K, MO, P, TEF); Mahanoro, Ambodibonara, forêt Nakinà, 20 ° 21 ’ 34 ’’ S 48 ° 36 ’ 21 ’’ E, 13. XI. 2003, fl., Rabevohitra et al. 4753 (G, K, MO, NY, P, TAN, TEF); Soanieran’Ivongo, NW Tanambao, 16 ° 45 ’ 12 ’’ S 49 ° 42 ’ 56 ’’ E, 31. I. 2004, fr., Rabevohitra et al. 4980 (MO, NY, P, TEF); Brickaville, Andranokoditra, Vohibola, 18 ° 33 ’ 23 ’’ S 49 ° 14 ’ 58 ’’ E, 5 m, 10. XII. 2001, fr., Rabenantoandro et al. 871 (MO, P, TAN, TEF); Tampolo forest station, 0 - 5 m, 31. VII. 1996, ster., A. Randrianasolo 457 (MO, P, TAN); Ambila Lemaitso, 18 ° 49 ’ S 49 ° 08 ’ E, 0 - 3 m, 28. X. 1996, fr., A. Randrianasolo 512 (G, K, MO, P, TAN); ibid. loco, 18 ° 51 ’ 38 ’’ S 49 ° 08 ’ 34 ’’ E, 0 - 5 m, 9. X. 1999, ster., A. Randrianasolo et al. 586 (G, MO, P, TAN); Maroantsetra, Nosy Mangabe, 15 ° 30 ’ S 49 ° 46 ’ E, 0 - 330 m, 13 - 23. IV. 1988, ster., Schatz 2131 (MO, TAN); Mahanoro, [19 ° 44 ’ 00 ’’ S 48 ° 48 ’ 30 ’’ E], 29. I. 1951, fr., Service Forestier 2998 (P, TAN, TEF); Ambila Lemaitso, [18 ° 49 ’ S 49 ° 08 ’ E], 11. XII. 1952, fl., Service Forestier 6448 (P [2 sheets], TEF); ibid. loco, 12. XII. 1952, fl., Service Forestier 6486 (P, TEF); ibid. loco, 24. I. 1953, fr., Service Forestier 7253 (MO, P, TEF). Prov. Toliara: Fort-Dauphin, Mahatalaky St Luce S 7, 2 km S d’Ambadrika, 24 ° 47 ’ 47 ’’ S 47 ° 08 ’ 37 ’’ E, 20 m, 25. XI. 2011, fl., Ratovoson 1793 (MO, P, TAN).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA1FFB9FE4383651C0E0E06.taxon	materials_examined	Typus: MADAGASCAR. Prov. Antsiranana: Rég. Diana, Nossi-Bé, 1840, fl., Richard 366 (holo-: P [P 00722663]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA1FFB9FE4383651C0E0E06.taxon	description	Description Trees 10 - 20 m tall, 23 - 25 cm DBH; young twigs very often triangular, lenticellate, with ferruginous indument. Leaves subopposite to opposite; blades obovate to oblanceolate or narrowly oblong to elliptic, 4 - 17 3 1.5 - 5.5 cm, chartaceous to subcoriaceous, apex retuse, rounded, or emarginate, margin entire, minutely thickened and pale in color on abaxial surface, base cuneate or cuneate-obtuse, adaxial surface glabrous, somewhat waxy, abaxial surface pubescent when young, later nearly glabrous, venation craspedodromous (rarely a few secondary veins not quite reaching the margin), midvein prominent abaxially, secondary veins 35 - 40 pairs, 2 - 6 mm apart, more or less parallel, prominent on both surfaces but more so abaxially, tertiary veins slightly raised on abaxial surface; petiole 5 - 15 mm long, canaliculate on adaxial surface, glabrous, waxy. Inflorescence axillary or terminal, a panicle, 3 - 5 cm long, the axes terete or flattened, with ferruginous indument. Male and female flowers 5 - merous (rarely 6 - merous); pedicel 0 - 0.5 mm long, ferruginous pubescent or strigose; calyx lobes ovate, c. 1 3 1 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent or ferruginous pubescent; corolla lobes ovate, 2 3 1 mm, imbricate, adaxial surface glabrous, abaxial surface ferruginous pubescent or scabrous; stamens 5 or 6. Male flowers with filaments 1 - 1.5 mm long, glabrous, anthers c. 0.5 mm long, subglobose, glabrous; disk cupuliform, sometimes shallowly so, 1 - 1.5 mm in diam., glabrous; ovary rudimentary. Female flowers with filaments 0.5 - 0.7 mm long, glabrous, anthers c. 0.3 mm long, subglobose to slightly ovoid, glabrous; disk annular, cupuliform, sometimes shallowly so, 1.5 mm in diam., sterile, glabrous; ovary very widely ovoid, slightly asymmetric, 1 3 1 mm, glabrous, placentation apical; style simple, very short, inserted more sub-apically than laterally, 1 mm long, glabrous, stigma capitate, 3 - 4 - lobed. Fruits c. 2 3 1 cm, ellipsoid, slightly asymmetrical, glabrous, with obscure longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA1FFB9FE4383651C0E0E06.taxon	vernacular_names	Vernacular names “ Hazombarorana’ ’, “ Manavodrevo’ ’, “ Tarata’ ’, “ Tsiramiramy’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA1FFB9FE4383651C0E0E06.taxon	distribution	Distribution, ecology and phenology Abrahamia oblongifolia occurs in far northern and northwestern Madagascar (Map 6), from sea level to an elevation of c. 300 m, usually along rivers or at the base of hills or mountains. Flowering material has been recorded in October and November, and fruits have been collected in December and January.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA1FFB9FE4383651C0E0E06.taxon	conservation	Conservation status With an EOO of 3,220 km 2, a minimum AOO of 32 km 2, and estimated maximum AOO of <500 km 2, and 7 recorded localities representing 7 locations with respect to the threat of forest exploitation, and with subpopulations in 2 protected areas (Analamera and Montagne des Français) and projected decline outside of protected areas, A. oblongifolia is assigned a preliminary conservation status of “ Vulnerable’ ’ [VU B 1 ab (iii) + 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA1FFB9FE4383651C0E0E06.taxon	discussion	Notes Abrahamia oblongifolia can be distinguished from all other members of the genus by the combination of its numerous secondary veins (35 - 40), both terminal and axillary inflorescences, and glabrous fruits.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA1FFB9FE4383651C0E0E06.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Antsiranana: Analamera RS, Betsieka, [12 ° 42 ’ S 49 ° 20 ’ E], 10 - 20 m, I. 1938, ster., Humbert 19091 ter (P); Mahavanona, Montagne des Français, Ambatosarolo, 12 ° 22 ’ 27 ’’ S 49 ° 21 ’ 26 ’’ E, 204 m, 8. X. 2004, y. fr., Randriambololomamonjy et al. 42 (MO, P, TAN); Sakaramy, près de Diego, [12 ° 27 ’ S 49 ° 13 ’ E], bud, Perrier de la Bâthie 17672 (P); Ramena, Montagne des Français, Andavakoera, 12 ° 20 ’ 48 ’’ S 49 ° 21 ’ 28 ’’ E, 88 m, 7. IX. 2004, fl., Randrianaivo 1081 (CNARP, G, MO, P, TAN, TEF); Mahavanona, Montagne des Français, campement Antafiankoroka, 12 ° 21 ’ 39 ’’ S 49 ° 21 ’ 34 ’’ E, 281 m, 24. IX. 2004, y. fr., Randrianarivelo et al. 139 (G, K, MO, P, TAN); Nosy Be, [13 ° 19 ’ S 48 ° 16 ’ E], 1840, fr., Richard 315 (P); Montagnes des Français, [12 ° 22 ’ S 49 ° 21 ’ E], 31. XI. 1952, bud, Service Forestier 5885 (MO, P [2 sheets], TEF); ibid. loco, 23. XI. 1955, bud, Service Forestier 15355 (G, MO, P [2 sheets], TEF); Forêt de Sahankazo, [12 ° 29 ’ S 49 ° 26 ’ E], 11. XII. 1955, fr., Service Forestier 15507 (G, MO, P [2 sheets], TEF); Montagnes des Français, [12 ° 22 ’ S 49 ° 21 ’ E], 21. IV. 1954, ster., Service Forestier 28 - R- 61 (P); ibid. loco, 21. V. 1951, ster., Service Forestier 51 - R- 150 (P); ibid. loco, 24. III. 1955, ster., Service Forestier 442 - R- 6 (P). Sine loco: 17. X. 1927, fl., Ursch 186 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA7FFBBFE4383651CFE0A85.taxon	materials_examined	Typus: MADAGASCAR. Prov. Antsiranana: Rég. SAVA, Vohémar, commune rurale de Daraina, forêt de Bobankora, partie N, 510 m, 13 ° 13 ’ 15 ’’ S 49 ° 45 ’ 42 ’’ E, 510 m, 29. I. 2005, fr., Ranirison & Nusbaumer 919 (holo-: G [G 00019652]!; iso-: MO- 6671434!, P [P 00580356]!, TEF [TEF 000886]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA7FFBBFE4383651CFE0A85.taxon	diagnosis	Diagnosis Abrahamia patrickii Randrian. & Lowry can easily be distinguished from other members of the genus by its sub-verticillate to subopposite leaves, narrowly oblanceolate leaf blade with very closely spaced secondary veins, and the presence of circular orange-brown scales on the abaxial leaf surface.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA7FFBBFE4383651CFE0A85.taxon	description	Description Trees 13.5 m tall, 20 cm in diam.; young twigs lenticellate. Leaves sub-verticillate or subopposite; blade oblanceolate, 3 - 5.5 3 1 - 1.5 cm, subcoriaceous, apex emarginate or round, margin undulate, base cuneate to acute, adaxial surface glabrous, abaxial surface mostly glabrous but with scattered short reddish indument along midvein and margin, as well as circular, orange-brown scales, venation craspedodromous, midvein prominent abaxially, secondary veins 24 - 28 pairs, 1 - 3 mm apart, very often closely spaced, raised on both surfaces, tertiary veins not visible on either surface; petiole 6 - 13 mm long, c. 0.8 - 1 mm in diam., glabrous, canaliculate on adaxial surface. Inflorescence and flowers unknown. Fruits 2.3 - 2.6 3 1.1 - 1.3 cm, slightly asymmetric, ellipsoid, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA7FFBBFE4383651CFE0A85.taxon	etymology	Etymology The species epithet honors Patrick Ranirison for his dedication to the study of the Malagasy flora, most notably his work in the Daraina region of northeastern Madagascar.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA7FFBBFE4383651CFE0A85.taxon	distribution	Distribution, ecology and phenology Abrahamia patrickii is known only from the type collection from the Daraina area and a second gathering made N of Sambava (Map 6), in semi-deciduous and evergreen humid forest. It has been collected with fruits in January and February.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA7FFBBFE4383651CFE0A85.taxon	conservation	Conservation status With a minimum AOO of 8 km 2 and 2 localities, both of which situated are within protected areas (Loky Manambato and Makirovona), A. patrickii is assigned a preliminary conservation status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA7FFBBFE4383651CFE0A85.taxon	discussion	Notes Abrahamia patrickii closely resembles A. nitida by its sub-verticillate leaves with an oblanceolate blade, similar number of craspedodromous secondary veins (24 - 28 in A. patrickii and 20 - 30 in A. nitida), similar petiole length (6 - 13 mm in A. patrickii and 5 - 15 mm in A. nitida), and glabrous fruit. However, the leaves of A. patrickii differ from those of A. nitida by their undulate (vs revolute) margins, orange-brown circular scales (vs glabrous) on the abaxial surface, narrowly spaced secondary veins (1 - 3 vs 3 mm apart), obscure (vs visible and raised) tertiary veins, and fruit with longitudinal striations (vs smooth) on the surface.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA7FFBBFE4383651CFE0A85.taxon	materials_examined	Paratypus MADAGASCAR. Prov. Antsiranana: Rég. SAVA, Sambava, forêt d’Antsahaha Kasaka, 2 km NW du village Ambavala, 14 ° 05 ’ 39 ’’ S 50 ° 02 ’ 42 ’’ E, 146 m, 16. II. 2006, fr., Andrianjafy et al. 1590 (MO, TAN, P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA5FFA5FE4383651DEF0F79.taxon	materials_examined	Lectotypus (designated here): MADAGASCAR. Prov. Antsiranana: Rég. Diana, Nossibe, I. 1850, fl. & y. fr., Boivin 2257 (P [P 00580352]!; isolecto-: G!, P [P 00364873, P 06774888, P 06774892]!, MO- 6684876!, TAN!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA5FFA5FE4383651DEF0F79.taxon	description	Description Trees 6 - 11 m tall, 11 - 15 cm DBH; young branch tips covered with short and appressed indument. Leaves opposite or subopposite; blade obovate, 3.9 - 9.2 3 1.8 - 4 cm, chartaceous, apex rounded to emarginate, margin undulate, base cuneate to attenuate, adaxial surface glabrous, abaxial surface with scattered short appressed indument, venation craspedodromous, midvein prominent abaxially, secondary veins 14 - 17 pairs, more or less parallel and ascending, 3 - 9 mm apart, widely spaced at leaf mid-section, raised on both surfaces or some impressed near margin, tertiary veins impressed, not very visible, especially on adaxial surface; petiole 8 - 15 mm, c. 1 mm in diam., canaliculate on adaxial surface, covered with short, appressed indument when young, glabrescent. Inflorescence always axillary, a panicle, very short, up to 1.5 cm long, with no more than 5 flowers, axes strigose. Male flowers unknown. Female flowers 5 - merous, small, c. 1 mm long, subtended by a triangular bract with indument; pedicel 1 mm long; calyx lobes depressed ovate, very small, 0.5 3 1 mm, glabrous on both surfaces; corolla lobes ovate, thick, c. 2 x 1 mm, glabrous on both surfaces; staminodes with filaments c. 0.8 mm long, glabrous, anthers c. 0.3 mm long, ovoid, sterile, glabrous; disk cupuliform, glabrous, c. 1.5 mm in diam.; ovary spherical glabrous, style short, thick, 0.5 mm long. Fruits unknown.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA5FFA5FE4383651DEF0F79.taxon	distribution	Distribution, ecology and phenology Abrahamia pauciflora is restricted to NW Madagascar, where it has been collected on the island of Nosy Be and in the Ambongo-Boina area (Map 6) in humid forest. Material with floral buds and flowers has been collected in November and January (the fruiting period is not known).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA5FFA5FE4383651DEF0F79.taxon	conservation	Conservation status With a minimum AOO of 4 km 2 and a single known extant subpopulation, which is within a protected area (Ampasindava), and considering that the historical locality at Maevarano was documented in 1908, well more than 3 generations ago, A. pauciflora is assigned a preliminary conservation status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA5FFA5FE4383651DEF0F79.taxon	discussion	Notes Abrahamia pauciflora can easily be distinguished from other members of the genus by the combination of its leaves with an attenuate base and very widely spaced secondary veins (to 9 mm apart in the median section of the blade), and very short inflorescence (not exceeding 1.5 cm long) usually bearing only 1 - 5 flowers. Contrary to what is indicated in many online databases, the name Protorhus pauciflora (≡ Abrahamia pauciflora) was not treated in ENGLER (1881: 421 or 422), but rather was published two years later by the same author in his treatment of Anacardiaceae for the Monographiae Phanerogamarum (ENGLER, 1883: 313). Curiously, ENGLER (1883) erroneously cited that name as having been published in his earlier work (ENGLER, 1881: 421). While one of the specimens of Boivin 2257 in the Paris herbarium has the handwritten word ‘ type’ on the label, this was probably added subsequent to the publication of Protorhus pauciflora. We have chosen to designate this specimen as the lectotype because it bears Boivin’s original label. Two additional specimens of Boivin 2257 have labels indicating that they came from Mayotte in the Comoro Islands, but as indicated subsequently by Perrier de la Bâthie (in herb.), they are without question part of the same gathering as the material from Nossi Be.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFA5FFA5FE4383651DEF0F79.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Antsiranana: Ampasindava, forêt d’Andranomatavy, 13 ° 40 ’ 40 ’’ S 47 ° 58 ’ 35 ’’ E, 215 m, 2. XII. 2009, bud, Ammann et al. 467 (G, MO, P, TEF); ibid. loco, 13 ° 40 ’ 04 ’’ S 47 ° 59 ’ 21 ’’ E, 295 m, 25. XI. 2009, fl., Madiomanana et al. 272 (G, MO, P, TEF). Prov. Mahajanga: Morarivo, Maevarano, près de Majunga, [15 ° 56 ’ S 46 ° 31 ’ E], VII. 1908, ster., Perrier de la Bâthie 2341 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBBFFA1FE4383651BE90F53.taxon	materials_examined	Typus: MADAGASCAR. Prov. Toliara: Rég. Androy, Andohahela PN, parcelle 2, 24 ° 50 ’ S 46 ° 32 ’ E, 100 m, 7. XII. 1988, fr., Phillipson 2853 (holo-: MO- 3767006!; iso-: K, P [P 06775412]!, TAN [TAN 001997]!, US- 3661611 image seen, WAG [WAG 1523550] image seen).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBBFFA1FE4383651BE90F53.taxon	diagnosis	Diagnosis Abrahamia phillipsonii Randrian. & Lowry most closely resembles A. lenticellata, with which its range overlaps, but can be easily distinguished by its craspedodromous (vs semicraspedodromous) venation, flowers borne in thyrses (vs panicles), and glabrous (vs velutinous) fruits.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBBFFA1FE4383651BE90F53.taxon	description	Description Trees 3 - 12 (- 20) m tall, 12 - 30 cm DBH, bark with white latex; young twigs glabrous or pubescent (sometimes ferruginous). Leaves alternate or subopposite; blades oblanceolate to obovate or narrowly elliptic to narrowly oblong, 2.6 - 7.8 3 0.6 - 2 (- 2.2) cm, chartaceous to subcoriaceous, apex emarginate, retuse or rounded, sometimes acute, margin entire, base acute or cuneate, adaxial surface glabrous, very often waxy, abaxial surface with very sparse indument, slightly more appressed on the midvein, nearly glabrous otherwise, venation craspedodromous, midvein prominent abaxially, secondary veins 11 - 19 (- 23) pairs, prominent abaxially, 1 - 5 mm apart, tertiary veins rarely visible on both surfaces; petiole c. 0.5 - 10 mm long, glabrous, canaliculate on adaxial surface. Inflorescence axillary or terminal, a thyrse, 1.5 - 5.5 cm long, the axes flattened, with short, appressed, ferruginous or reddish indument. Male flowers 5 - merous; calyx lobes widely depressed ovate to depressed ovate or triangular, 0.8 - 1 3 1 - 1.5 mm, valvate, glabrous, persistent on fruits; corolla lobes ovate, 2 - 2.2 3 1 - 1.5 mm, imbricate, glabrous, stamens 5, filaments c. 1.5 mm long, white, glabrous, anthers 0.8 - 1 mm long, ovoid to subglobose, yellow, glabrous; disk 1 - 1.2 mm in diam., glabrous; ovary rudimentary. Female flowers unknown. Fruits 1.5 - 2 3 1 - 1.3 cm, obovoid to ellipsoid, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBBFFA1FE4383651BE90F53.taxon	etymology	Etymology The species epithet honors our friend and colleague Peter. B. Phillipson, whose work has contributed so much to our knowledge of the Malagasy flora, including that of the drier parts of the island, where he has conducted most of his field work over the last three decades.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBBFFA1FE4383651BE90F53.taxon	vernacular_names	Vernacular names “ Manantsadrano’ ’, “ Tarantantily’ ’, “ Tsitaké’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBBFFA1FE4383651BE90F53.taxon	distribution	Distribution, ecology and phenology Abrahamia phillipsonii is restricted to sub-arid southern Madagascar (Map 8), where it occurs at elevations between 100 and 1100 m. Flowering material has been collected between September and November, and fruits have been gathered from December through February. However, in drought years flowering can be delayed, which accounts for the limited number of collection with flowers gathered in March and April.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBBFFA1FE4383651BE90F53.taxon	conservation	Conservation status With an EOO of 64,857 km 2, a minimum AOO of 112 km 2, and 29 recorded localities, with subpopulations in 4 protected areas (Analavelona, Andohahela, Isalo, and Zombitsy), A. phillipsonii is assigned a preliminary conservation status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBBFFA1FE4383651BE90F53.taxon	discussion	Notes Abrahamia phillipsonii vegetatively most closely resembles A. deflexa, but it differs by its oblanceolate to obovate, narrowly elliptic to narrowly oblong (vs ovate, oblong or elliptic) leaf blades and its axillary or terminal (vs only terminal) inflorescence.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBBFFA1FE4383651BE90F53.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Fianarantsoa: Ihosy, Analalava forest, 2 - 3 km W of western boundary of Isalo PN, [22 ° 35 ’ 38 ’’ S 45 ° 08 ’ 20 ’’ E], 770 m, 18. V. 2004, fr., Rogers et al. 456 (G, K, MO, P, TAN); massif de l’Isalo, [22 ° 08 ’ S 45 ° 10 ’ E], 20. XII. 1968, fr., Service Forestier 28504 (P, TEF); Ranohira, PK 712, [22 ° 38 ’ S 45 ° 20 ’ E], 23. VII. 1955, ster., Service Forestier 13889 (MO, P, TEF); Isalo, 20. VII. 1954, ster., Service Forestier 135 - R- 141 (P); Isalo, Benenitra, 8. VIII. 1954, ster., Service Forestier 180 - R- 18 (P); Isalo, Ranohira, 23. VII. 1955, ster., Service Forestier 288 - R- 239 (P); Ihosy, 30. X. 1953, ster., Service Forestier 777 - R- 1 (P). Prov. Toliara: Tolagnaro, Andohahela PN, N du mont Apiky, [24 ° 47 ’ 02 ’’ S 46 ° 43 ’ 12 ’’ E], 708 m, 18. I. 2007, fr., Andriamihajarivo 1156 (MO, P, TAN, TEF); Amboasary, fourré d’Ambatofotsy Masay, 24 ° 18 ’ 16 ’’ S 46 ° 08 ’ 00 ’’ E, 350 m, 11. XI. 2008, fr., Andriamihajarivo et al. 1574 (G, MO, P, TAN); Sakaraha, forêt de Zombitsy, [22 ° 48 ’ S 44 ° 41 ’ E], XII. 1959, fr., Bosser 13402 (MO, P [2 sheets], TAN); Andohahela PN, parcelle 3, [25 ° 00 ’ S 46 ° 40 ’ E], 17. XI. 1990, fr., Dumetz 1413 (MO, P, TAN); vallée de Fiherenana, 300 - 500 m, 1 - 2. VIII. 1928, fr., Humbert & Swingle 5063 (= Service Forestier 50) (P); Betroka, Mont Vohipolaka, 1200 - 1400 m, XI. 1933, ster., Humbert 11694 ter (P, TAN); Amboasary, entre Andohahela et l’Elakelaka, [24 ° 45 ’ S 46 ° 43 ’ E], 700 m, I-II. 1934, fr., Humbert 13770 (P); entre Andohahela et l’Elakelaka, mont Apiky, [24 ° 45 ’ 30 ’’ S 46 ° 43 ’ 30 ’’ E], 800 - 900 m, I-II. 1934, fr., Humbert 13850 (G, P [2 sheets]); Beroroha, bassin de la Malio, [21 ° 58 ’ S 45 ° 15 ’ E], 400 - 450 m, 23 - 27. XI. 1946, bud, Humbert 19403 (P); forêt de Zombitsy, [22 ° 48 ’ S 44 ° 41 ’ E], 600 - 850 m, 26 - 29. III. 1955, fl. & bud, Humbert 29595 (G, MO [2 sheets], P); Beronono-Makay, 21 ° 18 ’ 48 ’’ S 45 ° 08 ’ 55 ’’ E, 531 m, 16. I. 2010, fr., Rakotovao et al. 5117 (MO, P, TAN); PN Andohahela, Isaka Ivondro, 24 ° 48 ’ S 46 ° 52 ’ E, 100 - 500 m, 16 - 24. III. 1993, Randriamampionona 202 (G, MO, P, TAN); PN Andohahela, parcel 3, 25 ° 04 ’ S 46 ° 41 ’ E, 100 - 300 m, 8 - 10. IV. 1993, y. fr., Randriamampionona 285 (G, MO, P, TAN); Andohahela PN, parcel 1, Tsimelahy, 24 ° 50 ’ 20 ’’ S 46 ° 32 ’ 17 ’’ E, 6. IV. 1996, ster., Randriamampionona 1259 (MO, TAN); Sakaraha, Mahaboboka, Ambinanintelo, forêt d’Analavelona, 22 ° 41 ’ 14 ’’ S 44 ° 09 ’ 32 ’’ E, 961 m, 3. XII. 2016, fr., Randrianarivony et al. 745 (MO, P, TAN); PN Andohahela, parcelle n ° 3, [25 ° 04 ’ S 46 ° 41 E], 100 - 300 m, 19. X. 1990, fl., A. Randrianasolo 181 (MO [2 sheets], P, TAN, TEF); Sakaraha, at start of Kinanamboribory valley, 22 ° 39 ’ 44 ’’ S 44 ° 11 ’ 02 ’’ E, 1208 m, 8. XII. 2009, fr., A. Randrianasolo 1246 (G, MO, P, TAN); Ambatoabo, 5 km N d’Imonty, 24 ° 46 ’ 44 ’’ S 46 ° 42 ’ 56 ’’ E, 500 m, 25. XI. 2009, fr., Ratovoson 1534 (G, K, MO, P, TAN); Betorabato, Abotorabatorano, 21 ° 34 ’ 04 ’’ S 45 ° 34 ’ 34 ’’ E, 296 m, 13. I. 2011, fr., Razakamalala 6000 (G, MO, P, TAN); Ankomaka, [22 ° 36 ’ S 44 ° 02 ’ E], 26. I. 1952, fr., Service Forestier 4556 (P, TEF); forêt de Zombitsy, [22 ° 46 ’ S 44 ° 42 ’ E], 24. IV. 1953, fl., Service Forestier 7757 (P, TEF); env. de Tsimela, S d’Ambatoabo, [24 ° 56 ’ S 46 ° 38 ’ E], 28. IX. 1953, fl., Service Forestier 8523 bis (G, NY, MO, P, TEF); forêt de Zombitsy, [22 ° 46 ’ 30 ’’ S 44 ° 38 ’ 00 ’’ E], III. 1931, fl., Service Forestier 11931 (G, MO, P); Analavelona, [22 ° 40 ’ S 44 ° 10 ’ E], 25. XI. 1954, fr., Service Forestier 12089 (P, TEF); ibid. loco, 28. XI. 1954, fr., Service Forestier 12140 (P, TEF); Betioky, vallée Sakamarakely, [22 ° 11 ’ S 45 ° 10 ’ E], 14. XI. 1954, fr., Service Forestier 12362 (P); Zombitsy, [22 ° 46 ’ S 44 ° 42 ’ E], 7. XI. 1954, fl., Service Forestier 12428 (P [2 sheets], TEF); Ambovombe, Beanantara, [25 ° 03 ’ 30 ’’ S 46 ° 12 ’ 30 ’’ E], 17. IX. 1955, fl., Service Forestier 15656 (P, TEF); Ampolitsy, Beroroha, [21 ° 40 ’ 30 ’’ S 45 ° 10 ’ E], 16. IV. 1956, y. fr., Service Forestier 15800 (P, TEF); Amboasary, bush entre Bevilany et Ranomainty, [25 ° 00 ’ S 46 ° 34 ’ E], 1. XII. 1961, fr., Service Forestier 20481 (P [2 sheets], TEF); crête du massif de l’Angavo, [24 ° 50 ’ 30 ’’ S 45 ° 48 ’ 00 ’’ E], c. 500 m, 21. I. 1963, fr., Service Forestier 22487 (P, TEF); Zombitse, Andranolava, 13. V. 1954, ster., Service Forestier 27 - R- 224 (P); Kitranga, Ihosy, 25. VIII. 1955, ster., Service Forestier 43 - R- 230 (P); Analavelona, 7. IV. 1954, ster., Service Forestier 76 - R- 18 (P); forêt de Zombitsy, 26. II. 1955, ster., Service Forestier 86 - R- 161 (P); forêt d’Ihera, 25. IV. 1955, ster., Service Forestier 104 - R- 161 (P); ibid. loco, 9. IV. 1954, ster., Service Forestier 108 - R- 18 (P); Bejangoa, Ankazoabo, 25. V. 1954, ster., Service Forestier 112 - R- 13 (P); env. d’Ampandrandava (entre Bekily et Tsivory), [24 ° 05 ’ S 45 ° 42 ’ E], 1100 m, XI. 1942, fl., Seyrig 265 (P [2 sheets]); ibid. loco, Seyrig 265 B (MO, P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBFFFA0FE4383651A120CA6.taxon	materials_examined	Typus: MADAGASCAR. Prov. Antsiranana: Rég. SAVA, Vohémar, forêt littoral au S de Vohémar, [13 ° 21 ’ S 50 ° 00 ’ E], 1 - 19. XII. 1966, fr., Service Forestier 24318 A (holo: P [P 06168913]!; iso-: MO- 05056713!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBFFFA0FE4383651A120CA6.taxon	diagnosis	Diagnosis Abrahamia revoluta Randrian. & Lowry can be distinguished from its congeners by its glabrous leaves with undulate and revolute margins and semicraspedodromous secondary veins.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBFFFA0FE4383651A120CA6.taxon	description	Description Trees 15 - 20 m tall, 21.6 cm DBH; young twigs glabrous, lenticellate. Leaves opposite or subopposite; blade obovate, 3.2 - 9 3 1.4 - 3.8 cm, subcoriaceous, apex retuse, sometimes emarginate, margin revolute, undulate, base cuneate, both surfaces glabrous, venation semicraspedodromous, midvein prominent abaxially, secondary veins 15 - 30 pairs, more or less parallel, 2 - 4 (- 5) mm apart, raised on both leaf surfaces, tertiary veins not visible on both surfaces; petiole 5 - 13 mm long, 0.5 - 1 mm in diam., canaliculate on adaxial surface, usually glabrous. Inflorescence and flowers unknown. Infrutescence terminal. Fruits 2.5 - 2.7 3 1.3 - 1.5 cm, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBFFFA0FE4383651A120CA6.taxon	distribution	Distribution, ecology and phenology Abrahamia revoluta occurs in NE Madagascar from S of Vohémar to the southern part of the Masoala peninsula (Map 9). Fruiting material has been gathered between October and December.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBFFFA0FE4383651A120CA6.taxon	conservation	Conservation status Abrahamia revoluta has a minimum AOO of 12 km 2 and only one of its three extant subpopulation occurs within a protected area (Masoala) whereas the other two were collected in areas where future decline is expected in terms of quality of habitat. It has therefore been assigned a preliminary status of “ Endangered’ ’ [EN B 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBFFFA0FE4383651A120CA6.taxon	discussion	Notes Abrahamia revoluta most closely resembles A. ditimena, with which it shares obovate leaves with a retuse apex that are similar in size and the number of secondary veins. However, A. revoluta differs by its subcoriaceous (vs coriaceous) leaves, semicraspedodromous (vs craspedodromous) venation, revolute and undulate (vs entire and flat) leaf margins, obscure (vs raised and visible) tertiary venation on the lower leaf surface, terminal (vs terminal or axillary) infrutescences, and glabrous (vs velutinous) fruits.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBFFFA0FE4383651A120CA6.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Antsiranana: versant NW de la Montagne d’Ambre, forêt d’Ankarefo (Ankazobe), 19 - 21. X. 1954, y. fr., Service Forestier 11357 bis (P, MO). Prov. Toamasina: Maroantsetra, Masoala Peninsula, S of Ambanizana, 15 ° 40 ’ 24 ’’ S 49 ° 57 ’ 51 ’’ E, 110 - 260 m, 29. XI. 1994, ster., Vasey et al. 176 (MO, TAN).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBDFFADFE4383651B550856.taxon	materials_examined	Typus: MADAGASCAR. Prov. Mahajanga: Rég. Diana, Ambato forêt classée, c. 40 km N d’Ambanja, 13 ° 26 ’ S 48 ° 33 ’ E, 90 - 459 m, 23. XI. 1996, fr., Randrianaivo et al. 29 (holo-: MO- 6671436!; iso-: G!, MO- 6671435!, P [P 06775160]!, TAN [TAN 001998]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBDFFADFE4383651B550856.taxon	diagnosis	Diagnosis Abrahamia sambiranensis Randrian. & Lowry can be distinguished from other members of the genus by the combination of large (10 - 19 3 2.5 - 5.6 cm), coriaceous leaves with numerous (50 - 60) distinctly raised secondary veins and a folded apex in dry material, its long petiole (30 - 65 mm), and its large, glabrous fruits (2.5 - 3 cm long).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBDFFADFE4383651B550856.taxon	description	Description Trees 10 - 15 m tall, 15 - 18 cm DBH, bark with white latex; young twigs pubescent towards the tips. Leaves alternate, clustered toward the branch ends; blade oblanceolate or narrowly elliptic to lanceolate, 10 - 19.4 3 3.2 - 5.5 cm, coriaceous, apex acuminate, folded in dry material, margin undulate, base cuneate to broadly cuneate, adaxial surface glabrous, often glaucous, abaxial surface sparsely sericeous, more so on midvein, venation craspedodromous, midvein prominent abaxially, secondary veins 50 - 60 pairs, more or less parallel, 2 - 4 mm apart, raised on both surfaces, tertiary veins not visible on both surfaces; petiole (20) - 25 - 58 mm long, 2 - 3.5 mm in diam., canaliculate on adaxial surface, tomentose. Inflorescence axillary, a panicle, 2 - 9.5 cm long, very few flowers reaching maturity, axes pubescent. Male flowers 5 - merous; pedicel c. 0.5 - 1 mm long, pubescent; calyx lobes triangular to ovate, 1 3 1 mm, connate at base, adaxial surface glabrous, abaxial surface pubescent; corolla lobes ovate, 2.5 3 1.5 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent; stamens 5, filaments 1 mm long, anthers c. 1 3 1 mm, obloid to ovoid; disk cupuliform, glabrous; rudimentary ovary absent. Female flowers unknown. Fruits 2.5 - 3 3 1.5 - 1.8 cm, ellipsoid, pubescent when young, glabrescent, with fine longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBDFFADFE4383651B550856.taxon	distribution	Distribution, ecology and phenology Abrahamia sambiranensis is restricted to humid forests in the Sambirano region of NW Madagascar and extends into the northern part of the island (Map 9). It flowers in July and August, and fruits have been collected in October and November.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBDFFADFE4383651B550856.taxon	conservation	Conservation status With an EOO of 1,717 km 2, a minimum AOO of 28 km 2, and seven localities, with only a single subpopulation within a protected area (Galoko), A. sambiranensis exists at 7 locations, six of which are subjected to pressure from forest exploitation and land clearing; it is therefore assigned a preliminary conservation status of “ Vulnerable ” [VU B 1 ab (iii) + 2 ab (iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBDFFADFE4383651B550856.taxon	discussion	Notes Abrahamia sambiranensis vegetatively resembles A. elongata, but differs by having leaves that lack tertiary venation (vs present and raised on both surfaces in A. elongata) and whose apex is usually folded in pressed material (vs flat), and by its fruits that are glabrescent when mature (vs glabrous in A. elongata).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFBDFFADFE4383651B550856.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Antsiranana: Rég. Diana, Ambilomangodra, 13 ° 02 ’ 42 ’’ S 49 ° 09 ’ 11 ’’ E, 50 m, 14. VII. 2005, fl., Randrianaivo et al. 1201 (CNARP, MO, P, TAN); forêt de Kalabenono, 13 ° 37 ’ 27 ’’ S 48 ° 40 ’ 05 ’’ E, 369 m, 10. XI. 2006, fr., Razafitsalama et al. 1064 (G, MO, P, TAN); Galoko, [13 ° 33 ’ 45 ’’ S 48 ° 45 ’ 00 ’’ E], 29. VII. 1955, y. fr., Service Forestier 15748 (P, TEF); forêts à l’E d’Ampondrabe, 28. X. 1954, fr., Service Forestier 18931 (MO, P [2 sheets]); massif de l’Andavakoera, N d’Angodromena, [13 ° 06 ’ 30 ’’ S 49 ° 18 ’ 30 ’’ E], 1. X. 1958, fr., Service Forestier 18947 (G, MO, P [2 sheets], TEF); Ampandrikoera, 12. X. 1951, ster., Service Forestier 181 - R- 64 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFB3FFAAFE4383651B0D0D79.taxon	materials_examined	Typus: MADAGASCAR: sine loco, s. d., y. fr., Thouars s. n. (holo-: P [P 00580351]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFB3FFAAFE4383651B0D0D79.taxon	description	Description Trees 5 - 25 m tall, 11 - 50 cm DBH, bark with white or translucid latex; young twigs fer-ruginous-pubescent or with short, yellow indument, lenticellate, waxy. Leaves alternate or subopposite, sometimes verticellate; blades oblanceolate to obovate, sometimes obo-vate, 5 - 15.8 3 2.7 - 4.5 cm, subcoriaceous, apex emarginate, retuse or shortly acuminate, sometimes truncate, margin entire, base acute or cuneate, adaxial surface glabrous, abax-ial surface pubescent when young, later glabrescent, venation craspedodromous, midvein prominent abaxially, secondary veins 40 - 70 pairs, more or less parallel, 0.5 - 3 mm apart, prominent on both surfaces, tertiary veins raised on adaxial surface; petiole 8 - 20 mm long, canaliculate on adaxial surface and very often flattened at base, pubescent when young, glabrescent. Inflorescence terminal, sometimes axillary, a panicle or thyrse, 5 - 10 cm long, the axes flattened, covered with yellowish, ferruginous strigose indument. Male and female flowers 5 - or 6 - merous; pedicel 0 - 0.3 mm long, strigose or pubescent; calyx lobes deltate or narrowly deltate, 1 - 1.2 3 1 - 1.5 mm, imbricate, rarely valvate, adaxial surface glabrous, abaxial surface pubescent or ferruginous, persistent on fruit; corolla lobes ovate, 2 - 3 3 1 - 1.8 mm, imbricate or quincuncial, adaxial surface glabrous, abaxial surface pubescent; stamens 5 or 6. Male flowers with filaments 1 - 1.5 mm long, glabrous, anthers 0.8 - 1 mm long, ovoid, glabrous; disk 1 - 2 mm in diam., pubescent; ovary rudimentary. Female flowers with filaments c. 1 mm long, glabrous, anthers 0.2 - 0.5 mm long, ovoid, sterile, glabrous; disk 1.5 - 2 mm in diam., pubescent; ovary very widely ovoid, 1 - 1.5 3 1 - 1.5 mm, covered with sericeous indument, placentation subapical; style very shortly trifid (apparently sim-ple), apical, 0.5 - 1 mm long, pubescent, stigmas capitate. Fruits 2 - 3 3 0.8 - 1.5 cm, ellip-soid, slightly asymmetrical, velutinous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFB3FFAAFE4383651B0D0D79.taxon	vernacular_names	Vernacular names “ Ditimena’ ’, “ Hazombarorana’ ’, “ Rodrano’ ’ (Tanala), “ Sandramy’ ’, “ Sosoka’ ’, “ Tsimalazo’ ’, “ Varoa’ ’, “ Vodihazo’ ’ (Tanala).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFB3FFAAFE4383651B0D0D79.taxon	distribution	Distribution, ecology and phenology Abrahamia sericea occurs in eastern humid domain forest, extending from coastal areas to the escarpment (Map 8), where it can reach 800 - 1200 m elevation. It flowers from September through December, and sets fruit from December until May.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFB3FFAAFE4383651B0D0D79.taxon	conservation	Conservation status With an EOO of 178,062 km 2, a minimum AOO of 236 km 2, and 64 known localities, with subpopulations in numerous protected areas, A. sericea is assigned a preliminary conservation status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFB3FFAAFE4383651B0D0D79.taxon	discussion	Notes Abrahamia sericea and A. grandidieri are distinctive within the genus in having very closely spaced, parallel secondary veins, but A. sericea differs by the presence of indument on its disk and ovary, a character absent in A. grandidieri but shared with another species, A. thouvenotii, which, however, has very widely spaced, arcuate secondary veins.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FFB3FFAAFE4383651B0D0D79.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Antsiranana: Rég. Diana, Tsaratanana RNI, Mandrizavona village, 13 ° 52 ’ 33 ’’ S 48 ° 48 ’ 44 ’’ E, 700 m, 27. III. 2000, fr., Antilahimena et al. 429 (G, K, MO, NY, P, TAN); Ambodimanga Ramena, Tsaratanana Reserve, 13 ° 52 ’ 37 ’’ S 48 ° 51 ’ 02 ’’ E, 547 m, 15. XI. 2001, y. fr., Antilahimena et al. 817 (G, K, MO, P, TAN); Antalaha, forêt littorale Ambodipont, Masoala PN, 15 ° 44 ’ 45 ’’ S 50 ° 19 ’ 25 ’’ E, 3. X. 1996, bud, Bernard 360 (MO, P, TAN); Ambodimanga, Antsahabe, 13 ° 49 ’ 30 ’’ S 48 ° 46 ’ 59 ’’ E, 500 m, 7. X. 1998, fl., Birkinshaw et al. 527 (MO, P, TAN); Manongarivo RS, bras gauche de l’Ambatra, 13 ° 59 ’ S 48 ° 26 ’ E, 470 m, 5. VI. 2000, old fr., Gautier et al. 375 1 (G, MO, P, TAN); vallée de la Lokoho, [14 ° 26 ’ S 49 ° 49 ’ E], 75 m, 13. XII. 1948, bud, Humbert 22383 (G, MO, P); Sambava, vallée de l’Andalangy, [14 ° 18 ’ S 49 ° 43 ’ E], 200 - 800 m, 12 - 14. XI. 1950, bud & fl., Humbert 24164 (MO, P); Ambilobe, massif de Marivorahona au SW de Manambato, [13 ° 46 ’ 40 ’’ S 48 ° 59 ’ 30 ’’ E], 1750 - 2000 m, 18 - 26. III. 1951, fr., Humbert 25713 (MO, P); E of Ankaramy, Manongarivo RS, Antsatrotro, 14 ° 05 ’ S 48 ° 23 ’ E, 600 - 900 m, 26 - 30. III. 1993, fr., Malcomber et al. 2291 (G, K, MO, NY, P, TAN); Sambirano, forêt de l’Ankaizina, [14 ° 30 ’ S 48 ° 55 ’ E], 1000 m, VII. 1908, bud, Perrier de la Bâthie 5859 (P [2 sheets]); Antalaha, Tampolo, 2 km S of Befotaka, 14 ° 44 ’ S 50 ° 13 ’ E, 27. I. 2007, fr., Ranarivelo 1255 (CAS, G, MO); Ambodimanga-Ramena, le long de la piste vers Antsaharatsy, 13 ° 48 ’ 33 ’’ S 48 ° 45 ’ 07 ’’ E, 460 m, 18. IV. 2000, y. fr. & fr., Ratovoson 236 (P); Masoala PN, Ambohitralanana, [15 ° 33 ’ S 50 ° 07 ’ E], 24. I. 1953, fl., Réserves Naturelles 4942 (P, TEF); Masoala PN, Ampanavoana, [15 ° 33 ’ S 50 ° 07 ’ E], 6. XII. 1954, fl., Réserves Naturelles 6808 (P [2 sheets], TEF); bord d’un affluent de l’Andalangy, Sambava, [14 ° 18 ’ S 49 ° 43 ’ E], 350 m, 12. XII. 1950, bud, Service Forestier 810 (MO, P [3 sheets], TEF); Masoala PN, Ambohitralanana, 15 ° 33 ’ S 50 ° 07 ’ E, 10. II. 1954, fl., Réserves Naturelles 6618 (P, TEF); massif du Marivorahona, Ankijabe, [13 ° 46 ’ 40 ’’ S 49 ° 00 ’ 00 ’’ E], 1050 m, 26. III. 1951, fr., Service Forestier 3070 (P [2 sheets], TEF); Tsiroanimandidy-Ambanja, [13 ° 52 ’ 30 ’’ S 48 ° 21 ’ 00 ’’ E], 200 m, 26. IX. 1952, fl., Service Forestier 6113 (P [3 sheets], TEF); Andrambavato, [14 ° 15 ’ 00 ’’ S 50 ° 01 ’ 30 ’’ E], 100 m, 26. XI. 1954, fl., Service Forestier 12232 (P [2 sheets], TEF); forêt de Betafononana, Maromandia, [14 ° 53 ’ S 50 ° 16 ’ E], 0 - 50 m, 16. XI. 1954, fl., Service Forestier 12449 (P, TEF); Ambohitralanana, Antalaha, [former Masoala RN], [15 ° 33 ’ S 50 ° 07 ’ E], 0 - 50 m, 23. X. 1956, fl., Service Forestier 17438 [= Réserves Naturelles 8592] (P, TEF). Prov. Fianarantsoa: Fort-Carnot, [21 ° 51 ’ 30 ’’ S 47 ° 26 ’ 30 ’’ E], 1986, bud, Beaujard 12 (P); Rég. Tanala, [21 ° 51 ’ S 47 ° 26 ’ E], 1994, ster., Beaujard 441 (P); Mananjary, entre les bassins du Matilana et du Mananjary, [22 ° 26 ’ S 47 ° 55 ’ E], 800 m, X. 1911, fl., Perrier de la Bâthie 5851 (P [2 sheets]); Vohipeno, Ampasina-Maroala, [22 ° 21 ’ S 48 ° 11 ’ E], 0 - 100 m, 20. X. 1952, bud & fl., Service Forestier 6431 (P [2 sheets], TEF); Sahavana, Ft. Carnot, [21 ° 51 ’ 30 ’’ S 47 ° 26 ’ 30 ’’ E], 23. XII. 1952, fr., Service Forestier 6664 (P, TEF); Vangaindrano, Vohipaho, [23 ° 33 ’ 30 ’’ S 47 ° 30 ’ 30 ’’ E], 10 m, 15. XI. 1952, fl., Service Forestier 7107 (MO, P, TEF); Tolongoina, Mandriandry, [21 ° 34 ’ 30 ’’ S 47 ° 31 ’ 00 ’’ E], 500 m, 28. IX. 1952, bud, Service Forestier 7113 (P, TEF); Maromiandra, Manampatroana, [21 ° 41 ’ S 47 ° 31 ’ E], 300 m, 29. XI. 1952, bud, Service Forestier 7150 (P, TEF); Nosy Varika, Nosiarivo, [20 ° 30 ’ 30 ’’ S 48 ° 00 ’ 00 ’’ E], 500 m, 28. I. 1953, fr., Service Forestier 7302 (P, TEF); rive droite de Sahanana Ampasinambo, [20 ° 31 ’ S 48 ° 00 ’ E], 530 m, 14. XI. 1953, bud & fl., Service Forestier 9955 (P, TEF); Sakahana, Tolongoina, Ft. Carnot, [21 ° 31 ’ 30 ’’ S 47 ° 45 ’ 30 ’’ E], 878 - 1000 m, 17. XI. 1954, fl., Service Forestier 13653 (P, TEF); Sahavana, Ft. Carnot, [21 ° 51 ’ 30 ’’ S 47 ° 26 ’ 30 ’’ E], 31. XI. 1954, fl., Service Forestier 14511 (P, TEF); Ambalangavo, Vohipeno, [22 ° 21 ’ S 47 ° 40 ’ E], 0 - 100 m, 15. X. 1954, fr., Service Forestier 14929 (MO, P, TEF). Prov. Toamasina: Moramanga, Analamazaotra, 18 ° 55 ’ 11 ’’ S 48 ° 25 ’ 17 ’’ E, 100 - 1200 m, 28. II. 2000, fr., Andrianjafy 27 (MO, P, TAN); Vavatenina, Manakambahiny I, 17 ° 39 ’ 07 ’’ S 48 ° 58 ’ 14 ’’ E, 550 - 600 m, 21. X. 2002, bud, Andrianjafy et al. 293 (CNARP, MO, P, TAN); Maroantsetra, Rantabe, Mafaipoza, 15 ° 39 ’ 24 ’’ S 49 ° 34 ’ 53 ’’ E, 80 m, 21. II. 2002, fr., Antilahimena 872 (G, MO, P, TAN); Maroantsetra, along Andramofotsy river, 15 ° 07 ’ 30 ’’ S 49 ° 49 ’ 13 ’’ E, 99 m, 6. XI. 2003, bud, Antilahimena et al. 2264 (G, K, MO, P, TAN); Toamasina II, Fotsimavo, Ambodiriana, [17 ° 55 ’ S 49 ° 13 ’ E], 300 - 400 m, XII. 1962, fr., Bosser 16913 (MO, P, TAN); Nosy Mangabe, 15 ° 29 ’ S 49 ° 45 ’ E, 15 m, IV. 1990, ster., Carlson 149 (MO); ibid loco, Carlson 195 (MO); Ambatondrazaka, Ampitanonoka à Fotsialanana, [17 ° 46 ’ S 48 ° 54 ’ E], 1200 m, 19. I. 1945, fr., Cours 2465 (MO, P); Brickaville, itinéraire de Didy, [18 ° 07 ’ 00 ’’ S 48 ° 32 ’ 40 ’’ E], 100 - 1100 m, bud, Cours 4907 (P [4 sheets]); Moramanga, PK 163 - 164, W of Andasibe (Périnet), [18 ° 56 ’ S 47 ° 57 ’ E], 1105 m, 19. I. 1986, fr., Dorr et al. 4582 (MO, P, TAN); forêt d’Analamazaotra, [18 ° 55 ’ S 48 ° 25 ’ E], 1000 m, fl., Louvel 3 (P); Andasibe, Mantadia NP, [18 ° 49 ’ 30 ’’ S 48 ° 28 ’ 00 ’’ E], 930 - 1040 m, 10. I. 2003, y. fr., Pell et al. 621 (MO, NY, P, TAN); forêt d’Analamazotra, [18 ° 55 ’ 48 ’’ S 48 ° 25 ’ 48 ’’ E], s. d., fl. & fr., Perrier de la Bâthie 3 (P); bords des rivières, Ambatovola, [18 ° 53 ’ S 48 ° 34 ’ E], I. 1928, fr., Perrier de la Bâthie 18365 (P [2 sheets]); Zahamena PN, entre rivières Laindrotra et Saharahara, 17 ° 45 ’ 34 ’’ S 48 ° 42 ’ 04 ’’ E, 900 m, 9. III. 1990, fr., Rakotondrajaona 168 (MO, P, TAN); Analamazaotra RS, 18 ° 56 ’ S 48 ° 26 ’ E, 960 m, 15. VII. 1996, ster., A. Randrianasolo 404 (MO, P, TAN); Tampolo forestry station, 17 ° 17 ’ S 49 ° 23 ’ E, 5 m, 1. VIII. 1996, ster., A. Randrianasolo 468 (MO, P); Analamazaotra RS, 18 ° 56 ’ S 48 ° 26 ’ E, c. 900 m, 28. X. 1996, bud, A. Randrianasolo 513 (G, K, MO, NY, P, TAN); Mantadia PN, c. 11 km N of village of Andasibe, 18 ° 49 ’ 47 ’’ S 48 ° 25 ’ 56 ’’ E, 939 m, 10. I. 2003, ster., A. Randrianasolo 783 (MO, P, TEF); Vatomandry, Ambalabe, N of Marotoho, 19 ° 10 ’ 19 ’’ S 48 ° 37 ’ 26 ’’ E, 367 m, 26. XI. 2004, y. fr., A. Randrianasolo et al. 969 (G, K, MO, NY, P, TAN, WAG); Ambalabe, Antsofotra forest, 1 km NW of Mahatsara village, 19 ° 13 ’ 48 ’’ S 48 ° 37 ’ 57 ’’ E, 226 m, 3. II. 2011, ster., A. Randrianasolo 1341 (G, K, MO, P, TAN); Manakambahiny-Est, Amparafarabe, Zahamena PN, 17 ° 43 ’ 42 ’’ S 48 ° 40 ’ 27 ’’ E, 875 m, 2. XI. 2001, fl., S. Randrianasolo et al. 251 (CNARP, MO, P, TAN, TEF); Ambatovy, Andranovery, 18 ° 51 ’ 51 ’’ S 48 ° 18 ’ 49 ’’ E, 1005 m, 14. II. 2008, ster., S. Randrianasolo et al. 670 (MO, P, TAN); Manakambahiny-Est, Zahamena PN, forêt Alaindrotra, 17 ° 45 ’ 07 ’’ S 48 ° 43 ’ 01 ’’ E, 2. III. 2001, fr., Randrianjanaka et al. 595 (CNARP, MO, P, TAN, TEF); Zahamena PN, W de Mitanonoka, 17 ° 45 ’ 53 ’’ S 48 ° 54 ’ 01 ’’ E, 606 m, 26. XI. 2001, fl., Randrianjanaka et al. 654 (CNARP, G, MO, NY, P, TAN, TEF); Maroseranana, bord d’Iampanga, 18 ° 23 ’ 53 ’’ S 48 ° 49 ’ 50 ’’ E, 554 m, 3. II. 2012, fr., Ravelonarivo et al. 4323 (MO, P, TAN); Ambalabe, forêt de Vohibe, 19 ° 09 ’ 31 ’’ S 48 ° 34 ’ 40 ’’ E, 536 m, 19. II. 2009, ster., Razanatsima et al. 660 (G, MO, P, TAN); Ambendrantsara, Manakambahiny-Est, [17 ° 46 ’ S 48 ° 39 ’ E], 750 - 1512 m, 29. III. 1949, fr., Réserves Naturelles 1923 (P, TAN); Zahamena PN, Mangabe, 15. II. 1952, fr., Réserves Naturelles 3728 (P); Toamasina I, Zahamena PN, Ampasimalaza, [17 ° 42 ’ S 48 ° 54 ’ E], 22. V. 1952, fr., Réserves Naturelles 3732 (P [2 sheets]); Ambodiriana, Tamatave, [Betampona RN], [17 ° 56 ’ S 49 ° 17 ’ E], 20. XI. 1960, fr., Réserves Naturelles 11656 (P [2 sheets], TEF); Nosy Mangabe, 15 ° 30 ’ S 49 ° 46 ’ E, 0 - 330 m, 12 - 23. IV. 1988, ster., Schatz et al. 2131 (MO, TAN); ibid. loco, fl., Schatz et al. 2492 (MO, P, TAN); ibid. loco, 23 - 28. II. 1989, fr., Schatz 2639 (MO, P, TAN); Analamazaotra, [18 ° 56 ’ S 48 ° 26 ’ E], 1000 m, 21. III. 1949, fr., Service Forestier 561 (P, TEF); ibid. loco, 3. IV. 1957, fr., Service Forestier 3723 (P, TEF); bords d’un cours d’eau entre Amboditavolo et le col d’Antandrokolaka, [15 ° 46 ’ S 49 ° 32 ’ E], 350 m, 13. II. 1954, fr., Service Forestier 9015 (MO, P, TEF); Andriantantely, [18 ° 44 ’ S 48 ° 38 ’ E], 200 - 300 m, 19. X. 1954, bud, Service Forestier 12620 (P, TEF); Farankaraina, Maroantsetra, [15 ° 25 ’ S 49 ° 52 ’ E], 20 - 25 m, 2. IV. 1955, fr., Service Forestier 14361 (P [2 sheets], TEF); Fénérive-Est, Ambodiampana, [17 ° 03 ’ S 49 ° 21 ’ E], 100 m, 16. X. 1955, bud, Service Forestier 15645 (P, TEF); forêt d’Analamazaotra, [18 ° 56 ’ S 48 ° 26 ’ E], 6. XI. 1957, fl., Service Forestier 18397 (MO, P, TEF); ibid. loco, 1000 m, 22. IX. 1966, ster., Service Forestier 26018 (P); canton Manakambahiny-Est, Mandrivo Morafeno, [17 ° 46 ’ S 48 ° 39 ’ E], 1000 m, 9. II. 1967, fr., Service Forestier 26286 (P, TEF). Prov. Toliara: Commune d’Enaniliha, forêt entre Eminiminy et le pic Trafonaomby, Andohahela PN, parcelle 1, 24 ° 38 ’ 03 ’’ S 46 ° 45 ’ 56 ’’ E, 445 m, 25. X. 2009, fl., Bernard et al. 1474 (MO, P, TAN); Ampitanonoka à Fotsialana, [17 ° 46 ’ S 48 ° 54 ’ E], 1200 m, 19. I. 1945, fr., Homolle 2465 (P); Andohahela PN, parcelle 1, NW of Eminiminy, 24 ° 38 ’ S 46 ° 46 ’ E, 500 - 1000 m, 6 - 13. II. 1993, fr., Malcomber et al. 2174 (G, K, MO, NY, P, TAN, WAG); Andohahela PN, parcelle 1, 24 ° 37 ’ 33 ’’ S 46 ° 45 ’ 55 ’’ E, 440 m, 25. X. 1995, fl., Rakotomalaza et al. 471 (G, K, MO, NY, P, TAN); PN Andohahela, parcelle 1, à proximité d’Eminiminy, 24 ° 40 ’ S 46 ° 48 ’, 200 - 700 m, 13 - 25. I. 1993, fr., Randriamampionona 37 (MO, P, TAN); Commune Emagnobo, Malamamay, [24 ° 36 ’ 27 ’’ S 47 ° 00 ’ 52 ’’ E], 20. X. 2016, fl., A. Randrianasolo 1665 (G, K, MO, NY, P, TAN); Commune Bevoay, forêt Mahabo, 24 ° 30 ’ 00 ’’ S 47 ° 05 ’ 14 ’’ E, 144 m, 18. XI. 2009, fl. & bud, Razakamalala 4706 (G, MO, P, TAN); Commune Bevoay, Androangabe, forêt d’Ampitanitanony, [24 ° 30 ’ 39 ’’ S 47 ° 05 ’ 08 ’’ E], 238 m, 12. I. 2016, fr., Razanatsima et al. 1442 (MO, P, TAN).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4AFF51FE4383651AB50DD6.taxon	materials_examined	Typus: MADAGASCAR. Prov. Antsiranana: Rég. Diana, NW, près de Ramena, 12 ° 14 ’ S 49 ° 21 ’ E, 9. XI. 1961, Service Forestier 20083 (holo-: P [P 06169101]!; iso-: G!, MO- 6684137!, NY!, P [P 00722667]!, TEF [TEF 000887]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4AFF51FE4383651AB50DD6.taxon	diagnosis	Diagnosis Abrahamia suarezensis Randrian. & Lowry can easily be distinguished from other members of the genus by its small, narrow oblanceolate leaves (2.5 - 6.5 3 0.7 - 2.5 cm) with ascending secondary veins that form an acute angle with the midvein at the leaf base.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4AFF51FE4383651AB50DD6.taxon	description	Description Shrubs or trees 1.5 - 20 m tall, 6 - 40 cm DBH; young twigs waxy, lenticellate and pubescent at the very tip. Leaves opposite to subopposite; blades oblanceolate to obovate, 2.5 - 7.5 3 0.7 - 3.5 cm, subcoriaceous to chartaceous, apex rounded, retuse, or emarginate, margin entire, undulate, base acute to cuneate, adaxial surface glabrous, abaxial surface with sparse sericeous indument when young, glabrescent, venation craspedodromous, midvein slightly prominent abaxially, secondary veins 9 - 12 pairs, 1 - 5 mm apart, not very prominent abaxially but well defined, tertiary veins obscure on both surfaces; petiole 4 - 10 mm long, canaliculate on adaxial surface, sericeous when young, glabrescent. Inflorescence axillary or terminal, a panicle (sometimes depauperate), 1.5 - 3.5 cm long, the axes flattened, sericeous. Male and female flowers with a pedicel 0.5 - 1 mm long, pubescent; calyx lobes very widely ovate, c. 1 3 1 mm, glabrous on both surface; corolla lobes ovate, 2.5 - 3 3 2 mm, imbricate, glabrous on both surfaces, yellow green or light yellow; stamens 4 or 5. Male flowers 5 - merous, filaments 1.5 - 2 mm long, glabrous, white, anthers 1 mm long, obloid to slightly ovoid, yellow, glabrous; disk 1 mm in diam., glabrous; ovary rudimentary. Female flowers 4 - merous, filaments c. 1.5 mm long, glabrous, white, anthers c. 0.8 mm long, ovoid, sterile, yellow, glabrous; disk annular, cupuliform, c. 1.5 mm in diam., glabrous; ovary very widely ovoid, 1 3 1 mm, glabrous, placentation apical or slightly subapical; style shortly trifid, latero-subapical or apical, 0.7 - 0.8 mm long, glabrous, stigmas capitate. Fruits 1.8 - 2.7 3 1 - 1.5 cm, ellipsoid, slightly asymmetrical, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4AFF51FE4383651AB50DD6.taxon	distribution	Distribution, ecology and phenology Abrahamia suarezensis is known only from far northern and northeastern Madagascar, from the vicinity of Antsiranana to the area around Daraina and extending along the east coast to S of Vohémar (Map 8). It appears to occur primarily in coastal sandy dune forests although several collections have been made in dry forest on other substrates in the Daraina area (i. e., to the W of Vohémar). Flowering material has been collected in October and November, and fruits are known from November and December.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4AFF51FE4383651AB50DD6.taxon	conservation	Conservation status With an EOO of 9,014 km 2, a minimum AOO of 116 km 2, and 35 recorded localities, with subpopulations in 3 protected areas (Loky Manambato, Oronjia, and Montagne des Français), representing c. 23 locations, A. suarezensis is assigned a preliminary conservation status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4AFF51FE4383651AB50DD6.taxon	discussion	Notes Abrahamia suarezensis most closely resembles A. patrickii, from which it differs by having leaves with fewer secondary veins (9 - 12 vs 24 - 28) that are widely (vs closely) spaced, and shorter petioles (4 - 10 vs 6 - 13 mm). Typical material of this species from dunes in the area around Antsiranana has narrowly obovate to oblanceolate leaves up to 5 cm long, but collections from farther South near Daraina and Mahavanona have more broadly obovate leaves that can reach 5 - 7 cm in length, as well as slightly larger fruits. While these differences appear to show some geographic structuring, we have chosen to refrain from recognizing the larger-leaved material from the South as a distinct taxon because no clear limits can be seen between the forms and certain specimens bear leaves spanning the range of variation in both shape and size.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4AFF51FE4383651AB50DD6.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Antsiranana: Vohémar, Ampisikina, Misoribe, 12 ° 58 ’ 10 ’’ S 49 ° 34 ’ 58 ’’ E, 179 m, 28. IX. 2013, fl., Bernard 2170 (MO, P, TAN); Antsiranana II, Ramena, Orangea, Mamelon vert, 12 ° 14 ’ 38 ’’ S 49 ° 22 ’ 04 ’’ E, 39 m, 17. XII. 2008, fr., Claude Christian et al. 14 (MO, P, TAN); Antsiranana, Ramena, forêt d’Orangea, 12 ° 15 ’ S 49 ° 23 ’ 30 ’’ E, 24. I. 2002, fr., De Block et al. 1378 (BR, G, K, MO, P, TAN); Vohémar, Daraina, forêt d’Ambohitsitondroina, 13 ° 08 ’ S 49 ° 27 ’ E, 180 m, 20. III. 2004, fr., Gautier et al. 4655 (G, MO, TAN); Vohémar, Ampasiramena, 12 ° 50 ’ 34 ’’ S 49 ° 49 ’ 44 ’’ E, 7 m, 12. XI. 2005, fl., Jullet Be et al. 260 (CNARP, MO, P, TAN); Daraina, Ambilondambo, 13 ° 09 ’ 55 ’’ S 49 ° 38 ’ 52 ’’ E, 292 m, 17. XI. 2005, fl., Jullet Be et al. 294 (CNARP, MO, P, TAN); Nosibe, forêt d’Analabe, Andavakabe, 13 ° 03 ’ 49 ’’ S 49 ° 54 ’ 02 ’’ E, 25 m, 4. XII. 2004, y. fr., Manjakahery et al. 58 (MO, P, TAN); ibid. loco, 13 ° 04 ’ 06 ’’ S 49 ° 54 ’ 24 ’’ E, 29 m, 8. XII. 2004, y. fr., Manjakahery et al. 76 (MO, P, TAN); ibid. loco, 13 ° 04 ’ 20 ’’ S 49 ° 54 ’ 04 ’’ E, 18 m, 15. XII. 2004, y. fr., Manjakahery et al. 94 (MO, P, TAN); Sakalava Bay, 12 ° 14 ’ S 49 ° 22 ’ E, 20 m, 26. XI. 1989, fl., McPherson 14563 (MO, P, TAN); Antsahalalina, 12 km E of Daraina, 14 ° 07 ’ S 49 ° 46 ’ E, 205 - 607 m, 24. II. 1990, fr., Meyers 45 (MO, P, TAN); Analamazava, SW of Daraina, [14 ° 73 ’ S 49 ° 38 ’ E], 200 - 1180 m, 23. III. 1990, ster., Meyers 73 (MO, TAN); ibid. loco, 26. IV. 1990, ster., Meyers 93 (MO, P, TAN); ibid. loco, 24. XI. 1990, fl., Meyers 204 (MO, P, TAN); ibid. loco, 19. II. 1991, fr., Meyers 273 (P, MO, TAN); Manatamena, 7 km NE of Daraina, 14 ° 60 ’ S 49 ° 42 ’ E, 112 - 330 m, IV. 1990, ster., Meyers et al. 106 (MO, P, TAN); Daraina, forêt d’Ampondrabe, 12 ° 58 ’ S 49 ° 42 ’ E, 199 m, 15. II. 2005, fr., Nusbaumer et al. 1474 (G, MO, TAN); Daraina, forêt d’Ambohitsitondroina, 13 ° 08 ’ S 49 ° 27 ’ E, 272 m, 20. I. 2006, fr., Nusbaumer et al. 1966 (G, MO, TAN); Ramena, forêt de l’Orangea, 12 ° 15 ’ 13 ’’ S 49 ° 21 ’ 44 ’’ E, 13 - 25 m, 3. IX. 2001, fl., Rabenantoandro et al. 627 (MO, P, TAN); forêt d’Anaborano, près du Lac Sahaka, 13 ° 04 ’ 42 ’’ S 49 ° 54 ’ 13 ’’ E, 25 m, 2. XI. 2002, fl., Rabenantoandro et al. 1095 (G, K, MO, P, TAN, TEF); Tsarabaria, forêt d’Ambondrobe, 13 ° 43 ’ 05 ’’ S 50 ° 05 ’’ 52 ’’ E, 13 m, 3. III. 2003, fr., Rabenantoandro et al. 1412 (G, MO, P, TEF); Daraina, E du village d’Ambondrombe, 13 ° 41 ’ 48 ’’ S 50 ° 05 ’ 18 ’’ E, 14. III. 2004, fr., Rabevohitra et al. 5149 (G, K, MO, P, TEF [2 sheets]); forêt d’Orangea, 12 ° 15 ’ S 49 ° 20 ’ E, 3. X. 2006, fl., Rakotonandrasana et al. 1026 (CNARP, MO, P, TAN); Ambolobozobe, Ambovomavo, 12 ° 31 ’ 13 ’’ S 49 ° 31 ’ 21 ’’ E, 24. XI. 2007, fr., Rakotonandrasana et al. 1208 (CNARP, G, MO, P, TAN); Mahavanona, forêt d’ Analabe, 12 ° 24 ’ 46 ’’ S 49 ° 22 ’ 25 ’’ E, 219 m, 15. II. 2005, fr., Rakotondrafara 295 (CNARP, G, MO, P, TAN); Andrafiabe, Anjialava, 12 ° 26 ’ 57 ’’ S 49 ° 30 ’ 48 ’’ E, 2 m, 13. XII. 2007, fr., Rakotondrafara et al. 519 (CNARP, G, MO, P, TAN); forêt E de Diego-Suarez, [12 ° 16 ’ 55 ’’ S 49 ° 17 ’ 40 ’’ E], X. 1975, fl., Rakotozafy 1529 (MO, TAN); Baie de Sakalava, 12 ° 16 ’ 39 ’’ S 49 ° 22 ’ 49 ’’ E, 21 m, 5. XI. 2006, fl., Ranaivojaona 1480 (G, MO, P, TAN); Orangea, 12 ° 14 ’ 12 ’’ S 49 ° 22 ’ 08 ’’ E, 33 m, 7. XI. 2006, fr., Ranaivojaona 1571 (G, MO, P, TAN); Ankorangana, forêt d’Andramadana, 12 ° 39 ’ 18 ’’ S 49 ° 32 ’ 11 ’’ E, 15 m, 22. XI. 2001, fr., Randrianaivo et al. 803 (MO, P, TAN); forêt d’Orangea, 12 ° 15 ’ 01 ’’ S 49 ° 21 ’ 40 ’’ E, 76 m, 8. XII. 2004, fr., Randrianaivo et al. 1131 (G, MO, P, TAN); Daraina, forêt de Bakaraoka Sud, 13 ° 11 ’ 13 ’’ S 49 ° 42 ’ 40 ’’ E, 191 m, 15. XI. 2005, fl., S. Randrianasolo et al. 551 (CNARP, MO, P, TAN); Daraina, forêt d’Ambilondomba, [13 ° 10 ’ S 49 ° 39 ’ E], 471 m, 27. I. 2004, fr., Ranirison et al. 326 (G, MO, TAN); Daraina, Befarafara, Solanampilaka, 13 ° 05 ’ 56 ’’ S 49 ° 35 ’ 29 ’’ E, 110 m, 16. XI. 2005, fl., Ratovoson et al. 1109 (CNARP, G, MO, P, TAN); fourrée d’Orangea, 12 ° 16 ’ 59 ’’ S 49 ° 23 ’ 03 ’’ E, 46 m, 30. XI. 2005, fl., Razafitsalama et al. 868 (CNARP, G, MO, P, TAN); Vohémar, Tsarabaria, forêt E du village Ambondrombe, 13 ° 43 ’ 08 ’’ S 50 ° 05 ’ 50 ’’ E, 13. III. 2004, fl., Razakamalala et al. 1006 (G, K, MO, P, TAN); Near village of Ankorikakely, 2 - 3 km NE Sakalava bay, 12 ° 16 ’ 40 ’’ S 49 ° 23 ’ 05 ’’ E, 48 m, 4. XI. 2006, fr., Rogers et al. 1089 (MO, P, TAN); Orangea, Ramena, 12 ° 14 ’ 13 ’’ S 49 ° 22 ’ 08 ’’ E, 10 m, 12. II. 2005, fr., Schatz et al. 4190 (MO, P, TAN); Vohémar, 27. XII. 1951, ster., Service Forestier 4348 (P); forêt d’Andranotsimaka, [12 ° 23 ’ 00 ’’ S 49 ° 07 ’ 40 ’’ E], 4. X. 1952, fl. & fr., Service Forestier 5788 (MO, P, TEF); forêt d’Orangea, [12 ° 16 ’ 55 ’’ S 49 ° 17 ’ 40 ’’ E], 18. X. 1954, fl., Service Forestier 11337 (G, MO, NY, P [2 sheets], TEF); Analabe, Vohémar, [14 ° 03 ’ S 49 ° 43 ’ E], 13. IX. 1955, fl., Service Forestier 15498 (G, MO, NY, P, TEF); dunes littorales, près de Ramena, [12 ° 14 ’ S 49 ° 21 ’ E], 9. XI. 1961, fl., Service Forestier 20368 (G, K, MO, MY, P [2 sheets], TEF); ibid. loco, Service Forestier 20369 (MO, P, TEF); forêt littorale, S de Vohémar, [13 ° 21 ’ S 50 ° 00 ’ E], 11 - 19. XII. 1966, bud & fl., Service Forestier 27279 (MO, P, TEF); ibid. loco, fr., Service Forestier 27280 (P); versant E du massif de l’Ankerana, N de Vohémar, [13 ° 17 ’ S 49 ° 51 ’ E], 50 - 450 m, 17. XII. 1966, fl., Service Forestier 27352 (P, TEF); forêt d’Orangea area, 12 ° 14 ’ S 49 ° 21 ’ E, 55 m, 23. X. 1993, fl., van Nek 1978 (TAN); ibid. loco, 12 ° 14 ’ S 49 ° 22 ’ E, 65 m, 23. X. 1993, fl., van Nek 1984 (G, MO, P, TAN).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4FFF5CFE4383631B7E0D26.taxon	materials_examined	Lectotypus (designated here): MADAGASCAR. Prov. Toamasina: Rég. Alaotra-Mangoro, Moramanga, Forêt d’Analamazaotra, [18 ° 56 ’ S 48 ° 26 ’ E], 1000 m, s. d., fr., Thouvenot 80 (P [P 00580346]!; isolecto-: MO- 6684138!, P [P 00580347]!, TAN [TAN 000076]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4FFF5CFE4383631B7E0D26.taxon	description	Description Trees 5 - 20 m tall, 5 - 54 cm DBH, bark with white milky latex; young twigs ferruginous pubescent. Leaves alternate; blades oblanceolate to obovate or elliptic to narrowly elliptic or narrowly oblong, 3 - 15 3 1.5 - 5 cm, coriaceous, apex emarginate, retuse, or shortly acuminate, sometimes truncate, margin entire, base acute to cuneate or obtuse, adaxial surface ferruginous pubescent when young, later glabrescent or pubescent only on the midvein, abaxial surface pubescent, venation craspedodromous, midvein prominent abaxially, secondary veins 10 - 20 pairs, arcuate, 2 - 10 (- 15) mm apart, prominent on both surfaces, tertiary veins raised on both surfaces; petiole 5 - 15 mm long, canaliculate on adaxial surface, pubescent, sometimes glabrescent. Inflorescence axillary or terminal, a panicle, 3 - 8 cm long, axes terete, tomentose. Male and female flowers 5 - merous; pedicel 0 - 0.3 mm long, pubescent; calyx lobes deltate to ovate, 1.2 - 1.5 3 1 - 1.5 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent, persistent on fruit; corolla lobes ovate, 2 - 3 3 1 - 1.2 mm, imbricate, adaxial surface pale yellow, glabrous, abaxial surface brown, pubescent; stamens 5. Male flowers with filaments 1 mm long, glabrous, white, anthers 0.6 - 0.8 mm long, ovoid, yellow, glabrous; disk 1 - 1.2 mm in diam., pubescent; ovary rudimentary. Female flowers with filaments c. 0.8 - 1 mm long, glabrous, white, anthers c. 0.5 - 0.6 mm long, subglobose or ovoid, sterile, yellow, glabrous; disk 1.5 - 2 mm in diam., pubescent; ovary widely to very widely ovoid, c. 1 - 1.2 3 1 mm, pubescent, placentation subapical; style very shortly trifid, apical, 0.3 - 0.4 mm long, covered with indument. Fruits 1.5 - 2.5 3 0.8 - 1.5 cm, ellipsoid, very often glabrate or glabrescent, rarely completely glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4FFF5CFE4383631B7E0D26.taxon	vernacular_names	Vernacular names “ Ditimena’ ’, “ Menavahatra’ ’, “ Tandraviravy’ ’ (North East).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4FFF5CFE4383631B7E0D26.taxon	distribution	Distribution, ecology and phenology Abrahamia thouvenotii occurs mainly in eastern Madagascar and the Sambirano region (Map 10). It is found principally in mid-elevation rainforests at an elevation between 800 and 900 m, although a few subpopulations have been recorded at much higher elevations. Flowers have generally been collected from August to October, although some individuals may flower as early as July. Fruits have been recorded between November and January.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4FFF5CFE4383631B7E0D26.taxon	conservation	Conservation status With an EOO of 116,472 km 2, a minimum AOO of 180 km 2, and 46 recorded localities, with subpopulations in at least 10 protected areas (Anjanaharibe-Sud, Makira, Manon-garivo, Mantadia, Marojejy, Masoala, Montagne d’Ambre, Analamazaotra, Ranomafana, and Zahamena), A. thouvenotii is assigned a preliminary conservation status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4FFF5CFE4383631B7E0D26.taxon	discussion	Notes Abrahamia thouvenotii can be distinguished from other members of the genus by the presence of widely spaced secondary leaf venation and indument on the ovary and disk. Three collections from northern and northeastern Madagascar (Miller 3625, Service Forestier 15067 and Ursch 195) differ from typical material by having flowers with a gla-brous disk. Additional collections from this area, especially from female individuals, will be needed, however, to determine whether they may represent a distinct taxon. Until such time, we have chosen to include them in our circumscription of A. thouvenotii. PERRIER DE LA BâTHIE (1944) cited two collections in the protologue of Protorhus thouvenotii, one in bud (Thouvenot 7) and the other in fruit (Thouvenot 80). We have designated a sheet of Thouvenot 80 as the lectotype because it is represented by two additional specimens, whereas there is only a single sheet of Thouvenot 7. Protorhus lecomtei (≡ Abrahamia lecomtei) was based on two collections that belong to different taxa, one of which (Perrier de la Bâthie 5283) we have placed in A. thouvenotii. As indicated above under A. lecomtei, we have designated a specimen of the other syntype gathering, Perrier de la Bâthie 14301, as the lectotype in order to retain a circum-scription that includes the additional material assigned by PERRIER DE LA BâTHIE (1944) to Protorhus lecomtei.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF4FFF5CFE4383631B7E0D26.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Antananarivo: Anjozorobe, 2 km NNE d’Andranomay, 18 ° 28 ’ 08 ’’ S 47 ° 57 ’ 03 ’’ E, 1340 - 1400 m, 20. XII. 1996, fr., Birkinshaw et al. 373 (G, K, MO, P, TAN). Prov. Antsiranana: Ambilobe, forêt de Kalabenono, 13 ° 38 ’ 43 ’’ S 48 ° 40 ’ 41 ’’ E, 971 m, 27. XI. 2006, fr., Callmander et al. 643 (G, NY, MO, P, TAN); Marojejy PN, versant S de Beondroka, 14 ° 25 ’ S 49 ° 50 ’ E, 600 m, 14. XI. 1989, bud, Deroin & Badré 96 (MO, P, TAN); Ambanja, Antsatrotro (Manongarivo), [14 ° 05 ’ S 48 ° 23 ’ E], 800 m, XI. 1954, fr., Service Forestier 11505 (P [3 sheets], TEF); Andapa, SW edge of Anjanaharibe-Sud Reserve, 14 ° 48 ’ 15 ’’ S 49 ° 26 ’ 45 ’’ E, 1000 - 1100 m, 6. VIII. 1997, bud, McPherson 17266 (G, K, MO, P, TAN); Marojejy PN, NW Manantenina, Campement n ° 3, 14 ° 26.2 ’ S 49 ° 44.5 ’ E, 1200 m, 1. XI. 1996, fl., Messmer et al. 305 (G, MO, P, TAN); Montagne d’Ambre, 12 ° 32 ’ S 49 ° 10 ’ E, 1050 m, 21. X. 1988, fl., Miller 3625 (MO [2 sheets], P, TAN); Ambilobe, Antanambao Belinta village, Kalabinono, 13 ° 39 ’ 22 ’’ S 48 ° 40 ’ 11 ’’ E, 1004 m, 3. XI. 2007, fr., Rakotovao 3897 (G, MO, P, TAN); Marojejy PN, env. d’Andohan’Antsahamberokely et Marojejybe, 14 ° 25 ’ 45 ’’ S 49 ° 40 ’ 20 ’’ E, 1672 m, 7. III. 1994, fr., Rasoavimbahoaka 176 (MO, P, TAN); Andapa-Bealanana, Befingotra, 14 ° 12 ’ S 49 ° 27 ’ E, 1100 m, 13 - 18. IX. 1996, fl., Ravelonarivo et al. 1029 (G, K, MO, P, TAN); Marojejy PN, between Campsite II and Campsite III, 14 ° 25 ’ 09 ’’ - 14 ° 25 ’ 24 ’’ S 49 ° 44 ’ 46 ’’ - 49 ° 45 ’ 29 ’’ E, 1280 - 1350 m, 23. XI. 2003, fl., Schmidt et al. 4308 (G, K, MO, P, TAN); Antalaha, Ambodigavo-Maromandia, [14 ° 51 ’ S 50 ° 07 ’ E], 120 m, 11. IX. 1955, bud, Service Forestier 15067 (P [2 sheets], TEF); forêt d’Ambre, [12 ° 36 ’ S 49 ° 09 ’ E], 16. VI. 1954, ster., Service Forestier 160 - R- 152 (P); Diego Suarez, [12 ° 16 ’ S 49 ° 17 ’ E], 17. X. 1927, bud, Ursch 195 (P). Prov. Fianarantsoa: Ifanadiana, 7 km W of Ranomafana, 21 ° 16 ’ S 47 ° 25 ’ E, 1000 m, 19. X. 1987, ster., Daniels 95 (MO, P, UMO); Ranomafana PN, parcelle 3, Talatakely, Piste C, [21 ° 15 ’ S 47 ° 27 ’ E], 1645 m, 3 - 8. IX. 1993, fl., Kotozafy et al. 230 (G, K, MO, NY, P, TAN, WAG); Ranomafana PN, Vatoharanana, Parcelle 3, 21 ° 16 ’ S 47 ° 26 ’ E, 900 - 1100 m, ster., Kotozafy et al. 544 (MO, P, TAN); Ranomafana PN, parcelle 3, 21 ° 16 ’ S 47 ° 26 ’ E, 980 m, 16 - 19. II. 1994, ster., Rakoto et al. 514 (MO, P, TAN); Ranomafana PN, Talatakely area, 21 ° 16 ’ S 47 ° 25 ’ E, 800 - 900 m, 8. X. 1996, ster., A. Randrianasolo 497 (MO, P, TAN); Ranomafana NP, parcelle 3, 21 ° 15 ’ 30 ’’ S 47 ° 25 ’ E, 900 - 1100 m, 23. IX. 1994, fl., J. Randrianasolo et al. 154 (MO, P, TAN); ibid. loco, J. Randrianasolo et al. 155 (MO, P, TAN); ibid. loco, J. Randrianasolo et al. 157 (MO, P, TAN); Andrambovato, E de Fianarantsoa, [21 ° 31 ’ S 47 ° 25 ’ E], 878 - 1000 m, 24 - 25. I. 1955, ster., Service Forestier 11587 (P, TEF); ibid. loco, Service Forestier 13260 (P); Ranomafana PN, Talatakely trail system, 21 ° 15 ’ 30 ’’ S 47 ° 25 ’ 00 ’’ E, 950 - 1150 m, 23. IX. 1993, fl., Solo & J. Randrianasolo 8 (MO, P, TAN); ibid. loco, 16. VIII. 1993, fl., Turk et al. 527 (G, K, MO, NY, P, TAN, WAG); Ranomafana PN, parcelle I, [21 ° 14 ’ S 14 ° 27 ’ E], 850 m, 29. IX. 1994, fl., Turk et al. 637 (MO, P, TAN). Prov. Mahajanga: Anjanaharibe-Sud RS, [14 ° 42 ’ S 49 ° 27 ’ E], 1100 m, 14. XII. 1994, fr., Ravelonarivo et al. 552 (MO, P, TAN). Prov. Toamasina: Vavatenina, Zahamena PN, 17 ° 38 ’ 27 ’’ S 48 ° 52 ’ 32 ’’ E, 700 - 1040 m, 4. X. 2001, fl., Andrianjafy et al. 194 (CNARP, MO, P, TAN); Masoala PN, Ambohitsitondroina, 15 ° 25 ’ 37 ’’ S 45 ° 57 ’ 24 ’’ E, 1158 m, 10. X. 2003, bud, Antilahimena et al. 2169 (G, MO, P, TAN); Commune Vinanibe, Makira forest, 15 ° 27 ’ 40 ’’ S 49 ° 17 ’ 46 ’’ E, 1153 m, 28. XI. 2003, fl., Antilahimena 2421 (G, K, MO, NY, P, TAN); Maroantsetra, Ambatofotsy, Andampy forest, 15 ° 19 ’ 03 ’’ S 49 ° 24 ’ 06 ’’ E, 800 m, 4. IX. 2004, fr., Antilahimena 2746 (G, MO, P, TAN); Ambatovy, Sahavarina forest, 18 ° 51 ’ 37 ’’ S 48 ° 20 ’ 25 ’’ E, 1020 m, 24. XI. 2005, fr., Antilahimena et al. 4230 (G, MO, P, TAN); Ambatovy, Ampangadiantrandraka Forest, 18 ° 51 ’ 43 ’’ S 48 ° 17 ’ 39 ’’ E, 1049 m, 9. XII. 2006, fr., Antilahimena & Edmond 5087 (G, MO, P, TAN); forêt d’Analamay, 18 ° 49 ’ 37 ’’ S 48 ° 19 ’ 11 ’’ E, 1153 m, 6. VI. 2007, ster., Bernard et al. 529 (MO, P, TAN); Mantadia PN, 700 m N from point PK 13, 18 ° 48 ’ 09 ’’ S 48 ° 25 ’ 39 ’’ E, 870 - 930 m, 23. VI. 1996, y. fl., Birkinshaw et al. 313 (MO, P, TAN); moyenne vallée du Mangoro, [18 ° 53 ’ S 48 ° 07 ’ E], 10. IX. 1942, fl., Decary 18516 (MO, P); Analamazaotra, [18 ° 56 ’ S 48 ° 26 ’ E], X. 1925, bud, Louvel 44 (P); ibid. loco, 1925, bud, Louvel 112 (P); Mantadia PN, 10. I. 2003, ster., Pell et al. 633 (MO, NY, P, TAN); forêt d’Analamazaotra, [18 ° 56 ’ S 48 ° 26 ’ E], 800 m, s. d., fr., Perrier de la Bâthie 5283 (P [4 sheets]); Ambatondrazaka, Commune Didy, Fokontany Antsevabe, 17 ° 59 ’ S 48 ° 36 ’ E, 1165 m, 28. IX. 2005, fl., Rakotonasolo 1066 (G, K, MO, P, TAN); Amparafaravola, E d’Ambodisaina, 17 ° 16 ’ 49 ’’ S 48 ° 49 ’ 18 ’’ E, 850 m, 11. XII. 2001, fr., Rakotondrafara et al. 102 (CNARP, MO, P, TAN); 1.5 km E du campement du PK 14, piste de Mantadia PN, 18 ° 49 ’ S 48 ° 26 ’ E, 940 - 1012 m, 21. IX. 1996, fl., Ralimanana et al. 10 (G, MO, P, TAN); Lakato, village Agnalatsara, 19 ° 07 ’ 29 ’’ S 48 ° 23 ’ 32 ’’ E, 815 m, 1. VI. 2007, ster., Ranaivojaona et al. 1724 (MO, P, TAN); Analamazaotra RS, [18 ° 56 ’ S 48 ° 26 ’ E], 950 m, 15. VII. 1996, ster., A. Randrianasolo 413 (MO, P, TAN); Analamazaotra RS, along the Indri trail, about 300 m from observation point, 18 ° 56 ’ 12 ’’ S 48 ° 25 ’ 10 ’’ E, 1000 m, 10. I. 2003, ster., A. Randrianasolo 779 (MO, TEF); Analamazaotra forestry station, 18 ° 56 ’ S 48 ° 26 ’ E, 950 m, 19. X. 1996, fl., A. Randrianasolo 506 (MO, P, TAN); ibid. loco, 18. VII. 1996, ster., A. Randrianasolo 433 (MO, P, TAN); ibid. loco, 16. VII. 1996, ster., A. Randrianasolo 414 (MO, P, TAN); Zahamena PN, forêt communautaire d’Antenina, 17 ° 32 ’ 32 ’’ S 48 ° 43 ’ 20 ’’ E, 1250 m, 6. XII. 2002, fr., Randrianjanaka et al. 762 (CNARP, G, MO, NY, P, TAN, TEF); Ambodimangavolo, Ambatoharanana, Moango, 17 ° 33 ’ 57 ’’ S 48 ° 53 ’ 58 ’’ E, 740 - 1200 m, 25. X. 2005, fl., Ratovoson et al. 336 (CNARP, G, MO, P, TAN, TEF); Ambohibary, Ambatovy, 18 ° 52 ’ 28 ’’ S 48 ° 19 ’ 19 ’’ E, 1036 m, 12. XII. 2011, fr., Ravelonarivo et al. 3972 (G, K, MO, NY, P, TAN); entre Analamay et Ambatovy, 18 ° 49 ’ 52 ’’ S 48 ° 18 ’ 45 ’’ E, 1192 m, 22. XI. 2005, fl., Razafindraibe et al. 104 (MO, P, TAN); Ambatovy, forêt de cuirasse de Vatovy, 18 ° 51 ’ 21 ’’ S 48 ° 18 ’ 29 ’’ E, 1104 m, 1. XII. 2005, fr., Razanatsoa et al. 194 (MO, NY, P, TAN); Masoala Peninsula, SE of Ambanizana, 15 ° 38 ’ S 49 ° 59 ’ E, 150 - 170 m, 1. XI. 1992, bud, Schatz et al. 3401 (G, MO, P, TAN); Sahamaloto, Périnet, [18 ° 56 ’ S 48 ° 26 ’ E], 1000 m, 11. X. 1952, fl., Service Forestier 6052 (P [2 sheets], TEF); massif de l’Ambohitsitondroina de Mahalevona, NW de la presqu’île Masoala, [15 ° 25 ’ S 49 ° 58 ’ E], 900 m, 5. XII. 1953, bud, Service Forestier 8714 (G, MO, P, TEF); Sahamaloto, Périnet, [19 ° 03 ’ S 48 ° 12 ’ E], 1054 m, 20. IV. 1954, y. fr., Service Forestier 10359 (P, TEF); Amparafaravola, forêt d’Andranobe, W de Manohilahy, [17 ° 46 ’ S 47 ° 58 ’ E], 1000 - 1200 m, 31. VIII. 1958, fl., Service Forestier 18772 (MO, P [2 sheets], TEF); vestiges de forêts près du village d’Ankarahara (PK 100 - 102 Antananarivo-Moramanga), Mangoro, [18 ° 54 ’ S 48 ° 09 ’ E], 800 m, 8. VIII. 1961, bud, Service Forestier 20337 (MO, P [3 sheets], TEF); Analamazaotra, [18 ° 56 ’ S 48 ° 26 ’ E], 1000 m, 21. V. 1962, ster., Service Forestier 21142 (MO, P); ibid. loco, 25. V. 1962, ster., Service Forestier 21143 (MO, P); Andasitsimanga, Périnet, [18 ° 53 ’ 30 ’’ S 48 ° 22 ’ 30 ’’ E], 800 - 1000 m, 2. VIII. 1962, bud, Service Forestier 21224 (MO, P); Perinet, [18 ° 56 ’ S 48 ° 26 ’ E], 1000 m, II. 1966, ster., Service Forestier 25720 (P); ibid. loco, Service Forestier 25721 (P); ibid. loco, Service Forestier 25722 (P); ibid. loco, 29. VI. 1966, ster., Service Forestier 25960 (P); ibid. loco, N. de la pisciculture, III. 1966, ster., Service Forestier 25778 (P); Ankazomanitra, PK 45 route Moramanga-Anosibe, [18 ° 56 ’ 20 ’’ S 48 ° 13 ’ 40 ’’ E], 900 m, 9. XI. 1968, y. fr., Service Forestier 26830 (P, TEF); forêt W d’Antanandava (PK 45 de la route Moramanga-Anosibe), [19 ° 17 ’ S 48 ° 12 ’ E], 900 m, XI. 1968, fr., Service Forestier 28443 (P); Sahamaloto, Périnet, [19 ° 03 ’ S 48 ° 12 ’ E], 1054 m, 20. VII. 1952, bud, Service Forestier 45 B-R- 172 (P, TEF); ibid. loco, 27. IX. 1952, bud, Service Forestier 165 B-R- 172 (P, TEF); Ampatsakambe, 28. V. 1950, ster., Service Forestier 189 B-R- 56 (P).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF41FF59FE4383651A9D0F39.taxon	materials_examined	Typus: MADAGASCAR. Prov. Fianarantsoa: Rég. Vatovavy-Fitovinany, Ranomafana National Park, parcelle 3, S of National road 25 at 7 km W of Ranomafana, Talatakely trail system, 21 ° 15 ’ 30 ’’ S 47 ° 25 ’ 00 ’’ E, 950 - 1150 m, 20. X. 1993, fl., Solo & J. Randrianasolo 35 (holo-: MO- 6671437!; iso-: G, K, MO- 6671438!, P [P 06168911]!, TAN [TAN 002002]!, WAG).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF41FF59FE4383651A9D0F39.taxon	diagnosis	Diagnosis Abrahamia turkii Randrian. & Lowry can be distinguished from other members of the genus by its coriaceous leaves, inflorescence with thick axes and ferruginous indument, and fruits that are small in size (1.8 - 2 3 1 cm) and glabrous.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF41FF59FE4383651A9D0F39.taxon	description	Description Trees 11 - 30 m tall, 18 - 100 cm DBH, bark with white latex; young twigs sericeous on tips, lenticellate farther from the tip. Leaves usually alternate, sometimes subopposite; blade obovate, 2.5 - 8 3 1.5 - 3.5 cm, coriaceous, apex usually emarginate or retuse, sometimes rounded, margin slightly revolute, sometimes undulate, base cuneate, adaxial surface sparsely pubescent, abaxial surface sericeous or strigose, venation craspedodromous, midvein prominent abaxially, secondary veins 14 - 24 pairs, 1 - 4 mm apart, raised on both surfaces, tertiary veins visible on both surfaces; petiole 0.5 - 1 cm long, 1 - 1.5 mm in diam., pubescent or scabrous, canaliculate on adaxial surface. Inflorescence axillary or terminal, a panicle, erect, axes thick, 1 - 5 cm long, reddish brown pubescent. Male flowers 5 - merous, c. 3 - 4 mm long; pedicel c. 0.3 - 0.5 mm long, reddish brown pubescent; calyx lobes triangular, 1.5 - 2 3 1 - 2 mm, adaxial surface glabrous, abaxial surface, pubescent; corolla lobes ovate, 2 - 4 3 2 mm, adaxial surface glabrous, abaxial surface pubescent; stamens 5, filament 1 - 2 mm long, glabrous, anthers 1 mm long, ovoid to obloid, glabrous; disk slightly cupuliform, c. 2 mm in diam., glabrous; ovary very rudimentary or absent. Female flowers unknown. Fruits 1.8 - 2 3 1 cm, ovoid, glabrous, wrinkled when dry (longitudinal striations not as evident as in other species).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF41FF59FE4383651A9D0F39.taxon	distribution	Distribution, ecology and phenology Abrahamia turkii is distributed mainly in south-central and southern Madagascar, although one collection has been made at the Ambohitantely forest, far to the north of the other recorded subpopulations (Map 10). Flowering material has been recorded in October and November, and fruits have been collected in January.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF41FF59FE4383651A9D0F39.taxon	conservation	Conservation status With an EOO of 85,992 km 2, a minimum AOO of 40 km 2 but an estimated value well exceeding 2,000 km 2, and 9 localities, with subpopulations in 6 protected areas (Ambohitantely, Andohahela, Betampona, Kalambatritra, Pic d’Ivohibe, and Ranomafana), representing 9 locations, only some of which may be subjected to pressure from forest exploitation and land clearing, A. turkii is assigned a preliminary conservation status of “ Least Concern ” [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF41FF59FE4383651A9D0F39.taxon	discussion	Notes Abrahamia turkii closely resembles A. ditimena by its medium sized, coriaceous leaves with craspedodromous venation, usually emarginate or retuse apex, and tertiary venation that is visible on both surfaces, and its paniculate inflorescence, but it can easily be distinguished by its short (1 - 5 vs 2 - 10 cm) inflorescence covered with reddish brown pubescence (vs glabrescent or tomentose), and its glabrous (vs velutinous) fruits.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF41FF59FE4383651A9D0F39.taxon	materials_examined	Paratypi MADAGASCAR. Prov. Antananarivo: forêt d’Ambohitantely, 18 km NW of Ankazobe, [18 ° 10 ’ S 47 ° 18 ’ E], 1400 - 1646 m, 21. XI. 1985, bud, Barnett et al. 533 (MO, P, TAN). Prov. Fianarantsoa: Ivohibe RS, piste II et III, 22 ° 29 ’ 00 ’’ S 46 ° 58 ’ 06 ’’ E, 1275 m, 20. X. 1997, fl., Rakotomalaza et al. 1434 (G, MO, P, TAN); Ivohibe RS, 5.5 km SE d’Angondongodona, camp 4, 22 ° 25 ’ 18 ’’ S 46 ° 53 ’ 54 ’’ E, 1150 m, 4. XI. 1997, fl., Rakotomalaza et al. 1494 (G, K, MO, P, TAN); Haute Matsiatra, Ialatsara forest, 21 ° 03 ’ 07 ’’ S 47 ° 12 ’ 42 ’’ E, 12. X. 2005, fl., Ranarivelo et al. 719 (CAS, MO); Ranomafana PN, parcelle 3, Vatoharana trail system, 21 ° 17 ’ S 47 ° 26 ’ E, 1100 - 1250 m, 18. XI. 1993, fl., Turk & J. Randrianasolo 585 (G, MO, P, TAN); Andrambovato, E de Fianarantsoa, [21 ° 31 ’ S 47 ° 25 ’ E], 878 - 1000 m, 24. I. 1955, fr., Service Forestier 11590 (G, MO, P, TEF). Prov. Toamasina: RN 1 [Betampona], s. d., fl. & y. fr., Anonymous s. n. (G, MO, P [2 sheets]). Prov. Toliara: Betroka, Kalambatritra RS, forêt d’Analamaro, 23 ° 28 ’ 28 ’’ S 46 ° 25 ’ 15 ’’ E, 1423 m, 9. I. 2004, fl., Andrianjafy et al. 601 (MO, P, TAN); Andohahela PN, parcelle 1, 13.5 km NW d’Eminiminy, 24 ° 35 ’ S 46 ° 44 ’ 1 ’’ E, 1200 m, 15. XI. 1995, fl., Messmer et al. 105 (G, MO).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF47FF5BFE4383651B3D0C48.taxon	materials_examined	Lectotypus (designated here): MADAGASCAR. Prov. Toamasina: Rég. Alaotra-Mangoro, Moramanga, forêt d’Analamazaotra, [18 ° 56 ’ S 48 ° 26 ’ E], 1000 m, 19. X. 1912, fl., Humbert & Viguier 815 (P [P 00580348]!; iso-: K, MO- 6684877!, P [P 00580349, P 02141537, P 06242027]!).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF47FF5BFE4383651B3D0C48.taxon	description	Description Trees 2 - 25 m tall, 10 - 15 cm DBH; young twigs lanate. Leaves alternate, sometimes verticellate; blades oblanceolate, 8 - 30 3 2 - 6 cm, chartaceous, apex acuminate, margin slightly revolute, base acute or cuneate, adaxial surface lanate, abaxial surface pubescent or lanate only on midvein, venation craspedodromous, midvein prominent abaxially, secondary veins 30 - 40 pairs, 2 - 15 mm apart, prominent abaxially, tertiary veins visible on both surfaces; petiole 5 - 20 mm long, canaliculate on adaxial surface, lanate. Inflorescence axillary, a panicle, 3 - 11 cm long, axes terete, villous or lanate. Male and female flowers 5 - merous, glabrous; pedicel 0.2 - 0.3 mm long, villous; calyx lobes triangular, 0.5 3 0.5 - 0.7 mm, imbricate, adaxial surface glabrous, abaxial surface pubescent to lanate, persistent on fruit; corolla lobes ovate, 2 3 1 mm, imbricate or quincuncial, adaxial surface glabrous, abaxial surface pubescent on the longitudinal main axis; stamens 5; disk c. 1 - 1.2 mm in diam., pubescent. Male flowers with filaments 1.5 mm long, anthers 0.5 mm long, ovoid to subglobose, yellow, glabrous; ovary completely absent. Female flowers with filaments 0.5 - 0.7 mm long, anthers 0.3 - 0.5 mm long, ovoid, sterile, yellow, glabrous; ovary widely depressed ovate, 1 - 1.2 3 0.8 - 1 mm, glabrous, placentation subapical, style simple (or with a minute trace of division), apically inserted, 0.5 - 0.7 mm long, glabrous; stigma 4 - lobed, capitate. Fruits 3 3 1.5 cm, ellipsoidal, symmetrical or very slightly asymmetrical, glabrous, with longitudinal striations.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF47FF5BFE4383651B3D0C48.taxon	etymology	Etymology The species is dedicated to Dan Turk, a forest botanist who has contributed tremendously to our knowledge of the trees of Ranomafana National Park.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF47FF5BFE4383651B3D0C48.taxon	vernacular_names	Vernacular names “ Lambafohala’ ’, “ Natoboka’ ’, “ Sefana’ ’, “ Tsitsebona’ ’.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF47FF5BFE4383651B3D0C48.taxon	distribution	Distribution, ecology and phenology Abrahamia viguieri ranges from east coast humid littoral forests on sand and laterite at low elevation to premontane forests along the eastern escarpment at an elevation of up to c. 900 m (Map 9). Material has been collected with flowers in October and November; very young fruits have been recorded in December and mature fruits in late February.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF47FF5BFE4383651B3D0C48.taxon	conservation	Conservation status With an EOO of 28,641 km 2, a minimum AOO of 60 km 2, and 17 localities, with subpopulations in six protected areas (Ambalabe, Analalava, Betampona, Makira, Analamazaotra, and Zahamena), A. viguieri is assigned a preliminary conservation status of “ Least Concern’ ’ [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF47FF5BFE4383651B3D0C48.taxon	discussion	Notes Abrahamia viguieri is easily distinguished from other members of the genus by the lanate upper surface of its leaves and its male flowers with a pubescent disk and completely lacking even a rudimentary or vestigial ovary.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF47FF5BFE4383651B3D0C48.taxon	description	PERRIER DE LA BâTHIE (1944) cited two collections in the protologue of Protorhus viguieri, one with male flowers (Humbert & Viguier 815) and the other with fruits (Perrier de la Bâthie 4387, erroneously cited as 4397). We have selected a specimen of Humbert & Viguier 815 as the lectotype because it is the best preserved among the syntypes and because an isolectotype was available for distribution to MO.	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
038887C6FF47FF5BFE4383651B3D0C48.taxon	materials_examined	Additional material examined MADAGASCAR. Prov. Fianarantsoa: Fort Carnot, exploitation Charlier, [21 ° 51 ’ 30 ’’ S 47 ° 26 ’ 30 ’’ E], 23. II. 1954, fr., Service Forestier 14652 (MO, P, TEF). Prov. Toamasina: Vatomandry, Ambalabe, Ambinanindrano II, forêt de Vohibe, Sahamanahirana, 19 ° 09 ’ 32 ’’ S 48 ° 34 ’ 39 ’’ E, 635 m, 28. X. 2012, bud & fl., Andriamiarinoro 304 (MO, P, TAN); Bricka-ville, Sahatsara, 18 ° 26 ’ 23 ’’ S 48 ° 49 ’ 40 ’’ E, 698 m, 28. X. 2005, fl., Andriamihajarivo et al. 620 (MO, P, TAN, TEF); Brickaville, Anivorano-Sud, Ankerana, 18 ° 26 ’ 03 ’’ S 48 ° 49 ’ 53 ’’ E, 575 m, 31. X. 2005, bud, Andriantiana 297 (P, MO, TAN); Maroantsetra, Rantabe, Beho-vana village, 15 ° 47 ’ 34 ’’ S 49 ° 32 ’ 45 ’’ E, 521 m, 31. X. 2003, fl., Antilahimena 2237 (G, K, MO, NY, P, TAN, WAG); Brickaville, Didy à Brickaville, [18 ° 07 ’ S 48 ° 32 ’ E], s. d., fl., Cours 4823 (MO, P [3 sheets]); Moramanga, station forestière d’Andasibe, [18 ° 56 ’ S 48 ° 26 ’ E], 900 m, 8. XII. 1989, bud & fl., Evrard 11232 (P, TAN); Betampona RN, 17 ° 31 ’ S 49 ° 07 ’ E, 275 - 650 m, 26. IX. 1993, bud, Lewis et al. 619 (G, K, MO [2 sheets], NY, P, TAN, WAG); forêt d’Analamazaotra, [18 ° 55 ’ 48 ’’ S 48 ° 25 ’ 48 ’’ E], 800 m, s. d., fr., Perrier de la Bâthie 4387 (P); Zahamena PN, NE à partir de la rivière de Moango, 17 ° 33 ’ 49 ’’ S 48 ° 54 ’ 01 ’’ E, 794 m, 20. X. 2000, fl., Rakotonandrasana et al. 441 (CNARP, MO, P, TAN); Vavatenina, Zahamena PN, Besakay, 17 ° 39 ’ 20 ’’ S 48 ° 54 ’ 22 ’’ E, 780 m, 7. X. 2001, bud, Rakotonandrasana et al. 573 (CNARP, MO, P, TAN); Analamazaotra RS, along the Indri trail, about 300 m from observation point, 18 ° 56 ’ 12 ’’ S 48 ° 25 ’ 10 ’’ E, 1000 m, 10. I. 2003, ster., A. Randrianasolo 776 (MO, TEF); Fotsimavo, canton Tamatave, [17 ° 56 ’ S 49 ° 13 ’ E], 10. XI. 1955, bud & fl., Réserves Naturelles 7712 (MO, P, TEF); Ambodiriana, [17 ° 55 ’ S 49 ° 13 ’ E], 100 - 200 m, 12. IV. 1959, bud, Réserves Naturelles 10847 (MO, P); Sahatavy, [Zahamena PN], [17 ° 38 ’ S 48 ° 50 ’ E], 750 - 1912 m, 18. X. 1960, bud, Réserves Naturelles 11360 (MO, P [2 sheets], TEF); Fénérive-Est, forêt d’Analalava, Foulpointe, [17 ° 42 ’ S 49 ° 29 ’ E], 0 - 50 m, 30. X. 1963, bud & fl., Service Forestier 22781 (P, TEF); Ambatonandrohina, Sahatavy, [17 ° 27 ’ S 49 ° 00 ’ E], 13. X. 1966, bud & fl., Service Forestier 26261 (P, TEF).	en	Randrianasolo, Armand, Lowry II, Porter P., Schatz, George E. (2017): Taxonomic treatment of Abrahamia Randrian. & Lowry, a new genus of Anacardiaceae from Madagascar. Boissiera 71: 1-152, ISBN: 978-2-8277-0087-5, DOI: 10.5281/zenodo.7618113
