taxonID	type	description	language	source
0388E84058351D73FF13F8861D1AF8F6.taxon	description	(Figs 1 A, 4 A, 7 A, 10 A, 11 A, B) Description. Male. Measurements (mm). Body length 3.20 – 5.52, LH 0.80 – 1.20, WH 0.82 – 1.24, WF 0.50 – 0.72, WOT 0.20 – 0.30. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe angulate; gena not visible in dorsal view; mandible with one distinct ventralmost tooth and three remaining teeth fused; hypostoma strongly curved medially; frons punctured and smooth; frontal line absent; eye glabrous; anterior ocellus anterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus narrow, incomplete anteriorly, diverging anterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, incomplete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin incurved. Genitalia. Harpe with medial-dorsal corner partially arched, apical margin with small, well-delimited excavation, dorsal margin outcurved; cuspis divided in dorsal and ventral arms; dorsal arm laminar, apex acute and equally sized to ventral arm apex; ventral arm laminar and sharp, apex setose and anterior to apex of digitus; digitus with posterior margin smooth; aedeagus with apical lobe not gibbous, to such an extent that dorsal surface is flat, aedeagal apex posterior to apex of ventral arm of cuspis, with three pairs of apical lobes, outer lobe truncate, with apex posterior to inner lobe apex and inner lobe progressively narrowing apicad. Female. Unknown. Variations. Third metapostnotal lateral carina of some paratypes of the type locality can be absent.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058351D73FF13F8861D1AF8F6.taxon	diagnosis	Differential diagnosis. This species is morphologically similar to C. paraensis by having the harpe with medial-dorsal corner partially arched, the apical margin with an emargination less than 0.15 × the harpe length, the dorsal margin out curved and cuspis with ventral arm setose. However, C. awaguaja sp. nov. differs from C. paraensis by having the median clypeal lobe trapezoidal, the frontal line present and the anteromesoscutum with notaulus complete, whereas C. paraensis has the median clypeal lobe angulate, the frontal line absent and the anteromesoscutum with notaulus incomplete.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058351D73FF13F8861D1AF8F6.taxon	etymology	Etymology. The specific epithet awaguaja refers to the indigenous people “ Awá-Guajá ” from Maranhão, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058351D73FF13F8861D1AF8F6.taxon	materials_examined	Material examined. Holotype, male: BRAZIL, MA [ranhão], Carolina, PAR [que] NA [cional] Chapada das Mesas, Riacho Sucuruiu, 07 ° 07 ' 05.6 '' S 47 ° 18 ' 31.6 '' W, 240 m, Malaise trap, 20 - 30. xi. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); Paratypes. Same data as the holotype, except: Cachoeira de Prata, 25 - 30. vi. 2009, A. L. Costa, M. B. Aguiar Neto, P. A. M. Moraes cols., 2 males, Malaise trap, 1 male, Light trap (NESB); 1 male, Riacho Cancela, 07 ° 06 ' 44.2 '' S 47 ° 17 ' 56.8 '' W, 225 m, Suspended trap, 14 - 30. vi. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. A. Silva cols. (CZMA); Suspended Water-Surface trap, 1 male, 10 - 20. viii. 2013 (CZMA); 1 male, 10 - 20. ix. 2013 (CZMA); 1 male, Malaise trap, 14 - 30. vi. 2013 (CZMA); 1 male, 1 - 10. ix. 2013 (CZMA); 1 male, 20 - 30. ix. 2013 (CZMA); 1 male, 01 - 10. x. 2013 (CZMA); 1 male, 20 - 31. xii. 2013 (NESB); 1 male, Riacho Corrente, 298 m, 07 ° 04 ' 29.2 '' S 47 ° 05 ' 42.2 '' W, Malaise trap, 20 - 31. viii. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 4 males, Riacho Estiva, 07 ° 06 ' 59.8 '' S 47 ° 21 ' 21.0 '' W, 265 m, Double Suspended trap, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA, NESB); 1 male, 01 - 15. vii. 2013 (CZMA); 1 male, 01 - 10. viii. 2013 (CZMA); 2 males, 10 - 20. ix. 2013 (CZMA); 1 male, Suspended Water-Surface trap, 15 - 31. vii. 2013 (CZMA); 1 male, 01 - 10. xi. 2013 (NESB); 1 male, Riacho Sucuruiu, 07 ° 07 ' 05.6 '' S 47 ° 18 ' 31.6 '' W, 240 m, 22 - 01. x. 2013, Malaise trap, J. A. Rafael, F. Limeira-de-Oliveira, S. Pereira cols. (CZMA); 1 male, 14 - 30. vi. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 1 male, 01 - 15. vii. 2013 (CZMA); 1 male, 15 - 31. vii. 2013 (CZMA); 1 male, 01 - 10. viii. 2013 (CZMA); 1 male, 10 - 20. viii. 2013 (CZMA); 2 males, 20 - 31. viii. 2013 (CZMA); 1 male, 01 - 10. ix. 2013 (CZMA); 3 males, 10 - 20. ix. 2013 (CZMA, NESB); 2 males, 01 - 10. x. 2013 (CZMA); 2 males, 10 - 20. x. 2013 (CZMA); 1 male, 20 - 31. x. 2013 (CZMA); 2 males, 01 - 10. xi. 2013 (CZMA); 1 male, 10 - 20. xi. 2013 (CZMA); 1 male, 13 - 20. xi. 2013 (CZMA); 2 males, 20 - 30. xi. 2013 (CZMA); 6 males, 01 - 10. xii. 2013 (CZMA); 2 males, 10 - 20. xii. 2013 (CZMA); 1 male, 10 - 20. i. 2014 (CZMA); 1 male, 04. iii. 2014 (CZMA); 1 male, Simple Suspended trap 20 m, 20 - 31. i. 2014, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (NESB); 1 male, Mirador, Parq [ue] Estadual Mirador, Base da Geraldina, 06 ° 37 ' 25 '' S 45 ° 52 ' 08 '' W, Malaise trap, 08 - 14. iii. 2013, L. S. Santos, F. Limeira-de-Oliveira, L. L. M. Santos cols. (CZMA); 3 males, Riachão, Fazenda Altos, Malaise trap, 18 - 22. viii. 2009, F. Limeira-de-Oliveira, M. B. Aguiar Neto cols. (CZMA, NESB).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058351D73FF13F8861D1AF8F6.taxon	distribution	Distribution. Brazil (Maranhão).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058341D74FF13F8821D9CFAFE.taxon	description	(Figs 1 B, 4 B, 7 B)	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058341D74FF13F8821D9CFAFE.taxon	description	Redescription. Male. Measurements (mm). Body length 2.58 – 3.52, LH 0.78 – 0.84, WH 0.71 – 0.75, WF 0.39 – 0.42, WOT 0.35 – 0.21. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe angulate; gena not visible in dorsal view; mandible with five distinct apical teeth; hypostoma curved medially; frons punctured and smooth; frontal line strong and carinate; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus complete, broad, diverging anterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, incomplete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin incurved. Genitalia. Harpe with its surface strongly arched longitudinally to such an extent that it looks like bilaminar, apical margin slightly incurved, dorsal margin outcurved, divided in dorsal and ventral arms, dorsal arm bilaminar, apex rounded and posterior to ventral arm apex, ventral arm laminar and rounded, apex setose and anterior to apex of digitus; digitus with posterior margin smooth; aedeagus with apical lobe acute apically to such an extent that dorsal surface is slightly convex, aedeagal apex posterior to apex of ventral arm of cuspis, with two pairs of apical lobes, outer lobe apex posterior to inner lobe apex both acute apically. Female. Unknown.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058341D74FF13F8821D9CFAFE.taxon	materials_examined	Material examined. Type material. Holotype, male: BRAZIL, Paraná, Telêmaco-Borba, Reserva Samuel Klabin, 21. iii. 1998, Malaise trap, Profapar survey (DZUP). New material. BRAZIL, MA [ranhão], 1 male, PAR [que] NA [cional] Chapada das Mesas, Riacho Sucuruiu, 240 m, 07 ° 07 ' 05.6 '' S 47 ° 18 ' 31.6 '' W, Malaise trap, 10 - 20. x. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols (CZMA); Mirador, Parque Estadual Mirador, Base da Geraldina, Malaise trap, F. Limeira-de-Oliveira, L. L. M. Santos and T. L. Rocha cols., 1 male, 06 ° 37 ' 48 '' S 45 ° 52 ' 46 '' W, 428 m, 20 - 30. xi. 2013 (NESB); 1 male, 06 ° 35 ' 58 '' S 45 ° 50 ' 49 '' W, 402 m, 01 - 10. xii. 2013 (CZMA); Pi [auí], Caracol, PAR [que] NA [cional] Serra das Confusões, Malaise trap, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols, 1 male, 15 - 31. vii. 2013 (CZMA); 1 male, 01 - 11. viii. 2013 (CZMA); 1 male, 01 - 10. ix. 2013 (CZMA); 2 males, 10 - 20. ix. 2013 (CZMA); 1 male, 20 - 30. ix. 2013 (CZMA); 3 males, Riacho dos Bois, 09 ° 13 ' 11.9 '' S 43 ° 29 ' 26.2 '' W, 575 m, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 4 males, 01 - 11. viii- 2013 (CZMA, NESB); Guaribas, 2 males, PAR [que] NA [cional] Serra das Confusões, Andorinhas, 515 m, 09 ° 08 ' 27.8 ” S 43 ° 33 ' 42.1 ” W, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A Silva cols. (CZMA); 1 male, 01 - 10. ix. 2013 (CZMA); 1 male, 01 - 10. i. 2014 (CZMA); Piracuruca, PAR [que] NA [cional] de Sete Cidades, Posto do ICMBio, 04 ° 05 ' 57 '' S 41 ° 42 ' 34 '' W, Suspended trap, 1 male, 15 - 28. ii. 2013, F. Limeira-de-Oliveira, J. S. Pinto Júnior cols. (NESB); 1 male, 19 - 28. ii. 2013, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 1 male, Malaise trap, 12 - 27. iii. 2013 (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058341D74FF13F8821D9CFAFE.taxon	distribution	Distribution. Brazil (Bahia, Espírito Santo, Maranhão, Piauí, Paraná, Santa Catarina, São Paulo).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058341D74FF13F8821D9CFAFE.taxon	discussion	Remarks. This species was recorded from Bahia, Espírito Santo, Pará, Santa Catarina and São Paulo and for the first time is recorded from Maranhão, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058331D75FF13FA9A1DA3FAC2.taxon	description	(Figs 1 C, 4 C, 7 C, 10 B, 11 C, D) Description. Male. Measurements (mm). Body length 4.72 – 5.76, LH 0.98 – 1.15, WH 1.06 – 1.28, WF 0.54 – 0.72, WOT 0.28 – 0.30. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe trapezoidal; gena not visible in dorsal view; mandible with four distinct apical teeth; hypostoma angled medially; frons punctured and smooth; frontal line absent; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus complete, narrow, diverging posterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, incomplete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin incurved. Genitalia. Harpe with medial-dorsal corner partially arched, apical margin slightly incurved, dorsal margin outcurved, divided in dorsal and ventral arms; dorsal arm laminar, apex straight and anterior to ventral arm apex; ventral arm laminar and rounded, apex setose and posterior to apex of cuspis; aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat, aedeagal apex posterior to apex of ventral arm of cuspis, with one pair of apical lobes, lobe progressively narrowing apically. Female. Unknown. Variations. Posterior ocellus crossing supra-ocular line is present in species from Maranhão and Piauí.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058331D75FF13FA9A1DA3FAC2.taxon	diagnosis	Differential diagnosis. This species is morphologically similar to C. spinipes by having the genitalia harpe with medial-dorsal corner partially arched, the apical margin of harpe slightly incurved, the dorsal margin of harpe outcurved and apical projection of basivolsella setose and posterior to the apex of digitus. However, C. guajajara sp. nov. differs from C. spinipes by having the transverse posterior carina absent, the paraspiracular carina absent, the metapostnotal median carina incomplete posteriorly, the aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat and with one pair of apical lobes, whereas C. spinipes has the transverse posterior carina present, the paraspiracular carina present, the metapostnotal median carina complete, the aedeagus with slightly gibbous to such an extent that dorsal surface is slightly convex with two pairs of apical lobes.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058331D75FF13FA9A1DA3FAC2.taxon	etymology	Etymology. The specific epithet guajajara refers to the indigenous people " Guajajara " from Maranhão, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058331D75FF13FA9A1DA3FAC2.taxon	materials_examined	Material examined. Holotype, male: BRAZIL, PI [auí], Guaribas, PAR [que] NA [cional] Serra das Confusões, Andorinha [s], 515 m, 09 ° 08 ' 27.8 '' S 43 ° 33 ' 42.1 '' W, Malaise trap, 01 - 10. x. 2013, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA); Paratypes. MA [ranhão], C [ândido] Mendes, Fazenda Sete irmãos, 1 male, 01 ° 51 ' 37 '' S 45 ° 46 ' 10 '' W, Malaise trap, 03 - 06. x. 2008, F. Limeira-de-Oliveira, J. A. Rafael, PAM Moraes cols. (CZMA); Carolina, PAR [que] NA [cional] Chapada das Mesas, Riacho Sucuruiu, 07 ° 07 ' 05.6 '' S 47 ° 18 ' 31.6 '' W, Malaise trap, 1 male, 20 - 30. ix. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols (CZMA); 2 males, 20 {02} - 13. xi. 2013 (CZMA, NESB); 1 male, Mirador, Parq [ue] Estadual Mirador, Base da Geraldina, Malaise trap, 23 - 28. ix. 2006, F. L. O. col. (CZMA); PI [auí], Caracol, PAR [que] NA [cional] Serra das Confusões, Malaise trap, 1 male, 15 - 31. vii. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 2 males, 20 - 31. viii. 2013 (CZMA); 3 males, 01 - 10. ix. 2013 (CZMA); 1 male, 10 - 20. ix. 2013 (CZMA); 1 male, 20 - 30. ix. 2013 (CZMA); 1 male, Riacho dos Bois, Suspended trap, 01 - 10. xi. 2013 (CZMA); 1 male, Malaise trap, 19. i. 2014 (CZMA); 1 male, 23. i. 2014 (CZMA); 1 male, 29. i. 2014 (CZMA); 1 male, 01. iii. 2014 (CZMA); 1 male, 575 m, 09 ° 13 ' 11.9 '' S 43 ° 29 ' 26.2 '' W, Suspended trap, 01 - 10. x. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols (CZMA); 1 male, Malaise trap, 10 - 20. x. 2013 (CZMA); 2 males, 20 - 31. x. 2013 (CZMA); 1 male, 01 - 10. xi. 2013 (CZMA); 3 males, 10 - 20. xi. 2013 (CZMA); 1 male, 20 - 30. xi. 2013 (CZMA); 1 male, 20 - 30. xii. 2013 (CZMA); 1 male, 20 - 31. xii. 2013 (CZMA); 3 males, 20 - 31. i. 2014 (CZMA); 2 males, 01. ii. 2014 (CZMA); 1 male, 01 - 10. ii. 2014 (CZMA); 5 males, 20 - 28. ii. 2014 (CZMA); 1 male, Saco da Cachorra, Malaise trap, 27. xii. 2013 (CZMA); 1 male, 522 m, 09 ° 12 ' 42.7 '' S 43 ° 30 ' 04.2 '' W, Malaise trap, 11 - 20. viii. 2013 J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 1 male, 20 - 31. x. 2013 (CZMA); 5 males, 01 - 10. xi. 2013 (CZMA, NESB); 3 males, 10 - 20. xi. 2013 (CZMA); 1 male, 01 - 10. i. 2014 (CZMA); 4 males, 20 - 31. i. 2014 (CZMA); 1 male, 10 - 20. ii. 2014 (CZMA); 1 male, Suspended trap, 5 m, 10 - 20. x. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. A. Silva cols. (CZMA); 1 male, Guaribas, PAR [que] NA [cional] Serra das Confusões, Malaise trap, 01 - 10. ix. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. A. Silva. (CZMA); 1 male, Andorinhas, 09 ° 08 ' 27.8 '' S 43 ° 33 ' 42.1 '' W, 515 m, 01 - 10. x. 2013, J. A. Rafael, T. T. A. Silva and F. Limeira-de-Oliveira cols. (CZMA); 3 males, Riacho dos Bois, 575 m, 20 - 30. xi. 2013 (CZMA, NESB).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058331D75FF13FA9A1DA3FAC2.taxon	distribution	Distribution. Brazil (Maranhão, Piauí).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058321D77FF13FA4B1C25FE22.taxon	description	(Figs 1 D, 4 D, 7 D)	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058321D77FF13FA4B1C25FE22.taxon	description	Redescription. Male. Measurements (mm). Body length 2.56 – 4.48, LH 0.56 – 0.85, WH 0.59 – 0.92, WF 0.34 – 0.52, WOT 0.15 – 0.32. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe trapezoidal; gena not visible in dorsal view; mandible with three distinct apical teeth; hypostoma angled medially; frons punctured and coriaceous; frontal line narrow and carinate; eye glabrous; anterior ocellus anterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and coriaceous; notaulus complete, broad, diverging posterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, complete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin straight. Genitalia. Harpe flat, apical margin slightly incurved, dorsal margin straight, divided in dorsal and ventral arms; dorsal arm laminar, apex rounded and anterior to ventral arm apex; ventral arm laminar and rounded, apex glabrous and posterior to apex of digitus; digitus with posterior margin smooth; aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat, aedeagal apex posterior to apex of ventral arm of cuspis, with one pair of apical lobes, rounded apically. Female. Unknown.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058321D77FF13FA4B1C25FE22.taxon	materials_examined	Material examined. Type material. Holotype, male: PERU, Upper io Pachitea, 21, July, 1920 (Cornell Univ. Exped.) (CU [IC], Nº 3880). New material. BRAZIL, MA [ranhão], 3 males, Carolina, PAR [que] NA [cional] Chapada das Mesas, Malaise trap (CZMA, NESB); 1 male, Pov [oado] Canto Grande, Balneário Urupuxete, Light trap, 23 - 27. ii. 2009, A. L. Costa, F. Limeira-de-Oliveira, M. M. Abreu cols. (CZMA); 1 male, Mirador, Parq [ue] Estadual Mirador, Base da Geraldina, arm [adilha] Malaise, 05. iv. 2007, F. L. O. col. (CZMA); Pi [auí], 2 males, Piracuruca, PAR [que] NA [cional] de Sete Cidades, Posto do ICMBio, 04 ° 05 ' 57 '' S 41 ° 42 ' 34 '' W, Malaise trap, 16 - 31. i. 2013, F. Limeira-de-Oliveira, J. S. Pinto Júnior cols. (CZMA); 1 male, 01 - 14. ii. 2013, J. A. Rafael, F. Limeira-de-Oliveira cols. (CZMA); 2 males, 16 - 28. ii. 2013, J. S. Pinto Júnior (CZMA); 7 males, 01 - 10. iii. 2013, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 1 male, F. Limeira-de-Oliveira, J. S. Pinto Júnior cols (CZMA); 44 males, 12 - 27. iii. 2013, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 3 males, 17 - 27. iii. 2013 (CZMA); 1 male, 28. iii- 12. iv. 2013 (CZMA); 1 male, 15 - 30. vi. 2013 (CZMA); 3 males, 15 - 30. vii. 2013 (CZMA); 14 males, Suspended trap, 15 - 28. ii. 2013 (CZMA, NESB); 10 males, 19 - 28. ii. 2013 (CZMA); 2 males, 01 - 06. vii. 2013, varredura [sweeping], J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058321D77FF13FA4B1C25FE22.taxon	distribution	Distribution. Brazil (Amazonas, Amapá, Maranhão, Piauí, Paraná, Santa Catarina, São Paulo).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058321D77FF13FA4B1C25FE22.taxon	discussion	Remarks. This species was recorded from Amazonas, Amapá, Pará, Santa Catarina and São Paulo and for the first time is recorded from Maranhão and Piauí, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583F1D79FF13FF5E1DA3FBE2.taxon	description	(Figs 1 E, 4 E, 7 E, 10 C, 11 E, F) Description. Male. Measurements (mm). Body length 2.16 – 3.12, LH 0.54 – 0.72, WH 0.52 – 0.76, WF 0.33 – 0.68, WOT 0.17 – 24. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe angulate; gena not visible in dorsal view; mandible with five distinct apical teeth; hypostoma curved medially; frons punctured and smooth; frontal line narrow and carinate; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus complete, broad, diverging posterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, incomplete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin incurved. Genitalia. Harpe with its surface strongly arched longitudinally to such an extent that it looks like bilaminar, apical margin slightly incurved, dorsal margin outcurved, divided in dorsal and ventral arms; dorsal arm bilaminar, apex rounded and posterior to ventral arm apex; ventral arm laminar and acute, apex setose and anterior to apex of digitus; aedeagus with apical lobe gibbous to such an extent that dorsal surface is convex, aedeagal apex posterior to apex of ventral arm of cuspis, with three pairs of apical lobes, lobe progressively narrowing apicad. Female. Unknown. Variations. There are no variations.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583F1D79FF13FF5E1DA3FBE2.taxon	diagnosis	Differential diagnosis. This species is morphologically similar to C. ocullatus by having the harpe whole arched to such an extent that is bilaminar, the apical harpal margin slightly incurved, the dorsal harpal margin outcurved, apical projection of basivolsella round and setose. However, C. kanela sp. nov. sp. nov. differs from C. ocullatus, by having the anterior ocellus posterior to supra-ocular line, the second metapostnotal lateral carina present, the posterior hypopygeal margin incurved, the genitalia cuspis apex aligned with apex of digitus and aedeagus with apical lobe gibbous to such an extent that dorsal surface is convex, whereas C. ocullatus has the anterior ocellus anterior to supra-ocular line, the second metapostnotal lateral carina absent, the posterior hypopygeal margin straight, the genitalia cuspis apex anterior to apex of digitus and aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583F1D79FF13FF5E1DA3FBE2.taxon	etymology	Etymology. The specific epithet kanela refers to the indigenous people “ Kanela ” from Maranhão and Piauí, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583F1D79FF13FF5E1DA3FBE2.taxon	materials_examined	Material examined. Holotype, male: BRAZIL, PI [auí], Caracol, PAR [que] NA [cional] Serra das Confusões, Malaise trap, 15 - 31. vii. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. A. Silva cols (CZMA); Paratypes. MA [ranhão], Carolina, PAR [que] NA [cional] Chapada das Mesas, 07 ° 04 ' 24.2 '' S 47 ° 05 ' 25.2 '' W, 288 m, 1 male, Riacho Corrente, Malaise trap, 01 - 10. x. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (NESB); 1 male, Caxias, Re [serva]. Ecol. [ógica]. Inhamum. Povoado Coités, 04 ° 54 ' 43 '' S 43 ° 25 ' 30 '' W, Malaise trap, 27 - 29. xii. 2012, F. Limeira-de-Oliveira, L. S. Santos, cols; 1 male, Mirador, Parq [ue] Estadual Mirador, Base da Geraldina, Malaise trap, 27 - 30. ix. 2006, F. Limeira-de-Oliveira col. (CZMA); PI [auí], 3 males, same data as holotype (CZMA, NESB); 1 male, Malaise trap (CZMA); 1 male, 01 - 11. viii. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 4 males, 11 - 20. viii. 2013 (CZMA); 2 males, 20 - 31. viii. 2013 (CZMA); 1 male, 01 - 10. ix. 2013 (CZMA); 3 males, 01 - 10. ix. 2013 (CZMA); 1 male, Suspended trap, 15 - 30. vi. 2013 (CZMA); 3 males, Andorinhas, 09 ° 08 ' 27.8 '' S 43 ° 33 ' 42.1 '' W, 515 m, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 6 + 1 males, Riacho dos Bois, 09 ° 13 ' 11.9 '' S 43 ° 29 ' 26.2 '' W, 575 m, Malaise trap, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols (CZMA, NESB); 3 males, 01 - 11. viii. 2013 (CZMA); 2 males, 01 - 15. viii. 2013 (CZMA); 1 male, Saco da Cachorra, 522 m, 09 ° 12 ' 42.7 '' S 43 ° 30 ' 04.2 '' W, Malaise trap, 20 - 31. vii. 2013, J. A. Rafael, F. Limeira-de-Oliveira, A. Silva cols (CZMA); 1 male, 01 - 10. ix. 2013 (CZMA); 1 male, 10 - 20. xi. 2013 (CZMA); 1 male, 01 - 10. ix. 2014 (CZMA); 1 male, Guararibas, PAR [que] NA [cional] Serra das Confusões, Andorinhas, 09 ° 08 ' 27.8 ” S 43 ° 33 ' 42.1 ” W, 515 m, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A Silva cols. (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583F1D79FF13FF5E1DA3FBE2.taxon	distribution	Distribution. Brazil (Maranhão, Piauí).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583E1D7AFF13FBAB1FC6FF26.taxon	description	(Figs 1 F, 4 F, 7 F)	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583E1D7AFF13FBAB1FC6FF26.taxon	description	Redescription. Male. Measurements (mm). Body length 3.44, LH 0.76 – 0.88 WH 0.78 – 0.88, WF 0.46 – 0.50, WOT 0.20 – 0.26. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe angulate; gena not visible in dorsal view; mandible with five distinct apical teeth; hypostoma curved medially; frons punctured and smooth; frontal line narrow and sulcate; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus complete, narrow, diverging posterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, complete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin straight. Genitalia. Harpe with medial-dorsal corner partially arched, apical margin strongly emarginated, dorsal margin outcurved, divided in dorsal and ventral arms; dorsal arm laminar, apex rounded and anterior to ventral arm apex; ventral arm laminar and acute, apex glabrous and posterior to apex of digitus; aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat, aedeagal apex anterior to apex of ventral arm of cuspis, with one pair of apical lobes, lobe progressively narrowing apically. Female. Unknown.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583E1D7AFF13FBAB1FC6FF26.taxon	materials_examined	Material examined. Type material. Holotype, male: BRAZIL, Acre, Parque Nacional da Serra do Divisor, ponto norte 9, 20 - 21. XI. 1996, Malaise trap, E. F. Morato col. (UFES). New material. BRAZIL, MA [ranhão], Carolina, PAR [que] NA [cional] Chapada das Mesas, Riacho Corrente, 07 ° 04 ' 24.2 '' S 47 ° 05 ' 25.2 '' W, 288 m, 1 male, Malaise trap, 20 - 30. ix. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); PI [auí], Caracol, PAR [que] NA [cional], Serra das Confusões, 1 male, Malaise trap, 20 - 31. viii. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583E1D7AFF13FBAB1FC6FF26.taxon	distribution	Distribution. Brazil (Acre, Maranhão, Pará).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583E1D7AFF13FBAB1FC6FF26.taxon	discussion	Remarks. This species was recorded from Acre and Pará and for the first time is recorded from Maranhão, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583B1D7DFF13F9831EF6FC56.taxon	description	(Figs 2 A, 5 A, 8 A)	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583B1D7DFF13F9831EF6FC56.taxon	description	Redescription. Male. Measurements (mm). Body length 3.20 – 4.08, LH 0.80 – 0.90, WH 0.88 – 1, WF 0.40 – 0.58, WOT 0.26 – 0.28. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna light brown; legs dark brown to light brown. Head. Malar space absent; median clypeal lobe angulate; gena not visible in dorsal view; mandible with three distinct apical teeth; hypostoma curved medially; frons punctured and coriaceous; frontal line narrow and carinate; eye glabrous; anterior ocellus anterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and coriaceous; notaulus complete, broad, diverging anterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, complete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior hypopygeal margin incurved. Genitalia. Harpe with its surface strongly arched longitudinally to such an extent that it looks like bilaminar, apical margin outcurved, dorsal margin straight, divided in dorsal and ventral arms; dorsal arm laminar, apex rounded and posterior to ventral arm apex; ventral arm laminar and rounded, apex setose and aligned to apex of digitus; digitus with posterior margin smooth; aedeagus with apical lobe gibbous to such an extent that dorsal surface is convex, aedeagal apex posterior to apex of ventral arm of cuspis, with two pairs of apical lobes, outer lobe apex posterior to inner lobe apex, outer and inner lobe narrowing apicad. Female. Unknown.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583B1D7DFF13F9831EF6FC56.taxon	materials_examined	Material examined. Type material. Holotype, male: BRAZIL, Pará, Baker col. (CASC # 17057). New material. BRAZIL, CE [ará], Ubajara, PAR [que] NA [cional] Ubajara, Cachoeira do Cafundó, 03 ° 50 ' 13 '' S 40 ° 54 ' 35 '' W, 1 male, Malaise trap, 01 - 14. ii. 2013, J. A. Rafael and F. Limeira-de-Oliveira cols. (CZMA); 1 male, varredura, 01 - 06. vii. 2013, [J. A.] Rafael F. Limeira-de-Oliveira, T. T. A Silva cols. (CZMA); MA [ranhão], Mirador, Parq [ue] Estadual Mirador, Base da Geraldina, 06 ° 37 ' 25 '' S 45 ° 52 ' 08 '' W, Malaise trap, 1 male, 14 - 18. viii. 2012, D. W. A Marques and F. Limeira-de-Oliveira, J. S. P. Júnior cols. (CZMA); 2 males, 08 - 14. iii. 2013, L. S. Santos, F. Limeira-de-Oliveira, L. L. M. Santos cols. (CZMA, NESB); 1 male, 06 ° 37 ' 06 '' S 45 ° 51 ' 51 '' W, 01 - 12. viii. 2013 (CZMA); PI [auí], Guaribas, PAR [que] NA [cional] Serra das Confusões, Andorinhas, 1 male, 09 ° 08 ' 27.8 '' S 43 ° 33 ' 42.1 '' W, Malaise trap, 515 m, 01 - 10. xi. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583B1D7DFF13F9831EF6FC56.taxon	distribution	Distribution. Brazil (Espírito Santo, Ceará, Maranhão, Pará, Piauí).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583B1D7DFF13F9831EF6FC56.taxon	discussion	Remarks. This species was recorded from Espírito Santo and Pará and for the first time is recorded from Ceará, Maranhão and Piauí, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583A1D7EFF13FC221B39FF23.taxon	description	(Figs 2 B, 5 B, 8 B, 12 A)	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583A1D7EFF13FC221B39FF23.taxon	description	Redescription. Male. Measurements (mm). Body length 3.44 – 4.08, LH 0.87 – 1.20, WH 0.88 – 1.20, WF 0.50 – 0.68, WOT 0.20 – 0.26. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe angulate; gena not visible in dorsal view; mandible with five distinct apical teeth; hypostoma curved medially; frons punctured and smooth; frontal line strong and sulcate; eye glabrous; anterior ocellus anterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus complete, broad, diverging posterad parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, incomplete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium posterior margin straight. Genitalia. Harpe with its surface strongly arched longitudinally to such an extent that it looks like bilaminar, apical margin slightly incurved, dorsal margin outcurved, divided in dorsal and ventral arms; dorsal arm bilaminar, apex rounded and posterior to ventral arm apex; ventral arm laminar and rounded, apex setose and anterior to apex of digitus; aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat, aedeagal apex posterior to apex of ventral arm of cuspis, with one pair of apical lobes, lobes rounded apically. Female. Unknown.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583A1D7EFF13FC221B39FF23.taxon	materials_examined	Material examined. Type material. Holotype, male: BRAZIL, Acre, Parque Nacional da Serra do Divisor, ponto norte 5, 14 - 15. XI. 1996, Malaise trap, E. F. Morato col. (UFES). New material. BRAZIL, MA [ranhão], Carolina, PAR [que] NA [cional] Chapada das Mesas, Riacho Cancela, 07 ° 06 ' 44.2 '' S 47 ° 17 ' 58.3 '' W, 225 m, 1 male, Suspended trap, 01 - 10. viii. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. A. Silva cols. (CZMA); 1 male, 07 ° 06 ' 44.2 '' S 47 ° 17 ' 56.8 '' W, suspensa lâmina d'água, 225 m, 01 - 10. x. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. A. Silva cols. (CZMA); 1 male, Riacho Corrente, 288 m, 06 - 29. ii. 2014 (CZMA); 1 male, Riacho Estiva, 07 ° 06 ' 59.8 '' S 47 ° 21 ' 21.0 '' W, 265 m, Suspended trap, 20 - 31. x. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. T. A. {T. T. A.} Silva cols. (NESB); 1 male, Riacho Sucuruiu, 01 - 10. ix. 2013, arm [adilha] Malaise (CZMA); Mirador, Parque Estadual Mirador, Base da Geraldina, 06 ° 35 ' 58 '' S 45 ° 50 ' 49 '' W, 402 m, Malaise trap, F. Limeira-de-Oliveira, L. L. M. Santos, T. L. Rocha cols., 1 male, 10 - 20. x. 2013 (CZMA); 1 male, 20 - 31. xii. 2013 (CZMA); 2 males, 02 - 12. viii. 2013, L. S. Santos, F. Limeira-de-Oliveira, L. L. M. Santos cols. (CZMA, NESB).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583A1D7EFF13FC221B39FF23.taxon	distribution	Distribution. Brazil (Acre, Maranhão).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E840583A1D7EFF13FC221B39FF23.taxon	discussion	Remarks. This species is recorded for the first time from Maranhão, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058391D7FFF13F8FB1DBFF990.taxon	description	(Figs 2 C, 5 C, 8 C, 12 B)	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058391D7FFF13F8FB1DBFF990.taxon	description	Redescription. Male. Measurements (mm). Body length 3.28 – 5.36, LH 0.83 – 1.20, WH 0.88 – 1.30, WF 0.54 – 0.76, WOT 0.20 – 0.38. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe trapezoidal; gena not visible in dorsal view; mandible with one distinct ventralmost tooth and remaining teeth indistinctly fused; hypostoma strongly curved medially; frons punctured and smooth; frontal line narrow and sulcate; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus complete, narrow, diverging posterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, incomplete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin incurved. Genitalia. Harpe with medial-dorsal corner partially arched, apical margin with small, well delimited excavation, dorsal margin outcurved, divided in dorsal and ventral arms; dorsal arm laminar, apex rounded and anterior to ventral arm apex; ventral arm laminar and sharp, apex setose and posterior to apex of digitus; aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat, aedeagal apex posterior to apex of ventral arm of cuspis, with two pairs of apical lobes, outer lobe apex posterior to inner lobe apex, rounded apically. Female. Unknown.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058391D7FFF13F8FB1DBFF990.taxon	materials_examined	Material examined. Type material. Holotype, male: BRAZIL, Pará (USNM). New material. BRAZIL, Ce [ará], 1 male, Ubajara, PAR [que] NA [cional] Ubajara, Cachoeira do Cafundó, 03 ° 50 ' 13 '' S / 40 ° 54 ' 35 '' W, Suspended trap, 15 - 28. ii. 2012, J. S. Pinto Júnior and F. Limeira-de-Oliveira cols. (CZMA); 1 male, 16 - 31. xii. 2012 (CZMA); 1 male, Malaise trap, 01 - 15. i. 2013 (CZMA); 1 male, 01 - 14. ii. 2013 (CZMA); 1 male, 19 - 28. ii. 2013 (CZMA); MA [ranhão], 1 male, C [ândido] Mendes, Fazenda Sete Irmãos, 01 ° 51 ' 37 '' S 45 ° 46 ' 10 '' W, Malaise trap, 03 - 06. x. 2008, F Limeira-de-Oliveira, JA Rafael, PAM Moraes cols. (CZMA); 1 male, Carolina, PAR [que] NA [cional] Chapada das Mesas, Malaise trap (CZMA); 1 male, Riacho Corrente, 14 - 22. ii. 2014 (CZMA); Caxias, 1 male, Res [erva] Ecol [ógica] Inhamum, Malaise trap, 20 - 24. iv. 2006, G. A. Cunha col. (CZMA); 1 male, Zona Rural, Riacho do Ouro, 04 ° 43 ' 05 '' S 43 ° 14 ' 16 '' W, varredura, 24. vii. 2012, JA Rafael, F. Limeira-de-Oliveira, JS Pinto Júnior cols (CZMA); 2 males, REBIO, Centro Novo, Malaise trap, 7 - 11. i. 2011 (CZMA, NESB); 1 male, Mirador, Parq [ue] Estadual Mirador, Base da Geraldina, 06 ° 37 ' 25 '' S 45 ° 52 ' 08 '' W, Malaise trap, 14 - 18. viii. 2012, D. W. A. Marques, F. Limeira-de-Oliveira and J. S. P. Júnior cols (CZMA); 3 males, 13 - 19. x. 2012, L. S. Santos, F. Limeira-de-Oliveira, L. L. M. Santos cols. (CZMA, NESB); 2 males, 08 - 14. iii. 2013 (CZMA); 3 males, 10 - 20. x. 2013, F Limeira-de-Oliveira, L. L. M. Santos, T. L. Rocha cols (CZMA); 1 male, 20 - 31. i. 2014, F. Limeira-de-Oliveira, L. L. M. Santos and L. S. Santos cols. (CZMA). 1 male, 411 m, 06 ° 37 ' 06 '' S 45 ° 51 ' 51 '' W, 02 - 12. viii. 2013, F. Limeira-de-Oliveira, L. L. M. Santos and T. L. Rocha cols. (CZMA); Pi [auí], Piracuruca, PAR [que] NA [cional] de Sete Cidades, Posto do ICMBio, 04 ° 05 ' 57 '' S 41 ° 42 ' 34 '' W, Suspended trap, F. Limeira-de-Oliveira, J. S. Pinto Júnior cols., 1 male, 01 - 14. ii. 2013 (CZMA); 1 male, 15 - 28. ii. 2013 (CZMA); Malaise trap, 1 male, 15 - 28. ii. 2013 (CZMA); 1 male, 16 - 28. ii. 2013 (CZMA); 1 male, 01 - 15. xii. 2013 (CZMA); F. Limeira-de-Oliveira, T. T. A. Silva cols, 1 male, 19 - 28. ii. 2013 (CZMA); 12 males, 12 - 27. iii. 2013 (CZMA, NESB); 1 male 17 - 27. iii. 2013 (CZMA); 3 males, 15 - 30. vi. 2013 (CZMA, NESB); 2 males, 01 - 06. vii. 2013 (CZMA); 1 male 15 - 30. xi. 2013 (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058391D7FFF13F8FB1DBFF990.taxon	distribution	Distribution. Brazil (Acre, Amazonas, Ceará, Maranhão, Pará, Piauí, Roraima).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058391D7FFF13F8FB1DBFF990.taxon	discussion	Remarks. This species was recorded from Acre, Amazonas, Maranhão, Pará and Roraima and for the first time is recorded from Ceará and Maranhão, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058381D61FF13F97D1D22FE5E.taxon	description	(Figs 2 D, 5 D, 8 D, 10 D, 11 G, H) Description. Male. Measurements (mm). Body length 3.20 – 3.44, LH 0.80, WH 0.84 – 0.86, WF 0.42 – 0.50, WOT 0.20 – 0.22. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe angulate; gena not visible in dorsal view; mandible with one distinct ventralmost tooth and three remaining teeth fused; hypostoma angled medially; frons punctured and smooth; frontal line absent; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus incomplete anteriorly, narrow, diverging posterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, incomplete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin incurved. Genitalia. Harpe with medial-dorsal corner partially arched, apical margin strongly emarginated, dorsal margin ondulated, divided in dorsal and ventral arms; dorsal arm laminar, apex acute and equally sized to ventral arm apex; ventral arm laminar and acute, apex setose posteriorly and aligned to apex of digitus; digitus with posterior margin smooth; aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat, aedeagal apex posterior to apex of ventral arm of cuspis, with three pairs of apical lobes, lobes acute apically. Female. Unknown. Variations. There are no variations	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058381D61FF13F97D1D22FE5E.taxon	diagnosis	Differential diagnosis. The genitalia of this species is morphologically distinct from all other species by having the dorsal harpal margin ondulate. Etymology. The specific epithet piripiri refers to the indigenous people “ Piripiri ” from Piauí, Brazil. Material examined. Holotype, male: BRAZIL, PI [auí], Piracuruca, P [arque] N [acional] de Sete Cidades, Posto do ICMBio, 04 ° 05 ' 57 '' S 41 ° 42 ' 34 '' W, Malaise trap, 12 - 27. iii. 2013, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); Paratypes. PI [auí], 1 male, same data as the holotype (NESB); 1 male, Suspended trap, 15 - 28. ii. 2013, F. Limeira-de-Oliveira, J. S. Pinto Júnior cols. (CZMA). Distribution. Brazil (Piauí).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058261D61FF13FE371C2DF8A8.taxon	description	(Figs 2 E, 5 E, 8 E)	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058261D61FF13FE371C2DF8A8.taxon	description	Redescription. Male. Measurements (mm). Body length 3.68 – 4.16, LH 0.85 – 0.90, WH 0.90 – 0.96, WF 0.56 – 0.60, WOT 0.22 – 0.24. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe trapezoidal; gena not visible in dorsal view; mandible with one distinct ventralmost tooth and remaining teeth indistinctly fused; hypostoma strongly curved medially; frons punctured and smooth; frontal line narrow and carinate; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus complete, narrow, diverging anterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, incomplete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin incurved. Genitalia. Harpe with medial-dorsal corner partially arched, apical margin with small, well-delimited excavation, dorsal margin outcurved, divided in dorsal and ventral arms; dorsal arm laminar, apex rounded and anterior to ventral arm apex; ventral arm laminar and sharp, apex glabrous and posterior to apex of digitus; aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat, aedeagal apex posterior to apex of ventral arm of cuspis, with two pairs of apical lobes, outer lobe apex posterior to inner lobe apex, outer lobe rounded and inner lobe acute apically. Female. Measurements. Body length 4.20 mm. Color. Light castaneous. Head. Subquadrate in dorsal view, LH 1.00 × WH, globoid in lateral view, height 0.79 × LH; median clypeal lobe large, extending beyond the antennal rim apical margin straight with median concavity, distinctly long setae apically, median carina spine-shaped; frons punctured and smooth; vertex convex with parallel sides. Mesosoma. Dorsal pronotal area subquadrate in dorsal view, with straight lateral margins, 1.60 × longer than wide, with pronounced anterior constriction, anterior width 0.51 × wider than posterior one; mesoscutellum triangular with concave anterior margin; metasternum triangular, with median sulcus progressively widening posterad; coxae dilated, especially the mesocoxa; protibia unarmed; mesotibia spinose; metatibia spinose. Metasoma. Anterior area of the second abdominal sternum distinctly divided and abruptly vertical, tergum longer than wide.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058261D61FF13FE371C2DF8A8.taxon	materials_examined	Material examined. Type material. Holotype, male: BRAZIL, Diamantina, Minas Gerais, 14 - 18. Nov. 1919 (Cornell Univ. Exped) (CU [IC] Nº 3881). Revised material. Female: BRAZIL, Distrito Federal: Brasília (Reserva Ecológica do IBGE, 15 ° 55 ' 58 '' S, 47 ° 51 ' 02 '' W), campo limpo, Malaise [trap], 18 - 25. xi. 1982. New material. BRAZIL, PI [auí], Piracuruca, P [arque] N [acional] de Sete Cidades, Posto do ICMBio, 04 ° 05 ' 57 '' S 41 ° 42 ' 34 '' W, 2 males, Suspended trap, 15 - 28. ii. 2013, F. Limeira-de-Oliveira, J. S. Pinto Júnior cols. (CZMA, UFES); 1 male, Malaise trap, 12 - 27. iii. 2013, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058261D61FF13FE371C2DF8A8.taxon	distribution	Distribution. Brazil (Amazonas, Distrito Federal, Espírito Santo, Minas Gerais, Piauí).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058261D61FF13FE371C2DF8A8.taxon	discussion	Remarks. This species was recorded from Amazonas, Distrito Federal, Espírito Santo and Minas Gerais and for the first time this species is recorded from Piauí, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058251D62FF13FF5F1AB0F923.taxon	description	(Figs 2 F, 5 F, 8 F)	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058251D62FF13FF5F1AB0F923.taxon	description	Redescription. Male. Measurements (mm). Body length 3.28 – 5.52, LH 0.78 – 1.23, WH 0.80 – 1.28, WF 0.46 – 0.72 WOT 0.24 – 0.34. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe angulate; gena not visible in dorsal view; mandible with one distinct ventralmost tooth and three remaining teeth fused; hypostoma curved medially; frons punctured and smooth; frontal line narrow and carinate; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus complete narrow, diverging posterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, incomplete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin incurved. Genitalia. Harpe with medial-dorsal corner partially arched, apical margin with large excavation, dorsal margin outcurved, divided in dorsal and ventral arms; dorsal arm laminar, apex rounded and equally sized to ventral arm apex; ventral arm laminar and sharp, apex setose and anterior to apex of digitus; aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat, aedeagal apex posterior to apex of ventral arm of cuspis, with two pairs of apical lobes, outer lobe apex posterior to inner lobe apex, outer lobe acute and inner lobe progressively narrowing. Female. Unknown.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058251D62FF13FF5F1AB0F923.taxon	materials_examined	Material examined. Type material. Holotype, male: BRAZIL, Pará: Serra Norte (Estrada Serraria), 07. VII. 1984, Suspended trap (MPEG). New material. BRAZIL, MA [ranhão], C [ândido] Mendes, Fazenda Sete irmãos, 01 ° 51 ' 37 '' S 45 ° 46 ' 10 '' W, 1 male, Malaise trap, 06. x. 2008, F. Limeira-de-Oliveira, J. A. Rafael, P. A. M. Moraes, col. (CZMA); 1 male, Carolina, PAR [que] NA [cional] Chapada das Mesas, Riacho Cancela, Suspended Water-Surface trap, 01 - 10. xii. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. A. Silva cols. (CZMA); 1 male, 07 ° 06 ' 44.2 '' S 47 ° 17 ' 56.8 '' W, 225 m, Malaise trap, 1 - 15. vii. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. A. Silva cols. (CZMA); 1 male, 20 - 30. ix. 2013 (CZMA); 1 male, 20 - 31. xii. 2013 (CZMA); 1 male, Suspended trap, 14 - 30. vi. 2013 (CZMA); 1 male, 01 - 10. viii. 2013 (CZMA); 1 male, Suspended Water-Surface trap, 01 - 10. viii. 2013 (CZMA); 1 male, 01 - 10. viii. 2013 (CZMA); 1 male, 20 - 30. ix. 2013 (CZMA); 1 male, 01 - 10. x. 2013 (CZMA); 2 males, 20 - 31. x. 2013 (CZMA, NESB); 2 males, 01 - 10. xi. 2013 (CZMA); 1 male, Riacho Sucuruiu, 07 ° 07 ' 05.6 '' S 47 ° 18 ' 31.6 '' W, 20 m, Simple Suspended trap, 20 - 31. i. 2014, J. A. Rafael, F. Limeira-de-Oliveira, T. L. Rocha, S. Pereira cols. (CZMA); 1 male, 07 ° 07 ' 05.6 '' S 47 ° 18 ' 31.6 '' W, 240 m, Malaise trap, 21 - 29. i. 2014, J. A. Rafael, F. Limeira-de-Oliveira, T. L. Rocha, S. Pereira cols (CZMA); 2 males, Mirador, Parq [ue] Estadual Mirador, Base dos Cágados, 06 ° 48 ' 29 '' S 45 ° 06 ' 34 '' W, Malaise trap, 29. ix- 02. x. 2011, D. W. A. Marques and F. Limeira-de-Oliveira cols. (CZMA); 1 male, 06 ° 37 ' 25 '' S 45 ° 52 ' 08 '' W, Varredura, 14 - 18. viii. 2012, J. S. Pinto Júnior, F. L. - de-Oliveira and D. W. A. Marques cols. (CZMA); 3 males, Malaise trap, 14 - 18. viii. 2012 (CZMA); 3 males, 08 - 14. iii. 2013, L. S. Santos, F. Limeira-de-Oliveira and L. L. M. Santos cols. (CZMA, NESB); 3 males, 10 - 16. v. 2013 (CZMA); 1 male, 06 ° 35 ' 48 '' S 45 ° 52 ' 49 '' W, 402 m, Malaise trap, 20 - 30. xi. 2013, F. Limeira-de-Oliveira, L. L. M. Santos and T. L. Rocha cols. (CZMA); 1 male, 06 ° 37 ' 06 '' S 45 ° 51 ' 51 '' W, 411 m, 02 - 12. viii. 2013 (CZMA); 1 male, 06 ° 37 ' 48 '' S 45 ° 52 ' 46 '' W, 428 m, 19 - 28. ii. 2013, F. Limeira-de-Oliveira, L. L. M. Santos, L. S. Santos cols. (CZMA); 1 male, 06 ° 37 ' 48 '' S 45 ° 52 ' 49 '' W, 416 m, 10 - 20. xi. 2013, F. Limeira-de-Oliveira, L. L. M. Santos and T. L. Rocha cols. (CZMA); 1 male, RE [serva] BIO [lógica] Centro Novo, Malaise trap, 7 - 11. i. 2011 (CZMA); 1 male, Riachão, Fazenda da Altos, Malaise trap, F. Limeira-de-Oliveira, M. B. Aguiar cols. (CZMA); 2 males, 18 - 22. viii. 2009 (CZMA, NESB).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058251D62FF13FF5F1AB0F923.taxon	distribution	Distribution. Brazil (Maranhão, Pará).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058251D62FF13FF5F1AB0F923.taxon	discussion	Remarks. This species was only recorded from Pará and for the first time is recorded from Maranhão, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058251D65FF13F8DE1D3FF82B.taxon	description	(Figs 3 A, 6 A, 9 A)	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058251D65FF13F8DE1D3FF82B.taxon	description	Redescription. Male. Measurements (mm). Body length 2.24, LH 0.60, WH 0.62, WF 0.36, WOT 0.16. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe trapezoidal; gena not visible in dorsal view; mandible with one distinct ventralmost tooth and remaining teeth indistinctly fused; hypostoma angled medially; frons punctured and coriaceous; frontal line absent; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and coriaceous; notaulus complete, broad, diverging anterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc irregular, complete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin straight. Genitalia. Harpe flat, apical margin slightly incurved, dorsal margin straight, divided in dorsal and ventral arms; dorsal arm laminar, apex rounded and equally sized to ventral arm apex; ventral arm laminar and straight, apex glabrous and aligned to apex of digitus; aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat, aedeagal apex posterior to apex of ventral arm of cuspis, with one pair of apical lobes, lobes truncate apicad. Female. Unknown.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058251D65FF13F8DE1D3FF82B.taxon	materials_examined	Material examined. Type material. Holotype, male: BRAZIL, Santa Catarina, Nova Teutonia, 7 May 1937 (Fritz Plaumann) (BMNH). New material. BRAZIL, CE [ará], PAR [que] NA [cional] Ubajara, Cachoeira do Cafundó, 03 º 50 ' 13 '' S 40 º 54 ' 35 '' W, 1 male, Malaise trap, 01 - 15. xii. 2012, J. S. Pinto Júnior and F. Limeira-de-Oliveira cols. (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058251D65FF13F8DE1D3FF82B.taxon	distribution	Distribution. Brazil (Ceará, Santa Catarina).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058251D65FF13F8DE1D3FF82B.taxon	discussion	Remarks. This species was only recorded from Santa Catarina and for the first time is recorded from Ceará, Brazil..	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058211D67FF13FF5E1EF6FE96.taxon	description	(Figs 3 B, 6 B, 9 B, 12 C)	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058211D67FF13FF5E1EF6FE96.taxon	description	Redescription. Male. Measurements (mm). Body length 3.48 – 6.24, LH 0.88 – 1.18, WH 0.94 – 1.26, WF 0.58 – 0.74, WOT 0.25 – 0.30. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna and legs light brown. Head. Malar space absent; median clypeal lobe trapezoidal; gena not visible in dorsal view; mandible with four distinct apical teeth; hypostoma angled medially; frons punctured and smooth; frontal line absent; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus complete, narrow, diverging posterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc straight, incomplete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin incurved. Genitalia. Harpe with medial-dorsal corner partially arched, apical margin slightly incurved, dorsal margin outcurved, divided in dorsal and ventral arms; dorsal arm laminar, apex rounded and equally sized to ventral arm apex; ventral arm laminar and rounded, apex setose and posterior to apex of digitus; aedeagus with apical lobe slightly gibbous to such an extent that dorsal surface is slightly convex, aedeagal apex posterior to apex of ventral arm of cuspis, with two pairs of apical lobes, lobes apex aligned, outer lobe acute and inner lobe rounded apicad. Female. Unknown.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058211D67FF13FF5E1EF6FE96.taxon	materials_examined	Material examined. Type material. Holotype, male: ARGENTINA, Tucumán, 11 km W of Las Cejas, 3 - 19 June 1966 (FIML). New Material. BRAZIL. MA [ranhão], Carolina, PAR [que] NA [cional], 1 male, Chapada da Mesas, Riacho Cancela, 225 m, 07 ° 06 ' 44.2 '' S 47 ° 17 ' 56.8 '' W, Suspended trap, 14 - 30. vi. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. A. Silva cols. (CZMA); Riacho Corrente, 07 ° 04 ' 24.2 '' S 47 ° 05 ' 25.2 '' W, 288 m, Malaise trap, 1 male, 01 - 10. ix. 2013 (CZMA); 1 male, 01 - 10. xii. 2013; 1 male, 01 - 10. i. 2014, J. A. Rafael, F. Limeira-de-Oliveira, T. L. Rocha, S. Pereira cols. (CZMA); 1 male, Riacho Estiva, 265 m, 07 ° 06 ' 59.8 '' S 47 ° 21 ' 21.0 '' W, Double Suspended trap, 01 - 10. ix. 2013, J. A. Rafael, F. Limeira-de-Oliveira and T. T. A. Silva cols. (CZMA); 1 male, Riacho Sucuruiu, 240 m, 07 ° 07 ' 05.6 '' S 47 ° 18 ' 31.6 '' W, Malaise trap, 20 - 30. ix. 2013 (CZMA); Caxias, Res [erva] Ecol [ógica] Inhamum, G. A. Cunha, col., 1 male, 25 - 28. xi. 2005 (CZMA); 1 male, 10. xii. 2005 (CZMA); Mirador, Parq [ue] Estadual Mirador, Base dos Cágados, 2 males, 29. ix- 02. x. 2011, D. W. A. Marques and F Limeira-de-Oliveira cols. (CZMA); 1 male, Base da Geraldina, Malaise trap, 09. v. 2014 (CZMA); 06 ° 37 ' 25 '' S 45 ° 52 ' 08 '' W, F. Limeira-de-Oliveira and D. W. A. Marques cols., 3 males, 30. vi- 06. viii. 2011 (CZMA); 1 male, 30. vii- 06. viii. 2011 (CZMA); 2 males, 14 - 18. viii. 2012, J. S. Pinto Junior, F. L. - de-Oliveira and D. W. A. Marques cols. (CZMA); 5 males, 13 - 19. x. 2012, L. S Santos, F. Limeira-de-Oliveira and L. L. M. Santos cols. (CZMA); 9 males, 08 - 14. iii. 2013 (CZMA, NESB); 6 males, 10 - 16. v. 2013 (CZMA); 2 males, 419 m, 01 - 10. x. 2013, F. Limeira-de-Oliveira, L. L. M. Santos, T. L. Rocha cols.; 1 male, 402 m, 06 ° 35 ' 58 '' S 45 ° 50 ' 49 '' W, Malaise trap, 01 - 10. xii. 2013, F. Limeira-de-Oliveira, L. L. M. Santos and T. L. Rocha cols. (CMZA); 2 males, 10 - 20. ii. 2014 (CZMA); 1 male, 416 m, 06 ° 37 ' 48 '' S 45 ° 52 ' 49 '' W, 12 - 20. viii. 2013, L. S. Santos, F. Limeira-de-Oliveira and L. L. M. Santos cols. (CMZA); 2 males, 20 - 31. viii. 2013 (CZMA); 3 males, 10 - 20. x. 2013, T. L. Rocha, F. Limeira-de-Oliveira, L. L. M. Santos cols. (CZMA); 4 males, 10 - 20. xi. 2013 (CZMA, NESB); 4 males, 20 - 31. xi. 2013 (CZMA); 3 males, 411 m, 06 ° 37 ' 06 '' S 45 ° 51 ' 51 '' W, 01 - 12. viii. 2013, L. S. Santos, F. Limeira-de-Oliveira and L. L. M. Santos (CZMA); 3 males, 02 - 12. viii. 2013, F. Limeira-de-Oliveira, L. L. M. Santos and T. L. Rocha cols. (CZMA); 1 male, 06 ° 37 ' 48 '' S 45 ° 52 ' 49 '' W, 416 m, Suspended trap, 01 - 10. xi. 2013, F. Limeira-de-Oliveira, L. L. M. Santos and T. L. Rocha cols. (CZMA); 1 male, 06 ° 35 ' 58 '' S 45 ° 50 ' 49 '' W, 20 - 31. xii. 2013, T. L. Rocha, F. Limeira-de-Oliveira, L. L. M. Santos cols. (CZMA); PI [auí], Caracol, PAR [que] NA [cional], Serra das Confusões, 3 males, Malaise trap, 15 - 31. vii. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA, NESB); 1 male, 01 - 11. viii. 2013 (CZMA); Riacho dos Bois, 09 º 13 ' 11.9 '' S 43 º 29 ' 26.2 '' W, 1 male, Malaise trap, 29. i. 2014 (CZMA); 1 male, 01 - 11. viii. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 1 male, 10 - 21. xi. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. L. Rocha cols. (CZMA); 1 male, 20 - 31. i. 2014, J. A. Rafael, F. Limeira-de-Oliveira, T. L. Rocha, S. Pereira cols. (CZMA); 1 male, 01. ii. 2014, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CMZA); 1 male, Saco da Cachorra, 09 ° 12 ' 42.7 '' S 43 ° 30 ' 04.2 '' W, 522 m, Malaise trap, 1 - 10. ix. 2024, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 5 males, Andorinhas, 515 m, 09 ° 08 ' 27.8 '' S 43 ° 33 ' 42.1 '' W, Malaise trap, 01 - 10. xi. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA, NESB); 1 male, Riacho dos Bois, 575 m, 09 ° 13 ' 11.9 '' S 43 ° 29 ' 26.2 '' W, Malaise trap, 20 - 30. xi. 2013, J. A. Rafael, T. T. A. Silva and F. Limeira-de-Oliveira cols. (CZMA); Piracuruca, P [arque] N [acional] de Sete Cidades, Posto do ICMBio, 04 ° 05 ' 57 '' S 41 ° 42 ' 34 '' W, Malaise trap, 1 male, 01 - 15. xii. 2012, F. Limeira-de-Oliveira, J. S. Pinto Júnior cols. (CZMA); 1 male, 01 - 10. iii. 2013, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); 1 male, 12 - 27. iii. 2013 (CZMA); 1 male, [no further data], Malaise trap, 18 - 25. iii. 2012, F. Limeira-de-Oliveira, DWA Marques cols. (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058211D67FF13FF5E1EF6FE96.taxon	distribution	Distribution. Argentina and Brazil (Alagoas, Distrito Federal, Maranhão, Pará, Piauí).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058211D67FF13FF5E1EF6FE96.taxon	discussion	Remarks. This species was recorded from Alagoas, Distrito Federal and Pará and for the first time from Maranhão and Piauí, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058201D67FF13FE621DA3F925.taxon	description	(Figs 3 C, 6 C, 9 C, 10 E, 11 I, J) Description. Male. Measurements (mm). Body length 2.48 – 4.48, LH 0.63 – 0.93, WH 0.62 – 0.94, WF 0.38 – 0.58, WOT 0.16 – 0.28. Color. Head and mesosoma black, metasoma dark brown; wings hyaline; antenna light brown; legs dark brown to light brown. Head. Malar space absent; median clypeal lobe trapezoidal; gena not visible in dorsal view; mandible with three distinct apical teeth; hypostoma angled medially; frons punctured and coriaceous; frontal line narrow and sulcate; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and coriaceous; notaulus complete, narrow, diverging posterad; parapsidal signum narrow; metapostnotal median carina of metapectal-propodeal disc straight, complete posteriorly. Hind wing thick, jugum not fused with remigium. Metasoma. Hypopygium with posterior margin outcurved. Genitalia. Harpe flat, apical margin slightly incurved, dorsal margin straight, divided in dorsal and ventral arms, dorsal arm laminar, apex rounded and converging ventrally, ventral arm straight and glabrous posteriorly, apex aligned to apex of digitus; aedeagus with apical lobe slightly gibbous, so that dorsal surface is slightly convex, aedeagal apex posterior to apex of ventral arm of cuspis, with two pairs of apical lobes, lobes rounded apicad. Female. Unknown. Variations. There are no variations.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058201D67FF13FE621DA3F925.taxon	diagnosis	Differential diagnosis. This species is morphologically similar to C. inca by having the harpe flat, the apical margin slightly incurved, the dorsal margin straight and the apical projection of basivolsella setose. However, C. tenetehara sp. nov. differs from C. inca by having the anterior ocellus anterior to supra-ocular line, the posterior hypopygeal margin straight, the cuspis apex rounded and posterior to apex of digitus, whereas C. inca has the anterior ocellus posterior to supra-ocular line, the posterior hypopygeal margin outcurved, the cuspis apex straight and aligned with apex of digitus.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058201D67FF13FE621DA3F925.taxon	etymology	Etymology. The specific epithet tenetehara refers to the indigenous people “ Tenetehara ” from Maranhão and Piauí, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058201D67FF13FE621DA3F925.taxon	materials_examined	Material examined. Holotype, 1 male: BRAZIL, MA [ranhão], Carolina, PAR [que] NA [cional] Chapada das Mesas, Riacho Sucuruiu, 07 ° 07 ' 05.6 '' S 47 ° 18 ' 31.6 '' W, 240 m, Malaise trap, 14 - 30. vi. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA); Paratypes. 1 male, Riacho Cancela, 255 m, 07 ° 06 ' 44.2 '' S 47 ° 17 ' 56.8 '' W, Malaise trap, 11 - 14. i. 2014, J. A. Rafael, F. Limeira-de-Oliveira, J. T. Câmara, A. A. Santos cols. (CZMA); 1 male, Riacho Corrente, 288 m, 07 ° 04 ' 24.2 '' S 47 ° 05 ' 25.2 '' W, Malaise trap, 01 - 10. i. 2014, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva, S. Pereira cols. (NESB); 3 males, Riacho Sucuruiu, 07 ° 07 ' 05.6 '' S 47 ° 18 ' 31.6 '' W, Malaise trap, 14 - 30. vi. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA, NESB); 1 male, 15 - 31. viii. 2013 (CZMA); 2 males, 01 - 10. x. 2013 (CZMA); 1 male, 01 - 10. xi. 2013 (CZMA); 1 male, 01 - 10. xii. 2013 (CZMA); 1 male, 10 - 20. xii. 2013 (CZMA); PI [auí], Caracol, PAR [que] NA [cional] Serra das Confusões, 2 males, 20 m, Double Suspended trap, 20 - 30. ix. 2013, J. A. Rafael, F. Limeira de-Oliveira and T. T. A. Silva cols. (CZMA, NESB); 1 male, Piracuruca, PAR [que] NA [cional] de Sete Cidades, Posto do ICMBio, 04 ° 05 ' 57 '' S 41 ° 42 ' 34 '' W, Malaise trap, 15 - 30. vi. 2013, J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva cols. (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058201D67FF13FE621DA3F925.taxon	distribution	Distribution. Brazil (Maranhão, Piauí).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058201D69FF13F8EE1DABFEEE.taxon	description	(Figs 3 D, 6 D, 9 D, 10 F, 11 K, L) Description. Male. Measurements (mm). length 2.88 – 4.48, LH 0.75 – 0.90, WH 0.78 – 0.94, WF 0.46 – 0.58, WOT 0.20 – 0.24. Color. Head and mesosoma black, prothorax brown, metasoma dark brown; wings hyaline; antenna brown; legs dark brown to brown. Head. Malar space absent; median clypeal lobe angulate; gena not visible in dorsal view; mandible with one distinct ventralmost tooth and three remaining teeth fused; hypostoma angled medially; frons punctured and smooth; frontal line narrow and carinate; eye glabrous; anterior ocellus posterior to supra-ocular line. Mesosoma. Dorsal pronotal area punctured and smooth; notaulus complete, narrow, diverging posterad; metapostnotal median carina of metapectal-propodeal disc irregular, incomplete posteriorly. Genitalia. Harpe with medial-dorsal corner partially arched, apical margin with large excavation, dorsal margin straight, divided in dorsal and ventral arms, dorsal arm laminar, apex acute and diverging, ventral arm straight and setose posteriorly, apex posterior to apex of digitus; digitus with posterior margin smooth; aedeagus with apical lobe not gibbous to such an extent that dorsal surface is flat, aedeagal apex posterior to apex of ventral arm of cuspis, with two pairs of apical lobes, outer lobe apex anterior to inner lobe apex, outer lobe acute and inner lobe progressively narrowing. Female. Unknown. Variations. There are no variations.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058201D69FF13F8EE1DABFEEE.taxon	diagnosis	Differential diagnosis. This species is morphologically similar to C. quelatus by having the harpe with medial-dorsal corner partially arched, the apical margin with large excavation and the apical projection of basivolsella glabrous. However, C. tremembe sp. nov. differs from C. quelatus by having the hypostoma angled medially, the mesopleural epicoxal sulcus absent, the hypopygeal anteromedial apodeme absent, the dorsal harpal margin straight, the cuspis glabrous, the cuspis apex posterior to apex of digitus, the cuspis apex straight and aedeagus with one pair of apical lobes, whereas C. quelatus has the hypostoma curved medially, the mesopleural epicoxal sulcus present, the hypopygeal anteromedial apodeme present, the dorsal harpal margin outcurved, the cuspis setose, the cuspis apex anterior to apex of digitus, the cuspis apex sharp and aedeagus with two pairs of apical lobes.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058201D69FF13F8EE1DABFEEE.taxon	etymology	Etymology. The specific epithet tremembe refers to the indigenous people “ Tremembé ” from Ceará and Maranhão, Brazil.	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058201D69FF13F8EE1DABFEEE.taxon	materials_examined	Material examined. Holotype, male: BRAZIL, MA [ranhão], Mirador, Parq [ue] Estadual Mirador, Base da Geraldina, 06 ° 37 ' 25 '' S 45 ° 52 ' 08 '' W, Malaise trap, 14 - 18. viii. 2012, J. S. Pinto Júnior, F. L. - de-Oliveira and D. W. A. Marques cols. (CZMA); Paratypes. 1 male, [no further data], Malaise trap, 18 - 25. iii. 2012, F. Limeira-de-Oliveira, D. W. A. Marques cols. (CZMA); CE [ará], Ubajara, PAR [que] NA [cional] Ubajara, Cachoeira do Cafundó, 03 ° 50 ' 13 '' S 40 ° 54 ' 35 '' W, Malaise trap, 1 male, 16 - 31. vii. 2012, J. S. Pinto Junior and F. Limeira-de-Oliveira cols. (NESB); 1 male, varredura, 20 - 22. vii. 2012, J. A. Rafael and F. Limeira-de-Oliveira cols. (CZMA); MA [ranhão], 2 males, same data as the holotype (CZMA, NESB); 1 male, 06 ° 37 ' 06 '' S 45 ° 51 ' 51 '' W, 411 m, Malaise trap, 02 - 12. viii. 2013, F. Limeira-de-Oliveira, L. L. M. Santos and T. L. Rocha cols. (CZMA); 1 male, 06 ° 37 ' 48 '' S 45 ° 52 ' 49 '' W, 416 m, 20 - 31. xi. 2013, T. L. Rocha, F. Limeira-de-Oliveira and L. L. M. Santos (CZMA); 1 male, 06 ° 37 ' 48 '' S 45 ° 52 ' 46 '' W, 428 m, 19 - 28. ii. 2013, F. Limeira-de-Oliveira, L. L. M. Santos, L. S. Santos cols. (CZMA); 1 male, 20 - 31. viii. 2013 (CZMA).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
0388E84058201D69FF13F8EE1DABFEEE.taxon	distribution	Distribution. Brazil (Ceará, Maranhão).	en	Marques Jr, Wilson J., Azevedo, Celso O., Colombo, Wesley D. (2025): Revision of the Brazilian Northeastern Cleistepyris (Hymenoptera: Bethylidae), with description of six new species. Zootaxa 5692 (1): 1-30, DOI: 10.11646/zootaxa.5692.1.1, URL: https://doi.org/10.11646/zootaxa.5692.1.1
