identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03880E0EC224815EFF5EF97C9D66FA18.text	03880E0EC224815EFF5EF97C9D66FA18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aptinoma Fisher	<div><p>Aptinoma Fisher gen. n.</p><p>Figures 1–5, 7 a</p><p>Type-species: Aptinoma mangabe sp. n., by present designation.</p><p>Diagnosis of worker. Dimorphic, with distinct minor and major workers. Major workers polymorphic. Lateral and anterior corner of hypostoma reduced to a thin sclerite, without expanded flange. Mandible with 4–6 teeth apically on masticatory margin; counting from the apex, third tooth smaller than fourth, fifth smaller than sixth. Teeth followed by a fine series of 4–8 denticles (effaced in some major workers); basal angle blunt, with a relatively uninterrupted curve between the masticatory and basal margins; basal margin with minute serrations. Antennae with 12 segments. Palpal formula 6:3 (confirmed with dissection of A. mangabe). Fourth maxillary palp segment subequal in length to segment 5. Fifth maxillary palp segment at the apical extreme of segment 4. Metanotal groove impressed but often weak. Propodeum unarmed, without any acute angle. Propodeal declivity almost straight in profile. Petiolar scale present, in the form of a node with low standing forward face. Gaster with only four plates on upper surface; fifth tergite (abdominal tergite 7) reflexed ventrally, and visible in ventral view where it forms a transverse plate abutting the fifth sternite. Mesosoma lacking erect hairs.</p><p>Diagnosis of queen. Based on one specimen of A. mangabe . Ergatoid with diagnostic characters of workers. Ocelli present. Scutellum distinct. Petiolar scale present, in the form of a node with a low standing forward face.</p><p>Diagnosis of male. Based on A. mangabe . Scape not reaching or surpassing posterior lateral margin of head in full-face view; scape length shorter than the length of segments 3 + 4. Second antenatal segment (pedicle) cone-shaped. Compound eyes relatively anterior on head. Anterior clypeal margin with a broad, shallow concavity. Posterior clypeal margin between the anterior and posterior surfaces of the antennal condyle. Palpal formula 6:3 (not confirmed with dissection). Mandible with single apical tooth followed by series of serrate teeth or denticles (&gt; 20) along masticatory margin; basal angle of mandible indistinct, with a relatively uninterrupted curve between the two margins. Petiole with distinct scale, angled dorsally and strongly inclined anteriorly, with anterior face shorter than posterior face. First gastral segment in dorsal view with a groove for reception of entire height of petiole.</p><p>Discussion of characters. In many respects Aptinoma workers are similar to Tapinoma workers. Both genera share a number of traits, including the shape of the clypeus, position of eyes, and shape and number of mandibular teeth and denticles. Aptinoma differs from Tapinoma in palpal formula (6: 3 in Aptinoma, 6: 4 in Tapinoma (confirmed with dissection; Fig. 7 c). Aptinoma also differs in having a more developed petiole node. The petiolar node in Tapinoma is flat, with almost no visible rise in the anterior face. In Aptinoma, a visible face is present. The face is more pronounced in major workers. The combination of palpal formula and form of petiolar node form an inclusive diagnosis that isolates Aptinoma from all other genera within the subfamily. Within the Malagasy region, the presence of dimorphic workers (with majors and minors), distinguish Aptinoma from all other dolichoderine genera. In males, the length of scape (shorter than the length of antennal segments 3 + 4), the presence of a petiole scale, and palpal formula 6:3 form an inclusive diagnosis.</p></div>	https://treatment.plazi.org/id/03880E0EC224815EFF5EF97C9D66FA18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fisher, Brian L.	Fisher, Brian L. (2009): Two new dolichoderine ant genera from Madagascar: Aptinoma gen. n. and Ravavy gen. n. (Hymenoptera: Formicidae). Zootaxa 2118: 37-52, DOI: 10.5281/zenodo.274902
03880E0EC227815EFF5EFA7099F0F8B9.text	03880E0EC227815EFF5EFA7099F0F8B9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aptinoma	<div><p>Key to workers of Malagasy Aptinoma</p><p>1 In profile, metanotal groove distinct and set in shallow V-shaped impression (Figs 1 a, e). Propodeal dorsum more or less convex, meeting steeply sloped declivity in blunt, narrowly rounded curve. In dorsal view, lateral margin of pronotum evenly convex, without blunt angle at widest point (Figs 1 c, 2c). Body light to dark brown............................. ......................................................................................................................................................... Aptinoma antongil</p><p>- In profile, metanotal groove not set in a V-shaped impression (Figs 3 a, 4a). Propodeum in absolute profile without differentiated dorsal and declivitous surfaces, forming single evenly rounded surface. In dorsal view, lateral margin of pronotum with blunt angle at widest point (Figs 3 c, g). Body yellow to orange. ...................................................... ........................................................................................................................................................ Aptinoma mangabe</p></div>	https://treatment.plazi.org/id/03880E0EC227815EFF5EFA7099F0F8B9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fisher, Brian L.	Fisher, Brian L. (2009): Two new dolichoderine ant genera from Madagascar: Aptinoma gen. n. and Ravavy gen. n. (Hymenoptera: Formicidae). Zootaxa 2118: 37-52, DOI: 10.5281/zenodo.274902
03880E0EC226815FFF5EFF079E92F940.text	03880E0EC226815FFF5EFF079E92F940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aptinoma antongil Fisher	<div><p>Aptinoma antongil Fisher sp. n.</p><p>Figures: worker 1, 2</p><p>Type material: Holotype: major worker, MADAGASCAR, Province Toamasina, Parc National de Masoala, 39.4 km 150° SSE Maroantsetra, 15.71°S, 49.97°E, 200m, 1 December 2001, under canopy moss and leaf litter, rainforest (coll. B.L.Fisher &amp; H.J.Ratsirarson) collection code: BLF04719, pin code: CASENT0418269 (CASC). Paratypes: 5 workers with same data as holotype but pins coded CASENT0 418267, CASENT0418268, CASENT0418270 (BMNH, CASC, MBCC, MCZC).</p><p>Major Worker measurements: maximum and minimum based on all specimens, n=10: HL 0.47–0.63, HW 0.48–0.63, CI 93–102, ED 0.10–0.13, SL 0.34–0.40, SI 63–73, WL 0.50–0.70, FL 0.39–0.49, PW 0.29–0.39.</p><p>Minor Worker measurements: maximum and minimum based on all specimens, n=2: HL 0.40–0.42, HW 0.39, CI 94–98, ED 0.09, SL 0.30–0.32, SI 77–80, WL 0.46, FL 0.34, PW 0.22–0.26.</p><p>With characters described in the genus diagnosis for major and minor workers: Anterior clypeal margin without a distinct shallow impression; medial clypeal margin with two or more pairs of setae directed anteriorly; additional pair of setae, slightly anterior of clypeal margin, directed anterodorsally. Scape not surpassing posterior margin of the head in full-face view in both major and minor . In dorsal view, lateral margin of pronotum evenly convex, without blunt angle at widest point. Metanotal groove in profile distinct and impressed. Dorsum of propodeum more or less convex, meeting the steeply sloped declivity in a blunt, narrowly rounded curve. In profile, propodeal spiracle along margin of declivitous face. Body light to dark brown.</p><p>Major workers are variable in size and are distinguished from the minor workers by shorter scapes relative to head width (SI 63–73). Since only two minor workers were collected, further samples are needed to evaluate if there is a continuum of worker sizes from minors to the larger majors.</p><p>A. antongil can be separated from A. mangabe by color and shape of propodeum and metanotal groove as outlined in the key to species. Minor workers of A. antongil are easily distinguished from minor workers of A. mangabe by length of scape. The scape surpasses the posterior margin of the head in minor workers in A. mangabe while in A. antongil it does not reach the margin.</p><p>QUEEN and MALE: unknown.</p><p>Distribution and biology. The distribution is limited to forests around Antongil Bay, Madagascar. Specimens have been collected from canopy habitat on the Masoala Peninsula and on low vegetation near the summit of Montagne d'Anjanaharibe (1100 m) just north of Maroansetra. Like A. mangabe, this species is arboreal. However, it may not nest in dead twigs as does A. mangabe . Collections suggest that the species nests under canopy litter and moss. The collections in Masoala were made by exploring the canopy using ropes. A. antongil workers were found foraging under moss and litter at the junction of branches.</p><p>Additional material examined for Aptinoma antongil: In addition to the type material, specimens from five additional collecting events from the following two localities were examined in this study. MADAGASCAR, Province Toamasina: Parc National de Masoala, 39.4 km 150° SSE Maroantsetra, 200 m, 15.71°S, 49.97°E, (B.L.Fisher &amp; H.J.Ratsirarson), 1 December 2001, rainforest, canopy moss and leaf litter collection codes: BLF04716, BLF04717, BLF04718 BLF04721. Montagne d'Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, 1100 m, 15.17833°S, 49.635°E, (Fisher et al.), 14 March 2003, montane rainforest, beating low vegetation, collection code: BLF08151.</p></div>	https://treatment.plazi.org/id/03880E0EC226815FFF5EFF079E92F940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fisher, Brian L.	Fisher, Brian L. (2009): Two new dolichoderine ant genera from Madagascar: Aptinoma gen. n. and Ravavy gen. n. (Hymenoptera: Formicidae). Zootaxa 2118: 37-52, DOI: 10.5281/zenodo.274902
03880E0EC226815AFF5EF8A99DE5F854.text	03880E0EC226815AFF5EF8A99DE5F854.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aptinoma mangabe Fisher	<div><p>Aptinoma mangabe Fisher sp. n.</p><p>Figures: worker 3, 7a, queen 4, male 5</p><p>Type Material: Holotype: major worker, MADAGASCAR, Province Toamasina, Nosy Mangabe, 7.43 km S Maroantsetra, 15.4973°S, 49.76223°E, 5 m. 27 July 2007, ex dead twig above ground, littoral rainforest edge (coll. J.J. Rafanomezantsoa.) collection code: BLF17925, pin code: CASENT0130146 (CASC). Paratypes: 1 minor worker, 1 major worker, ergatoid queen, male with same data as holotype but pins coded CASENT0130148, CASENT0192285, CASENT0175008, CASENT0130147, respectively (BMNH, CASC, MCZC).</p><p>Major worker measurements: maximum and minimum based on all specimens, n=5, HL 0.54–0.57, HW 0.53–0.56, CI 99–102, ED 0.10–0.11, SL 0.39–0.41, SI 70–75, WL 0.57–0.64, FL 0.47–0.49, PW 0.34–0.37.</p><p>Minor worker measurements: maximum and minimum based on all specimens, n=5, HL 0.42–0.46, HW 0.37–0.42, CI 86–96, ED 0.09, SL 0.38–0.41, SI 97–103, WL 0.48–0.54, FL 0.41–0.45, PW 0.22–0.29.</p><p>With characters described in the genus diagnosis for major and minor workers: Apical teeth on mandible followed by 4–6 denticles; basal margin with minute, serrate teeth. Anterior clypeal margin with distinct shallow median impression; medial clypeal margin with three pairs of setae arched anteriorly over mandibles; additional pair of setae on clypeus, posterior of clypeal margin, directed anterodorsally. Scape surpassing posterior margin of head in full-face view in minors, not surpassing margin in majors. In dorsal view, lateral margin of pronotum with blunt angle at widest point. Metanotal groove in profile not impressed; propodeum in profile with dorsal and declivitous surfaces not differentiated, forming single evenly rounded surface without trace of angle at junction with declivity. Body yellow to orange.</p><p>A. mangabe can be separated from A. antongil by color and shape of propodeum and metanotal groove as outlined in the key to species. Differences in minor workers are discussed under antongil .</p><p>Queen (ergatoid) measurements: n=1. HL 0.61, HW 0.59, CI 97, ED 0.15, SL 0.44, SI 75, WL 0.82, FL 0.57, PW 0.44.</p><p>All main morphological characters of the major worker are duplicated in the queen caste. In the ergatoid queen, ocelli are present and the mesoscutellum is distinct. Metanotum indistinguishable, but metanotal groove distinctly wider than that of workers.</p><p>Male measurements: maximum and minimum based on n=3: HL 0.40–0.43, HW 0.54–0.57, CI 127–137, ED 0.22–0.25, SL 0.21, SI 37–39, WL 0.74–0.78, FL 0.52–0.55.</p><p>In profile, inner margin of eye convex, not flat. Antenna with 13 articles. Scape not reaching posterior margin of head, shorter than flagellar segments 1+2+3. Pedicel conical. First flagellar article cylindrical, straight, articles 1 and 2 twice as long as broad. Anteromedial clypeal margin with central concavity. Anterior clypeal margin with several pairs of straight setae projecting anteriorly, central pair longer than the setae at lateral margins. Just posterior to anterior margin, a single pair of suberect setae projecting anterodorsally. Posterior clypeal margin convex between antennal socket cavities. Palpal formula 6:3 (not confirmed with dissection). Mandible with apical tooth followed by a series of 15 or more minute denticles that decrease in size towards the basal margin. An uninterrupted curve between masticatory and basal margins. Propodeal angle indistinct; declivitous and dorsal faces of propodeum forming a single convexity. Petiole in profile with distinct scale, angled dorsally and strongly inclined anteriorly with anterior margin shorter than posterior margin. Venter of petiole with weakly developed lobe. Pygostyles present.</p><p>Distribution and biology. Distribution is limited to forests around Antongil Bay including the island of Nosy Mangabe and the adjacent forest of the Masoala Peninsula. Specimens were collected from dead twigs or vines, or foraging in the canopy. Males were collected in colonies in April and July. The only queen collected was an ergatoid.</p><p>Additional material examined for Aptinoma mangabe: In addition to the type material, specimens from 8 additional collecting events from the following 4 localities were examined in this study. MADAGASCAR, Province Toamasina:</p><p>P.N. Masoala, 40 km 154° SSE Maroantsetra, 150 m, 15.72667°S, 49.95667°E, (A. Dejean &amp; B. Rajemison), 14 October 2001, rainforest, canopy. Tampolo, Masoala Peninsula, 40.4 km 154° SSE Maroantsetra, 30 m, 15.73°S, 49.96°E, (B.L.Fisher, &amp; H.J.Ratsirarson), 28 November 2001, rainforest, canopy collection codes: BLF04698 BLF04713, BLF04717. Nosy Mangabe, 7.43 km S Maroantsetra, 5 m, 15.4973°S, 49.76223°E, (C. Ranaivo &amp; C. E. Randrianandrasana), littoral rainforest edge, collection codes: BLF17931, 27 July 2007, on low vegetation; BLF18015, 31 August 2007, ex dead twig above ground. Nosy Mangabe, &lt;5 m, 15.5°S, 49.76667°E, (P.S.Ward), collection code: PSW10346, 20 April 1989, rainforest edge, ex dead twig of vine.</p></div>	https://treatment.plazi.org/id/03880E0EC226815AFF5EF8A99DE5F854	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fisher, Brian L.	Fisher, Brian L. (2009): Two new dolichoderine ant genera from Madagascar: Aptinoma gen. n. and Ravavy gen. n. (Hymenoptera: Formicidae). Zootaxa 2118: 37-52, DOI: 10.5281/zenodo.274902
03880E0EC22F8156FF5EFF719ED4FB24.text	03880E0EC22F8156FF5EFF719ED4FB24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ravavy Fisher	<div><p>Ravavy Fisher gen. n.</p><p>Figures 6, 7 b</p><p>Type species Ravavy miafina, sp. n., by present designation.</p><p>WORKER and QUEEN: unknown.</p><p>Diagnosis of male. Based on specimens of Ravavy miafina . Head longer than wide. Ventral margin of eye more or less flat. Antennae with 13 articles. Scape length twice that of flagellar segment 1. Scape not reaching posterior margin of head. Pedicel conical. First flagellar segment cylindrical, straight. Flagellar segments 1 and 2 three times as long as broad. Anteromedian clypeal margin entire, without central concavity. Posterior clypeal margin straight, not projecting between antennal socket cavities. Palpal formula 6:3 (confirmed with dissection). Mandible with single narrow apical tooth, located at tip of mandible; without series of serrate denticles or teeth along margin. Masticatory margin meeting basal margin in abrupt angle. Basal margin elongate, greater than twice length of masticatory margin. Propodeal angle indistinct; declivitous and dorsal faces of propodeum convex. In profile, petiole with distinct node; anterior and posterior margin subequal in height, dorsum evenly convex. Attachment to gaster broad. First gastral segment in dorsal view without grove for the reception of the entire height of the petiole.</p><p>Discussion of male characters. Ravavy is distinguished from all other described males in the dolichoderine genera by its unique mandibular shape. The elongate basal margin and reduced masticatory margin is considered apomorphic for the genus. The shape of the clypeus, length of scape and form of petiole, together form an inclusive diagnosis that isolates Ravavy from all other genera in the subfamily. No other described male is even superficially similar to Ravavy . The shape of the mandible in Bothriomyrmex Emery (see images on AntWeb CASENT0103279), however, is reminiscent in some respects. The mandible in Bothriomyrmex is not triangular as in Tapinoma for example, but like Ravavy, has a short masticatory margin compared to the basal margin. In Bothriomyrmex, however, the masticatory margin includes at least three teeth, while in Ravavy a single tooth is present.</p></div>	https://treatment.plazi.org/id/03880E0EC22F8156FF5EFF719ED4FB24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fisher, Brian L.	Fisher, Brian L. (2009): Two new dolichoderine ant genera from Madagascar: Aptinoma gen. n. and Ravavy gen. n. (Hymenoptera: Formicidae). Zootaxa 2118: 37-52, DOI: 10.5281/zenodo.274902
03880E0EC22F8151FF5EFA859E99FC83.text	03880E0EC22F8151FF5EFA859E99FC83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ravavy miafina Fisher	<div><p>Ravavy miafina Fisher sp. n.</p><p>Figures: male 6, 7b</p><p>Type material: Holotype: male, MADAGASCAR, Province Antsiranana, Ampasindava, Forêt d'Ambilanivy, 3.9 km 181° S Ambaliha, 13.79861°S, 48.16167°E, 600m, 4–9 March 2001, Malaise trap, rainforest (coll. B.L. Fisher et al.) collection code: BLF03251, pin code: CASENT0081523 (CASC). Paratypes: 6 males with same data as holotype but pins coded CASENT0 476953, CASENT0 476983, CASENT0 476947, CASENT0081525 (BMNH, CASC, MBCC, MCZC).</p><p>WORKER and QUEEN: unknown.</p><p>Male measurements: maximum and minimum based on n=10: HL 0.46–0.50, HW 0.43–0.47, CI 91–98, ED 0.20–0.23, SL 0.17–0.21, SI 39–47, WL 0.78–0.84, FL 0.60–0.70.</p><p>With characters of the genus, and the following. Head longer than wide, CI 91–98. Clypeus prominent; width of clypeus from posterior margin at antennal sockets to anterior margin equal in length to antennal segment 3 (flagellar segment 1). Anteromedial clypeal margin with single pair of setae projecting in line with the mandibles, posterior to this pair and more distal, single pair of suberect setae project anterodorsally above mandibles. Pair of erect setae posterior of clypeus between antennal sockets. Clypeus and head with short appressed setae. Mandibles smooth under dense matte of appressed setae. Dorsum of mesosoma, and gaster with dense pubescence. Mesosoma without erect setae; abdominal tergites A5–8 with at least one pair of erect setae. Petiolar node smooth and shiny, without pubescence. Venter of petiole with well-developed lobe. Propodeum elongate, dorsal and declivitous faces continuous without angle. Pygostyles present.</p><p>Distribution and biology. Males have been collected in Malaise traps sampled from January through August across western Madagascar. Habitats in which this species was collected include the spiny bush near Fort Dauphin, dry forest in Kirindy near Morandava, rainforest in the Sambirano region on the Ampasindava Peninsula, and coastal scrub on sand dunes near Antiranana in the northeastern tip of Madagascar.</p><p>The nesting habits of Ravavy are not known.</p><p>Additional material examined for Ravavy miafina: In addition to the type material, specimens from 9 additional collecting events from five localities were examined in this study. MADAGASCAR, Province Toliara: Forêt de Kirindy, 15.5 km 64° ENE Marofandilia, 100 m, 20.045°S, 44.66222°E, (Fisher et al.), tropical dry forest, Malaise trap collection code: BLF04601, 28 November 2001 – 03 December 2001, Malaise trap; PN Andohaela, 60 m, 24.83083°S, 46.53617°E, (Rin'Ha, Irwin), dry forest spiny forest, Malaise trap, collection code: MG-21-42, 17, December 2003– 19 December 2003. Province Antsiranana: Parc National Montagne d'Ambre [Petit Lac road], 1125 m, 12.52028°S, 49.17917°E, (R. Harin'Hala): rainforest, Malaise trap, collection code MA-01-01D-05, 0 4 March 2001 – 19 March 2001; collection code: MA-01-01D-05, 0 4 March 2001 – 19 March 2001, collection code: MA-01-01D-12, 30 May 2001 – 06 June 2001. Sakalava Beach [vegetated beach dunes], 10 m, 12.26278 °S, 49.3975°E, (R. Harin'Hala), across sandy trail in dwarf littoral forest, Malaise trap, collection code: MA-01-04B-10, 16 May 2001 – 31 May 2001; collection code: MA-01- 04B-17, 13 August 2001 – 20 August 2001; 7 km N Joffreville, 360 m, 12.33333°S, 49.25°E, (M.E. Irwin), in dry forest, hand netted, collection code: MA- 01-07-03, 26 January 2001; (R. Harin'Hala) collection code: MA- 01-07-06, 29 January 2001 – 15 February 2001.</p></div>	https://treatment.plazi.org/id/03880E0EC22F8151FF5EFA859E99FC83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fisher, Brian L.	Fisher, Brian L. (2009): Two new dolichoderine ant genera from Madagascar: Aptinoma gen. n. and Ravavy gen. n. (Hymenoptera: Formicidae). Zootaxa 2118: 37-52, DOI: 10.5281/zenodo.274902
